513 lines
94 KiB
XML
513 lines
94 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1079.76412" ID-GBIF-Dataset="90785384-12f1-40ef-aa18-5e08ff36e4bc" ID-PMC="PMC8741714" ID-Pensoft-Pub="1313-2970-1079-145" ID-Pensoft-UUID="C75BE4A1E65C51598D01C9EA871B90E5" ID-PubMed="35068961" ID-ZooBank="55F9DF83DE924EC3ACEC05AA4466F147" ModsDocID="1313-2970-1079-145" checkinTime="1641053224064" checkinUser="pensoft" docAuthor="Wittmann, Karl J. & Chevaldonne, Pierre" docDate="2021" docId="CD9C61C1BD695761931785C0ACCAE211" docLanguage="en" docName="ZooKeys 1079: 145-227" docOrigin="ZooKeys 1079" docPubDate="2021-12-31" docSource="http://dx.doi.org/10.3897/zookeys.1079.76412" docTitle="Pseudomma kryotroglodytum Wittmann & Chevaldonné 2021, sp. nov." docType="treatment" docUuid="5B212BFB-7ADE-41CE-A671-CFC363B95A1C" docUuidSource="ZooBank" docVersion="4" id="C75BE4A1E65C51598D01C9EA871B90E5" lastPageNumber="145" masterDocId="C75BE4A1E65C51598D01C9EA871B90E5" masterDocTitle="First report of the order Mysida (Crustacea) in Antarctic marine ice caves, with description of a new species of Pseudomma and investigations on the taxonomy, morphology and life habits of Mysidetes species" masterLastPageNumber="227" masterPageNumber="145" pageNumber="145" updateTime="1668151265394" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>First report of the order Mysida (Crustacea) in Antarctic marine ice caves, with description of a new species of Pseudomma and investigations on the taxonomy, morphology and life habits of Mysidetes species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Wittmann, Karl J.</mods:namePart>
|
||
<mods:affiliation>Department of Environmental Health, Medical University of Vienna, Kinderspitalgasse 15, 1090 Vienna, Austria. Corresponding author</mods:affiliation>
|
||
<mods:nameIdentifier type="email">karl.wittmann@meduniwien.ac.at</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Chevaldonne, Pierre</mods:namePart>
|
||
<mods:affiliation>IMBE, CNRS, IRD, Aix Marseille Universite, Avignon Universite, Station Marine d'Endoume, Rue de la Batterie des Lions, 13007 Marseille, France</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2021</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2021-12-31</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>1079</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>145</mods:start>
|
||
<mods:end>227</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/zookeys.1079.76412</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1079.76412</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">1313-2970-1079-145</mods:identifier>
|
||
<mods:identifier type="ZooBank">55F9DF83DE924EC3ACEC05AA4466F147</mods:identifier>
|
||
<mods:identifier type="Pensoft-UUID">C75BE4A1E65C51598D01C9EA871B90E5</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-GBIF-Taxon="191828690" LSID="urn:lsid:zoobank.org:act:5B212BFB-7ADE-41CE-A671-CFC363B95A1C" httpUri="http://treatment.plazi.org/id/CD9C61C1BD695761931785C0ACCAE211" lastPageNumber="145" pageId="0" pageNumber="145">
|
||
<subSubSection pageId="0" pageNumber="145" type="nomenclature">
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<taxonomicName LSID="http://zoobank.org/5B212BFB-7ADE-41CE-A671-CFC363B95A1C" authority="Wittmann & Chevaldonné, 2021" authorityName="Wittmann & Chevaldonné" authorityYear="2021" class="Malacostraca" family="Mysidae" genus="Pseudomma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pseudomma kryotroglodytum" order="Mysida" pageId="0" pageNumber="145" phylum="Schizopoda" rank="species" species="kryotroglodytum" status="sp. nov.">Pseudomma kryotroglodytum</taxonomicName>
|
||
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="description">
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pseudomma kryotroglodytum sp. nov. from ice cave of Bernard Island, Antarctica A female holotype, lateral B cephalothorax of female holotype, dorsal C physical aspect of the ice cave environment A, B living specimen in laboratory." figureDoi="10.3897/zookeys.1079.76412.figure2" httpUri="https://binary.pensoft.net/fig/629312" pageId="0" pageNumber="145">Figures 2</figureCitation>
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E) and paratype subadult female 21.5 mm (F) A terminal portion of telson, dorsal, details show scales on left disto-mesial spine (B) and barbs on left terminal seta (C) D anterior half of cephalothorax, lateral, arrow points to distolateral edge of carapace E sixth pleomere with tail fan, lateral F example for pores (three to the left) and coat of scales on tergite of first pleomere." figureDoi="10.3897/zookeys.1079.76412.figure3" httpUri="https://binary.pensoft.net/fig/629313" pageId="0" pageNumber="145">, 3</figureCitation>
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A, B, E, F) and paratype subadult female 21.5 mm (C, D). A right antennula, dorsal B antenna with antennal gland, ventral, setae omitted from antennal scale C eyeplate and carapace expanded on slide, short arrow points to distolateral edge of carapace, detail (D) pore group in pre-cardial position E mandibles with left palpus, caudal aspect F maxillula, caudal. Scales omitted from objects A-C, E, F but not from eyeplate in panel C; pore diameters not to scale in C, D." figureDoi="10.3897/zookeys.1079.76412.figure4" httpUri="https://binary.pensoft.net/fig/629314" pageId="0" pageNumber="145">, 4</figureCitation>
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm A labrum B labium C maxilla, caudal aspect D thoracopod 1 (caudal) with thoracic sternites 1, 2 (ventral) E detail of panel (D) showing dactylus 1 with nail F thoracic endopod 2, rostral G thoracic endopod 4, rostral H detail of panel (G) showing dactylus 4 with nail I inner face of oostegite 1, drawn above part of sympod 6. Scales omitted from objects A-D, F, G; setae from E, H." figureDoi="10.3897/zookeys.1079.76412.figure5" httpUri="https://binary.pensoft.net/fig/629315" pageId="0" pageNumber="145">, 5</figureCitation>
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E, H-N) and paratype subadult female 21.5 mm (F, G). A cardiac portion of foregut, dorsal view, dorsal wall omitted B-D details of panel (A) showing modified spines E details of panel (A) showing teeth emerging from a common basis F pleonite 1, cuticle detached and expanded on slide, dorsal and lateral faces on top, ventral face folded in, setae omitted from pleopods G detail of panel (F) showing pore group on left lateral face H-L series of pleopods 1 - 5, rostral = lateral face M uropods, ventral N telson. Scales omitted from objects F, M, N pore diameters not to scale in F, G." figureDoi="10.3897/zookeys.1079.76412.figure6" httpUri="https://binary.pensoft.net/fig/629316" pageId="0" pageNumber="145">, 6</figureCitation>
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Structure of eyeplates in Pseudomma kryotroglodytum sp. nov. and Michthyops parvus. Holotype adult female (D) and paratype subadult female (A, B) of P. kryotroglodytum and non-type adult female of M. parvus (C). A eyeplate expanded on slide, dorsal, to the right somewhat distorted B detail of panel (A) showing cyst connected with bottom of median cleft C homologous cyst in another species and genus, arrow points to tubular connection with anterior margin of eyeplate D series of denticles along sub-lateral section of anterior margin of eyeplate." figureDoi="10.3897/zookeys.1079.76412.figure23" httpUri="https://binary.pensoft.net/fig/629333" pageId="0" pageNumber="145">, 23A, B, D</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="type series">
|
||
<paragraph pageId="0" pageNumber="145">Type series.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
|
||
spent female (♀0-) BL 26.8 mm (on slides at NHMW 27296, GenBank nos. OK351330 and OK353694), East Antarctica,
|
||
<normalizedToken originalValue="Adélie">Adelie</normalizedToken>
|
||
Land, near Dumont
|
||
<normalizedToken originalValue="d’Urville">d'Urville</normalizedToken>
|
||
Station, NE of
|
||
<emphasis bold="true" pageId="0" pageNumber="145">Claude Bernard Island</emphasis>
|
||
,
|
||
<geoCoordinate degrees="66" direction="south" minutes="39.64" orientation="latitude" precision="9" value="-66.66067">66°39.64'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="140" direction="east" minutes="01.55" orientation="longitude" precision="9" value="140.02583">140°01.55'E</geoCoordinate>
|
||
, ice cave, dive #612, depth 10 m, diver-operated suction bottle, 15 Jan 2016, leg. P.
|
||
<normalizedToken originalValue="Chevaldonné">Chevaldonne</normalizedToken>
|
||
& S. Hourdez.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Paratype</emphasis>
|
||
subadult female (♀S-) BL 21.5 mm (on slides at NHMW 27297, GenBank nos. OK351329 and OK353693), dive #611, 13 Jan 2016, remaining sampling data as for holotype.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
Covers females only. Species of the genus
|
||
<taxonomicName authorityName="G. O. Sars" authorityYear="1870" class="Malacostraca" family="Mysidae" genus="Pseudomma" kingdom="Animalia" lsidName="Pseudomma" order="Mysidacea" pageId="0" pageNumber="145" phylum="Schizopoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Pseudomma</emphasis>
|
||
</taxonomicName>
|
||
G.O. Sars, 1870, with cornea-like lateral portions separated by sulci from main part of eyeplate (Figs
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pseudomma kryotroglodytum sp. nov. from ice cave of Bernard Island, Antarctica A female holotype, lateral B cephalothorax of female holotype, dorsal C physical aspect of the ice cave environment A, B living specimen in laboratory." figureDoi="10.3897/zookeys.1079.76412.figure2" httpUri="https://binary.pensoft.net/fig/629312" pageId="0" pageNumber="145">2B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A, B, E, F) and paratype subadult female 21.5 mm (C, D). A right antennula, dorsal B antenna with antennal gland, ventral, setae omitted from antennal scale C eyeplate and carapace expanded on slide, short arrow points to distolateral edge of carapace, detail (D) pore group in pre-cardial position E mandibles with left palpus, caudal aspect F maxillula, caudal. Scales omitted from objects A-C, E, F but not from eyeplate in panel C; pore diameters not to scale in C, D." figureDoi="10.3897/zookeys.1079.76412.figure4" httpUri="https://binary.pensoft.net/fig/629314" pageId="0" pageNumber="145">4C</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Structure of eyeplates in Pseudomma kryotroglodytum sp. nov. and Michthyops parvus. Holotype adult female (D) and paratype subadult female (A, B) of P. kryotroglodytum and non-type adult female of M. parvus (C). A eyeplate expanded on slide, dorsal, to the right somewhat distorted B detail of panel (A) showing cyst connected with bottom of median cleft C homologous cyst in another species and genus, arrow points to tubular connection with anterior margin of eyeplate D series of denticles along sub-lateral section of anterior margin of eyeplate." figureDoi="10.3897/zookeys.1079.76412.figure23" httpUri="https://binary.pensoft.net/fig/629333" pageId="0" pageNumber="145">23A</figureCitation>
|
||
), no visual elements. Disto-median fissure penetrates one third of eyeplate. Distal margin of eyeplates with series of minute teeth along sublateral sector (
|
||
<normalizedToken originalValue="‘shoulders’">'shoulders'</normalizedToken>
|
||
, Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Structure of eyeplates in Pseudomma kryotroglodytum sp. nov. and Michthyops parvus. Holotype adult female (D) and paratype subadult female (A, B) of P. kryotroglodytum and non-type adult female of M. parvus (C). A eyeplate expanded on slide, dorsal, to the right somewhat distorted B detail of panel (A) showing cyst connected with bottom of median cleft C homologous cyst in another species and genus, arrow points to tubular connection with anterior margin of eyeplate D series of denticles along sub-lateral section of anterior margin of eyeplate." figureDoi="10.3897/zookeys.1079.76412.figure23" httpUri="https://binary.pensoft.net/fig/629333" pageId="0" pageNumber="145">23A, D</figureCitation>
|
||
). Basal segment of antennular trunk without medio-ventral carina. Antennal scale (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A, B, E, F) and paratype subadult female 21.5 mm (C, D). A right antennula, dorsal B antenna with antennal gland, ventral, setae omitted from antennal scale C eyeplate and carapace expanded on slide, short arrow points to distolateral edge of carapace, detail (D) pore group in pre-cardial position E mandibles with left palpus, caudal aspect F maxillula, caudal. Scales omitted from objects A-C, E, F but not from eyeplate in panel C; pore diameters not to scale in C, D." figureDoi="10.3897/zookeys.1079.76412.figure4" httpUri="https://binary.pensoft.net/fig/629314" pageId="0" pageNumber="145">4B</figureCitation>
|
||
) with setose apical lobe contributing about 1/4 scale length. Mandibular palp (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A, B, E, F) and paratype subadult female 21.5 mm (C, D). A right antennula, dorsal B antenna with antennal gland, ventral, setae omitted from antennal scale C eyeplate and carapace expanded on slide, short arrow points to distolateral edge of carapace, detail (D) pore group in pre-cardial position E mandibles with left palpus, caudal aspect F maxillula, caudal. Scales omitted from objects A-C, E, F but not from eyeplate in panel C; pore diameters not to scale in C, D." figureDoi="10.3897/zookeys.1079.76412.figure4" httpUri="https://binary.pensoft.net/fig/629314" pageId="0" pageNumber="145">4E</figureCitation>
|
||
) 3-segmented, very large, about as long as antennal scale. Three pairs of oostegites (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm A labrum B labium C maxilla, caudal aspect D thoracopod 1 (caudal) with thoracic sternites 1, 2 (ventral) E detail of panel (D) showing dactylus 1 with nail F thoracic endopod 2, rostral G thoracic endopod 4, rostral H detail of panel (G) showing dactylus 4 with nail I inner face of oostegite 1, drawn above part of sympod 6. Scales omitted from objects A-D, F, G; setae from E, H." figureDoi="10.3897/zookeys.1079.76412.figure5" httpUri="https://binary.pensoft.net/fig/629315" pageId="0" pageNumber="145">5I</figureCitation>
|
||
) contributing to wall of brood pouch. Pleopods (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E, H-N) and paratype subadult female 21.5 mm (F, G). A cardiac portion of foregut, dorsal view, dorsal wall omitted B-D details of panel (A) showing modified spines E details of panel (A) showing teeth emerging from a common basis F pleonite 1, cuticle detached and expanded on slide, dorsal and lateral faces on top, ventral face folded in, setae omitted from pleopods G detail of panel (F) showing pore group on left lateral face H-L series of pleopods 1 - 5, rostral = lateral face M uropods, ventral N telson. Scales omitted from objects F, M, N pore diameters not to scale in F, G." figureDoi="10.3897/zookeys.1079.76412.figure6" httpUri="https://binary.pensoft.net/fig/629316" pageId="0" pageNumber="145">6H-L</figureCitation>
|
||
) reduced to setose rods with residual differentiation of endopod (pseudobranchial lobes). Telson (Figs
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E) and paratype subadult female 21.5 mm (F) A terminal portion of telson, dorsal, details show scales on left disto-mesial spine (B) and barbs on left terminal seta (C) D anterior half of cephalothorax, lateral, arrow points to distolateral edge of carapace E sixth pleomere with tail fan, lateral F example for pores (three to the left) and coat of scales on tergite of first pleomere." figureDoi="10.3897/zookeys.1079.76412.figure3" httpUri="https://binary.pensoft.net/fig/629313" pageId="0" pageNumber="145">3A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E, H-N) and paratype subadult female 21.5 mm (F, G). A cardiac portion of foregut, dorsal view, dorsal wall omitted B-D details of panel (A) showing modified spines E details of panel (A) showing teeth emerging from a common basis F pleonite 1, cuticle detached and expanded on slide, dorsal and lateral faces on top, ventral face folded in, setae omitted from pleopods G detail of panel (F) showing pore group on left lateral face H-L series of pleopods 1 - 5, rostral = lateral face M uropods, ventral N telson. Scales omitted from objects F, M, N pore diameters not to scale in F, G." figureDoi="10.3897/zookeys.1079.76412.figure6" httpUri="https://binary.pensoft.net/fig/629316" pageId="0" pageNumber="145">6N</figureCitation>
|
||
) trapezoid, as long as ultimate pleonite. Its length twice maximum width at basis and four times width at apex. Lateral margins of telson without setae and spines, only minute scales present. Transversely truncate terminal margin with only two pairs of spines, both hispid due to minute scales (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E) and paratype subadult female 21.5 mm (F) A terminal portion of telson, dorsal, details show scales on left disto-mesial spine (B) and barbs on left terminal seta (C) D anterior half of cephalothorax, lateral, arrow points to distolateral edge of carapace E sixth pleomere with tail fan, lateral F example for pores (three to the left) and coat of scales on tergite of first pleomere." figureDoi="10.3897/zookeys.1079.76412.figure3" httpUri="https://binary.pensoft.net/fig/629313" pageId="0" pageNumber="145">3B</figureCitation>
|
||
) along more than proximal 2/3 spine length. Large latero-apical and same-sized submedio-apical spines flank median pair of closely set setae (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E) and paratype subadult female 21.5 mm (F) A terminal portion of telson, dorsal, details show scales on left disto-mesial spine (B) and barbs on left terminal seta (C) D anterior half of cephalothorax, lateral, arrow points to distolateral edge of carapace E sixth pleomere with tail fan, lateral F example for pores (three to the left) and coat of scales on tergite of first pleomere." figureDoi="10.3897/zookeys.1079.76412.figure3" httpUri="https://binary.pensoft.net/fig/629313" pageId="0" pageNumber="145">3C</figureCitation>
|
||
) with twice spine length. Margin with short, well-rounded indentation between each spine, median indentation largest. Disto-lateral edge without tooth, with spine only.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="description">
|
||
<paragraph pageId="0" pageNumber="145">Description of the holotype.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
All features of the diagnosis. Female with body length 26.8 mm. Cephalothorax measures 39% body length, pleon without telson 48%, telson 13% and carapace 32%. Large parts of the body, particularly carapace, pleon, telson, and uropods scaly-hispid; most appendages and eyeplates only to a minor degree. However, with 30
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
episcopic microscopy, the entire body appears smooth (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E) and paratype subadult female 21.5 mm (F) A terminal portion of telson, dorsal, details show scales on left disto-mesial spine (B) and barbs on left terminal seta (C) D anterior half of cephalothorax, lateral, arrow points to distolateral edge of carapace E sixth pleomere with tail fan, lateral F example for pores (three to the left) and coat of scales on tergite of first pleomere." figureDoi="10.3897/zookeys.1079.76412.figure3" httpUri="https://binary.pensoft.net/fig/629313" pageId="0" pageNumber="145">3D-E</figureCitation>
|
||
) due to small size of scales. With 600
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
transmitted phase contrast microscopy, large areas of the (artificially shed) dorsal cuticle of the animal resembles fish skin (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E) and paratype subadult female 21.5 mm (F) A terminal portion of telson, dorsal, details show scales on left disto-mesial spine (B) and barbs on left terminal seta (C) D anterior half of cephalothorax, lateral, arrow points to distolateral edge of carapace E sixth pleomere with tail fan, lateral F example for pores (three to the left) and coat of scales on tergite of first pleomere." figureDoi="10.3897/zookeys.1079.76412.figure3" httpUri="https://binary.pensoft.net/fig/629313" pageId="0" pageNumber="145">3F</figureCitation>
|
||
) due to dense scale cover. Ventral portions of pleomeres less densely covered, thoracic sternites smooth.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Antennula</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pseudomma kryotroglodytum sp. nov. from ice cave of Bernard Island, Antarctica A female holotype, lateral B cephalothorax of female holotype, dorsal C physical aspect of the ice cave environment A, B living specimen in laboratory." figureDoi="10.3897/zookeys.1079.76412.figure2" httpUri="https://binary.pensoft.net/fig/629312" pageId="0" pageNumber="145">2B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A, B, E, F) and paratype subadult female 21.5 mm (C, D). A right antennula, dorsal B antenna with antennal gland, ventral, setae omitted from antennal scale C eyeplate and carapace expanded on slide, short arrow points to distolateral edge of carapace, detail (D) pore group in pre-cardial position E mandibles with left palpus, caudal aspect F maxillula, caudal. Scales omitted from objects A-C, E, F but not from eyeplate in panel C; pore diameters not to scale in C, D." figureDoi="10.3897/zookeys.1079.76412.figure4" httpUri="https://binary.pensoft.net/fig/629314" pageId="0" pageNumber="145">4A</figureCitation>
|
||
). Epi-antennular process triangular, projecting in median position beyond eyeplate like a small arrowhead (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pseudomma kryotroglodytum sp. nov. from ice cave of Bernard Island, Antarctica A female holotype, lateral B cephalothorax of female holotype, dorsal C physical aspect of the ice cave environment A, B living specimen in laboratory." figureDoi="10.3897/zookeys.1079.76412.figure2" httpUri="https://binary.pensoft.net/fig/629312" pageId="0" pageNumber="145">2B</figureCitation>
|
||
). Antennular trunk with three sparsely setose segments, separated by transverse articulations. Basal segment 45%, median segment 16% and terminal segment 39% trunk length. Length of basal segment is only 2/3 width; mid-dorsally with deep antennular depression leading down to a striated pad at the bottom (Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A, B, E, F) and paratype subadult female 21.5 mm (C, D). A right antennula, dorsal B antenna with antennal gland, ventral, setae omitted from antennal scale C eyeplate and carapace expanded on slide, short arrow points to distolateral edge of carapace, detail (D) pore group in pre-cardial position E mandibles with left palpus, caudal aspect F maxillula, caudal. Scales omitted from objects A-C, E, F but not from eyeplate in panel C; pore diameters not to scale in C, D." figureDoi="10.3897/zookeys.1079.76412.figure4" httpUri="https://binary.pensoft.net/fig/629314" pageId="0" pageNumber="145">4A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Pseudomma kryotroglodytum sp. nov., paratype subadult female BL 21.5 mm (A, B) and Dactylamblyops sp., adult female 21.8 mm (C, D); A ventrally orientated depression mid-dorsally on basal segment of right antennula, dorsal aspect B detail of panel (A), arrow points to striated pad on bottom of depression, dorsal C depression as in panel (A) for left antennula in another genus, lateral aspect, the short arrow points to mineral particles D detail of panel (C), the long arrow points to striated pad, lateral. A, antennular depression somewhat caudally tilted by the pressure exerted by the cover glass." figureDoi="10.3897/zookeys.1079.76412.figure24" httpUri="https://binary.pensoft.net/fig/629334" pageId="0" pageNumber="145">24A, B</figureCitation>
|
||
) as described below. Basal segment not produced at outer distal corner. Terminal segment with the usual dorsal lobe on distal margin. This lobe without spiniform extension, disto-laterally with four barbed setae, mid-terminally and disto-medially with thickened, rugged margin. Flagella large, width of outer flagellum measured near basis with 1.1-1.2 times width of inner flagellum. Trunk with scales over major portions of its surface, not so the flagella.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Antenna</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A, B, E, F) and paratype subadult female 21.5 mm (C, D). A right antennula, dorsal B antenna with antennal gland, ventral, setae omitted from antennal scale C eyeplate and carapace expanded on slide, short arrow points to distolateral edge of carapace, detail (D) pore group in pre-cardial position E mandibles with left palpus, caudal aspect F maxillula, caudal. Scales omitted from objects A-C, E, F but not from eyeplate in panel C; pore diameters not to scale in C, D." figureDoi="10.3897/zookeys.1079.76412.figure4" httpUri="https://binary.pensoft.net/fig/629314" pageId="0" pageNumber="145">4B</figureCitation>
|
||
). Antennal scale large, 1.8 times length of antennular trunk and 1.8 antennal peduncle. Scale extends 0.4 times its length beyond antennular trunk and 0.7 beyond eyeplate (taking into account that antennulae insert more rostrally). Scale unsegmented, 2.9 times longer than wide. Scale dorsally and ventrally scaly-hispid all over. The smooth portion (not considering minute cuticle scales) of its outer margin ends in a strong tooth; setose apical lobe extends 26-27% scale length beyond this tooth. Antennal peduncle three-segmented. Basal segment contributes 21%, median segment 42% and terminal segment 36% peduncle length. Sympod angular on disto-lateral edge, not forming a tooth-like projection. Sympod with hispid lateral face.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Eyes</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pseudomma kryotroglodytum sp. nov. from ice cave of Bernard Island, Antarctica A female holotype, lateral B cephalothorax of female holotype, dorsal C physical aspect of the ice cave environment A, B living specimen in laboratory." figureDoi="10.3897/zookeys.1079.76412.figure2" httpUri="https://binary.pensoft.net/fig/629312" pageId="0" pageNumber="145">2B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E) and paratype subadult female 21.5 mm (F) A terminal portion of telson, dorsal, details show scales on left disto-mesial spine (B) and barbs on left terminal seta (C) D anterior half of cephalothorax, lateral, arrow points to distolateral edge of carapace E sixth pleomere with tail fan, lateral F example for pores (three to the left) and coat of scales on tergite of first pleomere." figureDoi="10.3897/zookeys.1079.76412.figure3" httpUri="https://binary.pensoft.net/fig/629313" pageId="0" pageNumber="145">3D</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A, B, E, F) and paratype subadult female 21.5 mm (C, D). A right antennula, dorsal B antenna with antennal gland, ventral, setae omitted from antennal scale C eyeplate and carapace expanded on slide, short arrow points to distolateral edge of carapace, detail (D) pore group in pre-cardial position E mandibles with left palpus, caudal aspect F maxillula, caudal. Scales omitted from objects A-C, E, F but not from eyeplate in panel C; pore diameters not to scale in C, D." figureDoi="10.3897/zookeys.1079.76412.figure4" httpUri="https://binary.pensoft.net/fig/629314" pageId="0" pageNumber="145">4C</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Structure of eyeplates in Pseudomma kryotroglodytum sp. nov. and Michthyops parvus. Holotype adult female (D) and paratype subadult female (A, B) of P. kryotroglodytum and non-type adult female of M. parvus (C). A eyeplate expanded on slide, dorsal, to the right somewhat distorted B detail of panel (A) showing cyst connected with bottom of median cleft C homologous cyst in another species and genus, arrow points to tubular connection with anterior margin of eyeplate D series of denticles along sub-lateral section of anterior margin of eyeplate." figureDoi="10.3897/zookeys.1079.76412.figure23" httpUri="https://binary.pensoft.net/fig/629333" pageId="0" pageNumber="145">23A, B, D</figureCitation>
|
||
). Eyeplate extending 0.9 times the length of terminal segment of antennular trunk along mid-line beyond anterior margin of carapace. Length of eyeplate, including its dorsally covered portion, 1.3 times the length of terminal segment. In dorsal view, superimposed dorsal and ventral sulci separate cornea-like lateral portions from main part of eyeplate (Figs
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pseudomma kryotroglodytum sp. nov. from ice cave of Bernard Island, Antarctica A female holotype, lateral B cephalothorax of female holotype, dorsal C physical aspect of the ice cave environment A, B living specimen in laboratory." figureDoi="10.3897/zookeys.1079.76412.figure2" httpUri="https://binary.pensoft.net/fig/629312" pageId="0" pageNumber="145">2B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A, B, E, F) and paratype subadult female 21.5 mm (C, D). A right antennula, dorsal B antenna with antennal gland, ventral, setae omitted from antennal scale C eyeplate and carapace expanded on slide, short arrow points to distolateral edge of carapace, detail (D) pore group in pre-cardial position E mandibles with left palpus, caudal aspect F maxillula, caudal. Scales omitted from objects A-C, E, F but not from eyeplate in panel C; pore diameters not to scale in C, D." figureDoi="10.3897/zookeys.1079.76412.figure4" httpUri="https://binary.pensoft.net/fig/629314" pageId="0" pageNumber="145">4C</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Structure of eyeplates in Pseudomma kryotroglodytum sp. nov. and Michthyops parvus. Holotype adult female (D) and paratype subadult female (A, B) of P. kryotroglodytum and non-type adult female of M. parvus (C). A eyeplate expanded on slide, dorsal, to the right somewhat distorted B detail of panel (A) showing cyst connected with bottom of median cleft C homologous cyst in another species and genus, arrow points to tubular connection with anterior margin of eyeplate D series of denticles along sub-lateral section of anterior margin of eyeplate." figureDoi="10.3897/zookeys.1079.76412.figure23" httpUri="https://binary.pensoft.net/fig/629333" pageId="0" pageNumber="145">23A</figureCitation>
|
||
). Eyeplate containing tear-shaped cyst narrowing distally up to conjunction with eyeplate cleft (Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Structure of eyeplates in Pseudomma kryotroglodytum sp. nov. and Michthyops parvus. Holotype adult female (D) and paratype subadult female (A, B) of P. kryotroglodytum and non-type adult female of M. parvus (C). A eyeplate expanded on slide, dorsal, to the right somewhat distorted B detail of panel (A) showing cyst connected with bottom of median cleft C homologous cyst in another species and genus, arrow points to tubular connection with anterior margin of eyeplate D series of denticles along sub-lateral section of anterior margin of eyeplate." figureDoi="10.3897/zookeys.1079.76412.figure23" httpUri="https://binary.pensoft.net/fig/629333" pageId="0" pageNumber="145">23B</figureCitation>
|
||
). Sub-lateral portions of dorsal face with cover of minute scales (as in Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E) and paratype subadult female 21.5 mm (F) A terminal portion of telson, dorsal, details show scales on left disto-mesial spine (B) and barbs on left terminal seta (C) D anterior half of cephalothorax, lateral, arrow points to distolateral edge of carapace E sixth pleomere with tail fan, lateral F example for pores (three to the left) and coat of scales on tergite of first pleomere." figureDoi="10.3897/zookeys.1079.76412.figure3" httpUri="https://binary.pensoft.net/fig/629313" pageId="0" pageNumber="145">3F</figureCitation>
|
||
; visualised with 400
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
microscopy), series of 15 minute teeth (not all in focus in Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Structure of eyeplates in Pseudomma kryotroglodytum sp. nov. and Michthyops parvus. Holotype adult female (D) and paratype subadult female (A, B) of P. kryotroglodytum and non-type adult female of M. parvus (C). A eyeplate expanded on slide, dorsal, to the right somewhat distorted B detail of panel (A) showing cyst connected with bottom of median cleft C homologous cyst in another species and genus, arrow points to tubular connection with anterior margin of eyeplate D series of denticles along sub-lateral section of anterior margin of eyeplate." figureDoi="10.3897/zookeys.1079.76412.figure23" httpUri="https://binary.pensoft.net/fig/629333" pageId="0" pageNumber="145">23D</figureCitation>
|
||
), closely set along anterior margin of this portion. Brilliant red cornea-like portions of eyeplate feign functional eyes in living specimens (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pseudomma kryotroglodytum sp. nov. from ice cave of Bernard Island, Antarctica A female holotype, lateral B cephalothorax of female holotype, dorsal C physical aspect of the ice cave environment A, B living specimen in laboratory." figureDoi="10.3897/zookeys.1079.76412.figure2" httpUri="https://binary.pensoft.net/fig/629312" pageId="0" pageNumber="145">2B</figureCitation>
|
||
). Eyeplates become transparent (Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A, B, E, F) and paratype subadult female 21.5 mm (C, D). A right antennula, dorsal B antenna with antennal gland, ventral, setae omitted from antennal scale C eyeplate and carapace expanded on slide, short arrow points to distolateral edge of carapace, detail (D) pore group in pre-cardial position E mandibles with left palpus, caudal aspect F maxillula, caudal. Scales omitted from objects A-C, E, F but not from eyeplate in panel C; pore diameters not to scale in C, D." figureDoi="10.3897/zookeys.1079.76412.figure4" httpUri="https://binary.pensoft.net/fig/629314" pageId="0" pageNumber="145">4C</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Structure of eyeplates in Pseudomma kryotroglodytum sp. nov. and Michthyops parvus. Holotype adult female (D) and paratype subadult female (A, B) of P. kryotroglodytum and non-type adult female of M. parvus (C). A eyeplate expanded on slide, dorsal, to the right somewhat distorted B detail of panel (A) showing cyst connected with bottom of median cleft C homologous cyst in another species and genus, arrow points to tubular connection with anterior margin of eyeplate D series of denticles along sub-lateral section of anterior margin of eyeplate." figureDoi="10.3897/zookeys.1079.76412.figure23" httpUri="https://binary.pensoft.net/fig/629333" pageId="0" pageNumber="145">23A</figureCitation>
|
||
) after expansion on slide, embedding in Swan-medium and resultant bleaching; neither functional nor both vestigial ommatidia and neuronal structures visible.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1079.76412.figure3" httpUri="https://binary.pensoft.net/fig/629313" pageId="0" pageNumber="145" start="Figure 3" startId="F3">
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">Figure 3.</emphasis>
|
||
<taxonomicName authorityName="Wittmann & Chevaldonné" authorityYear="2021" class="Malacostraca" family="Mysidae" genus="Pseudomma" kingdom="Animalia" lsidName="Pseudomma kryotroglodytum" order="Mysidacea" pageId="0" pageNumber="145" phylum="Schizopoda" rank="species" species="kryotroglodytum">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Pseudomma kryotroglodytum</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype adult female BL 26.8 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="145">A-E</emphasis>
|
||
) and paratype subadult female 21.5 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="145">F</emphasis>
|
||
)
|
||
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
|
||
terminal portion of telson, dorsal, details show scales on left disto-mesial spine (
|
||
<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis>
|
||
) and barbs on left terminal seta (
|
||
<emphasis bold="true" pageId="0" pageNumber="145">C</emphasis>
|
||
)
|
||
<emphasis bold="true" pageId="0" pageNumber="145">D</emphasis>
|
||
anterior half of cephalothorax, lateral, arrow points to distolateral edge of carapace
|
||
<emphasis bold="true" pageId="0" pageNumber="145">E</emphasis>
|
||
sixth pleomere with tail fan, lateral
|
||
<emphasis bold="true" pageId="0" pageNumber="145">F</emphasis>
|
||
example for pores (three to the left) and coat of scales on tergite of first pleomere.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Carapace</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E) and paratype subadult female 21.5 mm (F) A terminal portion of telson, dorsal, details show scales on left disto-mesial spine (B) and barbs on left terminal seta (C) D anterior half of cephalothorax, lateral, arrow points to distolateral edge of carapace E sixth pleomere with tail fan, lateral F example for pores (three to the left) and coat of scales on tergite of first pleomere." figureDoi="10.3897/zookeys.1079.76412.figure3" httpUri="https://binary.pensoft.net/fig/629313" pageId="0" pageNumber="145">3D</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A, B, E, F) and paratype subadult female 21.5 mm (C, D). A right antennula, dorsal B antenna with antennal gland, ventral, setae omitted from antennal scale C eyeplate and carapace expanded on slide, short arrow points to distolateral edge of carapace, detail (D) pore group in pre-cardial position E mandibles with left palpus, caudal aspect F maxillula, caudal. Scales omitted from objects A-C, E, F but not from eyeplate in panel C; pore diameters not to scale in C, D." figureDoi="10.3897/zookeys.1079.76412.figure4" httpUri="https://binary.pensoft.net/fig/629314" pageId="0" pageNumber="145">4C</figureCitation>
|
||
) with broadly rounded anterior margin, disto-lateral edges well rounded. No typical rostral plate present, but a frontal bulge dorsally covered by the carapace; bulge best seen in lateral view (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E) and paratype subadult female 21.5 mm (F) A terminal portion of telson, dorsal, details show scales on left disto-mesial spine (B) and barbs on left terminal seta (C) D anterior half of cephalothorax, lateral, arrow points to distolateral edge of carapace E sixth pleomere with tail fan, lateral F example for pores (three to the left) and coat of scales on tergite of first pleomere." figureDoi="10.3897/zookeys.1079.76412.figure3" httpUri="https://binary.pensoft.net/fig/629313" pageId="0" pageNumber="145">3D</figureCitation>
|
||
). Antero-lateral edge of carapace with rounded protrusion (marked by arrows in Figs
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E) and paratype subadult female 21.5 mm (F) A terminal portion of telson, dorsal, details show scales on left disto-mesial spine (B) and barbs on left terminal seta (C) D anterior half of cephalothorax, lateral, arrow points to distolateral edge of carapace E sixth pleomere with tail fan, lateral F example for pores (three to the left) and coat of scales on tergite of first pleomere." figureDoi="10.3897/zookeys.1079.76412.figure3" httpUri="https://binary.pensoft.net/fig/629313" pageId="0" pageNumber="145">3D</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A, B, E, F) and paratype subadult female 21.5 mm (C, D). A right antennula, dorsal B antenna with antennal gland, ventral, setae omitted from antennal scale C eyeplate and carapace expanded on slide, short arrow points to distolateral edge of carapace, detail (D) pore group in pre-cardial position E mandibles with left palpus, caudal aspect F maxillula, caudal. Scales omitted from objects A-C, E, F but not from eyeplate in panel C; pore diameters not to scale in C, D." figureDoi="10.3897/zookeys.1079.76412.figure4" httpUri="https://binary.pensoft.net/fig/629314" pageId="0" pageNumber="145">4C</figureCitation>
|
||
). By forcing the detached carapace in a plane (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A, B, E, F) and paratype subadult female 21.5 mm (C, D). A right antennula, dorsal B antenna with antennal gland, ventral, setae omitted from antennal scale C eyeplate and carapace expanded on slide, short arrow points to distolateral edge of carapace, detail (D) pore group in pre-cardial position E mandibles with left palpus, caudal aspect F maxillula, caudal. Scales omitted from objects A-C, E, F but not from eyeplate in panel C; pore diameters not to scale in C, D." figureDoi="10.3897/zookeys.1079.76412.figure4" httpUri="https://binary.pensoft.net/fig/629314" pageId="0" pageNumber="145">4C</figureCitation>
|
||
), this protrusion becoming shifted caudally compared with its position
|
||
<emphasis italics="true" pageId="0" pageNumber="145">in situ</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E) and paratype subadult female 21.5 mm (F) A terminal portion of telson, dorsal, details show scales on left disto-mesial spine (B) and barbs on left terminal seta (C) D anterior half of cephalothorax, lateral, arrow points to distolateral edge of carapace E sixth pleomere with tail fan, lateral F example for pores (three to the left) and coat of scales on tergite of first pleomere." figureDoi="10.3897/zookeys.1079.76412.figure3" httpUri="https://binary.pensoft.net/fig/629313" pageId="0" pageNumber="145">3D</figureCitation>
|
||
). Carapace with cervical sulcus and cardial sulcus distinct; posterior margin concave, terminal indentation widely triangular. Two submedian groups of 8-10 pores symmetrically arranged directly in front of cardial sulcus (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A, B, E, F) and paratype subadult female 21.5 mm (C, D). A right antennula, dorsal B antenna with antennal gland, ventral, setae omitted from antennal scale C eyeplate and carapace expanded on slide, short arrow points to distolateral edge of carapace, detail (D) pore group in pre-cardial position E mandibles with left palpus, caudal aspect F maxillula, caudal. Scales omitted from objects A-C, E, F but not from eyeplate in panel C; pore diameters not to scale in C, D." figureDoi="10.3897/zookeys.1079.76412.figure4" httpUri="https://binary.pensoft.net/fig/629314" pageId="0" pageNumber="145">4D</figureCitation>
|
||
shows only seven pores in the smaller paratype). Carapace leaving posterior 1.5 thoracomeres dorsally exposed.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Mouthparts</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A, B, E, F) and paratype subadult female 21.5 mm (C, D). A right antennula, dorsal B antenna with antennal gland, ventral, setae omitted from antennal scale C eyeplate and carapace expanded on slide, short arrow points to distolateral edge of carapace, detail (D) pore group in pre-cardial position E mandibles with left palpus, caudal aspect F maxillula, caudal. Scales omitted from objects A-C, E, F but not from eyeplate in panel C; pore diameters not to scale in C, D." figureDoi="10.3897/zookeys.1079.76412.figure4" httpUri="https://binary.pensoft.net/fig/629314" pageId="0" pageNumber="145">4E, F</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm A labrum B labium C maxilla, caudal aspect D thoracopod 1 (caudal) with thoracic sternites 1, 2 (ventral) E detail of panel (D) showing dactylus 1 with nail F thoracic endopod 2, rostral G thoracic endopod 4, rostral H detail of panel (G) showing dactylus 4 with nail I inner face of oostegite 1, drawn above part of sympod 6. Scales omitted from objects A-D, F, G; setae from E, H." figureDoi="10.3897/zookeys.1079.76412.figure5" httpUri="https://binary.pensoft.net/fig/629315" pageId="0" pageNumber="145">5A-C</figureCitation>
|
||
). Labrum normal (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm A labrum B labium C maxilla, caudal aspect D thoracopod 1 (caudal) with thoracic sternites 1, 2 (ventral) E detail of panel (D) showing dactylus 1 with nail F thoracic endopod 2, rostral G thoracic endopod 4, rostral H detail of panel (G) showing dactylus 4 with nail I inner face of oostegite 1, drawn above part of sympod 6. Scales omitted from objects A-D, F, G; setae from E, H." figureDoi="10.3897/zookeys.1079.76412.figure5" httpUri="https://binary.pensoft.net/fig/629315" pageId="0" pageNumber="145">5A</figureCitation>
|
||
), rostrally forming a broad, rounded bulge; most caudal portions with strong lamellae and cover of scale-like fringes. Basal segment of mandibular palp (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A, B, E, F) and paratype subadult female 21.5 mm (C, D). A right antennula, dorsal B antenna with antennal gland, ventral, setae omitted from antennal scale C eyeplate and carapace expanded on slide, short arrow points to distolateral edge of carapace, detail (D) pore group in pre-cardial position E mandibles with left palpus, caudal aspect F maxillula, caudal. Scales omitted from objects A-C, E, F but not from eyeplate in panel C; pore diameters not to scale in C, D." figureDoi="10.3897/zookeys.1079.76412.figure4" httpUri="https://binary.pensoft.net/fig/629314" pageId="0" pageNumber="145">4E</figureCitation>
|
||
) contributing 9-10%, median segment 55-56% and apical segment 35-37% to total palp length. Length of median segment 3.1-3.2 times maximum width; its mesial margin convex, lateral margin sigmoid. Length of apical segment 3.7-3.8 times maximum width. Palp not hispid, its basal segment without setae, remaining segments densely setose along mesial and lateral margins. Caudal face of median segment with dense field of fine hairs near basis. Masticatory part of mandibles strong, asymmetrical. Left mandible as normal in
|
||
<taxonomicName authorityName="Haworth" authorityYear="1825" class="Malacostraca" family="Mysidae" kingdom="Animalia" lsidName="" order="Mysida" pageId="0" pageNumber="145" phylum="Arthropoda" rank="family">Mysidae</taxonomicName>
|
||
. Pars incisiva of the new species with three large teeth and digitus mobilis with four strong teeth. Spine row with four spines
|
||
<normalizedToken originalValue="‘serrated’">'serrated'</normalizedToken>
|
||
by numerous stiff bristles; processus molaris with grinding lamellae not ending in teeth and with dense cover of stiff bristles. Right mandible as normal in the genus
|
||
<taxonomicName authorityName="G. O. Sars" authorityYear="1870" class="Malacostraca" family="Mysidae" genus="Pseudomma" kingdom="Animalia" lsidName="Pseudomma" order="Mysidacea" pageId="0" pageNumber="145" phylum="Schizopoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Pseudomma</emphasis>
|
||
</taxonomicName>
|
||
; in the new species with four large teeth on pars incisiva; digitus mobilis with only one large apical tooth serrated by secondary teeth. Right spine row present as series of nine medium-sized smooth teeth plus a few small ones, rather than a smaller number of
|
||
<normalizedToken originalValue="‘serrated’">'serrated'</normalizedToken>
|
||
spines present on the left mandible as otherwise usual for both mandibles in
|
||
<taxonomicName authorityName="Haworth" authorityYear="1825" class="Malacostraca" family="Mysidae" kingdom="Animalia" lsidName="" order="Mysida" pageId="0" pageNumber="145" phylum="Arthropoda" rank="family">Mysidae</taxonomicName>
|
||
. Right processus molaris with strong masticatory lamellae, each with small, tooth-like, apical projection; processus with cover of stiff bristles less dense than that of left mandible.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
Paired labia (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm A labrum B labium C maxilla, caudal aspect D thoracopod 1 (caudal) with thoracic sternites 1, 2 (ventral) E detail of panel (D) showing dactylus 1 with nail F thoracic endopod 2, rostral G thoracic endopod 4, rostral H detail of panel (G) showing dactylus 4 with nail I inner face of oostegite 1, drawn above part of sympod 6. Scales omitted from objects A-D, F, G; setae from E, H." figureDoi="10.3897/zookeys.1079.76412.figure5" httpUri="https://binary.pensoft.net/fig/629315" pageId="0" pageNumber="145">5B</figureCitation>
|
||
) with stiff setae, lacking spines or teeth. Distal segment of maxillula (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A, B, E, F) and paratype subadult female 21.5 mm (C, D). A right antennula, dorsal B antenna with antennal gland, ventral, setae omitted from antennal scale C eyeplate and carapace expanded on slide, short arrow points to distolateral edge of carapace, detail (D) pore group in pre-cardial position E mandibles with left palpus, caudal aspect F maxillula, caudal. Scales omitted from objects A-C, E, F but not from eyeplate in panel C; pore diameters not to scale in C, D." figureDoi="10.3897/zookeys.1079.76412.figure4" httpUri="https://binary.pensoft.net/fig/629314" pageId="0" pageNumber="145">4F</figureCitation>
|
||
) with 11-12 weakly serrated, strong spines on terminally truncate margin; subterminally with 5-6 barbed setae. Holotype with 5-9 pores on the surface between setae bases and spines; potential additional pores may be covered by these setae (no pores identified in the paratype). Endite of maxillula with numerous normal setae; distally with three large, modified setae, armed with stiff bristles near apex¸ more proximally with a shorter seta bearing an apical brush of long bristles.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1079.76412.figure4" httpUri="https://binary.pensoft.net/fig/629314" pageId="0" pageNumber="145" start="Figure 4" startId="F4">
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">Figure 4.</emphasis>
|
||
<taxonomicName authorityName="Wittmann & Chevaldonné" authorityYear="2021" class="Malacostraca" family="Mysidae" genus="Pseudomma" kingdom="Animalia" lsidName="Pseudomma kryotroglodytum" order="Mysidacea" pageId="0" pageNumber="145" phylum="Schizopoda" rank="species" species="kryotroglodytum">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Pseudomma kryotroglodytum</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype adult female BL 26.8 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="145">A, B, E, F</emphasis>
|
||
) and paratype subadult female 21.5 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="145">C, D</emphasis>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
|
||
right antennula, dorsal
|
||
<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis>
|
||
antenna with antennal gland, ventral, setae omitted from antennal scale
|
||
<emphasis bold="true" pageId="0" pageNumber="145">C</emphasis>
|
||
eyeplate and carapace expanded on slide, short arrow points to distolateral edge of carapace, detail (
|
||
<emphasis bold="true" pageId="0" pageNumber="145">D</emphasis>
|
||
) pore group in pre-cardial position
|
||
<emphasis bold="true" pageId="0" pageNumber="145">E</emphasis>
|
||
mandibles with left palpus, caudal aspect
|
||
<emphasis bold="true" pageId="0" pageNumber="145">F</emphasis>
|
||
maxillula, caudal. Scales omitted from objects
|
||
<emphasis bold="true" pageId="0" pageNumber="145">A-C, E, F</emphasis>
|
||
but not from eyeplate in panel
|
||
<emphasis bold="true" pageId="0" pageNumber="145">C</emphasis>
|
||
; pore diameters not to scale in
|
||
<emphasis bold="true" pageId="0" pageNumber="145">C, D</emphasis>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Maxilla</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm A labrum B labium C maxilla, caudal aspect D thoracopod 1 (caudal) with thoracic sternites 1, 2 (ventral) E detail of panel (D) showing dactylus 1 with nail F thoracic endopod 2, rostral G thoracic endopod 4, rostral H detail of panel (G) showing dactylus 4 with nail I inner face of oostegite 1, drawn above part of sympod 6. Scales omitted from objects A-D, F, G; setae from E, H." figureDoi="10.3897/zookeys.1079.76412.figure5" httpUri="https://binary.pensoft.net/fig/629315" pageId="0" pageNumber="145">5C</figureCitation>
|
||
) with well-developed exopod, two-segmented endopod (palp) and four setose endites. Exopod normal-sized, shortly extending beyond basal segment of endopod. Outer margin and apex of exopod with dense series of plumose setae distally increasing in size. Length of apical segment of endopod 1.5-1.6 times its maximum width and 1.2-1.3 times length of the basal segment. Basal segment with total of eight barbed setae, namely 3-4 on caudal and 4-5 on rostral face. Apical segment with setae all around, except for proximal third of lateral margin.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Foregut</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E, H-N) and paratype subadult female 21.5 mm (F, G). A cardiac portion of foregut, dorsal view, dorsal wall omitted B-D details of panel (A) showing modified spines E details of panel (A) showing teeth emerging from a common basis F pleonite 1, cuticle detached and expanded on slide, dorsal and lateral faces on top, ventral face folded in, setae omitted from pleopods G detail of panel (F) showing pore group on left lateral face H-L series of pleopods 1 - 5, rostral = lateral face M uropods, ventral N telson. Scales omitted from objects F, M, N pore diameters not to scale in F, G." figureDoi="10.3897/zookeys.1079.76412.figure6" httpUri="https://binary.pensoft.net/fig/629316" pageId="0" pageNumber="145">6A-E</figureCitation>
|
||
). Lateralia, infoldings and superomedianum of the cardiac chamber densely covered by smooth, slender setae and spines. Lateralia anteriorly with dense series of slender, apically coronate spines (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E, H-N) and paratype subadult female 21.5 mm (F, G). A cardiac portion of foregut, dorsal view, dorsal wall omitted B-D details of panel (A) showing modified spines E details of panel (A) showing teeth emerging from a common basis F pleonite 1, cuticle detached and expanded on slide, dorsal and lateral faces on top, ventral face folded in, setae omitted from pleopods G detail of panel (F) showing pore group on left lateral face H-L series of pleopods 1 - 5, rostral = lateral face M uropods, ventral N telson. Scales omitted from objects F, M, N pore diameters not to scale in F, G." figureDoi="10.3897/zookeys.1079.76412.figure6" httpUri="https://binary.pensoft.net/fig/629316" pageId="0" pageNumber="145">6B</figureCitation>
|
||
) of different length, more caudally with slender acute spines (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E, H-N) and paratype subadult female 21.5 mm (F, G). A cardiac portion of foregut, dorsal view, dorsal wall omitted B-D details of panel (A) showing modified spines E details of panel (A) showing teeth emerging from a common basis F pleonite 1, cuticle detached and expanded on slide, dorsal and lateral faces on top, ventral face folded in, setae omitted from pleopods G detail of panel (F) showing pore group on left lateral face H-L series of pleopods 1 - 5, rostral = lateral face M uropods, ventral N telson. Scales omitted from objects F, M, N pore diameters not to scale in F, G." figureDoi="10.3897/zookeys.1079.76412.figure6" httpUri="https://binary.pensoft.net/fig/629316" pageId="0" pageNumber="145">6C</figureCitation>
|
||
). The latter spines with minute teeth on and close to apex. Both coronate and acute spines hispid due to minute scales along distal 50-70% of shaft. Posterior part of lateralia with powerful complex of many blunt teeth arising from common base. Twelve teeth differentiated (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E, H-N) and paratype subadult female 21.5 mm (F, G). A cardiac portion of foregut, dorsal view, dorsal wall omitted B-D details of panel (A) showing modified spines E details of panel (A) showing teeth emerging from a common basis F pleonite 1, cuticle detached and expanded on slide, dorsal and lateral faces on top, ventral face folded in, setae omitted from pleopods G detail of panel (F) showing pore group on left lateral face H-L series of pleopods 1 - 5, rostral = lateral face M uropods, ventral N telson. Scales omitted from objects F, M, N pore diameters not to scale in F, G." figureDoi="10.3897/zookeys.1079.76412.figure6" httpUri="https://binary.pensoft.net/fig/629316" pageId="0" pageNumber="145">6E</figureCitation>
|
||
) with translucent microscopy, additional teeth not excluded. Dorsolateral infoldings with two smooth, bent spines (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E, H-N) and paratype subadult female 21.5 mm (F, G). A cardiac portion of foregut, dorsal view, dorsal wall omitted B-D details of panel (A) showing modified spines E details of panel (A) showing teeth emerging from a common basis F pleonite 1, cuticle detached and expanded on slide, dorsal and lateral faces on top, ventral face folded in, setae omitted from pleopods G detail of panel (F) showing pore group on left lateral face H-L series of pleopods 1 - 5, rostral = lateral face M uropods, ventral N telson. Scales omitted from objects F, M, N pore diameters not to scale in F, G." figureDoi="10.3897/zookeys.1079.76412.figure6" httpUri="https://binary.pensoft.net/fig/629316" pageId="0" pageNumber="145">6D</figureCitation>
|
||
, subapically slightly serrate only in paratype). About 2/3 of comparatively large storage volume of foregut contained masticated, unidentifiable organic materials and mineral particles, also observed in paratype.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Thorax</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pseudomma kryotroglodytum sp. nov. from ice cave of Bernard Island, Antarctica A female holotype, lateral B cephalothorax of female holotype, dorsal C physical aspect of the ice cave environment A, B living specimen in laboratory." figureDoi="10.3897/zookeys.1079.76412.figure2" httpUri="https://binary.pensoft.net/fig/629312" pageId="0" pageNumber="145">2A, B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm A labrum B labium C maxilla, caudal aspect D thoracopod 1 (caudal) with thoracic sternites 1, 2 (ventral) E detail of panel (D) showing dactylus 1 with nail F thoracic endopod 2, rostral G thoracic endopod 4, rostral H detail of panel (G) showing dactylus 4 with nail I inner face of oostegite 1, drawn above part of sympod 6. Scales omitted from objects A-D, F, G; setae from E, H." figureDoi="10.3897/zookeys.1079.76412.figure5" httpUri="https://binary.pensoft.net/fig/629315" pageId="0" pageNumber="145">5D-I</figureCitation>
|
||
). At least tergites 6-8 covered by minute scales, no pores. Sternites 1-8 without pores, scales and also without ventrally projecting median processes (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm A labrum B labium C maxilla, caudal aspect D thoracopod 1 (caudal) with thoracic sternites 1, 2 (ventral) E detail of panel (D) showing dactylus 1 with nail F thoracic endopod 2, rostral G thoracic endopod 4, rostral H detail of panel (G) showing dactylus 4 with nail I inner face of oostegite 1, drawn above part of sympod 6. Scales omitted from objects A-D, F, G; setae from E, H." figureDoi="10.3897/zookeys.1079.76412.figure5" httpUri="https://binary.pensoft.net/fig/629315" pageId="0" pageNumber="145">5D</figureCitation>
|
||
). Sternite 1 with short anterior lobe projecting between left and right, first thoracic endopods. Basal plates of thoracic exopods (3-4) times longer than wide (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm A labrum B labium C maxilla, caudal aspect D thoracopod 1 (caudal) with thoracic sternites 1, 2 (ventral) E detail of panel (D) showing dactylus 1 with nail F thoracic endopod 2, rostral G thoracic endopod 4, rostral H detail of panel (G) showing dactylus 4 with nail I inner face of oostegite 1, drawn above part of sympod 6. Scales omitted from objects A-D, F, G; setae from E, H." figureDoi="10.3897/zookeys.1079.76412.figure5" httpUri="https://binary.pensoft.net/fig/629315" pageId="0" pageNumber="145">5D</figureCitation>
|
||
), not widening distally or only minimally so; lateral portions scaly-hispid, mesially smooth; disto-lateral edge unevenly rounded. Exopods 1, 8 with 15-segmented (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm A labrum B labium C maxilla, caudal aspect D thoracopod 1 (caudal) with thoracic sternites 1, 2 (ventral) E detail of panel (D) showing dactylus 1 with nail F thoracic endopod 2, rostral G thoracic endopod 4, rostral H detail of panel (G) showing dactylus 4 with nail I inner face of oostegite 1, drawn above part of sympod 6. Scales omitted from objects A-D, F, G; setae from E, H." figureDoi="10.3897/zookeys.1079.76412.figure5" httpUri="https://binary.pensoft.net/fig/629315" pageId="0" pageNumber="145">5D</figureCitation>
|
||
), remaining exopods with (17-20)-segmented, setose flagellum. Basis of endopod 1 with setose endite (below drawing plane in Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm A labrum B labium C maxilla, caudal aspect D thoracopod 1 (caudal) with thoracic sternites 1, 2 (ventral) E detail of panel (D) showing dactylus 1 with nail F thoracic endopod 2, rostral G thoracic endopod 4, rostral H detail of panel (G) showing dactylus 4 with nail I inner face of oostegite 1, drawn above part of sympod 6. Scales omitted from objects A-D, F, G; setae from E, H." figureDoi="10.3897/zookeys.1079.76412.figure5" httpUri="https://binary.pensoft.net/fig/629315" pageId="0" pageNumber="145">5D</figureCitation>
|
||
), remaining segments without clear endite. Endopods 1, 2 with six segments (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm A labrum B labium C maxilla, caudal aspect D thoracopod 1 (caudal) with thoracic sternites 1, 2 (ventral) E detail of panel (D) showing dactylus 1 with nail F thoracic endopod 2, rostral G thoracic endopod 4, rostral H detail of panel (G) showing dactylus 4 with nail I inner face of oostegite 1, drawn above part of sympod 6. Scales omitted from objects A-D, F, G; setae from E, H." figureDoi="10.3897/zookeys.1079.76412.figure5" httpUri="https://binary.pensoft.net/fig/629315" pageId="0" pageNumber="145">5D, F</figureCitation>
|
||
), remaining endopods with eight segments counting from basis to dactylus (basis omitted in Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm A labrum B labium C maxilla, caudal aspect D thoracopod 1 (caudal) with thoracic sternites 1, 2 (ventral) E detail of panel (D) showing dactylus 1 with nail F thoracic endopod 2, rostral G thoracic endopod 4, rostral H detail of panel (G) showing dactylus 4 with nail I inner face of oostegite 1, drawn above part of sympod 6. Scales omitted from objects A-D, F, G; setae from E, H." figureDoi="10.3897/zookeys.1079.76412.figure5" httpUri="https://binary.pensoft.net/fig/629315" pageId="0" pageNumber="145">5G</figureCitation>
|
||
). Endopods 3-8 long and slender; length and slenderness increase caudally; ischium shows the strongest relative increase (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pseudomma kryotroglodytum sp. nov. from ice cave of Bernard Island, Antarctica A female holotype, lateral B cephalothorax of female holotype, dorsal C physical aspect of the ice cave environment A, B living specimen in laboratory." figureDoi="10.3897/zookeys.1079.76412.figure2" httpUri="https://binary.pensoft.net/fig/629312" pageId="0" pageNumber="145">2A</figureCitation>
|
||
). All endopods with hispid carpopropodus and dactylus, endopods 2-8 in addition with hispid merus, to a minor degree, if any, also ischium. Carpopropodus 3-8 three-segmented; more than half its length contributed by basal segment. Setae patterns might feign additional subdivision of carpopropodus (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm A labrum B labium C maxilla, caudal aspect D thoracopod 1 (caudal) with thoracic sternites 1, 2 (ventral) E detail of panel (D) showing dactylus 1 with nail F thoracic endopod 2, rostral G thoracic endopod 4, rostral H detail of panel (G) showing dactylus 4 with nail I inner face of oostegite 1, drawn above part of sympod 6. Scales omitted from objects A-D, F, G; setae from E, H." figureDoi="10.3897/zookeys.1079.76412.figure5" httpUri="https://binary.pensoft.net/fig/629315" pageId="0" pageNumber="145">5G</figureCitation>
|
||
). Suture between median and terminal segment transverse in carpopropodus 3, weakly (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm A labrum B labium C maxilla, caudal aspect D thoracopod 1 (caudal) with thoracic sternites 1, 2 (ventral) E detail of panel (D) showing dactylus 1 with nail F thoracic endopod 2, rostral G thoracic endopod 4, rostral H detail of panel (G) showing dactylus 4 with nail I inner face of oostegite 1, drawn above part of sympod 6. Scales omitted from objects A-D, F, G; setae from E, H." figureDoi="10.3897/zookeys.1079.76412.figure5" httpUri="https://binary.pensoft.net/fig/629315" pageId="0" pageNumber="145">5G</figureCitation>
|
||
), but not always distinctly oblique in carpopropodus 4, weakly and distinctly oblique in carpopropodites 5-8. Dactylus 3-8 small. Dactylus 1, 3-8 with short, smooth claw (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm A labrum B labium C maxilla, caudal aspect D thoracopod 1 (caudal) with thoracic sternites 1, 2 (ventral) E detail of panel (D) showing dactylus 1 with nail F thoracic endopod 2, rostral G thoracic endopod 4, rostral H detail of panel (G) showing dactylus 4 with nail I inner face of oostegite 1, drawn above part of sympod 6. Scales omitted from objects A-D, F, G; setae from E, H." figureDoi="10.3897/zookeys.1079.76412.figure5" httpUri="https://binary.pensoft.net/fig/629315" pageId="0" pageNumber="145">5E, H</figureCitation>
|
||
). No claw detected in dense jungle of setae on dactylus 2.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Marsupium</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm A labrum B labium C maxilla, caudal aspect D thoracopod 1 (caudal) with thoracic sternites 1, 2 (ventral) E detail of panel (D) showing dactylus 1 with nail F thoracic endopod 2, rostral G thoracic endopod 4, rostral H detail of panel (G) showing dactylus 4 with nail I inner face of oostegite 1, drawn above part of sympod 6. Scales omitted from objects A-D, F, G; setae from E, H." figureDoi="10.3897/zookeys.1079.76412.figure5" httpUri="https://binary.pensoft.net/fig/629315" pageId="0" pageNumber="145">5I</figureCitation>
|
||
) empty in this specimen. Basal to median portions of dorsal margin without setae in oostegite 1, without setae from basal to subapical portions in oostegites 2 and 3. A narrow
|
||
<normalizedToken originalValue="‘fur’">'fur'</normalizedToken>
|
||
of densely set tiny hairs forming a ribbon along dorsal margin of oostegite 1 (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm A labrum B labium C maxilla, caudal aspect D thoracopod 1 (caudal) with thoracic sternites 1, 2 (ventral) E detail of panel (D) showing dactylus 1 with nail F thoracic endopod 2, rostral G thoracic endopod 4, rostral H detail of panel (G) showing dactylus 4 with nail I inner face of oostegite 1, drawn above part of sympod 6. Scales omitted from objects A-D, F, G; setae from E, H." figureDoi="10.3897/zookeys.1079.76412.figure5" httpUri="https://binary.pensoft.net/fig/629315" pageId="0" pageNumber="145">5I</figureCitation>
|
||
), no such hairs in oostegites 2 and 3. Scales on large portions of outer face in oostegites 2 and 3; no scales seen on oostegite 1. Ventral and anterior margins plus part of posterior margin with dense series of barbed setae, together with bilaterally opposite oostegite forming gate contributing to the ventral and caudal closure of marsupium (this configuration not impeding respiratory water flow through marsupium). Numbers of barbs per seta increase distally, albeit not reaching those of typical plumose seta. Oostegite 1 with barbed setae also on distal half of dorsal margin, suggesting pervious anterior closure of marsupium. Posterior parts of oostegites 1-3 on inner face with comparatively long setae microserrated on their distal half. Outer face of oostegites 2 and 3 with slender whip setae characterised by barbed shaft bearing longer thin flagellum. These setae implanted on distal portions of oostegites 2 and 3; additional whip setae along ventral margin only in oostegite 3. No whip setae in oostegite 1.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1079.76412.figure5" httpUri="https://binary.pensoft.net/fig/629315" pageId="0" pageNumber="145" start="Figure 5" startId="F5">
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">Figure 5.</emphasis>
|
||
<taxonomicName authorityName="Wittmann & Chevaldonné" authorityYear="2021" class="Malacostraca" family="Mysidae" genus="Pseudomma" kingdom="Animalia" lsidName="Pseudomma kryotroglodytum" order="Mysidacea" pageId="0" pageNumber="145" phylum="Schizopoda" rank="species" species="kryotroglodytum">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Pseudomma kryotroglodytum</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype adult female BL 26.8 mm
|
||
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
|
||
labrum
|
||
<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis>
|
||
labium
|
||
<emphasis bold="true" pageId="0" pageNumber="145">C</emphasis>
|
||
maxilla, caudal aspect
|
||
<emphasis bold="true" pageId="0" pageNumber="145">D</emphasis>
|
||
thoracopod 1 (caudal) with thoracic sternites 1, 2 (ventral)
|
||
<emphasis bold="true" pageId="0" pageNumber="145">E</emphasis>
|
||
detail of panel (
|
||
<emphasis bold="true" pageId="0" pageNumber="145">D</emphasis>
|
||
) showing dactylus 1 with nail
|
||
<emphasis bold="true" pageId="0" pageNumber="145">F</emphasis>
|
||
thoracic endopod 2, rostral
|
||
<emphasis bold="true" pageId="0" pageNumber="145">G</emphasis>
|
||
thoracic endopod 4, rostral
|
||
<emphasis bold="true" pageId="0" pageNumber="145">H</emphasis>
|
||
detail of panel (
|
||
<emphasis bold="true" pageId="0" pageNumber="145">G</emphasis>
|
||
) showing dactylus 4 with nail
|
||
<emphasis bold="true" pageId="0" pageNumber="145">I</emphasis>
|
||
inner face of oostegite 1, drawn above part of sympod 6. Scales omitted from objects
|
||
<emphasis bold="true" pageId="0" pageNumber="145">A-D, F, G</emphasis>
|
||
; setae from
|
||
<emphasis bold="true" pageId="0" pageNumber="145">E, H</emphasis>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Pleon</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E) and paratype subadult female 21.5 mm (F) A terminal portion of telson, dorsal, details show scales on left disto-mesial spine (B) and barbs on left terminal seta (C) D anterior half of cephalothorax, lateral, arrow points to distolateral edge of carapace E sixth pleomere with tail fan, lateral F example for pores (three to the left) and coat of scales on tergite of first pleomere." figureDoi="10.3897/zookeys.1079.76412.figure3" httpUri="https://binary.pensoft.net/fig/629313" pageId="0" pageNumber="145">3E, F</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E, H-N) and paratype subadult female 21.5 mm (F, G). A cardiac portion of foregut, dorsal view, dorsal wall omitted B-D details of panel (A) showing modified spines E details of panel (A) showing teeth emerging from a common basis F pleonite 1, cuticle detached and expanded on slide, dorsal and lateral faces on top, ventral face folded in, setae omitted from pleopods G detail of panel (F) showing pore group on left lateral face H-L series of pleopods 1 - 5, rostral = lateral face M uropods, ventral N telson. Scales omitted from objects F, M, N pore diameters not to scale in F, G." figureDoi="10.3897/zookeys.1079.76412.figure6" httpUri="https://binary.pensoft.net/fig/629316" pageId="0" pageNumber="145">6F-J</figureCitation>
|
||
). Pleonites 1-5 are each 0.6 times length of pleonite 6 measured along dorsal mid-line. Pleopod size increases distally. Pleopods 4-5 reaching only up to 2/3 length of pleonites 5 or 6, respectively. Not considering setae, pleopod 1 smooth (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E, H-N) and paratype subadult female 21.5 mm (F, G). A cardiac portion of foregut, dorsal view, dorsal wall omitted B-D details of panel (A) showing modified spines E details of panel (A) showing teeth emerging from a common basis F pleonite 1, cuticle detached and expanded on slide, dorsal and lateral faces on top, ventral face folded in, setae omitted from pleopods G detail of panel (F) showing pore group on left lateral face H-L series of pleopods 1 - 5, rostral = lateral face M uropods, ventral N telson. Scales omitted from objects F, M, N pore diameters not to scale in F, G." figureDoi="10.3897/zookeys.1079.76412.figure6" httpUri="https://binary.pensoft.net/fig/629316" pageId="0" pageNumber="145">6H</figureCitation>
|
||
) all around, remaining pleopods completely smooth only on caudal face, though scaly-hispid at least near basis of rostral (outer) face. Relative coverage with scales increases in series of pleopods 2-5 (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E, H-N) and paratype subadult female 21.5 mm (F, G). A cardiac portion of foregut, dorsal view, dorsal wall omitted B-D details of panel (A) showing modified spines E details of panel (A) showing teeth emerging from a common basis F pleonite 1, cuticle detached and expanded on slide, dorsal and lateral faces on top, ventral face folded in, setae omitted from pleopods G detail of panel (F) showing pore group on left lateral face H-L series of pleopods 1 - 5, rostral = lateral face M uropods, ventral N telson. Scales omitted from objects F, M, N pore diameters not to scale in F, G." figureDoi="10.3897/zookeys.1079.76412.figure6" httpUri="https://binary.pensoft.net/fig/629316" pageId="0" pageNumber="145">6I-L</figureCitation>
|
||
). All six pleonites dorsally densely covered by minute scales. Only pleonite 1 showing pores, namely two symmetrical transverse linear series each with 17-18 discontinuously spaced pores, in submedian position on dorsal face; and one additional, non-linear cluster of 8-9 pores on each lateral face (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E, H-N) and paratype subadult female 21.5 mm (F, G). A cardiac portion of foregut, dorsal view, dorsal wall omitted B-D details of panel (A) showing modified spines E details of panel (A) showing teeth emerging from a common basis F pleonite 1, cuticle detached and expanded on slide, dorsal and lateral faces on top, ventral face folded in, setae omitted from pleopods G detail of panel (F) showing pore group on left lateral face H-L series of pleopods 1 - 5, rostral = lateral face M uropods, ventral N telson. Scales omitted from objects F, M, N pore diameters not to scale in F, G." figureDoi="10.3897/zookeys.1079.76412.figure6" httpUri="https://binary.pensoft.net/fig/629316" pageId="0" pageNumber="145">6F-G</figureCitation>
|
||
shows fewer pores for the smaller paratype). Scutellum paracaudale sinusoid, well rounded.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Tail fan</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E) and paratype subadult female 21.5 mm (F) A terminal portion of telson, dorsal, details show scales on left disto-mesial spine (B) and barbs on left terminal seta (C) D anterior half of cephalothorax, lateral, arrow points to distolateral edge of carapace E sixth pleomere with tail fan, lateral F example for pores (three to the left) and coat of scales on tergite of first pleomere." figureDoi="10.3897/zookeys.1079.76412.figure3" httpUri="https://binary.pensoft.net/fig/629313" pageId="0" pageNumber="145">3A-C, E</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E, H-N) and paratype subadult female 21.5 mm (F, G). A cardiac portion of foregut, dorsal view, dorsal wall omitted B-D details of panel (A) showing modified spines E details of panel (A) showing teeth emerging from a common basis F pleonite 1, cuticle detached and expanded on slide, dorsal and lateral faces on top, ventral face folded in, setae omitted from pleopods G detail of panel (F) showing pore group on left lateral face H-L series of pleopods 1 - 5, rostral = lateral face M uropods, ventral N telson. Scales omitted from objects F, M, N pore diameters not to scale in F, G." figureDoi="10.3897/zookeys.1079.76412.figure6" httpUri="https://binary.pensoft.net/fig/629316" pageId="0" pageNumber="145">6M, N</figureCitation>
|
||
). Telson, endopod and exopod of uropods 1.0, 1.1-1.3 or 1.5-1.8 times length of sixth pleonite, respectively. Uropods (Figs
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E) and paratype subadult female 21.5 mm (F) A terminal portion of telson, dorsal, details show scales on left disto-mesial spine (B) and barbs on left terminal seta (C) D anterior half of cephalothorax, lateral, arrow points to distolateral edge of carapace E sixth pleomere with tail fan, lateral F example for pores (three to the left) and coat of scales on tergite of first pleomere." figureDoi="10.3897/zookeys.1079.76412.figure3" httpUri="https://binary.pensoft.net/fig/629313" pageId="0" pageNumber="145">3E</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E, H-N) and paratype subadult female 21.5 mm (F, G). A cardiac portion of foregut, dorsal view, dorsal wall omitted B-D details of panel (A) showing modified spines E details of panel (A) showing teeth emerging from a common basis F pleonite 1, cuticle detached and expanded on slide, dorsal and lateral faces on top, ventral face folded in, setae omitted from pleopods G detail of panel (F) showing pore group on left lateral face H-L series of pleopods 1 - 5, rostral = lateral face M uropods, ventral N telson. Scales omitted from objects F, M, N pore diameters not to scale in F, G." figureDoi="10.3897/zookeys.1079.76412.figure6" httpUri="https://binary.pensoft.net/fig/629316" pageId="0" pageNumber="145">6M</figureCitation>
|
||
) long, margins setose all around, dorsal and ventral faces scaly-hispid. Exopod with slightly convex, almost straight lateral margin and with more strongly convex mesial margin. Both margins diverge up to maximum width of the exopod at 1/3 length from basis and then converge up to the U-shaped, well-rounded terminus. Distal 4/5 of endopod with straight margins converging in V-shaped manner up to narrowly blunt apex. Endopod 0.7 times length of exopod, extending 1/5 of its length beyond telson, exopod 2/5 its length. Both statoliths unevenly discoid, mean diameter 0.22-0.23 mm, thickness 0.18 mm; core unevenly discoid as well, diameter 0.14 mm. Statolith formula 2 + 3 + (5-7) + 10 + (12-14) = 34. Statoliths composed of fluorite. Lateral margins and dorsal face of telson (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E, H-N) and paratype subadult female 21.5 mm (F, G). A cardiac portion of foregut, dorsal view, dorsal wall omitted B-D details of panel (A) showing modified spines E details of panel (A) showing teeth emerging from a common basis F pleonite 1, cuticle detached and expanded on slide, dorsal and lateral faces on top, ventral face folded in, setae omitted from pleopods G detail of panel (F) showing pore group on left lateral face H-L series of pleopods 1 - 5, rostral = lateral face M uropods, ventral N telson. Scales omitted from objects F, M, N pore diameters not to scale in F, G." figureDoi="10.3897/zookeys.1079.76412.figure6" httpUri="https://binary.pensoft.net/fig/629316" pageId="0" pageNumber="145">6N</figureCitation>
|
||
) completely covered by scales as in Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pseudomma kryotroglodytum sp. nov., holotype adult female BL 26.8 mm (A-E) and paratype subadult female 21.5 mm (F) A terminal portion of telson, dorsal, details show scales on left disto-mesial spine (B) and barbs on left terminal seta (C) D anterior half of cephalothorax, lateral, arrow points to distolateral edge of carapace E sixth pleomere with tail fan, lateral F example for pores (three to the left) and coat of scales on tergite of first pleomere." figureDoi="10.3897/zookeys.1079.76412.figure3" httpUri="https://binary.pensoft.net/fig/629313" pageId="0" pageNumber="145">3F</figureCitation>
|
||
, whereas ventral face only on its distal third.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">Colour</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pseudomma kryotroglodytum sp. nov. from ice cave of Bernard Island, Antarctica A female holotype, lateral B cephalothorax of female holotype, dorsal C physical aspect of the ice cave environment A, B living specimen in laboratory." figureDoi="10.3897/zookeys.1079.76412.figure2" httpUri="https://binary.pensoft.net/fig/629312" pageId="0" pageNumber="145">2A, B</figureCitation>
|
||
). Live colour was documented only in the laboratory; no difference visible between the two type specimens. Body and appendages generally whitish transparent. Lateral portions of eyeplate, foregut and part of mouthparts brilliant red, hepatic caeca yellow-green. The anterior pair of caeca covers part of the foregut dorsally, leaving a red M-like sign on the foregut in dorsal view. Posterior part of ultimate pleomere and adjoining basal portions of tail fan tinged light-red.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1079.76412.figure6" httpUri="https://binary.pensoft.net/fig/629316" pageId="0" pageNumber="145" start="Figure 6" startId="F6">
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">Figure 6.</emphasis>
|
||
<taxonomicName authorityName="Wittmann & Chevaldonné" authorityYear="2021" class="Malacostraca" family="Mysidae" genus="Pseudomma" kingdom="Animalia" lsidName="Pseudomma kryotroglodytum" order="Mysidacea" pageId="0" pageNumber="145" phylum="Schizopoda" rank="species" species="kryotroglodytum">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Pseudomma kryotroglodytum</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype adult female BL 26.8 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="145">A-E, H-N</emphasis>
|
||
) and paratype subadult female 21.5 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="145">F, G</emphasis>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
|
||
cardiac portion of foregut, dorsal view, dorsal wall omitted
|
||
<emphasis bold="true" pageId="0" pageNumber="145">B-D</emphasis>
|
||
details of panel (
|
||
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
|
||
) showing modified spines
|
||
<emphasis bold="true" pageId="0" pageNumber="145">E</emphasis>
|
||
details of panel (
|
||
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
|
||
) showing teeth emerging from a common basis
|
||
<emphasis bold="true" pageId="0" pageNumber="145">F</emphasis>
|
||
pleonite 1, cuticle detached and expanded on slide, dorsal and lateral faces on top, ventral face folded in, setae omitted from pleopods
|
||
<emphasis bold="true" pageId="0" pageNumber="145">G</emphasis>
|
||
detail of panel (
|
||
<emphasis bold="true" pageId="0" pageNumber="145">F</emphasis>
|
||
) showing pore group on left lateral face
|
||
<emphasis bold="true" pageId="0" pageNumber="145">H-L</emphasis>
|
||
series of pleopods 1-5, rostral = lateral face
|
||
<emphasis bold="true" pageId="0" pageNumber="145">M</emphasis>
|
||
uropods, ventral
|
||
<emphasis bold="true" pageId="0" pageNumber="145">N</emphasis>
|
||
telson. Scales omitted from objects
|
||
<emphasis bold="true" pageId="0" pageNumber="145">F, M, N</emphasis>
|
||
pore diameters not to scale in
|
||
<emphasis bold="true" pageId="0" pageNumber="145">F, G</emphasis>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="etymology">
|
||
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
The species name is an adjective with Latinised neutral ending formed by adjectivation of the amalgamated Ancient Greek adjective
|
||
<normalizedToken originalValue="κρύος">κρύος</normalizedToken>
|
||
(cold) with the noun
|
||
<normalizedToken originalValue="τρωγλοδύτης">τρωγλοδύτης</normalizedToken>
|
||
(cave dweller). The adjectivation has precedence in the name of the butterfly
|
||
<taxonomicName authorityName="Boisduval" authorityYear="1875" class="Insecta" family="Sphingidae" genus="Macroglossum" kingdom="Animalia" lsidName="Macroglossum troglodytus" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="troglodytus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Macroglossum troglodytus</emphasis>
|
||
</taxonomicName>
|
||
Boisduval, l875, listed by
|
||
<bibRefCitation author="Kemal, M" journalOrPublisher="Priamus" pageId="0" pageNumber="145" pagination="31 - 128" refId="B37" refString="Kemal, M, Kizildag, S, Kocak, AOe, 2019. Further notes on the Heterocera (Lepidoptera) of the Philippines, with a description of a new species. Priamus 18 (2): 31 - 128" title="Further notes on the Heterocera (Lepidoptera) of the Philippines, with a description of a new species." volume="18" year="2019">Kemal et al. (2019)</bibRefCitation>
|
||
as
|
||
<taxonomicName lsidName="M. troglodytum" pageId="0" pageNumber="145" rank="species" species="troglodytum">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">M. troglodytum</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="type locality">
|
||
<paragraph pageId="0" pageNumber="145">Type locality.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
Marine ice cave NE of Claude Bernard Island,
|
||
<normalizedToken originalValue="Adélie">Adelie</normalizedToken>
|
||
Land, East Antarctica,
|
||
<geoCoordinate degrees="66" direction="south" minutes="39.64" orientation="latitude" precision="9" value="-66.66067">66°39.64'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="140" direction="east" minutes="01.55" orientation="longitude" precision="9" value="140.02583">140°01.55'E</geoCoordinate>
|
||
, depth 10 m.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="subfamily heteromysinae norman, 1892">
|
||
<paragraph pageId="0" pageNumber="145">
|
||
Subfamily
|
||
<taxonomicName authorityName="Norman" authorityYear="1892" lsidName="" pageId="0" pageNumber="145" rank="subFamily" subFamily="Heteromysinae">Heteromysinae</taxonomicName>
|
||
Norman, 1892
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">
|
||
Tribe
|
||
<taxonomicName authorityName="Holt & Tattersall" authorityYear="1906" family="Mysidae" kingdom="Animalia" lsidName="" order="Mysida" pageId="0" pageNumber="145" rank="tribe" tribe="Mysidetini">Mysidetini</taxonomicName>
|
||
Holt & Tattersall, 1906
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |