268 lines
30 KiB
XML
268 lines
30 KiB
XML
<document id="CD46766160B96154EEF8403574B59A8B" ID-CLB-Dataset="53387" ID-DOI="10.11646/zootaxa.5124.3.1" ID-GBIF-Dataset="de55f596-c349-4ca2-a826-0e2bad912aa3" ID-ISSN="1175-5326" ID-Zenodo-Dep="6410696" ID-ZooBank="67D4C618-C7C2-4FFC-925B-B260E11CD7B4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1649054363463" checkinUser="plazi" docAuthor="Jiang, Zhuo-Yin, Zhao, Shuang, Yang, Zhen-Yu, Jia, Feng-Long & Hájek, Jiří" docDate="2022" docId="291CAA4FBF54FF941DC5FE169D34FA09" docLanguage="en" docName="zootaxa.5124.3.1.pdf" docOrigin="Zootaxa 5124 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Copelatus japonicus Sharp 1884" docType="treatment" docVersion="9" lastPageNumber="260" masterDocId="D525D237BF5DFF9D1D52FF849836FF9E" masterDocTitle="A review of Copelatus Erichson, 1832 of Mainland China, with description of ten new species from the japonicus complex (Coleoptera: Dytiscidae: Copelatinae)" masterLastPageNumber="295" masterPageNumber="251" pageNumber="260" updateTime="1699319256616" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="27A8F0E585F99FD09206EB29F8063F49">A review of Copelatus Erichson, 1832 of Mainland China, with description of ten new species from the japonicus complex (Coleoptera: Dytiscidae: Copelatinae)</mods:title>
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<mods:namePart id="CB05416FC5FE43917814F5346F620C4C">Jiang, Zhuo-Yin</mods:namePart>
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<mods:affiliation id="D2A6B997F699B55C8B459458F0C04C83">State Key Laboratory of Biocontrol and Institute of Entomology, Sun Yat-sen University, Guangzhou 510275, China. zhuoyin _ jiang @ 126. com; https: // orcid. org / 0000 - 0001 - 6825 - 4283</mods:affiliation>
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<mods:namePart id="A853904BF5737FCD78A3CCE8ECD71F56">Zhao, Shuang</mods:namePart>
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<mods:affiliation id="16D7C5C07213D1C9C5B811DB73BC7887">Guangzhou Customs Technology Center, Nansha District, Guangzhou, China.</mods:affiliation>
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<mods:namePart id="6C6AAE398CD3AB13AA1559867E92F368">Yang, Zhen-Yu</mods:namePart>
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<mods:affiliation id="F48F9D515CBAE3BD58DA15B8A3E0C952">Brilliant Landscaping and Gardening Company Limited, Macau.</mods:affiliation>
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<mods:namePart id="59EE04FD9DBCC6E6E8A3997E720766EB">Jia, Feng-Long</mods:namePart>
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<mods:affiliation id="9616DF280D8F82A19492E98BC10F0A8C">State Key Laboratory of Biocontrol and Institute of Entomology, Sun Yat-sen University, Guangzhou 510275, China. fenglongjia @ aliyun. com; lssjfl @ mail. sysu. edu. cn; http: // orcid. org / 0000 - 0003 - 2391 - 5038</mods:affiliation>
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<mods:nameIdentifier id="25F19C3906AC96357794FC9B6EBFB00B" type="email">fenglongjia@aliyun.com</mods:nameIdentifier>
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<mods:namePart id="AEB8B4AB1BA6183EA76560F4F6284CF6">Hájek, Jiří</mods:namePart>
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<mods:affiliation id="F9A12330ECAB9086AC181E1296D8B428">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice, Czech Republic. jiri. hajek @ nm. cz; https: // orcid. org / 0000 - 0001 - 5779 - 1542</mods:affiliation>
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<mods:title id="80C9E4DD7249FA370234B6947F60F691">Zootaxa</mods:title>
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<mods:date id="8FF39DF68EFD6B85EF65E4295DED4863">2022</mods:date>
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<mods:number id="0100442FA5D412181E2762E569DAD825">2022-04-04</mods:number>
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<treatment id="291CAA4FBF54FF941DC5FE169D34FA09" ID-DOI="http://doi.org/10.5281/zenodo.6410760" ID-GBIF-Taxon="194444065" ID-Zenodo-Dep="6410760" LSID="urn:lsid:plazi:treatment:291CAA4FBF54FF941DC5FE169D34FA09" httpUri="http://treatment.plazi.org/id/291CAA4FBF54FF941DC5FE169D34FA09" lastPageNumber="260" pageId="9" pageNumber="260">
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<taxonomicName id="66B560DABF54FF941DC5FE169A1CFE32" ID-CoL="5ZX7L" authority="Sharp, 1884" authorityName="Sharp" authorityYear="1884" box="[151,554,402,429]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="260" phylum="Arthropoda" rank="species" species="japonicus">
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<emphasis id="93C1C74BBF54FF941DC5FE169A1CFE32" bold="true" box="[151,554,402,429]" pageId="9" pageNumber="260">
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<emphasis id="93C1C74BBF54FF941DC5FE1699BAFE33" bold="true" box="[151,396,402,429]" italics="true" pageId="9" pageNumber="260">Copelatus japonicus</emphasis>
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<bibRefCitation id="C52466A8BF54FF941CC6FE169A1CFE32" author="Sharp, D." box="[404,554,402,428]" pageId="9" pageNumber="260" pagination="439 - 464" refId="ref33527" refString="Sharp, D. (1884) The water-beetles of Japan. The Transactions of the Entomological Society of London, 1884, 439 - 464." type="journal article" year="1884">Sharp, 1884</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="E9AF48D2BF54FF941DC5FE339975FE4F" box="[151,323,439,465]" pageId="9" pageNumber="260" type="description">
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<paragraph id="A10A1B59BF54FF941DC5FE339975FE4F" blockId="9.[151,554,402,465]" box="[151,323,439,465]" pageId="9" pageNumber="260">
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(
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<figureCitation id="398E07DCBF54FF941DCDFE3398D1FE4F" box="[159,231,439,465]" captionStart="FIGURES 5–8" captionStartId="32.[151,264,1943,1967]" captionTargetBox="[239,1352,196,1907]" captionTargetId="figure-23@32.[221,1367,181,1920]" captionTargetPageId="32" captionText="FIGURES 5–8. Habitus of Copelatus in China. 5. C. diversistriatus sp. nov. (holotype ♂; Pu’er, Yunnan); 6. C. diversistriatus sp. nov. (♂; NE Mintaingbin Forest Camp, Myanmar); 7. C. felicis sp. nov. (holotype ♂; Baishui, Yunnan); 8. C. japonicus (♂; Shanghai). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6410704" httpUri="https://zenodo.org/record/6410704/files/figure.png" pageId="9" pageNumber="260">Figs 8</figureCitation>
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,
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<figureCitation id="398E07DCBF54FF941DA6FE33990DFE4F" box="[244,315,439,465]" captionStart="FIGURES 39–50" captionStartId="39.[151,264,1839,1863]" captionTargetBox="[210,1370,207,1804]" captionTargetId="figure-18@39.[192,1395,181,1815]" captionTargetPageId="39" captionText="FIGURES 39–50. Male genitalia of Copelatus. 39, 41, 43, 45, 47, 49. median lobe in lateral view; 40, 42, 44, 46, 48, 50. paramere. 39–40. C. japonicus (Shanghai); 41. C. malaisei (a. Gaoligongshan, Yunnan; b. Renjiafen, Yunnan); 42. C. malaisei (Renjiafen, Yunnan); 43–44. C. miaowangi sp. nov. (paratype, Liupanshui, Guizhou); 45–46. C. mopanshanensis sp. nov. (holotype, Mopanshan, Yunnan); 47–48. C. nanlingensis sp. nov. (holotype, Dadongshan, Guangdong); 49–50. C. oblitus (Macao). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6412781" httpUri="https://zenodo.org/record/6412781/files/figure.png" pageId="9" pageNumber="260">39–40</figureCitation>
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)
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</paragraph>
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<paragraph id="A10A1B59BF54FF941DC5FE7F9A14FD8D" blockId="9.[151,546,507,531]" box="[151,546,507,531]" pageId="9" pageNumber="260">
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<treatmentCitationGroup id="81A53C77BF54FF941DC5FE7F9A14FD8D" box="[151,546,507,531]" pageId="9" pageNumber="260">
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<treatmentCitation id="20143D48BF54FF941DC5FE7F9A28FD8D" author="Sharp, D." box="[151,542,507,531]" page="445" pageId="9" pageNumber="260" year="1884">
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<taxonomicName id="66B560DABF54FF941DC5FE7F9A28FD8D" ID-CoL="5ZX7L" authority="Sharp, 1884: 445" authorityName="Sharp" authorityPageNumber="445" authorityYear="1884" box="[151,542,507,531]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="260" phylum="Arthropoda" rank="species" species="japonicus">
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<emphasis id="93C1C74BBF54FF941DC5FE7F9951FD8D" box="[151,359,507,531]" italics="true" pageId="9" pageNumber="260">Copelatus japonicus</emphasis>
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<bibRefCitation id="C52466A8BF54FF941C3FFE7F9A28FD8D" author="Sharp, D." box="[365,542,507,531]" pageId="9" pageNumber="260" pagination="439 - 464" refId="ref33527" refString="Sharp, D. (1884) The water-beetles of Japan. The Transactions of the Entomological Society of London, 1884, 439 - 464." type="journal article" year="1884">Sharp, 1884: 445</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<subSubSection id="E9AF48D2BF54FF941DC5FDC599BBFCAD" pageId="9" pageNumber="260" type="materials_examined">
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<paragraph id="A10A1B59BF54FF941DC5FDC59B8BFDC2" blockId="9.[151,1437,577,1432]" box="[151,957,577,604]" pageId="9" pageNumber="260">
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<materialsCitation id="11DD1104BF54FF941DC5FDC59B8EFDC2" ID-GBIF-Occurrence="3746435316" box="[151,952,577,604]" country="Japan" location="Honshu Island" pageId="9" pageNumber="260" specimenCount="1" stateProvince="Wakayama" typeStatus="holotype">
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<emphasis id="93C1C74BBF54FF941DC5FDC59900FDC5" bold="true" box="[151,310,577,603]" pageId="9" pageNumber="260">
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<typeStatus id="7E0EA5FBBF54FF941DC5FDC598E4FDC5" box="[151,210,577,603]" pageId="9" pageNumber="260">Type</typeStatus>
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locality.
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</emphasis>
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“Kiishiu”
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<emphasis id="93C1C74BBF54FF941CFDFDC5998EFDC5" bold="true" box="[431,440,577,603]" pageId="9" pageNumber="260">[</emphasis>
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<collectingCountry id="D9A25BC9BF54FF941CEAFDC699CDFDC2" box="[440,507,578,604]" name="Japan" pageId="9" pageNumber="260">Japan</collectingCountry>
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,
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<location id="A46A4D82BF54FF941F54FDC59A9DFDC5" LSID="urn:lsid:plazi:treatment:291CAA4FBF54FF941DC5FE169D34FA09:A46A4D82BF54FF941F54FDC59A9DFDC5" box="[518,683,577,603]" country="Japan" name="Honshu Island" pageId="9" pageNumber="260" stateProvince="Wakayama">Honshu Island</location>
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,
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<collectingRegion id="6371D5BBBF54FF941FE4FDC59B84FDC2" box="[694,946,577,604]" country="Japan" name="Wakayama" pageId="9" pageNumber="260">Wakayama Prefecture</collectingRegion>
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]
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="A10A1B59BF54FF941D95FDE198D6FD72" blockId="9.[151,1437,577,1432]" pageId="9" pageNumber="260">
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<emphasis id="93C1C74BBF54FF941D95FDE19943FDE1" bold="true" box="[199,373,613,639]" pageId="9" pageNumber="260">Type material.</emphasis>
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<materialsCitation id="11DD1104BF54FF941C2EFDE19B2AFD3A" ID-GBIF-Occurrence="3746435384" collectionCode="BMNH" country="Japan" location="Japan" pageId="9" pageNumber="260" specimenCount="1" specimenCount-male="1" typeStatus="syntype">
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<typeStatus id="7E0EA5FBBF54FF941C2EFDE199EEFDE1" box="[380,472,613,639]" pageId="9" pageNumber="260" type="syntype">Syntype</typeStatus>
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<specimenCount id="B7B3D0D0BF54FF941C8DFDE299C2FD1E" box="[479,500,614,640]" pageId="9" pageNumber="260" type="male">♂</specimenCount>
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(
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<collectionCode id="C7A4839CBF54FF941F51FDE19A6BFDE1" box="[515,605,613,639]" pageId="9" pageNumber="260">BMNH</collectionCode>
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), labelled: “
|
||
<taxonomicName id="66B560DABF54FF941FB3FDE19B62FDE1" authorityName="Erichson" authorityYear="1832" box="[737,852,613,639]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="260" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
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/ japonicus. / Type D.S. /
|
||
<collectingCountry id="D9A25BC9BF54FF94192BFDE29C8AFD1E" box="[1145,1212,614,640]" name="Japan" pageId="9" pageNumber="260">Japan</collectingCountry>
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||
. Lewis [hw] // Type [round label with red frame, p] // Sharp Coll / 1905-313.”
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||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="11DD1104BF54FF941E75FD0D98EAFD72" ID-GBIF-Occurrence="3746435491" collectionCode="BMNH" collectorName="G. Lewis. & D. Sharp." country="Japan" location="Japan" pageId="9" pageNumber="260" specimenCount="2" specimenCount-female="2" typeStatus="syntype">
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<typeStatus id="7E0EA5FBBF54FF941E75FD0D9BB5FD3D" box="[807,899,649,675]" pageId="9" pageNumber="260" type="syntype">Syntype</typeStatus>
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<specimenCount id="B7B3D0D0BF54FF941EDBFD0E9BA8FD3A" box="[905,926,650,676]" pageId="9" pageNumber="260" type="female">♀</specimenCount>
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(
|
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<collectionCode id="C7A4839CBF54FF941EFFFD0D9C31FD3D" box="[941,1031,649,675]" pageId="9" pageNumber="260">BMNH</collectionCode>
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), labelled: “Kii, 1879 / Wada [hw] //
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<collectingCountry id="D9A25BC9BF54FF941DC5FD2A98ECFD56" box="[151,218,686,712]" name="Japan" pageId="9" pageNumber="260">Japan</collectingCountry>
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. /
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||
<collectorName id="0C407E8FBF54FF941DA0FD299969FD59" box="[242,351,685,711]" pageId="9" pageNumber="260">G. Lewis.</collectorName>
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/ 1910–320. [p] //
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||
<taxonomicName id="66B560DABF54FF941F79FD299AADFD59" authorityName="Erichson" authorityYear="1832" box="[555,667,685,711]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="260" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
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/ japonicus. [hw] //
|
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<taxonomicName id="66B560DABF54FF941E2AFD299BDEFD59" authorityName="Erichson" authorityYear="1832" box="[888,1000,685,711]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="260" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
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||
/ japonicus, /
|
||
<specimenCount id="B7B3D0D0BF54FF9419D3FD2A9CA0FD56" box="[1153,1174,686,712]" pageId="9" pageNumber="260" type="female">♀</specimenCount>
|
||
Sharp. / teste.
|
||
<collectorName id="0C407E8FBF54FF94186BFD299DABFD59" box="[1337,1437,685,711]" pageId="9" pageNumber="260">D.Sharp.</collectorName>
|
||
[hw]”
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="A10A1B59BF54FF941D95FD7199BBFCAD" blockId="9.[151,1437,577,1432]" pageId="9" pageNumber="260">
|
||
<emphasis id="93C1C74BBF54FF941D95FD719A05FC91" bold="true" box="[199,563,757,783]" pageId="9" pageNumber="260">Additional material examined.</emphasis>
|
||
<materialsCitation id="11DD1104BF54FF941F6BFD7199BFFCAD" ID-GBIF-Occurrence="3746435489" collectingDate="2021-09-10" collectionCode="SYSU" collectorName="D. Zhou" country="China" location="Jiading" municipality="Jiading District" pageId="9" pageNumber="260" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Shanghai">
|
||
<emphasis id="93C1C74BBF54FF941F6BFD719B29FC91" bold="true" box="[569,799,757,783]" pageId="9" pageNumber="260">
|
||
<collectingCountry id="D9A25BC9BF54FF941F6BFD719AA0FC91" box="[569,662,757,783]" name="China" pageId="9" pageNumber="260">CHINA</collectingCountry>
|
||
:
|
||
<collectingRegion id="6371D5BBBF54FF941FF4FD719B2CFC91" box="[678,794,757,783]" country="China" name="Shanghai" pageId="9" pageNumber="260">Shanghai</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="B7B3D0D0BF54FF941E74FD719B67FC8E" box="[806,849,757,784]" pageId="9" pageNumber="260" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="B7B3D0D0BF54FF941E0FFD719BB0FC8E" box="[861,902,757,784]" pageId="9" pageNumber="260" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectingMunicipality id="416E8123BF54FF941EC1FD719C77FC91" box="[915,1089,757,783]" pageId="9" pageNumber="260">Jiading District</collectingMunicipality>
|
||
,
|
||
<location id="A46A4D82BF54FF94191EFD719CABFC91" LSID="urn:lsid:plazi:treatment:291CAA4FBF54FF941DC5FE169D34FA09:A46A4D82BF54FF94191EFD719CABFC91" box="[1100,1181,757,783]" country="China" municipality="Jiading District" name="Jiading" pageId="9" pageNumber="260" stateProvince="Shanghai">Jiading</location>
|
||
new town,
|
||
<date id="D50B3D99BF54FF94184FFD719DAEFC91" box="[1309,1432,757,783]" pageId="9" pageNumber="260" value="2021-09-10">
|
||
<collectingDate id="C54FC471BF54FF94184FFD719DAEFC91" box="[1309,1432,757,783]" pageId="9" pageNumber="260" value="2021-09-10">10.ix.2021</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="0C407E8FBF54FF941DC5FC9E98C2FCAD" box="[151,244,793,820]" pageId="9" pageNumber="260">D. Zhou</collectorName>
|
||
leg. (
|
||
<collectionCode id="C7A4839CBF54FF941C66FC9D99B7FCAD" box="[308,385,793,819]" pageId="9" pageNumber="260">SYSU</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E9AF48D2BF54FF941D95FCB9999AFB05" pageId="9" pageNumber="260" type="diagnosis">
|
||
<paragraph id="A10A1B59BF54FF941D95FCB9999AFB05" blockId="9.[151,1437,577,1432]" pageId="9" pageNumber="260">
|
||
<emphasis id="93C1C74BBF54FF941D95FCB99974FCC9" bold="true" box="[199,322,829,855]" pageId="9" pageNumber="260">Diagnosis.</emphasis>
|
||
Larger (TL:
|
||
<quantity id="664DB6BCBF54FF941C8EFCB99A5CFCC6" box="[476,618,829,856]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="5.7" metricValueMin="5.3" pageId="9" pageNumber="260" unit="mm" value="5.5" valueMax="5.7" valueMin="5.3">5.3–5.7 mm</quantity>
|
||
), oblong-oval species. Head brown, with reddish clypeus and two oval spots on vertex; pronotum brown, laterally broadly orange, anterior and posterior margin orange translucent; elytra pale brown on disc, base and lateral sides yellowish brown. Pronotum with only several short longitudinal strioles laterally. Each elytron with six complete discal striae and one submarginal stria: all striae beginning at base; stria 1 longest, ending close to apex, odd striae somewhat longer than even ones; stria 6 shortest, ending at apical fourth; submarginal stria long, beginning before elytral mid-length and ending at apical fourth (
|
||
<figureCitation id="398E07DCBF54FF9419DCFC759CE2FB95" box="[1166,1236,1009,1035]" captionStart="FIGURES 5–8" captionStartId="32.[151,264,1943,1967]" captionTargetBox="[239,1352,196,1907]" captionTargetId="figure-23@32.[221,1367,181,1920]" captionTargetPageId="32" captionText="FIGURES 5–8. Habitus of Copelatus in China. 5. C. diversistriatus sp. nov. (holotype ♂; Pu’er, Yunnan); 6. C. diversistriatus sp. nov. (♂; NE Mintaingbin Forest Camp, Myanmar); 7. C. felicis sp. nov. (holotype ♂; Baishui, Yunnan); 8. C. japonicus (♂; Shanghai). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6410704" httpUri="https://zenodo.org/record/6410704/files/figure.png" pageId="9" pageNumber="260">Fig. 8</figureCitation>
|
||
). Median lobe of aedeagus in lateral view simple, sickle shaped, strongly curved; broad in basal half and slender in apical half; apex pointed (
|
||
<figureCitation id="398E07DCBF54FF941DA9FBBD9966FBCD" box="[251,336,1081,1107]" captionStart="FIGURES 39–50" captionStartId="39.[151,264,1839,1863]" captionTargetBox="[210,1370,207,1804]" captionTargetId="figure-18@39.[192,1395,181,1815]" captionTargetPageId="39" captionText="FIGURES 39–50. Male genitalia of Copelatus. 39, 41, 43, 45, 47, 49. median lobe in lateral view; 40, 42, 44, 46, 48, 50. paramere. 39–40. C. japonicus (Shanghai); 41. C. malaisei (a. Gaoligongshan, Yunnan; b. Renjiafen, Yunnan); 42. C. malaisei (Renjiafen, Yunnan); 43–44. C. miaowangi sp. nov. (paratype, Liupanshui, Guizhou); 45–46. C. mopanshanensis sp. nov. (holotype, Mopanshan, Yunnan); 47–48. C. nanlingensis sp. nov. (holotype, Dadongshan, Guangdong); 49–50. C. oblitus (Macao). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6412781" httpUri="https://zenodo.org/record/6412781/files/figure.png" pageId="9" pageNumber="260">Fig. 39</figureCitation>
|
||
). Parameres slender, ‘C’-shaped; apex short and broad; apical lobes moderately long, club-shaped (
|
||
<figureCitation id="398E07DCBF54FF941DF2FBD998C3FBE9" box="[160,245,1117,1143]" captionStart="FIGURES 39–50" captionStartId="39.[151,264,1839,1863]" captionTargetBox="[210,1370,207,1804]" captionTargetId="figure-18@39.[192,1395,181,1815]" captionTargetPageId="39" captionText="FIGURES 39–50. Male genitalia of Copelatus. 39, 41, 43, 45, 47, 49. median lobe in lateral view; 40, 42, 44, 46, 48, 50. paramere. 39–40. C. japonicus (Shanghai); 41. C. malaisei (a. Gaoligongshan, Yunnan; b. Renjiafen, Yunnan); 42. C. malaisei (Renjiafen, Yunnan); 43–44. C. miaowangi sp. nov. (paratype, Liupanshui, Guizhou); 45–46. C. mopanshanensis sp. nov. (holotype, Mopanshan, Yunnan); 47–48. C. nanlingensis sp. nov. (holotype, Dadongshan, Guangdong); 49–50. C. oblitus (Macao). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6412781" httpUri="https://zenodo.org/record/6412781/files/figure.png" pageId="9" pageNumber="260">Fig. 40</figureCitation>
|
||
). Female similar to male; strioles on pronotum more numerous, microreticulation of pronotum and elytra slightly more impressed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E9AF48D2BF54FF941D95FB219CDCFAED" pageId="9" pageNumber="260" type="discussion">
|
||
<paragraph id="A10A1B59BF54FF941D95FB219CDCFAED" blockId="9.[151,1437,577,1432]" pageId="9" pageNumber="260">
|
||
<emphasis id="93C1C74BBF54FF941D95FB219A79FB21" bold="true" box="[199,591,1189,1215]" pageId="9" pageNumber="260">Comments on the classification.</emphasis>
|
||
This Japanese species was recorded from several Chinese provinces by various authors (see
|
||
<bibRefCitation id="C52466A8BF54FF941CD4FB4D9A2BFB7A" author="Nilsson, A. N." box="[390,541,1225,1252]" pageId="9" pageNumber="260" pagination="35 - 96" refId="ref33005" refString="Nilsson, A. N. (1995) Noteridae and Dytiscidae: Annotated check list of the Noteridae and Dytiscidae of China (Coleoptera). In: Jach, M. A. & Ji, L. (Eds.), Water beetles of China. Vol. I. Zoologisch-Botanische Gesellschaft in Osterreich and Wiener Coleopterologenverein, Wien, pp. 35 - 96." type="book chapter" year="1995">Nilsson 1995</bibRefCitation>
|
||
for summary), also under the names
|
||
<taxonomicName id="66B560DABF54FF941E82FB4E9C6DFB7D" authorityName="Regimbart" authorityYear="1899" box="[976,1115,1225,1252]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="260" phylum="Arthropoda" rank="species" species="chinensis">
|
||
<emphasis id="93C1C74BBF54FF941E82FB4E9C6DFB7D" box="[976,1115,1225,1252]" italics="true" pageId="9" pageNumber="260">C. chinensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="66B560DABF54FF9419C5FB4E9D78FB7D" authorityName="Falkenstrom" authorityYear="1933" box="[1175,1358,1225,1252]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="260" phylum="Arthropoda" rank="species" species="collocallosus">
|
||
<emphasis id="93C1C74BBF54FF9419C5FB4E9D78FB7D" box="[1175,1358,1225,1252]" italics="true" pageId="9" pageNumber="260">C. collocallosus</emphasis>
|
||
</taxonomicName>
|
||
which were synonymized with
|
||
<taxonomicName id="66B560DABF54FF941CFCFB6A9A0BFA99" authorityName="Sharp" authorityYear="1884" box="[430,573,1262,1288]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="260" phylum="Arthropoda" rank="species" species="japonicus">
|
||
<emphasis id="93C1C74BBF54FF941CFCFB6A9A0BFA99" box="[430,573,1262,1288]" italics="true" pageId="9" pageNumber="260">C. japonicus</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="C52466A8BF54FF941F38FB699B2FFA96" author="Guignot, F." box="[618,793,1261,1288]" pageId="9" pageNumber="260" pagination="17 - 31" refId="ref32148" refString="Guignot, F. (1952) Description de dytiscides inedits de la collection Regimbart. Revue Francaise d'Entomologie, 19, 17 - 31." type="journal article" year="1952">Guignot (1952)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="C52466A8BF54FF941E01FB699BEEFA96" author="Sato, M." box="[851,984,1261,1288]" pageId="9" pageNumber="260" pagination="57 - 67" refId="ref33445" refString="Sato, M. (1985) The genus Copelatus of Japan (Coleoptera: Dytiscidae). Transactions of the Shikoku Entomological Society, 17, 57 - 67." type="journal article" year="1985">Satô (1985)</bibRefCitation>
|
||
respectively. However, the study of the
|
||
<typeStatus id="7E0EA5FBBF54FF941DC5FA9698F1FAB2" box="[151,199,1298,1324]" pageId="9" pageNumber="260">type</typeStatus>
|
||
material of
|
||
<taxonomicName id="66B560DABF54FF941C19FA9699E5FAB5" authorityName="Regimbart" authorityYear="1899" box="[331,467,1297,1324]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="260" phylum="Arthropoda" rank="species" species="chinensis">
|
||
<emphasis id="93C1C74BBF54FF941C19FA9699E5FAB5" box="[331,467,1297,1324]" italics="true" pageId="9" pageNumber="260">C. chinensis</emphasis>
|
||
</taxonomicName>
|
||
proved that it is a valid species separate from
|
||
<taxonomicName id="66B560DABF54FF941E9FFA969C6FFAB5" authorityName="Sharp" authorityYear="1884" box="[973,1113,1298,1324]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="260" phylum="Arthropoda" rank="species" species="japonicus">
|
||
<emphasis id="93C1C74BBF54FF941E9FFA969C6FFAB5" box="[973,1113,1298,1324]" italics="true" pageId="9" pageNumber="260">C. japonicus</emphasis>
|
||
</taxonomicName>
|
||
, while the extensive material collected recently in various localities in southern
|
||
<collectingCountry id="D9A25BC9BF54FF941F83FAB19B22FAD1" box="[721,788,1333,1359]" name="China" pageId="9" pageNumber="260">China</collectingCountry>
|
||
belongs to numerous taxa new to science. At present, we are aware of only a single reliable record of
|
||
<taxonomicName id="66B560DABF54FF941FD4FADE9B22FAED" authorityName="Sharp" authorityYear="1884" box="[646,788,1370,1396]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="260" phylum="Arthropoda" rank="species" species="japonicus">
|
||
<emphasis id="93C1C74BBF54FF941FD4FADE9B22FAED" box="[646,788,1370,1396]" italics="true" pageId="9" pageNumber="260">C. japonicus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="D9A25BC9BF54FF941E0AFADD9BADFAED" box="[856,923,1369,1395]" name="China" pageId="9" pageNumber="260">China</collectingCountry>
|
||
–
|
||
<collectingRegion id="6371D5BBBF54FF941EE5FADD9C14FAED" box="[951,1058,1369,1395]" country="China" name="Shanghai" pageId="9" pageNumber="260">Shanghai</collectingRegion>
|
||
, presented above.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E9AF48D2BF54FF941D95FAF99D34FA09" box="[199,1282,1405,1432]" pageId="9" pageNumber="260" type="distribution">
|
||
<paragraph id="A10A1B59BF54FF941D95FAF99D34FA09" blockId="9.[151,1437,577,1432]" box="[199,1282,1405,1432]" pageId="9" pageNumber="260">
|
||
<emphasis id="93C1C74BBF54FF941D95FAF99957FA09" bold="true" box="[199,353,1405,1431]" pageId="9" pageNumber="260">Distribution.</emphasis>
|
||
A species occurring in
|
||
<collectingCountry id="D9A25BC9BF54FF941F35FAFA9A9CFA06" box="[615,682,1406,1432]" name="Japan" pageId="9" pageNumber="260">Japan</collectingCountry>
|
||
,
|
||
<collectingCountry id="D9A25BC9BF54FF941FE6FAF99B77FA06" box="[692,833,1405,1432]" name="South Korea" pageId="9" pageNumber="260">South Korea</collectingCountry>
|
||
and eastern
|
||
<collectingCountry id="D9A25BC9BF54FF941E9CFAF99C27FA09" box="[974,1041,1405,1431]" name="China" pageId="9" pageNumber="260">China</collectingCountry>
|
||
(
|
||
<collectingRegion id="6371D5BBBF54FF94194DFAF99CBCFA06" box="[1055,1162,1405,1432]" country="China" name="Shanghai" pageId="9" pageNumber="260">Shanghai</collectingRegion>
|
||
) (
|
||
<figureCitation id="398E07DCBF54FF9419F0FAF99CC3FA09" box="[1186,1269,1405,1431]" captionStart="FIGURE 81" captionStartId="43.[151,250,2008,2032]" captionTargetBox="[229,1358,1062,1978]" captionTargetId="figure-47@43.[224,1364,1054,1983]" captionTargetPageId="43" captionText="FIGURE 81. Map of distribution of Copelatus in China Mainland." figureDoi="http://doi.org/10.5281/zenodo.6411120" httpUri="https://zenodo.org/record/6411120/files/figure.png" pageId="9" pageNumber="260">Fig. 81</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |