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<document ID-DOI="10.11646/phytotaxa.581.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7571550" approvalRequired="50" approvalRequired_for_taxonomicNames="15" approvalRequired_for_textStreams="30" approvalRequired_for_treatments="5" checkinTime="1674712547726" checkinUser="plazi" docAuthor="Dang, Van-Son &amp; Naiki, Akiyo" docDate="2023" docId="43284262FFA9FFE85AB6FE47441BC828" docLanguage="en" docName="phytotaxa.581.1.1.pdf" docOrigin="Phytotaxa 581 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.1:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="1" docTitle="Lasianthus eberhardtii Pit. 1924" docType="treatment" docVersion="2" lastPageNumber="1935" masterDocId="BF113A1AFF9FFFDE5A3EFFA54026CC48" masterDocTitle="A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam" masterLastPageNumber="1935" masterPageNumber="1921" pageNumber="1921" updateTime="1674713874756" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dang, Van-Son</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8681-4141</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. &amp; dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141</mods:affiliation>
<mods:nameIdentifier type="email">dvsonitb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; naiki @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0002 - 3235 - 2823 * Author for correspondence</mods:affiliation>
<mods:nameIdentifier type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>581</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1921</mods:start>
<mods:end>1935</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.581.1.1</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7571550</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:43284262FFA9FFE85AB6FE47441BC828" httpUri="http://treatment.plazi.org/id/43284262FFA9FFE85AB6FE47441BC828" pageId="54">
<subSubSection box="[136,980,482,509]" pageId="54" type="nomenclature">
<paragraph blockId="54.[136,1452,482,580]" box="[136,980,482,509]" pageId="54">
<emphasis bold="true" box="[136,171,482,508]" pageId="54">19.</emphasis>
<taxonomicName ID-CoL="3SCKJ" authority="Pit. (1924: 375)" authorityName="Pit." authorityPageNumber="375" authorityYear="1924" box="[178,625,482,509]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="54" phylum="Tracheophyta" rank="species" species="eberhardtii">
<emphasis bold="true" box="[178,442,482,509]" italics="true" pageId="54">Lasianthus eberhardtii</emphasis>
Pit. (1924: 375)
</taxonomicName>
<figureCitation box="[655,733,482,509]" captionStart="FIGURE 39" captionStartId="55.[136,229,1778,1800]" captionTargetBox="[168,1414,194,1748]" captionTargetId="figure-18@55.[165,1418,193,1754]" captionTargetPageId="55" captionText="FIGURE 39. Lasianthus eberhardtii Pit. (AD &amp; HI from TH16-10 and EG from Eberhardt 4646, VAFS and P): A. Habit, B. Branch, C. Close-up of young branch and leaves, D. Stipules, E. Abaxial leaf surfaces, FG. Inflorescence showing side view flowers and bracts, HI. Young fruits (AD &amp; HI: photos by Trinh Ngoc Bon)." figureDoi="http://doi.org/10.5281/zenodo.7571670" httpUri="https://zenodo.org/record/7571670/files/figure.png" pageId="54">Fig. 39</figureCitation>
(plate),
<figureCitation box="[827,906,482,508]" captionStart="FIGURE 37" captionStartId="52.[136,229,1789,1811]" captionTargetBox="[176,1405,205,1753]" captionTargetId="figure-25@52.[165,1418,193,1765]" captionTargetPageId="52" captionText="FIGURE 37. Distribution maps of Lasianthus dalatensis, L. dienkhanhensis, L. eberhartii and L. elevatineurus." figureDoi="http://doi.org/10.5281/zenodo.7571662" httpUri="https://zenodo.org/record/7571662/files/figure.png" pageId="54">Fig. 37</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection pageId="54" type="materials_examined">
<paragraph blockId="54.[136,1452,482,580]" pageId="54">
<emphasis bold="true" box="[136,194,522,543]" pageId="54">Type:</emphasis>
<materialsCitation country="Vietnam" location="Eberhardt" municipality="Ban Khuang" pageId="54" specimenCount="1" stateProvince="Bac Kan" typeStatus="holotype">
<collectingCountry box="[219,332,522,543]" name="Vietnam" pageId="54">VIETNAM</collectingCountry>
.
<collectingRegion box="[351,549,522,544]" country="Vietnam" name="Bac Kan" pageId="54">Bac Kan Province</collectingRegion>
,
<collectingMunicipality box="[569,702,522,544]" pageId="54">Ban Khuang</collectingMunicipality>
,
<emphasis box="[723,884,522,543]" italics="true" pageId="54">
<location LSID="urn:lsid:plazi:treatment:43284262FFA9FFE85AB6FE47441BC828:CE5EA5AFFFA9FFE858EDFDAF4312CE57" box="[723,820,522,543]" country="Vietnam" municipality="Ban Khuang" name="Eberhardt" pageId="54" stateProvince="Bac Kan">Eberhardt</location>
4646
</emphasis>
(
<typeStatus box="[907,993,522,544]" pageId="54">holotype</typeStatus>
P! [P02285249]; isotypes P! [P02285249, P02285249])
</materialsCitation>
.
</paragraph>
<paragraph blockId="54.[136,1452,626,1121]" pageId="54">
<emphasis bold="true" box="[136,394,626,652]" pageId="54">
<collectingRegion box="[136,262,626,652]" pageId="54">Specimens</collectingRegion>
examined:
</emphasis>
<materialsCitation collectingDate="2016-04-29" collectorName="Naiki A. &amp; Yahara T. &amp; Toyama H. &amp; Tagane S. &amp; Nagamasu H. &amp; Nguyen" country="Vietnam" elevation="870" location="Hoang Lien National Park" pageId="54" specimenCount="1" stateProvince="Lao Cai">
<collectingCountry box="[423,555,626,652]" name="Vietnam" pageId="54">VIETNAM</collectingCountry>
.
<emphasis bold="true" box="[565,686,626,652]" pageId="54">Northern:</emphasis>
<collectingRegion box="[694,893,626,653]" country="Vietnam" name="Lao Cai" pageId="54">Lao Cai Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFA9FFE85AB6FE47441BC828:CE5EA5AFFFA9FFE859B6FDD64495CEC4" box="[904,1203,626,653]" country="Vietnam" name="Hoang Lien National Park" pageId="54" stateProvince="Lao Cai">Hoang Lien National Park</location>
,
<quantity box="[1214,1285,626,653]" metricMagnitude="2" metricUnit="m" metricValue="8.7" pageId="54" unit="m" value="870.0">
<elevation box="[1214,1285,626,653]" metricMagnitude="2" metricUnit="m" metricValue="8.7" pageId="54" unit="m" value="870.0">870 m</elevation>
</quantity>
elev.,
<date pageId="54" value="2016-04-29">
<collectingDate pageId="54" value="2016-04-29">29 April 2016</collectingDate>
</date>
,
<emphasis box="[206,940,662,689]" italics="true" pageId="54">
<collectorName box="[206,298,662,689]" pageId="54">Naiki A.</collectorName>
,
<collectorName box="[310,419,662,688]" pageId="54">Yahara T.</collectorName>
,
<collectorName box="[430,554,663,688]" pageId="54">Toyama H.</collectorName>
,
<collectorName box="[565,675,663,689]" pageId="54">Tagane S.</collectorName>
,
<collectorName box="[687,843,663,688]" pageId="54">Nagamasu H.</collectorName>
,
<collectorName box="[854,940,663,688]" pageId="54">Nguyen</collectorName>
</emphasis>
</materialsCitation>
<materialsCitation collectionCode="DLU, FU" collectorName="Hoang T. B. &amp; Hoang T. S. V" country="Vietnam" location="V. N." pageId="54" specimenCount="1" stateProvince="Northern">
<emphasis box="[946,1371,662,689]" italics="true" pageId="54">
<taxonomicNameLabel box="[946,996,663,688]" pageId="54" rank="variety">
<collectingCountry box="[946,996,663,688]" name="Vietnam" pageId="54">V.N.</collectingCountry>
</taxonomicNameLabel>
,
<collectorName box="[1006,1136,663,688]" pageId="54">Hoang T.B.</collectorName>
,
<collectorName box="[1147,1294,662,689]" pageId="54">Hoang T.S. V</collectorName>
4669?
</emphasis>
(
<collectionCode box="[1385,1447,662,688]" pageId="54">DLU</collectionCode>
,
<collectionCode box="[136,176,698,724]" country="China" name="Fudan University, Department of Biology" pageId="54">FU</collectionCode>
)
</materialsCitation>
.
<materialsCitation box="[195,1203,698,725]" collectingDate="2016-06-17" collectionCode="VAFS" collectorName="Trinh N. B." country="Vietnam" location="Thuong Xuan" pageId="54" specimenCount="1" stateProvince="Thanh Hoa">
<emphasis bold="true" box="[195,296,698,724]" pageId="54">Central:</emphasis>
<collectingRegion box="[303,537,698,725]" country="Vietnam" name="Thanh Hoa" pageId="54">Thanh Hoa Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFA9FFE85AB6FE47441BC828:CE5EA5AFFFA9FFE8581AFD1F42E7CE9D" box="[548,705,698,725]" country="Vietnam" name="Thuong Xuan" pageId="54" stateProvince="Thanh Hoa">Thuong Xuan</location>
,
<date box="[717,869,698,725]" pageId="54" value="2016-06-17">
<collectingDate box="[717,869,698,725]" pageId="54" value="2016-06-17">17 June 2016</collectingDate>
</date>
,
<emphasis box="[880,1106,698,724]" italics="true" pageId="54">
<collectorName box="[880,998,698,724]" pageId="54">Trinh N.B.</collectorName>
TH16-10
</emphasis>
(
<collectionCode box="[1120,1195,698,724]" pageId="54">VAFS</collectionCode>
)
</materialsCitation>
;
</paragraph>
</subSubSection>
<subSubSection pageId="54" type="distribution">
<paragraph blockId="54.[136,1452,626,1121]" box="[189,1404,734,761]" pageId="54">
<emphasis bold="true" box="[189,345,734,760]" pageId="54">Distribution:</emphasis>
—Endemic to
<collectingCountry box="[508,603,734,760]" name="Vietnam" pageId="54">Vietnam</collectingCountry>
(currently known as
<collectingRegion box="[840,940,735,761]" country="Vietnam" name="Bac Kan" pageId="54">Bac Kan</collectingRegion>
,
<collectingRegion box="[952,1044,734,761]" country="Vietnam" name="Lao Cai" pageId="54">Lao Cai</collectingRegion>
,
<collectingRegion box="[1055,1204,734,761]" country="Vietnam" name="Thai Nguyen" pageId="54">Thai Nguyen</collectingRegion>
, and
<collectingRegion box="[1262,1392,734,760]" country="Vietnam" name="Thanh Hoa" pageId="54">Thanh Hoa</collectingRegion>
).
</paragraph>
<paragraph blockId="54.[136,1452,626,1121]" pageId="54">
<emphasis bold="true" box="[189,422,770,796]" pageId="54">Habitat &amp; Ecology:</emphasis>
—Growing in open shade, on less sandy ground in primary or secondary forest at
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.5" metricValueMin="0.7" pageId="54" unit="m" value="1100.0" valueMax="1500.0" valueMin="700.0">7001500 m</quantity>
elev.
</paragraph>
<paragraph blockId="54.[136,1452,626,1121]" box="[189,885,842,869]" pageId="54">
<emphasis bold="true" box="[189,323,842,868]" pageId="54">Phenology:</emphasis>
—Flowering and fruiting from June to November.
</paragraph>
<paragraph blockId="54.[136,1452,626,1121]" box="[189,667,878,907]" pageId="54">
<emphasis bold="true" box="[189,405,878,904]" pageId="54">Vernacular name:</emphasis>
—Xú hưƠng eberhardt.
</paragraph>
<paragraph blockId="54.[136,1452,626,1121]" pageId="54">
<emphasis bold="true" box="[189,265,914,940]" pageId="54">Notes:</emphasis>
<taxonomicName box="[294,552,914,940]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="54" phylum="Tracheophyta" rank="species" species="eberhardtii">
<emphasis box="[294,552,914,940]" italics="true" pageId="54">Lasianthus eberhardtii</emphasis>
</taxonomicName>
was described in 1924 from a Vietnamese specimen (
<emphasis box="[1194,1375,914,940]" italics="true" pageId="54">Eberhardt 4646</emphasis>
). This species can easily be recognized in the field in having yellow hirsute branches, oblong leaf blade, tip pointed
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="54" unit="mm" value="13.5" valueMax="15.0" valueMin="12.0">1215 mm</quantity>
long, secondary veins 912 on each side of midrib, bracts
<quantity box="[838,959,986,1013]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="54" unit="mm" value="15.0" valueMax="20.0" valueMin="10.0">1020 mm</quantity>
long, and fruits
<quantity box="[1143,1208,986,1013]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="54" unit="mm" value="4.0">4 mm</quantity>
long, pyrenes 6.
</paragraph>
<paragraph blockId="54.[136,1452,626,1121]" pageId="54">
<taxonomicName box="[189,443,1022,1048]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="54" phylum="Tracheophyta" rank="species" species="eberhardtii">
<emphasis box="[189,443,1022,1048]" italics="true" pageId="54">Lasianthus eberhardtii</emphasis>
</taxonomicName>
is closely related to
<taxonomicName authority="Pierre ex Pit. (1924: 376)" authorityName="Pit." authorityPageNumber="376" authorityYear="1924" box="[673,1153,1022,1049]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="54" phylum="Tracheophyta" rank="species" species="harmandianus">
<emphasis box="[673,864,1022,1048]" italics="true" pageId="54">L. harmandianus</emphasis>
Pierre ex Pit. (1924: 376)
</taxonomicName>
which is endemic to
<collectingCountry box="[1390,1447,1023,1049]" name="Laos" pageId="54">Laos</collectingCountry>
, but differs from the latter by having leaves long hairy along margin (vs. smooth margin), bracts
<quantity box="[1223,1345,1058,1085]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="54" unit="mm" value="15.0" valueMax="20.0" valueMin="10.0">1020 mm</quantity>
long (vs.
<quantity box="[136,222,1094,1121]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="54" unit="mm" value="6.5">6.5 mm</quantity>
long), flowers grouped by 14 (vs. 812) and calyx 6, erect (vs. 5, reflexed).
</paragraph>
</subSubSection>
</treatment>
</document>