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<document ID-DOI="10.11646/phytotaxa.581.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7571550" approvalRequired="50" approvalRequired_for_taxonomicNames="15" approvalRequired_for_textStreams="30" approvalRequired_for_treatments="5" checkinTime="1674712547726" checkinUser="plazi" docAuthor="Dang, Van-Son &amp; Naiki, Akiyo" docDate="2023" docId="43284262FFA1FFE15AB6FF3B458ECEF8" docLanguage="en" docName="phytotaxa.581.1.1.pdf" docOrigin="Phytotaxa 581 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.1:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="1" docTitle="Lasianthus formosensis Matsum. 1901" docType="treatment" docVersion="2" lastPageNumber="1935" masterDocId="BF113A1AFF9FFFDE5A3EFFA54026CC48" masterDocTitle="A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam" masterLastPageNumber="1935" masterPageNumber="1921" pageNumber="1921" updateTime="1674713874756" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dang, Van-Son</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8681-4141</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. &amp; dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141</mods:affiliation>
<mods:nameIdentifier type="email">dvsonitb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; naiki @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0002 - 3235 - 2823 * Author for correspondence</mods:affiliation>
<mods:nameIdentifier type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>581</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1921</mods:start>
<mods:end>1935</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.581.1.1</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7571550</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7571668" ID-Zenodo-Dep="7571668" LSID="urn:lsid:plazi:treatment:43284262FFA1FFE15AB6FF3B458ECEF8" httpUri="http://treatment.plazi.org/id/43284262FFA1FFE15AB6FF3B458ECEF8" lastPageId="63" pageId="62">
<subSubSection box="[136,1038,158,185]" pageId="62" type="nomenclature">
<paragraph blockId="62.[136,1451,158,328]" box="[136,1038,158,185]" pageId="62">
<emphasis bold="true" box="[136,171,158,184]" pageId="62">24.</emphasis>
<taxonomicName ID-CoL="3SCL9" ID-ENA="1406744" authority="Matsum. (1901: 17)" authorityName="Matsum." authorityPageNumber="17" authorityYear="1901" box="[178,683,158,185]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="62" phylum="Tracheophyta" rank="species" species="formosensis">
<emphasis bold="true" box="[178,452,158,185]" italics="true" pageId="62">Lasianthus formosensis</emphasis>
Matsum. (1901: 17)
</taxonomicName>
<emphasis bold="true" box="[683,711,158,184]" pageId="62">—</emphasis>
<figureCitation box="[711,790,158,185]" captionStart="FIGURE 45" captionStartId="62.[136,229,1970,1992]" captionTargetBox="[168,1414,393,1941]" captionTargetId="figure-83@62.[165,1418,392,1945]" captionTargetPageId="62" captionText="FIGURE 45. Lasianthus formosensis Matsum.(A, G &amp; H–J from Dang 282 and B–F from Dang 280, VNM and the herbarium of Iriomote Station (Japan)): A. Habit, B. Branch with flower buds, C. Stipules, D. Abaxial leaf surfaces, E–G. Inflorescence showing side view buds and flowers, H. young fruits, I–J. Mature fruits." figureDoi="http://doi.org/10.5281/zenodo.7571690" httpUri="https://zenodo.org/record/7571690/files/figure.png" pageId="62">Fig. 45</figureCitation>
(plate),
<figureCitation box="[884,964,158,184]" captionStart="FIGURE 42" captionStartId="58.[136,229,1789,1811]" captionTargetBox="[173,1409,203,1757]" captionTargetId="figure-25@58.[165,1419,193,1765]" captionTargetPageId="58" captionText="FIGURE 42. Distribution maps of Lasianthus fansipanensis, L. foetidissimus, L. fordii and L. formosensis." figureDoi="http://doi.org/10.5281/zenodo.7571678" httpUri="https://zenodo.org/record/7571678/files/figure.png" pageId="62">Fig. 42</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection lastPageId="63" pageId="62" type="materials_examined">
<paragraph blockId="62.[136,1451,158,328]" box="[136,798,198,220]" pageId="62">
<emphasis bold="true" box="[136,194,198,219]" pageId="62">Type:</emphasis>
—
<materialsCitation box="[219,794,198,220]" collectorName="Taiwan, Y &amp; S. Tanaka" country="China" location="CHINA" pageId="62" specimenCount="1" typeStatus="holotype">
<collectingCountry box="[219,297,198,219]" name="China" pageId="62">CHINA</collectingCountry>
.
<collectorName box="[306,402,198,220]" pageId="62">
Taiwan,
<emphasis box="[388,402,198,219]" italics="true" pageId="62">Y</emphasis>
</collectorName>
. Tashiro &amp;
<collectorName box="[514,608,198,219]" pageId="62">S. Tanaka</collectorName>
131 (
<typeStatus box="[662,748,198,220]" pageId="62">holotype</typeStatus>
TI!)
</materialsCitation>
.
</paragraph>
<paragraph blockId="62.[136,1451,158,328]" box="[136,1164,234,256]" pageId="62">
<taxonomicName authority="Merr. (1931: 324)" authorityName="Merr." authorityPageNumber="324" authorityYear="1931" box="[136,562,234,255]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="62" phylum="Tracheophyta" rank="species" species="tenuicaudatus">
<emphasis box="[136,382,234,255]" italics="true" pageId="62">Lasianthus tenuicaudatus</emphasis>
<materialsCitation box="[388,562,234,255]" collectionCode="IBSC" collectorName="Y. Tsiang" country="China" location="Guangdong" pageId="62" specimenCount="1" stateProvince="Guangdong" typeStatus="holotype">Merr. (1931: 324)</materialsCitation>
</taxonomicName>
. Type:—
<collectingCountry box="[650,729,234,255]" name="China" pageId="62">CHINA</collectingCountry>
.
<collectingRegion box="[737,851,234,256]" country="China" name="Guangdong" pageId="62">Guangdong</collectingRegion>
,
<emphasis box="[861,984,234,255]" italics="true" pageId="62">
<collectorName box="[861,947,234,255]" pageId="62">Y. Tsiang</collectorName>
822
</emphasis>
(
<typeStatus box="[996,1082,234,256]" pageId="62">holotype</typeStatus>
<collectionCode box="[1088,1150,234,255]" country="China" lsid="urn:lsid:biocol.org:col:15606" name="South China Botanical Garden" pageId="62" type="Herbarium">IBSC</collectionCode>
!).
</paragraph>
<paragraph blockId="62.[136,1451,158,328]" pageId="62">
<taxonomicName authority="Merr. ex H. L. Li (1943: 458)" authorityName="Merr. ex H. L. Li" authorityPageNumber="458" authorityYear="1943" box="[136,655,270,292]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="62" phylum="Tracheophyta" rank="species" species="kwangsiensis">
<emphasis box="[136,374,270,291]" italics="true" pageId="62">Lasianthus kwangsiensis</emphasis>
<materialsCitation box="[380,655,270,292]" collectingDate="1924-12-12" country="Thailand" location="Khao Soi Dao" municipality="Type" pageId="62" specimenCount="1" stateProvince="Chanthaburi" typeStatus="holotype">Merr. ex H.L.Li (1943: 458)</materialsCitation>
</taxonomicName>
.
<collectingMunicipality box="[663,713,270,291]" pageId="62">Type</collectingMunicipality>
:—
<collectingCountry box="[741,869,270,291]" name="Thailand" pageId="62">THAILAND</collectingCountry>
.
<collectingRegion box="[878,999,270,292]" country="Thailand" name="Chanthaburi" pageId="62">Chanthaburi</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFA1FFE15AB6FF3B458ECEF8:CE5EA5AFFFA1FFE059CFFEAB445DCD6B" box="[1009,1147,270,292]" country="Thailand" municipality="Type" name="Khao Soi Dao" pageId="62" stateProvince="Chanthaburi">Khao Soi Dao</location>
,
<date box="[1157,1342,270,292]" pageId="62" value="1924-12-12">
<collectingDate box="[1157,1342,270,292]" pageId="62" value="1924-12-12">12 December 1924</collectingDate>
</date>
,
<emphasis box="[1352,1451,270,291]" italics="true" pageId="62">Kerr 9618</emphasis>
(
<typeStatus box="[196,282,306,328]" pageId="62">holotype</typeStatus>
K! [K000777028]; isotype E! [E00327853]).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7571690" ID-Zenodo-Dep="7571690" httpUri="https://zenodo.org/record/7571690/files/figure.png" pageId="62" startId="62.[136,229,1970,1992]" targetBox="[168,1414,393,1941]" targetPageId="62">
<paragraph blockId="62.[136,1452,1970,2064]" pageId="62">
<emphasis bold="true" box="[136,265,1970,1992]" pageId="62">FIGURE 45.</emphasis>
<taxonomicName authority="Matsum." authorityName="Matsum." box="[269,585,1970,1992]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="62" phylum="Tracheophyta" rank="species" species="formosensis">
<emphasis box="[269,494,1970,1991]" italics="true" pageId="62">Lasianthus formosensis</emphasis>
Matsum.
</taxonomicName>
(A, G &amp; H–J from
<emphasis box="[765,859,1971,1992]" italics="true" pageId="62">Dang 282</emphasis>
and B–F from
<emphasis box="[1001,1096,1971,1992]" italics="true" pageId="62">Dang 280</emphasis>
, VNM and the herbarium of Iriomote Station (Japan)):
<emphasis bold="true" box="[301,324,2006,2027]" pageId="62">A.</emphasis>
Habit,
<emphasis bold="true" box="[394,417,2007,2028]" pageId="62">B.</emphasis>
Branch with flower buds,
<emphasis bold="true" box="[670,694,2006,2028]" pageId="62">C.</emphasis>
Stipules,
<emphasis bold="true" box="[788,812,2007,2028]" pageId="62">D.</emphasis>
Abaxial leaf surfaces,
<emphasis bold="true" box="[1029,1082,2006,2028]" pageId="62">E–G.</emphasis>
Inflorescence showing side view buds and flowers,
<emphasis bold="true" box="[261,286,2043,2064]" pageId="62">H.</emphasis>
young fruits,
<emphasis bold="true" box="[420,459,2043,2064]" pageId="62">I–J.</emphasis>
Mature fruits.
</paragraph>
</caption>
<paragraph blockId="63.[136,1452,158,688]" pageId="63">
<emphasis bold="true" box="[136,393,158,184]" pageId="63">
<collectingRegion box="[136,262,158,184]" pageId="63">Specimens</collectingRegion>
examined:
</emphasis>
—
<materialsCitation collectingDate="2018-11-17" collectingDateMax="2018-11-18" collectingDateMin="2018-11-17" collectionCode="VNM" collectorName="Dang V. S. &amp; Naiki A. &amp; Dang &amp; Japan" country="Vietnam" county="City" elevation="1267" location="Iriomote Station" municipality="Ba Vi National Park" pageId="63" specimenCount="1" stateProvince="Ha Noi">
<collectingCountry box="[422,554,158,184]" name="Vietnam" pageId="63">VIETNAM</collectingCountry>
.
<emphasis bold="true" box="[563,684,158,184]" pageId="63">Northern:</emphasis>
<collectingRegion box="[691,772,158,185]" country="Vietnam" name="Ha Noi" pageId="63">Ha Noi</collectingRegion>
<collectingCounty box="[778,827,158,184]" pageId="63">City</collectingCounty>
,
<collectingMunicipality box="[838,1064,158,185]" pageId="63">Ba Vi National Park</collectingMunicipality>
,
<quantity box="[1074,1159,158,185]" metricMagnitude="3" metricUnit="m" metricValue="1.267" pageId="63" unit="m" value="1267.0">
<elevation box="[1074,1159,158,185]" metricMagnitude="3" metricUnit="m" metricValue="1.267" pageId="63" unit="m" value="1267.0">1267 m</elevation>
</quantity>
elev.,
<date box="[1230,1447,158,184]" pageId="63" value="2018-11-17">
<collectingDate box="[1230,1447,158,184]" pageId="63" value="2018-11-17">17 November 2018</collectingDate>
</date>
,
<emphasis box="[136,492,194,221]" italics="true" pageId="63">
<collectorName box="[136,246,195,221]" pageId="63">Dang V.S.</collectorName>
&amp;
<collectorName box="[279,371,194,221]" pageId="63">Naiki A.</collectorName>
,
<collectorName box="[382,444,195,220]" pageId="63">Dang</collectorName>
280
</emphasis>
(
<collectionCode box="[506,576,194,220]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="63" type="Herbarium">VNM</collectionCode>
, the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FFA1FFE15AB6FF3B458ECEF8:CE5EA5AFFFA0FFE15936FF67439BCC94" box="[776,957,194,220]" country="Vietnam" county="City" municipality="Ba Vi National Park" name="Iriomote Station" pageId="63" stateProvince="Ha Noi">Iriomote Station</location>
(Japan)); in the same location,
<quantity box="[1303,1387,194,221]" metricMagnitude="3" metricUnit="m" metricValue="1.268" pageId="63" unit="m" value="1268.0">
<elevation box="[1303,1387,194,221]" metricMagnitude="3" metricUnit="m" metricValue="1.268" pageId="63" unit="m" value="1268.0">1268 m</elevation>
</quantity>
elev.,
<date box="[136,357,230,256]" pageId="63" value="2018-11-17">
<collectingDate box="[136,357,230,256]" pageId="63" value="2018-11-17">17 November 2018</collectingDate>
</date>
,
<emphasis box="[369,739,230,257]" italics="true" pageId="63">
<collectorName box="[369,482,231,257]" pageId="63">Dang V.S.</collectorName>
&amp;
<collectorName box="[519,613,230,257]" pageId="63">Naiki A.</collectorName>
, Dang 282
</emphasis>
(
<collectionCode box="[755,825,230,256]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="63" type="Herbarium">VNM</collectionCode>
, the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FFA1FFE15AB6FF3B458ECEF8:CE5EA5AFFFA0FFE15E37FF4344E7CD48" box="[1033,1217,230,256]" country="Vietnam" county="City" municipality="Ba Vi National Park" name="Iriomote Station" pageId="63" stateProvince="Ha Noi">Iriomote Station</location>
(Japan));
<collectingRegion country="Vietnam" name="Ninh Binh" pageId="63">Ninh Binh Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFA1FFE15AB6FF3B458ECEF8:CE5EA5AFFFA0FFE15AC5FEAF420ACD6C" box="[251,556,266,292]" country="Vietnam" county="City" municipality="Ba Vi National Park" name="Cuc Phuong National Park" pageId="63" stateProvince="Ha Noi">Cuc Phuong National Park</location>
,
<quantity box="[569,641,266,293]" metricMagnitude="2" metricUnit="m" metricValue="3.65" pageId="63" unit="m" value="365.0">
<elevation box="[569,641,266,293]" metricMagnitude="2" metricUnit="m" metricValue="3.65" pageId="63" unit="m" value="365.0">365 m</elevation>
</quantity>
elev.,
<date box="[716,937,266,292]" pageId="63" value="2018-11-18">
<collectingDate box="[716,937,266,292]" pageId="63" value="2018-11-18">18 November 2018</collectingDate>
</date>
,
<emphasis box="[949,1319,266,293]" italics="true" pageId="63">
<collectorName box="[949,1062,267,293]" pageId="63">Dang V.S.</collectorName>
&amp;
<collectorName box="[1099,1194,266,293]" pageId="63">Naiki A.</collectorName>
, Dang 287
</emphasis>
(
<collectionCode box="[1335,1405,266,292]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="63" type="Herbarium">VNM</collectionCode>
, the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FFA1FFE15AB6FF3B458ECEF8:CE5EA5AFFFA0FFE15B1FFE8B41FECD00" box="[289,472,302,328]" country="Vietnam" county="City" municipality="Ba Vi National Park" name="Iriomote Station" pageId="63" stateProvince="Ha Noi">Iriomote Station</location>
(Japan)); in the same location,
<quantity box="[828,899,302,329]" metricMagnitude="2" metricUnit="m" metricValue="3.6799999999999997" pageId="63" unit="m" value="368.0">
<elevation box="[828,899,302,329]" metricMagnitude="2" metricUnit="m" metricValue="3.6799999999999997" pageId="63" unit="m" value="368.0">368 m</elevation>
</quantity>
elev.,
<date box="[973,1193,302,328]" pageId="63" value="2018-11-18">
<collectingDate box="[973,1193,302,328]" pageId="63" value="2018-11-18">18 November 2018</collectingDate>
</date>
,
<emphasis italics="true" pageId="63">
<collectorName box="[1205,1316,303,329]" pageId="63">Dang V.S.</collectorName>
&amp;
<collectorName box="[1353,1446,302,329]" pageId="63">Naiki A.</collectorName>
,
<collectorName box="[136,198,339,364]" pageId="63">Dang</collectorName>
288
</emphasis>
(
<collectionCode box="[262,332,338,364]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="63" type="Herbarium">VNM</collectionCode>
, the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FFA1FFE15AB6FF3B458ECEF8:CE5EA5AFFFA0FFE15827FEF742E9CD24" box="[537,719,338,364]" country="Vietnam" county="City" municipality="Ba Vi National Park" name="Iriomote Station" pageId="63" stateProvince="Ha Noi">Iriomote Station</location>
(
<collectorName box="[734,803,338,364]" pageId="63">Japan</collectorName>
))
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="63" type="distribution">
<paragraph blockId="63.[136,1452,158,688]" pageId="63">
<emphasis bold="true" box="[189,345,374,400]" pageId="63">Distribution:</emphasis>
—Widespread in tropical and subtropical regions in Asia (
<collectingCountry box="[1023,1094,374,400]" name="China" pageId="63">China</collectingCountry>
,
<collectingCountry box="[1109,1176,375,401]" name="Japan" pageId="63">Japan</collectingCountry>
,
<collectingCountry box="[1190,1295,374,400]" name="Thailand" pageId="63">Thailand</collectingCountry>
). In
<collectingCountry box="[1350,1448,374,400]" name="Vietnam" pageId="63">Vietnam</collectingCountry>
,
<taxonomicName box="[136,400,410,436]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="63" phylum="Tracheophyta" rank="species" species="formosensis">
<emphasis box="[136,400,410,436]" italics="true" pageId="63">Lasianthus formosensis</emphasis>
</taxonomicName>
occurs in
<collectingRegion box="[516,600,410,437]" country="Vietnam" name="Ha Noi" pageId="63">Ha Noi</collectingRegion>
,
<collectingRegion box="[611,705,410,437]" country="Vietnam" name="Ha Tinh" pageId="63">Ha Tinh</collectingRegion>
,
<collectingRegion box="[716,833,410,436]" country="Vietnam" name="Ninh Binh" pageId="63">Ninh Binh</collectingRegion>
and Thua Thien Hue.
</paragraph>
<paragraph blockId="63.[136,1452,158,688]" pageId="63">
<emphasis bold="true" box="[189,420,446,472]" pageId="63">Habitat &amp; Ecology:</emphasis>
—Growing in secondary montane forest, the edge of the forest, and an open area at
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.0" metricValueMax="13.0" metricValueMin="3.0" pageId="63" unit="m" value="800.0" valueMax="1300.0" valueMin="300.0">300–1300 m</quantity>
elev.
</paragraph>
<paragraph blockId="63.[136,1452,158,688]" box="[189,882,518,545]" pageId="63">
<emphasis bold="true" box="[189,323,518,544]" pageId="63">Phenology:</emphasis>
—Flowering and fruiting from June to December.
</paragraph>
<paragraph blockId="63.[136,1452,158,688]" box="[189,670,554,583]" pageId="63">
<emphasis bold="true" box="[189,405,554,580]" pageId="63">Vernacular name:</emphasis>
—Xú hưƠng nhiễu hoa.
</paragraph>
<paragraph blockId="63.[136,1452,158,688]" pageId="63">
<emphasis bold="true" box="[189,265,590,616]" pageId="63">Notes:</emphasis>
—
<taxonomicName box="[294,561,590,616]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="63" phylum="Tracheophyta" rank="species" species="formosensis">
<emphasis box="[294,561,590,616]" italics="true" pageId="63">Lasianthus formosensis</emphasis>
</taxonomicName>
was clarified in the revisions by
<bibRefCitation author="Zhu, H." box="[952,1082,590,616]" pageId="63" pagination="116 - 150" refId="ref55444" refString="Zhu, H. (2001) A taxonomic revision of the genus Lasianthus Jack (Rubiaceae) from Thailand. Acta Phytotaxonomica Sinica 39: 116 - 150." type="journal article" year="2001">Zhu (2001)</bibRefCitation>
and
<bibRefCitation author="Zhu, H. &amp; Taylor, C. M." box="[1142,1388,590,616]" pageId="63" pagination="185 - 198" refId="ref55538" refString="Zhu, H. &amp; Taylor, C. M. (2011) Lasianthus. In: Wu, Z. Y., Raven, P. H. &amp; Hong, D. Y. (Eds.) Flora of China, vol. 19. Science Press, Beijing &amp; Missouri Botanical Garden Press, St. Louis, pp. 185 - 198." type="book chapter" year="2011">Zhu &amp; Taylor (2011)</bibRefCitation>
. This species can be recognized in the field in having oblong to ovate-elliptic leaves, secondary veins 6–7 on each side of midrib, flowers grouped by 6–9, lanceolate calyx lobes longer than the tube, and fruit ovoid-globose, blue when ripe.
</paragraph>
</subSubSection>
</treatment>
</document>
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