256 lines
34 KiB
XML
256 lines
34 KiB
XML
<document id="DC10BB2B307CB3024CA413EF0D322BA4" ID-CLB-Dataset="7835" ID-DOI="10.11646/zootaxa.4938.1.1" ID-GBIF-Dataset="045b347a-4a5b-474f-b69f-1cd3fedf7678" ID-ISSN="1175-5326" ID-Zenodo-Dep="4561258" ID-ZooBank="074E95CD-E7C0-4098-93F0-70DE03CAD0D0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614236395824" checkinUser="plazi" docAuthor="Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes & Almeida, Lúcia Massutti De" docDate="2021" docId="CD475E66EC175A686BBF18F2FDEBFAB0" docLanguage="en" docName="zootaxa.4938.1.1.pdf" docOrigin="Zootaxa 4938 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Liogenys foveata Cherman 2021, new species" docType="treatment" docUuid="DB19CD93-CCCD-4A3F-A80E-74114513BDAC" docUuidSource="ZooBank" docVersion="7" lastPageNumber="21" masterDocId="317E261EEC055A7C6B281C59FFAAFFBD" masterDocTitle="Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species" masterLastPageNumber="59" masterPageNumber="1" pageNumber="19" updateTime="1698931070950" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="ABDD62E11F4BAF5F5717F4117A06D20C">Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species</mods:title>
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<mods:namePart id="F8DDCC739E3633A496DF38E7FBC81B98">Cherman, Mariana Alejandra</mods:namePart>
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<mods:affiliation id="512EFFCB569F6ED9A2751CFBE7257F05">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil. & Laboratório de Scarabaeoidologia, Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, 78060 - 900, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
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<mods:namePart id="D1A6974068DEBB99C1B39DC916D9603A">Basílio, Daniel Silva</mods:namePart>
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<mods:affiliation id="025ED17870D377F66CC0DE2E1F0A0286">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil. & basilio. dsb @ gmail. com; https: // orcid. org / 0000 - 0003 - 3556 - 5880</mods:affiliation>
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<mods:nameIdentifier id="5A3AFD28B6FC9EC1164DE4382650C1C4" type="email">basilio.dsb@gmail.com</mods:nameIdentifier>
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<mods:namePart id="B35A4EDCA3E1AA77819D2687C0FCA030">Mise, Kleber Makoto</mods:namePart>
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<mods:affiliation id="0A476ABD2FBFEB88EC07E782AB5A1336">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil. & klebermise @ yahoo. com. br; https: // orcid. org / 0000 - 0003 - 4391 - 9589</mods:affiliation>
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<mods:nameIdentifier id="B17F0378CBA807419176A0175E09FAC3" type="email">klebermise@yahoo.com.br</mods:nameIdentifier>
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<mods:namePart id="9D3DEACCCA694308C9A63D8F05326D08">Frisch, Johannes</mods:namePart>
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<mods:affiliation id="24599CC43B48C23EA3A772C17F4186A7">Museum für Naturkunde Berlin, Invalidenstrasse 43, 10115 Berlin, Germany.</mods:affiliation>
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<mods:namePart id="66FD0A6EB58A9CAECD92E964336459F8">Almeida, Lúcia Massutti De</mods:namePart>
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<mods:affiliation id="BC75CDD2C5F410F590B5A1C3A46D4B14">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil. & lalmeida @ ufpr. br; https: // orcid. org / 0000 - 0003 - 4277 - 711 X</mods:affiliation>
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<mods:nameIdentifier id="5F1FDD898A7BCC92913E5C257028A049" type="email">lalmeida@ufpr.br</mods:nameIdentifier>
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<mods:title id="10FEA94C773969882455B29D2D44A702">Zootaxa</mods:title>
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<mods:date id="350772DB6C0851B39BFC0DEF1A06DA1F">2021</mods:date>
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<mods:number id="D560DB9E7506E63B0B05B4BE8F635A86">2021-02-25</mods:number>
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<mods:number id="C7F2F9350E94DAF10059D577F550A11D">4938</mods:number>
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<treatment id="CD475E66EC175A686BBF18F2FDEBFAB0" ID-DOI="http://doi.org/10.5281/zenodo.4561278" ID-GBIF-Taxon="180301834" ID-Zenodo-Dep="4561278" LSID="urn:lsid:zoobank.org:act:DB19CD93-CCCD-4A3F-A80E-74114513BDAC" httpUri="http://treatment.plazi.org/id/CD475E66EC175A686BBF18F2FDEBFAB0" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19">
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<emphasis id="779A3362EC175A6E6BBF18F2FD2AFB79" bold="true" box="[151,640,1194,1221]" pageId="18" pageNumber="19">
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<taxonomicName id="82EE94F3EC175A6E6BBF18F2FE4EFB79" authority="Cherman" authorityName="Cherman" authorityYear="2021" box="[151,484,1194,1221]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="foveata" status="sp. nov.">
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<emphasis id="779A3362EC175A6E6BBF18F2FECEFB79" bold="true" box="[151,356,1194,1221]" italics="true" pageId="18" pageNumber="19">Liogenys foveata</emphasis>
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Cherman
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,
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<taxonomicNameLabel id="6CA98E19EC175A6E6AD918F2FD2AFB79" box="[497,640,1194,1221]" pageId="18" pageNumber="19" rank="species">new species</taxonomicNameLabel>
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<figureCitation id="DDD5F3F5EC175A6E6BBF1896FF4EFB54" box="[151,228,1231,1257]" captionStart="FIGURE 8" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@19.[203,1385,181,1920]" captionTargetPageId="19" captionText="FIGURE 8. Liogenys foveata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) distal maxillary palp, D) lateral, E) mesotibia and metatibia, F) pygidium, and parameres in views: G) dorsal, and H) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561282" httpUri="https://zenodo.org/record/4561282/files/figure.png" pageId="18" pageNumber="19">Figs. 8</figureCitation>
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,
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<figureCitation id="DDD5F3F5EC175A6E6BD91896FEBAFB54" box="[241,272,1231,1257]" captionStart="FIGURE 27" captionStartId="55.[151,250,1459,1483]" captionTargetBox="[171,1416,192,1422]" captionTargetId="figure-20@55.[151,1435,181,1434]" captionTargetPageId="55" captionText="FIGURE 27. Geographical distribution of the Liogenys new species. Shaped points: new species; colors: Neotropical provinces (Morrone 2014) and Monte province from the South American Transition Zone (Roig-Juñent et al. 2018)." figureDoi="http://doi.org/10.5281/zenodo.4561332" httpUri="https://zenodo.org/record/4561332/files/figure.png" pageId="18" pageNumber="19">27</figureCitation>
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.
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<paragraph id="4551EF70EC175A6E6BBF18ABFC08FAB0" blockId="18.[151,930,1194,1293]" box="[151,930,1266,1293]" pageId="18" pageNumber="19">
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http://zoobank.org/
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<subSubSection id="0DF4BCFBEC175A6E6BBF1962FD86FA7C" pageId="18" pageNumber="19" type="materials_examined">
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<emphasis id="779A3362EC175A6E6BBF1962FE96FAE9" bold="true" box="[151,316,1338,1365]" pageId="18" pageNumber="19">Type material</emphasis>
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.
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<materialsCitation id="F586E52DEC175A6E6A611962FC99FA20" ID-GBIF-Occurrence="3047558439" collectionCode="IADIZA" collectorName="F. Ocampo G. San Blas & F. Campon." country="Argentina" latitude="-24.090834" location="Formosa" longLatPrecision="21" longitude="-61.946945" pageId="18" pageNumber="19" specimenCount="2" stateProvince="Formosa" typeStatus="holotype">
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<typeStatus id="9A5551D2EC175A6E6A611962FE1BFAE8" box="[329,433,1339,1365]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
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labeled (
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<collectionCode id="23FF77B5EC175A6E693D1962FDD7FAE8" box="[533,637,1339,1365]" pageId="18" pageNumber="19">IADIZA</collectionCode>
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): [white typeset] “
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:
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/ RP 39, 23 km S Ing. Juárez, /
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<geoCoordinate id="20DA89B7EC175A6E6B811939FE8CFAC6" box="[169,294,1376,1403]" degrees="24" direction="south" minutes="05" orientation="latitude" pageId="18" pageNumber="19" precision="15" seconds="27" value="-24.090834">24º05ʹ27ʺS</geoCoordinate>
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. / 13-17-XII-08.
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<collectorName id="E81B8AA6EC175A6E69571906FC39FAC4" box="[639,915,1375,1401]" pageId="18" pageNumber="19">F. Ocampo G. San Blas</collectorName>
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,
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<collectorName id="E81B8AA6EC175A6E688A1906FB86FAC4" box="[930,1068,1375,1401]" pageId="18" pageNumber="19">F. Campón.</collectorName>
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”, [red, typeset and handwritten] “
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<taxonomicName id="82EE94F3EC175A6E6B8919DAFE19FA20" authority="FOVEATA" authorityName="FOVEATA" box="[161,435,1411,1437]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">LIOGENYS FOVEATA</taxonomicName>
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/
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<typeStatus id="9A5551D2EC175A6E6AE019DAFDF6FA20" box="[456,604,1411,1437]" pageId="18" pageNumber="19" type="holotype">HOLOTYPE</typeStatus>
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/ Cherman M. A.”.
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</materialsCitation>
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<paragraph id="4551EF70EC175A6E6BEF19FEFD86FA7C" blockId="18.[151,1436,1338,2013]" box="[199,556,1447,1473]" pageId="18" pageNumber="19">
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<typeStatus id="9A5551D2EC175A6E6BEF19FEFE85FA7C" box="[199,303,1447,1473]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
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deposited at IADIZA.
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</paragraph>
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</subSubSection>
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<subSubSection id="0DF4BCFBEC175A6E6BEF1993FAB2F95C" pageId="18" pageNumber="19" type="diagnosis">
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<paragraph id="4551EF70EC175A6E6BEF1993FAB2F95C" blockId="18.[151,1436,1338,2013]" pageId="18" pageNumber="19">
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<emphasis id="779A3362EC175A6E6BEF1993FE90FA59" bold="true" box="[199,314,1482,1508]" pageId="18" pageNumber="19">Diagnosis</emphasis>
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. Body elongate; brown, elytra brown or light brown, pronotum purplish brown, head dark at vertex (
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<figureCitation id="DDD5F3F5EC175A6E6B8819B6FF51F9B4" box="[160,251,1519,1545]" captionStart="FIGURE 8" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@19.[203,1385,181,1920]" captionTargetPageId="19" captionText="FIGURE 8. Liogenys foveata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) distal maxillary palp, D) lateral, E) mesotibia and metatibia, F) pygidium, and parameres in views: G) dorsal, and H) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561282" httpUri="https://zenodo.org/record/4561282/files/figure.png" pageId="18" pageNumber="19">Fig. 8A</figureCitation>
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); clypeal emargination deep but not reaching the base of the teeth, angulate and wide; outer sides of anterior teeth parallel; clypeal lateral margin projection subangulate; outer side of anterior teeth and clypeal lateral projection forming an angle (
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<figureCitation id="DDD5F3F5EC175A6E6AF41A6EFD99F9EC" box="[476,563,1591,1617]" captionStart="FIGURE 8" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@19.[203,1385,181,1920]" captionTargetPageId="19" captionText="FIGURE 8. Liogenys foveata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) distal maxillary palp, D) lateral, E) mesotibia and metatibia, F) pygidium, and parameres in views: G) dorsal, and H) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561282" httpUri="https://zenodo.org/record/4561282/files/figure.png" pageId="18" pageNumber="19">Fig. 8B</figureCitation>
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); fovea of the distal maxillary palpomere shallow, occupying almost the entire surface of the palpomere (
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<figureCitation id="DDD5F3F5EC175A6E6AEA1A02FDB1F9C8" box="[450,539,1627,1653]" captionStart="FIGURE 8" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@19.[203,1385,181,1920]" captionTargetPageId="19" captionText="FIGURE 8. Liogenys foveata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) distal maxillary palp, D) lateral, E) mesotibia and metatibia, F) pygidium, and parameres in views: G) dorsal, and H) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561282" httpUri="https://zenodo.org/record/4561282/files/figure.png" pageId="18" pageNumber="19">Fig. 8C</figureCitation>
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); pronotal posterior corners subangulate (
|
||
<figureCitation id="DDD5F3F5EC175A6E68DC1A02FBE7F9C8" box="[1012,1101,1627,1653]" captionStart="FIGURE 8" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@19.[203,1385,181,1920]" captionTargetPageId="19" captionText="FIGURE 8. Liogenys foveata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) distal maxillary palp, D) lateral, E) mesotibia and metatibia, F) pygidium, and parameres in views: G) dorsal, and H) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561282" httpUri="https://zenodo.org/record/4561282/files/figure.png" pageId="18" pageNumber="19">Fig. 8B</figureCitation>
|
||
); mesofemural surface setose abundantly, long thick erect bristles, as well as the outer margin of the metafemur (
|
||
<figureCitation id="DDD5F3F5EC175A6E6F161A26FB12F924" box="[1086,1208,1663,1689]" captionStart="FIGURE 8" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@19.[203,1385,181,1920]" captionTargetPageId="19" captionText="FIGURE 8. Liogenys foveata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) distal maxillary palp, D) lateral, E) mesotibia and metatibia, F) pygidium, and parameres in views: G) dorsal, and H) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561282" httpUri="https://zenodo.org/record/4561282/files/figure.png" pageId="18" pageNumber="19">Fig. 8D, E</figureCitation>
|
||
); pygidial width exceeding distance between spiracles of propygidium; pygidial disc with long bristles at the apex (
|
||
<figureCitation id="DDD5F3F5EC175A6E6FE91AFAFABFF900" box="[1217,1301,1699,1725]" captionStart="FIGURE 8" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@19.[203,1385,181,1920]" captionTargetPageId="19" captionText="FIGURE 8. Liogenys foveata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) distal maxillary palp, D) lateral, E) mesotibia and metatibia, F) pygidium, and parameres in views: G) dorsal, and H) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561282" httpUri="https://zenodo.org/record/4561282/files/figure.png" pageId="18" pageNumber="19">Fig. 8F</figureCitation>
|
||
); parameres almost uniform along its length; outer margins slightly convex at the midline; apex spatulate (
|
||
<figureCitation id="DDD5F3F5EC175A6E6F991A9EFAA1F95C" box="[1201,1291,1735,1761]" captionStart="FIGURE 8" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@19.[203,1385,181,1920]" captionTargetPageId="19" captionText="FIGURE 8. Liogenys foveata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) distal maxillary palp, D) lateral, E) mesotibia and metatibia, F) pygidium, and parameres in views: G) dorsal, and H) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561282" httpUri="https://zenodo.org/record/4561282/files/figure.png" pageId="18" pageNumber="19">Fig. 8G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DF4BCFBEC175A686BEF1AB2FB3EFCE0" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19" type="materials_examined">
|
||
<paragraph id="4551EF70EC175A686BEF1AB2FB3EFCE0" blockId="18.[151,1436,1338,2013]" lastBlockId="20.[151,1437,151,1293]" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19">
|
||
<emphasis id="779A3362EC175A6E6BEF1AB2FE9EF8B8" bold="true" box="[199,308,1771,1797]" pageId="18" pageNumber="19">
|
||
<typeStatus id="9A5551D2EC175A6E6BEF1AB2FE9EF8B8" box="[199,308,1771,1797]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="779A3362EC175A6E6A6D1AB2FE28F8B8" bold="true" box="[325,386,1771,1797]" pageId="18" pageNumber="19">Male</emphasis>
|
||
. Length:
|
||
<quantity id="82164295EC175A6E6ADC1AB3FDF4F8B8" box="[500,606,1770,1797]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="18" pageNumber="19" unit="mm" value="10.5">10.5 mm</quantity>
|
||
; width:
|
||
<quantity id="82164295EC175A6E69961AB3FCBDF8B8" box="[702,791,1770,1797]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="18" pageNumber="19" unit="mm" value="5.6">5.6 mm</quantity>
|
||
. Brown.
|
||
<emphasis id="779A3362EC175A6E68AD1AB2FC61F8B8" box="[901,971,1771,1797]" italics="true" pageId="18" pageNumber="19">Head:</emphasis>
|
||
distance between eyes almost twice the width of one eye; clypeal emargination deep but not reaching the base of the teeth, angulate and wide; outer sides of anterior teeth parallel; outer margin of anterior teeth almost equal to the eye length; lateral margin slightly convex, forming a subangulate projection; distance between clypeal lateral projection and anterior margin of eye shorter than the eye; distance between clypeal lateral projection and apex of the adjacent anterior tooth longer than the basal width of the anterior tooth; outer side of anterior teeth and clypeal lateral projection not forming an angle; distal maxillary palpomere, maximum width more than twice the apical width; fovea shallow, extending past the transverse midline, occupying almost the entire surface of the palpomere; antenna with 10 antennomeres, club lighter in color and slightly longer of the funicle.
|
||
<emphasis id="779A3362EC115A6869E01CE2FC88FF68" box="[712,802,187,213]" italics="true" pageId="20" pageNumber="21">Thorax:</emphasis>
|
||
anterior margin of pronotum straight; pronotal disc glabrous, punctures coarse and sparse; posterior corners subangulate; hypomere with short bristles, mesepisternum scaly; sides of metaventrite with bristles; distance between mesocoxae and metacoxae up to twice the length of the metacoxa; scutellum ogival, sparsely and finely punctate throughout.
|
||
<emphasis id="779A3362EC115A6868B11D7EFC42FEFC" box="[921,1000,295,321]" italics="true" pageId="20" pageNumber="21">Elytra:</emphasis>
|
||
shiny, glabrous, brown, lighter in color than pronotum; elytral suture darker than elytron and weakly elevated; all four elytral ridges weakly defined.
|
||
<emphasis id="779A3362EC115A686E761D12FA37FED9" box="[1374,1437,331,356]" italics="true" pageId="20" pageNumber="21">Legs:</emphasis>
|
||
three protibial teeth, the middle and apical equal in size; the three teeth equally spaced; mesofemural surface setose abundantly, as well as the outer margin of the metafemur, bristles long, thick, erect; mesotibia cylindrical in cross section, surface finely sculptured; two transverse carinae, the apical one present posteriorly; metacoxa with sparse, long scales; basal apophysis of metacoxa not produced; metatibia carinate along the inner margin towards the apex, apical inner surface setose; a metatibial transverse carina present posteriorly and posterior discontinuous longitudinal carina (
|
||
<figureCitation id="DDD5F3F5EC115A686A3A1E7AFECDFD80" box="[274,359,547,573]" captionStart="FIGURE 8" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@19.[203,1385,181,1920]" captionTargetPageId="19" captionText="FIGURE 8. Liogenys foveata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) distal maxillary palp, D) lateral, E) mesotibia and metatibia, F) pygidium, and parameres in views: G) dorsal, and H) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561282" httpUri="https://zenodo.org/record/4561282/files/figure.png" pageId="20" pageNumber="21">Fig. 8E</figureCitation>
|
||
); metatibial apical spurs of different lengths; the larger equal in length to the diameter of the tibial apex; in males protarsomeres and mesotarsomeres I to IV enlarged; protarsomere II slightly longer than it is wide; metatarsomere I slightly shorter than metatarsomere II; claw bifid, symmetrical, superior tooth of a claw longer and slightly narrower than the inferior, distance between teeth equal to the length of the inferior tooth.
|
||
<emphasis id="779A3362EC115A686E011ED6FA36FD14" box="[1321,1436,655,681]" italics="true" pageId="20" pageNumber="21">Abdomen:</emphasis>
|
||
ventrites with sparse, long bristles, abundantly on ventrite V; propygidium visible, with bristles; pygidium slightly convex, subquadrate, as wide as it is long; pygidial width exceeding distance between spiracles of propygidium; pygidial disc with long bristles at the apex; pygidial apex rounded.
|
||
<emphasis id="779A3362EC115A6868561EA2FC52FCA9" box="[894,1016,763,788]" italics="true" pageId="20" pageNumber="21">Parameres</emphasis>
|
||
: basal region as wide as both sections of the parameres at its maximum width, parameral split at 2/3; inner margins straight, opened; outer margins slightly convex at the midline; apex spatulate; parameres slightly concave in lateral view (
|
||
<figureCitation id="DDD5F3F5EC115A686F051F1AFB2DFCE0" box="[1069,1159,835,861]" captionStart="FIGURE 8" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@19.[203,1385,181,1920]" captionTargetPageId="19" captionText="FIGURE 8. Liogenys foveata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) distal maxillary palp, D) lateral, E) mesotibia and metatibia, F) pygidium, and parameres in views: G) dorsal, and H) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561282" httpUri="https://zenodo.org/record/4561282/files/figure.png" pageId="20" pageNumber="21">Fig. 8H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="1191BFF8EC165A6F6BBF1BCEFAA6F869" ID-DOI="http://doi.org/10.5281/zenodo.4561282" ID-Zenodo-Dep="4561282" httpUri="https://zenodo.org/record/4561282/files/figure.png" pageId="19" pageNumber="20" startId="19.[151,250,1943,1967]" targetBox="[203,1385,181,1920]" targetPageId="19">
|
||
<paragraph id="4551EF70EC165A6F6BBF1BCEFAA6F869" blockId="19.[151,1436,1943,2004]" pageId="19" pageNumber="20">
|
||
<emphasis id="779A3362EC165A6F6BBF1BCEFEB9F812" bold="true" box="[151,275,1943,1967]" pageId="19" pageNumber="20">FIGURE 8.</emphasis>
|
||
<taxonomicName id="82EE94F3EC165A6F6A321BC1FD87F80D" authority="Cherman" authorityName="Cherman" authorityYear="2021" box="[282,557,1944,1968]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="foveata" status="sp. nov.">
|
||
<emphasis id="779A3362EC165A6F6A321BC1FE6DF80D" box="[282,455,1944,1968]" italics="true" pageId="19" pageNumber="20">Liogenys foveata</emphasis>
|
||
Cherman
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="779A3362EC165A6F691F1BC1FD1EF812" bold="true" box="[567,692,1943,1968]" pageId="19" pageNumber="20">
|
||
<taxonomicNameLabel id="6CA98E19EC165A6F691F1BC1FD1EF812" box="[567,692,1943,1968]" pageId="19" pageNumber="20" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype views: A) dorsal, B) head and pronotum, C) distal maxillary palp, D) lateral, E) mesotibia and metatibia, F) pygidium, and parameres in views: G) dorsal, and H) lateral.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="0DF4BCFBEC115A686BEF1F3EFD2CFC74" pageId="20" pageNumber="21" type="etymology">
|
||
<paragraph id="4551EF70EC115A686BEF1F3EFD2CFC74" blockId="20.[151,1437,151,1293]" pageId="20" pageNumber="21">
|
||
<emphasis id="779A3362EC115A686BEF1F3EFEE2FC3C" bold="true" box="[199,328,871,897]" pageId="20" pageNumber="21">Etymology</emphasis>
|
||
. Adjective in the nominative singular. From the Latin
|
||
<emphasis id="779A3362EC115A6868871F3EFBA6FC3C" box="[943,1036,871,897]" italics="true" pageId="20" pageNumber="21">foveatus</emphasis>
|
||
(
|
||
<emphasis id="779A3362EC115A686F351F3EFBF0FC3C" box="[1053,1114,871,897]" italics="true" pageId="20" pageNumber="21">fovea</emphasis>
|
||
“ditch, pit”): having a slight depression or pit in a bone or organ. In this species the fovea of the maxillary distal palpomere is shallow but quite broadly extended, almost reaching the apex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DF4BCFBEC115A686BEF1F8AFC85FB88" pageId="20" pageNumber="21" type="materials_examined">
|
||
<paragraph id="4551EF70EC115A686BEF1F8AFE87FBAC" blockId="20.[151,1437,151,1293]" pageId="20" pageNumber="21">
|
||
<emphasis id="779A3362EC115A686BEF1F8AFEDAFC51" bold="true" box="[199,368,978,1005]" pageId="20" pageNumber="21">Type locality.</emphasis>
|
||
<materialsCitation id="F586E52DEC115A686AA81F8AFE83FBAC" ID-GBIF-Occurrence="3047558492" collectorName="S Ingeniero Juarez" country="Argentina" latitude="-24.090834" location="Formosa" longLatPrecision="21" longitude="-61.946945" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Formosa">
|
||
<collectingCountry id="3DF9AFE0EC115A686AA81F8AFD8CFC50" box="[384,550,979,1005]" name="Argentina" pageId="20" pageNumber="21">ARGENTINA</collectingCountry>
|
||
,
|
||
<collectingRegion id="872A2192EC115A6869141F8AFD0BFC50" box="[572,673,979,1005]" country="Argentina" name="Formosa" pageId="20" pageNumber="21">Formosa</collectingRegion>
|
||
, Ruta Provincial 39, 23 km
|
||
<collectorName id="E81B8AA6EC115A686F301F8AFAAAFC50" box="[1048,1280,979,1005]" pageId="20" pageNumber="21">S Ingeniero Juárez</collectorName>
|
||
,
|
||
<geoCoordinate id="20DA89B7EC115A686E3E1F8BFA36FC50" box="[1302,1436,978,1005]" degrees="24" direction="south" minutes="05" orientation="latitude" pageId="20" pageNumber="21" precision="15" seconds="27" value="-24.090834">24º05’27’’S</geoCoordinate>
|
||
<geoCoordinate id="20DA89B7EC115A686BBF1FAFFE83FBAC" box="[151,297,1014,1041]" degrees="61" direction="west" minutes="56" orientation="longitude" pageId="20" pageNumber="21" precision="15" seconds="49" value="-61.946945">61º56’49’’W</geoCoordinate>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="4551EF70EC115A686BEF1843FC85FB88" blockId="20.[151,1437,151,1293]" box="[199,815,1050,1077]" pageId="20" pageNumber="21">
|
||
<emphasis id="779A3362EC115A686BEF1843FE54FB89" bold="true" box="[199,510,1050,1076]" pageId="20" pageNumber="21">Geographical distribution</emphasis>
|
||
.
|
||
<collectingCountry id="3DF9AFE0EC115A6869221842FD06FB88" box="[522,684,1051,1077]" name="Argentina" pageId="20" pageNumber="21">ARGENTINA</collectingCountry>
|
||
(
|
||
<collectingRegion id="872A2192EC115A6869931842FC8EFB88" box="[699,804,1051,1077]" country="Argentina" name="Formosa" pageId="20" pageNumber="21">Formosa</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DF4BCFBEC115A686BEF1866FDEBFAB0" pageId="20" pageNumber="21" type="discussion">
|
||
<paragraph id="4551EF70EC115A686BEF1866FDEBFAB0" blockId="20.[151,1437,151,1293]" pageId="20" pageNumber="21">
|
||
<emphasis id="779A3362EC115A686BEF1866FE9EFBE4" bold="true" box="[199,308,1087,1113]" pageId="20" pageNumber="21">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="82EE94F3EC115A686A6A1866FDAAFBE4" authorityName="Cherman" authorityYear="2021" box="[322,512,1087,1113]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="foveata">
|
||
<emphasis id="779A3362EC115A686A6A1866FDAAFBE4" box="[322,512,1087,1113]" italics="true" pageId="20" pageNumber="21">Liogenys foveata</emphasis>
|
||
</taxonomicName>
|
||
resembles
|
||
<taxonomicName id="82EE94F3EC115A6869571866FD5BFBE4" authorityName="Moser" authorityYear="1921" box="[639,753,1087,1113]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tarsalis">
|
||
<emphasis id="779A3362EC115A6869571866FD5BFBE4" box="[639,753,1087,1113]" italics="true" pageId="20" pageNumber="21">L. tarsalis</emphasis>
|
||
</taxonomicName>
|
||
, mainly in the size, body color, and clypeus deeply emarginate.
|
||
<taxonomicName id="82EE94F3EC115A686BE1183AFE20FBC0" authorityName="Cherman" authorityYear="2021" box="[201,394,1123,1149]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="foveata">
|
||
<emphasis id="779A3362EC115A686BE1183AFE20FBC0" box="[201,394,1123,1149]" italics="true" pageId="20" pageNumber="21">Liogenys foveata</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="82EE94F3EC115A686900183AFD36FBC0" authorityName="Moser" authorityYear="1921" box="[552,668,1123,1149]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tarsalis">
|
||
<emphasis id="779A3362EC115A686900183AFD36FBC0" box="[552,668,1123,1149]" italics="true" pageId="20" pageNumber="21">L. tarsalis</emphasis>
|
||
</taxonomicName>
|
||
(in parenthesis) mainly in the clypeus with emargination angulate and lateral margin with a weak subangulate projection (emargination subangulate, lateral margin with a distinct, rounded projection); in the large fovea on the maxillary distal palpomere, occupying almost the entire surface of the palpomere; in the pronotal posterior corners subangulate (rounded); and in the shape of the pygidium, tarsi and parameres. Females remain unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |