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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1041.64460" ID-GBIF-Dataset="5dba2b2e-dcd9-43f5-92e5-ac318c319307" ID-PMC="PMC8184736" ID-Pensoft-Pub="1313-2970-1041-27" ID-Pensoft-UUID="5AE03537388755CFAF1E06C3CC9EFA72" ID-PubMed="34140825" ID-ZooBank="EEE8490BB41D4A6CA963234C256C99BF" ModsDocID="1313-2970-1041-27" checkinTime="1622492499165" checkinUser="pensoft" docAuthor="Brunke, Adam J., Pentinsaari, Mikko &amp; Klimaszewski, Jan" docDate="2021" docId="CDA6F43EAA375C109DEA6ABBD17D3B65" docLanguage="en" docName="ZooKeys 1041: 27-99" docOrigin="ZooKeys 1041" docPubDate="2021-05-31" docSource="http://dx.doi.org/10.3897/zookeys.1041.64460" docTitle="Gnypeta impressicollis Klimaszewski, Brunke &amp; Pentinsaari 2021, sp. nov." docType="treatment" docUuid="C6DC72D8-A182-43D3-9691-A352F0AF5D0E" docUuidSource="ZooBank" docVersion="4" id="5AE03537388755CFAF1E06C3CC9EFA72" lastPageNumber="27" masterDocId="5AE03537388755CFAF1E06C3CC9EFA72" masterDocTitle="Integrative taxonomy of Nearctic and Palaearctic Aleocharinae: new species, synonymies, and records (Coleoptera, Staphylinidae)" masterLastPageNumber="99" masterPageNumber="27" pageNumber="27" updateTime="1668150356826" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Integrative taxonomy of Nearctic and Palaearctic Aleocharinae: new species, synonymies, and records (Coleoptera, Staphylinidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Brunke, Adam J.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1158-936X</mods:nameIdentifier>
<mods:affiliation>Agriculture and Agri-Food Canada, Canadian National Collection of Insects, Arachnids and Nematodes, 960 Carling Avenue, Ottawa, Ontario, K 1 A 0 C 6, Canada</mods:affiliation>
<mods:nameIdentifier type="email">adam.j.brunke@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pentinsaari, Mikko</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7241-3873</mods:nameIdentifier>
<mods:affiliation>Centre for Biodiversity Genomics, 50 Stone Road East, University of Guelph, Guelph, Ontario, N 1 G 2 W 1, Canada</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Klimaszewski, Jan</mods:namePart>
<mods:affiliation>Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, 1055 du PEPS, PO Box 10380, Stn. Sainte-Foy, Quebec, QC, G 1 V 4 C 7, Canada</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-05-31</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1041</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>27</mods:start>
<mods:end>99</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1041.64460</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1041.64460</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1041-27</mods:identifier>
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<treatment ID-GBIF-Taxon="192735633" LSID="urn:lsid:zoobank.org:act:C6DC72D8-A182-43D3-9691-A352F0AF5D0E" httpUri="http://treatment.plazi.org/id/CDA6F43EAA375C109DEA6ABBD17D3B65" lastPageNumber="27" pageId="0" pageNumber="27">
<subSubSection pageId="0" pageNumber="27" type="nomenclature">
<paragraph pageId="0" pageNumber="27">
<taxonomicName LSID="http://zoobank.org/C6DC72D8-A182-43D3-9691-A352F0AF5D0E" authority="Klimaszewski, Brunke &amp; Pentinsaari" authorityName="Klimaszewski, Brunke &amp; Pentinsaari" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Gnypeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnypeta impressicollis" order="Coleoptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="impressicollis" status="sp. nov.">Gnypeta impressicollis Klimaszewski, Brunke &amp; Pentinsaari</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="27">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Gnypeta impressicollis Klimaszewski, Brunke &amp; Pentinsaari, sp. nov. A habitus B median lobe of aedeagus in lateral view C median lobe of aedeagus in dorsal view D spermatheca E male tergite VIII F male sternite VIII G female tergite VIII H female sternite VIII. Scale bars: 1 mm (A); 0.2 mm (B-H)." figureDoi="10.3897/zookeys.1041.64460.figure13" httpUri="https://binary.pensoft.net/fig/550278" pageId="0" pageNumber="27">Fig. 13A-H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="type material">
<paragraph pageId="0" pageNumber="27">Type material.</paragraph>
<paragraph pageId="0" pageNumber="27">
<emphasis bold="true" pageId="0" pageNumber="27">
<emphasis italics="true" pageId="0" pageNumber="27">Holotype</emphasis>
.
</emphasis>
(male, CNC):
<emphasis bold="true" pageId="0" pageNumber="27">Canada</emphasis>
, Ontario, Hartington, Eel Lake, South Frontenac, Paul
<normalizedToken originalValue="Heberts">Hebert's</normalizedToken>
cottage property,
<geoCoordinate degrees="44.563" direction="north" orientation="latitude" precision="55" value="44.563">44.563°N</geoCoordinate>
,
<geoCoordinate degrees="76.549" direction="west" orientation="longitude" precision="55" value="-76.549">76.549°W</geoCoordinate>
, 6.13.2017, Mikko Pentinsaari, Barcode of Life DNA voucher specimen, Sample ID: BIOUG34206-H01, Process ID: MPCAN465-17.
<emphasis bold="true" italics="true" pageId="0" pageNumber="27">Paratypes</emphasis>
(3 CBG, 4 CNC):
<emphasis bold="true" pageId="0" pageNumber="27">Canada</emphasis>
, labelled as the holotype except: Sample ID: BIOUG34206-H02, Process ID: MPCAN466-17 (1 male, CBG); Sample ID: BIOUG34206-G12, Process ID: MPCAN464-17 (1 male, CBG); Sample ID: BIOUG34206-G11, Process ID: MPCAN463-17 (1 female, CBG).
<emphasis bold="true" pageId="0" pageNumber="27">United States: North Carolina</emphasis>
: Haywood Co., 3 mi N Dellwood, 19.VIII.1972, A. Smetana (3, CNC);
<emphasis bold="true" pageId="0" pageNumber="27">Maryland</emphasis>
: Patuxent Wildl. Res. Ctr., 5 km E Montpelier, treading pond vegetation, 16.VI.1982, Bousquet &amp; Davies (1, CNC).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="etymology">
<paragraph pageId="0" pageNumber="27">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="27">The species epithet refers to the longitudinal impression on the pronotum, most strongly developed in males.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="diagnosis">
<paragraph pageId="0" pageNumber="27">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="27">
<taxonomicName authorityName="Klimaszewski, Brunke &amp; Pentinsaari" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Gnypeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnypeta impressicollis" order="Coleoptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="impressicollis">
<emphasis italics="true" pageId="0" pageNumber="27">Gnypeta impressicollis</emphasis>
</taxonomicName>
can be easily distinguished from all Nearctic species of the genus (except eastern
<taxonomicName lsidName="G. baltifera" pageId="0" pageNumber="27" rank="species" species="baltifera">
<emphasis italics="true" pageId="0" pageNumber="27">G. baltifera</emphasis>
</taxonomicName>
(LeConte)) by the hexagonal pronotum with a longitudinal impression in the basal half (females) to nearly entire pronotal length (males). Males also have an impression on the vertex of the head. We have examined the female type of
<taxonomicName lsidName="G. baltifera" pageId="0" pageNumber="27" rank="species" species="baltifera">
<emphasis italics="true" pageId="0" pageNumber="27">G. baltifera</emphasis>
</taxonomicName>
and it is externally similar but differs by the shorter, less angulate hexagonal pronotum, reddish and longer elytra and spermatheca with an elongate stem (C-shaped in
<taxonomicName lsidName="G. impressicollis" pageId="0" pageNumber="27" rank="species" species="impressicollis">
<emphasis italics="true" pageId="0" pageNumber="27">G. impressicollis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="description">
<paragraph pageId="0" pageNumber="27">Description.</paragraph>
<paragraph pageId="0" pageNumber="27">
Body length 3.2-3.4 mm; colour dark brown, elytra brown with irregular rust-brown patches, first two or three basal tergites rust-brown with posterior edge yellow, apex of abdomen rust-brown, legs and antennae rust-brown; integument highly glossy (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Gnypeta impressicollis Klimaszewski, Brunke &amp; Pentinsaari, sp. nov. A habitus B median lobe of aedeagus in lateral view C median lobe of aedeagus in dorsal view D spermatheca E male tergite VIII F male sternite VIII G female tergite VIII H female sternite VIII. Scale bars: 1 mm (A); 0.2 mm (B-H)." figureDoi="10.3897/zookeys.1041.64460.figure13" httpUri="https://binary.pensoft.net/fig/550278" pageId="0" pageNumber="27">13A</figureCitation>
); pubescence yellowish grey, moderately long and moderately sparse; all antennomeres distinctly elongate; head round with short neck (visible only when head is distended from thorax), vertex in males with broad central impression, vertex of females with much smaller and narrower median impression, maximum width of head slightly less than maximum width of pronotum; pronotum hexagonal in shape, ca. as long as head, with a longitudinal impression in the basal half (females) to nearly entire pronotal length (males), pubescence on disc directed anteriad along midline and obliquely laterad elsewhere; elytra wider than either head or pronotum, at suture shorter than pronotum along midline, pubescence directed obliquely posteriad forming wavy pattern medially on each side; abdomen arcuate laterally, broadest in apical third, at base distinctly narrower than elytra; legs very long, hind tarsus with basal tarsomere ca. as long as the two following ones combined. MALE. Tergite VIII with apical margin truncate medially and arcuate laterally (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Gnypeta impressicollis Klimaszewski, Brunke &amp; Pentinsaari, sp. nov. A habitus B median lobe of aedeagus in lateral view C median lobe of aedeagus in dorsal view D spermatheca E male tergite VIII F male sternite VIII G female tergite VIII H female sternite VIII. Scale bars: 1 mm (A); 0.2 mm (B-H)." figureDoi="10.3897/zookeys.1041.64460.figure13" httpUri="https://binary.pensoft.net/fig/550278" pageId="0" pageNumber="27">13E</figureCitation>
); sternite VIII elongate, narrowed apically, apex truncate medially and oblique laterally (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Gnypeta impressicollis Klimaszewski, Brunke &amp; Pentinsaari, sp. nov. A habitus B median lobe of aedeagus in lateral view C median lobe of aedeagus in dorsal view D spermatheca E male tergite VIII F male sternite VIII G female tergite VIII H female sternite VIII. Scale bars: 1 mm (A); 0.2 mm (B-H)." figureDoi="10.3897/zookeys.1041.64460.figure13" httpUri="https://binary.pensoft.net/fig/550278" pageId="0" pageNumber="27">13F</figureCitation>
); median lobe of aedeagus in lateral view with tubus very short, triangular and gradually tapering to narrowly rounded apex, ventral margin broadly curved ventrad in apical half (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Gnypeta impressicollis Klimaszewski, Brunke &amp; Pentinsaari, sp. nov. A habitus B median lobe of aedeagus in lateral view C median lobe of aedeagus in dorsal view D spermatheca E male tergite VIII F male sternite VIII G female tergite VIII H female sternite VIII. Scale bars: 1 mm (A); 0.2 mm (B-H)." figureDoi="10.3897/zookeys.1041.64460.figure13" httpUri="https://binary.pensoft.net/fig/550278" pageId="0" pageNumber="27">13B</figureCitation>
); in dorsal view bulbus moderately large and tubus swelled basally and triangular apically (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Gnypeta impressicollis Klimaszewski, Brunke &amp; Pentinsaari, sp. nov. A habitus B median lobe of aedeagus in lateral view C median lobe of aedeagus in dorsal view D spermatheca E male tergite VIII F male sternite VIII G female tergite VIII H female sternite VIII. Scale bars: 1 mm (A); 0.2 mm (B-H)." figureDoi="10.3897/zookeys.1041.64460.figure13" httpUri="https://binary.pensoft.net/fig/550278" pageId="0" pageNumber="27">13C</figureCitation>
); internal sac with complex membranous structures (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Gnypeta impressicollis Klimaszewski, Brunke &amp; Pentinsaari, sp. nov. A habitus B median lobe of aedeagus in lateral view C median lobe of aedeagus in dorsal view D spermatheca E male tergite VIII F male sternite VIII G female tergite VIII H female sternite VIII. Scale bars: 1 mm (A); 0.2 mm (B-H)." figureDoi="10.3897/zookeys.1041.64460.figure13" httpUri="https://binary.pensoft.net/fig/550278" pageId="0" pageNumber="27">13B,C</figureCitation>
). FEMALE. Tergite VIII broadly arcuate apically (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Gnypeta impressicollis Klimaszewski, Brunke &amp; Pentinsaari, sp. nov. A habitus B median lobe of aedeagus in lateral view C median lobe of aedeagus in dorsal view D spermatheca E male tergite VIII F male sternite VIII G female tergite VIII H female sternite VIII. Scale bars: 1 mm (A); 0.2 mm (B-H)." figureDoi="10.3897/zookeys.1041.64460.figure13" httpUri="https://binary.pensoft.net/fig/550278" pageId="0" pageNumber="27">13G</figureCitation>
); sternite VIII rounded apically with very shallow median emargination (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Gnypeta impressicollis Klimaszewski, Brunke &amp; Pentinsaari, sp. nov. A habitus B median lobe of aedeagus in lateral view C median lobe of aedeagus in dorsal view D spermatheca E male tergite VIII F male sternite VIII G female tergite VIII H female sternite VIII. Scale bars: 1 mm (A); 0.2 mm (B-H)." figureDoi="10.3897/zookeys.1041.64460.figure13" httpUri="https://binary.pensoft.net/fig/550278" pageId="0" pageNumber="27">13H</figureCitation>
); spermatheca C-shaped, capsule subspherical with broad apical invagination, stem tubular and C-shaped (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Gnypeta impressicollis Klimaszewski, Brunke &amp; Pentinsaari, sp. nov. A habitus B median lobe of aedeagus in lateral view C median lobe of aedeagus in dorsal view D spermatheca E male tergite VIII F male sternite VIII G female tergite VIII H female sternite VIII. Scale bars: 1 mm (A); 0.2 mm (B-H)." figureDoi="10.3897/zookeys.1041.64460.figure13" httpUri="https://binary.pensoft.net/fig/550278" pageId="0" pageNumber="27">13D</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1041.64460.figure13" httpUri="https://binary.pensoft.net/fig/550278" pageId="0" pageNumber="27" start="Figure 13" startId="F13">
<paragraph pageId="0" pageNumber="27">
<emphasis bold="true" pageId="0" pageNumber="27">Figure 13.</emphasis>
<taxonomicName authorityName="Klimaszewski, Brunke &amp; Pentinsaari" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Gnypeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnypeta impressicollis" order="Coleoptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="impressicollis">
<emphasis italics="true" pageId="0" pageNumber="27">Gnypeta impressicollis</emphasis>
</taxonomicName>
Klimaszewski, Brunke &amp; Pentinsaari, sp. nov.
<emphasis bold="true" pageId="0" pageNumber="27">A</emphasis>
habitus
<emphasis bold="true" pageId="0" pageNumber="27">B</emphasis>
median lobe of aedeagus in lateral view
<emphasis bold="true" pageId="0" pageNumber="27">C</emphasis>
median lobe of aedeagus in dorsal view
<emphasis bold="true" pageId="0" pageNumber="27">D</emphasis>
spermatheca
<emphasis bold="true" pageId="0" pageNumber="27">E</emphasis>
male tergite VIII
<emphasis bold="true" pageId="0" pageNumber="27">F</emphasis>
male sternite VIII
<emphasis bold="true" pageId="0" pageNumber="27">G</emphasis>
female tergite VIII
<emphasis bold="true" pageId="0" pageNumber="27">H</emphasis>
female sternite VIII. Scale bars: 1 mm (
<emphasis bold="true" pageId="0" pageNumber="27">A</emphasis>
); 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="27">B-H</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="distribution">
<paragraph pageId="0" pageNumber="27">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="27">
<emphasis bold="true" pageId="0" pageNumber="27">Origin</emphasis>
: Nearctic.
<emphasis bold="true" pageId="0" pageNumber="27">Canada</emphasis>
: ON.
<emphasis bold="true" pageId="0" pageNumber="27">United States</emphasis>
: MD, NC.
<taxonomicName authorityName="Klimaszewski, Brunke &amp; Pentinsaari" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Gnypeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnypeta impressicollis" order="Coleoptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="impressicollis">
<emphasis italics="true" pageId="0" pageNumber="27">Gnypeta impressicollis</emphasis>
</taxonomicName>
is probably broadly distributed in eastern North America.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="bionomics">
<paragraph pageId="0" pageNumber="27">Bionomics.</paragraph>
<paragraph pageId="0" pageNumber="27">Specimens were collected by sifting leaf litter along a lake margin and by treading pond vegetation.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="comments">
<paragraph pageId="0" pageNumber="27">Comments.</paragraph>
<paragraph pageId="0" pageNumber="27">
It was challenging to place this species in either
<taxonomicName authorityName="C.G. Thomson" authorityYear="1858" class="Insecta" family="Staphylinidae" genus="Gnypeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnypeta" order="Coleoptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Gnypeta</emphasis>
</taxonomicName>
or
<taxonomicName class="Insecta" family="Staphylinidae" genus="Ischnopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ischnopoda" order="Coleoptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Ischnopoda</emphasis>
</taxonomicName>
Stephens based on the concepts of
<bibRefCitation author="Pasnik, G" journalOrPublisher="Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Krakow" pageId="0" pageNumber="27" refId="B58" refString="Pasnik, G, 2010. Phylogeny and Generic Classification of Tachyusini (Coleoptera, Staphylinidae: Aleocharinae). Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Krakow" title="Phylogeny and Generic Classification of Tachyusini (Coleoptera, Staphylinidae: Aleocharinae)." year="2010">
<normalizedToken originalValue="Paśnik">Pasnik</normalizedToken>
(2010)
</bibRefCitation>
. The extremely long legs, pronotal shape, C-shaped spermatheca and superficial punctation of the pronotum and abdomen are consistent with at least some Neotropical members of
<taxonomicName class="Insecta" family="Staphylinidae" genus="Ischnopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ischnopoda" order="Coleoptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Ischnopoda</emphasis>
</taxonomicName>
but the ligula of
<taxonomicName lsidName="G. impressicollis" pageId="0" pageNumber="27" rank="species" species="impressicollis">
<emphasis italics="true" pageId="0" pageNumber="27">G. impressicollis</emphasis>
</taxonomicName>
is divided to the base, which is considered to be a feature of
<taxonomicName authorityName="C.G. Thomson" authorityYear="1858" class="Insecta" family="Staphylinidae" genus="Gnypeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnypeta" order="Coleoptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Gnypeta</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Pašnik">Pasnik</normalizedToken>
2010). The C-shaped spermatheca of
<taxonomicName authorityName="Klimaszewski, Brunke &amp; Pentinsaari" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Gnypeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnypeta impressicollis" order="Coleoptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="impressicollis">
<emphasis italics="true" pageId="0" pageNumber="27">Gnypeta impressicollis</emphasis>
</taxonomicName>
also bears some similarity to the
<taxonomicName lsidName="G. crebrepunctata" pageId="0" pageNumber="27" rank="species" species="crebrepunctata">
<emphasis italics="true" pageId="0" pageNumber="27">G. crebrepunctata</emphasis>
</taxonomicName>
group of
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.2.4" author="Klimaszewski, J" journalOrPublisher="ZooKeys" pageId="0" pageNumber="27" pagination="11 - 84" refId="B33" refString="Klimaszewski, J, Savard, K, Pelletier, G, Webster, RP, 2008. Species review of the genus Gnypeta Thomson from Canada, Alaska and Greenland (Coleoptera, Staphylinidae, Aleocharinae): systematics, bionomics and distribution. ZooKeys 2: 11 - 84, DOI: https://doi.org/10.3897/zookeys.2.4" title="Species review of the genus Gnypeta Thomson from Canada, Alaska and Greenland (Coleoptera, Staphylinidae, Aleocharinae): systematics, bionomics and distribution." url="https://doi.org/10.3897/zookeys.2.4" volume="2" year="2008">Klimaszewski et al. (2008)</bibRefCitation>
but it is rather different in external morphology. We place this species in
<taxonomicName authorityName="C.G. Thomson" authorityYear="1858" class="Insecta" family="Staphylinidae" genus="Gnypeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnypeta" order="Coleoptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Gnypeta</emphasis>
</taxonomicName>
pending future systematic research.
</paragraph>
</subSubSection>
</treatment>
</document>