465 lines
65 KiB
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465 lines
65 KiB
XML
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<mods:title id="2ABD4A9BDB1114EDBCF1A807D1E671FE">Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species</mods:title>
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<mods:namePart id="84BA9F1557379986EB47092271567B00">Cherman, Mariana Alejandra</mods:namePart>
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<mods:affiliation id="A39B8BD8C6A27B0F71FF0021FF2D72FD">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil. & Laboratório de Scarabaeoidologia, Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, 78060 - 900, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
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<figureCitation id="DDD5F3F5EC0A5A736BBF1E1EFF4FFDDC" box="[151,229,583,609]" captionStart="FIGURE 6" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@16.[203,1385,181,1920]" captionTargetPageId="16" captionText="FIGURE 6. Liogenys forcipata Frey, 1970 male views: A) dorsal, B) head and pronotum, C) lateral, D) base of metatibia, white arrow indicates basal apophysis and metatrochanter, E) pygidium, and parameres in views: F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561276" httpUri="https://zenodo.org/record/4561276/files/figure.png" pageId="15" pageNumber="16">Figs. 6</figureCitation>
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;
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;
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<figureCitation id="DDD5F3F5EC0A5A736A491E1EFE2AFDDC" box="[353,384,583,609]" captionStart="FIGURE 23" captionStartId="51.[151,250,1451,1475]" captionTargetBox="[171,1416,192,1415]" captionTargetId="figure-20@51.[151,1435,181,1427]" captionTargetPageId="51" captionText="FIGURE 23. Geographical distribution of four known Liogenys species occurring in the Chacoan province and its boundaries. Shaped points: previously known species; colors: Neotropical provinces (Morrone 2014) and Monte province from the South American Transition Zone (Roig-Juñent et al. 2018)." figureDoi="http://doi.org/10.5281/zenodo.4561324" httpUri="https://zenodo.org/record/4561324/files/figure.png" pageId="15" pageNumber="16">23</figureCitation>
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<emphasis id="779A3362EC0A5A736BBF1EF3FEFFFD7F" box="[151,341,682,706]" italics="true" pageId="15" pageNumber="16">Liogenys forcipata</emphasis>
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:
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Cherman
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<emphasis id="779A3362EC0A5A736F471EF2FB08FD7F" box="[1135,1186,682,706]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
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2016: 764
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, 766 (systematics);
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|
||
<bibRefCitation id="217F9281EC0A5A736A231E91FDA9FD5C" author="Cherman, M. A. & Mise, K. M. & Moron, M. A. & Vaz-de-Mello, F. Z. & Almeida, L. M." box="[267,515,712,737]" pageId="15" pageNumber="16" pagination="1 - 120" refId="ref35643" refString="Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. & Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031" type="journal article" year="2017">
|
||
Cherman
|
||
<emphasis id="779A3362EC0A5A736A471E90FE08FD5D" box="[367,418,712,736]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
|
||
2017: 10
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 16, 77–78 (generic redescription).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DF4BCFBEC0A5A736BBF1F56FAFFFB2C" pageId="15" pageNumber="16" type="materials_examined">
|
||
<paragraph id="4551EF70EC0A5A736BBF1F56FB7DFB98" blockId="15.[151,1437,782,2033]" pageId="15" pageNumber="16">
|
||
<emphasis id="779A3362EC0A5A736BBF1F56FEEAFC95" bold="true" box="[151,320,782,809]" pageId="15" pageNumber="16">Type material</emphasis>
|
||
.
|
||
<taxonomicName id="82EE94F3EC0A5A736A781F56FD85FC94" authorityName="Frey" authorityYear="1970" box="[336,559,783,809]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="forcipatus">
|
||
<emphasis id="779A3362EC0A5A736A781F56FD85FC94" box="[336,559,783,809]" italics="true" pageId="15" pageNumber="16">Liogenys forcipatus</emphasis>
|
||
</taxonomicName>
|
||
male
|
||
<typeStatus id="9A5551D2EC0A5A73695F1F56FD73FC94" box="[631,729,783,809]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
|
||
(
|
||
<collectionCode id="23FF77B5EC0A5A7369C21F56FCEFFC94" box="[746,837,783,809]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="15" pageNumber="16">NHMB</collectionCode>
|
||
): [white typeset] “
|
||
<collectingCountry id="3DF9AFE0EC0A5A736F3D1F56FB10FC94" box="[1045,1210,783,809]" name="Argentina" pageId="15" pageNumber="16">ARGENTINA</collectingCountry>
|
||
/ Pronunciamento /
|
||
<collectingRegion id="872A2192EC0A5A736BBF1F6AFEA4FCF0" box="[151,270,819,845]" country="Argentina" name="Entre Rios" pageId="15" pageNumber="16">Entre Rios</collectingRegion>
|
||
/
|
||
<date id="3150C9B0EC0A5A736A0D1F6AFE24FCF0" box="[293,398,819,845]" pageId="15" pageNumber="16" value="1962-12">XII.1962</date>
|
||
, leg. Pena”, [white handwritten] “Type /
|
||
<taxonomicName id="82EE94F3EC0A5A7368741F6AFC6EFCF0" authorityName="Guerin-Meneville" authorityYear="1831" box="[860,964,819,845]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
|
||
/ forcipatus / n sp / [typeset] det. G.
|
||
<bibRefCitation id="217F9281EC0A5A736E4C1F6AFF79FCCC" author="Frey, G." pageId="15" pageNumber="16" pagination="281 - 287" refId="ref36196" refString="Frey, G. (1970) Beitrag zur kenntnis der Sudamerikanischen Melolonthiden (Col.). Entomologische Arbeiten aus dem Museum G. Frey, 21, 281 - 287." type="journal article" year="1970">Frey, 1970</bibRefCitation>
|
||
”, [red typeset] “TYPE”.
|
||
<typeStatus id="9A5551D2EC0A5A736AC51F0EFDF0FCCC" box="[493,602,855,881]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
|
||
(2): one male (
|
||
<collectionCode id="23FF77B5EC0A5A7368201F0EFCF4FCCC" box="[776,862,855,881]" pageId="15" pageNumber="16">CMNC</collectionCode>
|
||
) with the same locality label as the
|
||
<typeStatus id="9A5551D2EC0A5A736FDD1F0EFAF0FCCC" box="[1269,1370,855,881]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
|
||
, plus: [white handwritten] “13 / ♂ / Type /
|
||
<taxonomicName id="82EE94F3EC0A5A7369001F22FD3AFC28" authorityName="Guerin-Meneville" authorityYear="1831" box="[552,656,891,917]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
|
||
/ forcipatus / n sp / [typeset] det. G.
|
||
<bibRefCitation id="217F9281EC0A5A736F361F22FB32FC28" author="Frey, G." box="[1054,1176,891,917]" pageId="15" pageNumber="16" pagination="281 - 287" refId="ref36196" refString="Frey, G. (1970) Beitrag zur kenntnis der Sudamerikanischen Melolonthiden (Col.). Entomologische Arbeiten aus dem Museum G. Frey, 21, 281 - 287." type="journal article" year="1970">Frey, 1970</bibRefCitation>
|
||
”, [red typeset] “PARATYPE”, [white, outlined black, typeset] “Canadian Museum of / Musée Canadien de la / NATURE / CMNEN 00018923”, [white, outlined black, typeset] “H. & A. HOWDEN / COLLECTION /
|
||
<emphasis id="779A3362EC0A5A736F4C1F9AFB2EFC61" box="[1124,1156,963,988]" italics="true" pageId="15" pageNumber="16">ex.</emphasis>
|
||
A. Martinez coll.”. One male (
|
||
<collectionCode id="23FF77B5EC0A5A736BF51FBEFE92FBBC" box="[221,312,999,1025]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="15" pageNumber="16">NHMB</collectionCode>
|
||
): [white typeset] “
|
||
<collectingCountry id="3DF9AFE0EC0A5A73692D1FBEFD00FBBC" box="[517,682,999,1025]" name="Argentina" pageId="15" pageNumber="16">ARGENTINA</collectingCountry>
|
||
/ Pronunciamento /
|
||
<collectingRegion id="872A2192EC0A5A7368B81FBEFBADFBBC" box="[912,1031,999,1025]" country="Argentina" name="Entre Rios" pageId="15" pageNumber="16">Entre Rios</collectingRegion>
|
||
/
|
||
<date id="3150C9B0EC0A5A736F371FBEFBD4FBBC" box="[1055,1150,999,1025]" pageId="15" pageNumber="16" value="1963-11">XI.1963</date>
|
||
, leg. Pena”, [white handwritten] “
|
||
<taxonomicName id="82EE94F3EC0A5A736A281852FEC1FB98" authorityName="Guerin-Meneville" authorityYear="1831" box="[256,363,1035,1061]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
|
||
/ forcipatus / n sp / [typeset] det. G.
|
||
<bibRefCitation id="217F9281EC0A5A73682E1852FC28FB98" author="Frey, G." box="[774,898,1035,1061]" pageId="15" pageNumber="16" pagination="281 - 287" refId="ref36196" refString="Frey, G. (1970) Beitrag zur kenntnis der Sudamerikanischen Melolonthiden (Col.). Entomologische Arbeiten aus dem Museum G. Frey, 21, 281 - 287." type="journal article" year="1970">Frey, 1970</bibRefCitation>
|
||
”, [red typeset] “
|
||
<typeStatus id="9A5551D2EC0A5A736F1D1852FB60FB98" box="[1077,1226,1035,1061]" pageId="15" pageNumber="16" type="paratype">PARATYPE</typeStatus>
|
||
”.
|
||
</paragraph>
|
||
<paragraph id="4551EF70EC0A5A736BEF1876FAFFFB2C" blockId="15.[151,1437,782,2033]" pageId="15" pageNumber="16">
|
||
<emphasis id="779A3362EC0A5A736BEF1876FE0BFBF5" bold="true" box="[199,417,1070,1097]" pageId="15" pageNumber="16">Non-type material</emphasis>
|
||
(4).
|
||
<materialsCitation id="F586E52DEC0A5A736AFB1876FA32FBF4" ID-GBIF-Occurrence="3047558421" box="[467,1432,1071,1097]" collectingDate="1942-01" collectionCode="MZUC" country="Argentina" location="Entre Rios" pageId="15" pageNumber="16" specimenCount="1" specimenCount-male="1" stateProvince="Entre Rios">
|
||
<collectingCountry id="3DF9AFE0EC0A5A736AFB1876FDD0FBF4" box="[467,634,1071,1097]" name="Argentina" pageId="15" pageNumber="16">ARGENTINA</collectingCountry>
|
||
.
|
||
<collectingRegion id="872A2192EC0A5A7369AB1876FD54FBF4" box="[643,766,1071,1097]" country="Argentina" name="Entre Rios" pageId="15" pageNumber="16">Entre Ríos</collectingRegion>
|
||
: without locality,
|
||
<date id="3150C9B0EC0A5A7368EA1876FBA6FBF4" box="[962,1036,1071,1097]" pageId="15" pageNumber="16" value="1942-01">
|
||
<collectingDate id="21143058EC0A5A7368EA1876FBA6FBF4" box="[962,1036,1071,1097]" pageId="15" pageNumber="16" value="1942-01">I.1942</collectingDate>
|
||
</date>
|
||
, without collector,
|
||
<specimenCount id="53E824F9EC0A5A736FC81876FA80FBF4" box="[1248,1322,1071,1097]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
|
||
(
|
||
<collectionCode id="23FF77B5EC0A5A736E111876FA3AFBF4" box="[1337,1424,1071,1097]" country="Chile" name="Museo de Zoologia, Universidad de Concepcion" pageId="15" pageNumber="16">MZUC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="F586E52DEC0A5A736BBF180AFD76FBD0" ID-GBIF-Occurrence="3047558351" box="[151,732,1107,1133]" collectingDate="1987-11" collectionCode="UNSM" country="Argentina" location="Liebig" pageId="15" pageNumber="16" specimenCount="1" specimenCount-male="1" stateProvince="Entre Rios">
|
||
<location id="4031B9ABEC0A5A736BBF180AFF49FBD0" LSID="urn:lsid:plazi:treatment:CD475E66EC0A5A6E6BBF1E7AFE1BFBE4:4031B9ABEC0A5A736BBF180AFF49FBD0" box="[151,227,1107,1133]" country="Argentina" name="Liebig" pageId="15" pageNumber="16" stateProvince="Entre Rios">Liebig</location>
|
||
,
|
||
<date id="3150C9B0EC0A5A736BC6180AFEE7FBD0" box="[238,333,1107,1133]" pageId="15" pageNumber="16" value="1987-11">
|
||
<collectingDate id="21143058EC0A5A736BC6180AFEE7FBD0" box="[238,333,1107,1133]" pageId="15" pageNumber="16" value="1987-11">XI.1987</collectingDate>
|
||
</date>
|
||
, without collector,
|
||
<specimenCount id="53E824F9EC0A5A73690B180AFDC4FBD0" box="[547,622,1107,1133]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
|
||
(
|
||
<collectionCode id="23FF77B5EC0A5A736955180AFD7EFBD0" box="[637,724,1107,1133]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="15" pageNumber="16">UNSM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="F586E52DEC0A5A7369CF180AFECBFB2C" ID-GBIF-Occurrence="3047558486" collectingDate="2008-01" collectionCode="IADIZA" collectorName="J. Corbelli" country="Argentina" elevation="48" latitude="-31.955433" location="Ubajay" longLatPrecision="1" longitude="-58.2969" pageId="15" pageNumber="16" specimenCount="1" specimenCount-female="1" stateProvince="Entre Rios">
|
||
<location id="4031B9ABEC0A5A7369CF180AFC93FBD0" LSID="urn:lsid:plazi:treatment:CD475E66EC0A5A6E6BBF1E7AFE1BFBE4:4031B9ABEC0A5A7369CF180AFC93FBD0" box="[743,825,1107,1133]" country="Argentina" latitude="-31.955433" longLatPrecision="1" longitude="-58.2969" name="Ubajay" pageId="15" pageNumber="16" stateProvince="Entre Rios">Ubajay</location>
|
||
,
|
||
<quantity id="82164295EC0A5A73686C180AFCD5FBD0" box="[836,895,1107,1133]" metricMagnitude="1" metricUnit="m" metricValue="4.8" pageId="15" pageNumber="16" unit="m" value="48.0">
|
||
<elevation id="CEC30843EC0A5A73686C180AFCD5FBD0" box="[836,895,1107,1133]" metricMagnitude="1" metricUnit="m" metricValue="4.8" pageId="15" pageNumber="16" unit="m" value="48.0">48 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate id="20DA89B7EC0A5A7368A2180DFBB8FBD2" box="[906,1042,1108,1135]" degrees="31" direction="south" minutes="57.326" orientation="latitude" pageId="15" pageNumber="16" precision="1" value="-31.955433">31º57.326ʹS</geoCoordinate>
|
||
<geoCoordinate id="20DA89B7EC0A5A736F30180DFB01FBD2" box="[1048,1195,1108,1135]" degrees="58" direction="west" minutes="17.814" orientation="longitude" pageId="15" pageNumber="16" precision="1" value="-58.2969">58º17.814ʹW</geoCoordinate>
|
||
,
|
||
<date id="3150C9B0EC0A5A736F9E180AFAABFBD0" box="[1206,1281,1107,1133]" pageId="15" pageNumber="16" value="2008-01">
|
||
<collectingDate id="21143058EC0A5A736F9E180AFAABFBD0" box="[1206,1281,1107,1133]" pageId="15" pageNumber="16" value="2008-01">I.2008</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="E81B8AA6EC0A5A736E24180AFA2EFBD0" box="[1292,1412,1107,1133]" pageId="15" pageNumber="16">J. Corbelli</collectorName>
|
||
,
|
||
<specimenCount id="53E824F9EC0A5A736EA6180AFF49FB2C" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
|
||
(
|
||
<collectionCode id="23FF77B5EC0A5A736BDA182EFEF3FB2C" box="[242,345,1143,1169]" pageId="15" pageNumber="16">IADIZA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="F586E52DEC0A5A736A44182EFAFBFB2C" ID-GBIF-Occurrence="3047558338" box="[364,1361,1142,1169]" collectionCode="MNHN" collectorName="Rio de la Plata" country="Argentina" location="Buenos Aires" pageId="15" pageNumber="16" specimenCount="1" specimenCount-male="1" stateProvince="Ciudad Autonoma de Buenos Aires">
|
||
<collectingRegion id="872A2192EC0A5A736A44182EFDADFB2C" box="[364,519,1143,1169]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="15" pageNumber="16">Buenos Aires</collectingRegion>
|
||
:
|
||
<collectorName id="E81B8AA6EC0A5A736939182EFD12FB2C" box="[529,696,1143,1169]" pageId="15" pageNumber="16">Rio de la Plata</collectorName>
|
||
, [without month].1906, without collector,
|
||
<specimenCount id="53E824F9EC0A5A736FBA182EFB77FB2C" box="[1170,1245,1143,1169]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
|
||
(
|
||
<collectionCode id="23FF77B5EC0A5A736FC5182EFAE3FB2C" box="[1261,1353,1143,1169]" country="France" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="16">MNHN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DF4BCFBEC0A5A736BEF18C3FE1FFA33" pageId="15" pageNumber="16" type="diagnosis">
|
||
<paragraph id="4551EF70EC0A5A736BEF18C3FE1FFA33" blockId="15.[151,1437,782,2033]" pageId="15" pageNumber="16">
|
||
<emphasis id="779A3362EC0A5A736BEF18C3FE90FB09" bold="true" box="[199,314,1178,1204]" pageId="15" pageNumber="16">Diagnosis</emphasis>
|
||
. Body elongate, wider on the posterior third; elytra non-uniform brown in color that varies from light to dark brown, basal portion darker, as are the head and pronotum (
|
||
<figureCitation id="DDD5F3F5EC0A5A7368ED18E6FB83FB64" box="[965,1065,1215,1241]" captionStart="FIGURE 6" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@16.[203,1385,181,1920]" captionTargetPageId="16" captionText="FIGURE 6. Liogenys forcipata Frey, 1970 male views: A) dorsal, B) head and pronotum, C) lateral, D) base of metatibia, white arrow indicates basal apophysis and metatrochanter, E) pygidium, and parameres in views: F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561276" httpUri="https://zenodo.org/record/4561276/files/figure.png" pageId="15" pageNumber="16">Figs. 6A</figureCitation>
|
||
,
|
||
<figureCitation id="DDD5F3F5EC0A5A736F1D18E6FBF2FB64" box="[1077,1112,1215,1241]" captionStart="FIGURE 7" captionStartId="17.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@17.[203,1385,181,1920]" captionTargetPageId="17" captionText="FIGURE 7. Liogenys tarsalis Moser, 1921 female views: A) dorsal, C) head and pronotum, E) pygidium; compared with L. forcipata Frey, 1970 female views: B) dorsal, D) head and pronotum, F) pygidium." figureDoi="http://doi.org/10.5281/zenodo.4561280" httpUri="https://zenodo.org/record/4561280/files/figure.png" pageId="15" pageNumber="16">7B</figureCitation>
|
||
); clypeal emargination deep, rounded and wide; outer sides of anterior teeth subparallel; clypeal lateral margin rounded and produced (
|
||
<figureCitation id="DDD5F3F5EC0A5A736E1D18BAFA32FB40" box="[1333,1432,1251,1277]" captionStart="FIGURE 6" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@16.[203,1385,181,1920]" captionTargetPageId="16" captionText="FIGURE 6. Liogenys forcipata Frey, 1970 male views: A) dorsal, B) head and pronotum, C) lateral, D) base of metatibia, white arrow indicates basal apophysis and metatrochanter, E) pygidium, and parameres in views: F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561276" httpUri="https://zenodo.org/record/4561276/files/figure.png" pageId="15" pageNumber="16">Figs. 6B</figureCitation>
|
||
,
|
||
<figureCitation id="DDD5F3F5EC0A5A736BBF195EFF17FA9C" box="[151,189,1287,1313]" captionStart="FIGURE 7" captionStartId="17.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@17.[203,1385,181,1920]" captionTargetPageId="17" captionText="FIGURE 7. Liogenys tarsalis Moser, 1921 female views: A) dorsal, C) head and pronotum, E) pygidium; compared with L. forcipata Frey, 1970 female views: B) dorsal, D) head and pronotum, F) pygidium." figureDoi="http://doi.org/10.5281/zenodo.4561280" httpUri="https://zenodo.org/record/4561280/files/figure.png" pageId="15" pageNumber="16">7D</figureCitation>
|
||
); basal apophysis of metacoxa not produced (
|
||
<figureCitation id="DDD5F3F5EC0A5A736990195EFCBBFA9F" box="[696,785,1287,1314]" captionStart="FIGURE 6" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@16.[203,1385,181,1920]" captionTargetPageId="16" captionText="FIGURE 6. Liogenys forcipata Frey, 1970 male views: A) dorsal, B) head and pronotum, C) lateral, D) base of metatibia, white arrow indicates basal apophysis and metatrochanter, E) pygidium, and parameres in views: F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561276" httpUri="https://zenodo.org/record/4561276/files/figure.png" pageId="15" pageNumber="16">Fig. 6D</figureCitation>
|
||
); in males tarsi semi-opaque and enlarged, mainly the protarsomeres and mesotarsomeres; pygidium flat or slightly convex; trapezoidal; pygidial disc glabrous (
|
||
<figureCitation id="DDD5F3F5EC0A5A736E2C1972FAC9FAF8" box="[1284,1379,1323,1349]" captionStart="FIGURE 6" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@16.[203,1385,181,1920]" captionTargetPageId="16" captionText="FIGURE 6. Liogenys forcipata Frey, 1970 male views: A) dorsal, B) head and pronotum, C) lateral, D) base of metatibia, white arrow indicates basal apophysis and metatrochanter, E) pygidium, and parameres in views: F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561276" httpUri="https://zenodo.org/record/4561276/files/figure.png" pageId="15" pageNumber="16">Figs. 6E</figureCitation>
|
||
,
|
||
<figureCitation id="DDD5F3F5EC0A5A736E461972FA24FAF8" box="[1390,1422,1323,1349]" captionStart="FIGURE 7" captionStartId="17.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@17.[203,1385,181,1920]" captionTargetPageId="17" captionText="FIGURE 7. Liogenys tarsalis Moser, 1921 female views: A) dorsal, C) head and pronotum, E) pygidium; compared with L. forcipata Frey, 1970 female views: B) dorsal, D) head and pronotum, F) pygidium." figureDoi="http://doi.org/10.5281/zenodo.4561280" httpUri="https://zenodo.org/record/4561280/files/figure.png" pageId="15" pageNumber="16">7F</figureCitation>
|
||
); parameres setose at the subapical portion; inner margins strongly convergent, narrowed apically and apex bilobed, curved inwards (
|
||
<figureCitation id="DDD5F3F5EC0A5A736A7A192AFE02FA33" box="[338,424,1395,1422]" captionStart="FIGURE 6" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@16.[203,1385,181,1920]" captionTargetPageId="16" captionText="FIGURE 6. Liogenys forcipata Frey, 1970 male views: A) dorsal, B) head and pronotum, C) lateral, D) base of metatibia, white arrow indicates basal apophysis and metatrochanter, E) pygidium, and parameres in views: F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561276" httpUri="https://zenodo.org/record/4561276/files/figure.png" pageId="15" pageNumber="16">Fig. 6F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DF4BCFBEC0A5A6E6BEF19CFFEC4FDDF" lastPageId="18" lastPageNumber="19" pageId="15" pageNumber="16" type="description">
|
||
<paragraph id="4551EF70EC0A5A6E6BEF19CFFEC4FDDF" blockId="15.[151,1437,782,2033]" lastBlockId="18.[151,1437,151,1113]" lastPageId="18" lastPageNumber="19" pageId="15" pageNumber="16">
|
||
<emphasis id="779A3362EC0A5A736BEF19CFFEC7FA0D" bold="true" box="[199,365,1430,1456]" pageId="15" pageNumber="16">Redescription</emphasis>
|
||
. Length 11.4–12.0 mm; width: 6.0–
|
||
<quantity id="82164295EC0A5A73683A19CEFCC4FA0C" box="[786,878,1431,1457]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="15" pageNumber="16" unit="mm" value="6.6">6.6 mm</quantity>
|
||
. Brown.
|
||
<emphasis id="779A3362EC0A5A7368F119CEFBB5FA0C" box="[985,1055,1431,1457]" italics="true" pageId="15" pageNumber="16">Head:</emphasis>
|
||
distance between eyes twice the width of one eye; clypeal emargination deep, rounded, and wide; outer sides of anterior teeth parallel; outer margin of anterior teeth slightly shorter than the eye; lateral margin convex and produced, sometimes forming a rounded projection; distance between clypeal lateral projection and anterior margin of eye as long as one eye or shorter; distance between clypeal lateral projection and apex of the adjacent anterior tooth equal to the basal width of the anterior tooth; angle between outer side of anterior teeth and clypeal lateral projection approximately 90º; distal maxillary palpomere, maximum width slightly wider than the apex; sensorial region smooth or forming a shallow fovea, extending past the transverse midline of the palpomere; antenna with 10 antennomeres, club lighter in color and 1.5 times the length of the funicle in males, slightly longer in females.
|
||
<emphasis id="779A3362EC0A5A7368DD1AEEFBE5F96C" box="[1013,1103,1719,1745]" italics="true" pageId="15" pageNumber="16">Thorax:</emphasis>
|
||
anterior margin of pronotum slightly produced medially; pronotal disc sometimes sulcate medially, glabrous, punctures coarse and sparse, denser near the anterior margin; pronotal posterior corners rounded; hypomere with short bristles and scattered scales, mesepisternum scaly; sides of metaventrite with sparse erect bristles; distance between mesocoxae and metacoxae up to twice the length of the metacoxa; scutellum ogival to rounded, sparsely and coarsely punctate.
|
||
<emphasis id="779A3362EC0A5A736E2C1B1EFAF9F8DC" box="[1284,1363,1863,1889]" italics="true" pageId="15" pageNumber="16">Elytra:</emphasis>
|
||
shiny, glabrous, brown to dark brown, sometimes apparently burnt; elytral suture darker than elytron and elevated, pair of inner ridges more defined than the three outer pairs.
|
||
<emphasis id="779A3362EC0A5A7368381BD6FCE5F815" box="[784,847,1935,1960]" italics="true" pageId="15" pageNumber="16">Legs:</emphasis>
|
||
three protibial teeth, the middle and apical equal in size; the three teeth equally spaced; mesofemural surface setose, thick erect bristles on posterior margin; mesotibia cylindrical in cross section, surface coarsely sculptured; two transverse carinae, the apical one complete; metacoxa with sparse bristles near the base of the leg; basal apophysis of metacoxa not produced (
|
||
<figureCitation id="DDD5F3F5EC175A6E6F401CCEFB6BFF0F" box="[1128,1217,151,178]" captionStart="FIGURE 6" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@16.[203,1385,181,1920]" captionTargetPageId="16" captionText="FIGURE 6. Liogenys forcipata Frey, 1970 male views: A) dorsal, B) head and pronotum, C) lateral, D) base of metatibia, white arrow indicates basal apophysis and metatrochanter, E) pygidium, and parameres in views: F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561276" httpUri="https://zenodo.org/record/4561276/files/figure.png" pageId="18" pageNumber="19">Fig. 6D</figureCitation>
|
||
); metatibia carinate along the inner margin, apex not produced, apical inner surface setose; two metatibial transverse carinae present posteriorly; metatibial apical spurs of different lengths; the larger equal in length to the diameter of the tibial apex; in males tarsi opaque, with tarsomeres I to IV enlarged in every leg; protarsomeres and mesotarsomeres almost twice the width of the metatarsomeres; protarsomere II wide; metatarsomere I shorter than metatarsomere II; claw bifid, symmetrical, superior tooth of a claw longer than the inferior and equal in width, distance between teeth more than twice the length of the inferior tooth; inferior tooth oblique to the axis of the superior tooth.
|
||
<emphasis id="779A3362EC175A6E6FA31D36FB54FE34" box="[1163,1278,367,393]" italics="true" pageId="18" pageNumber="19">Abdomen:</emphasis>
|
||
ventrites with scattered bristles on disc; propygidium slightly visible, glabrous; pygidium flat or slightly convex, subtrapezoidal, wide; pygidial width not exceeding distance between spiracles of propygidium; pygidial disc glabrous; pygidial apex rounded.
|
||
<emphasis id="779A3362EC175A6E6A141D82FE1CFE49" box="[316,438,475,500]" italics="true" pageId="18" pageNumber="19">Parameres</emphasis>
|
||
: basal region narrower than both sections of the parameres at its maximum width, parameral split at the third portion, inner margins convergent; apex bilobed and curved inwards, inner margins overlapped; parameres abundantly setose on subapical portion; parameres in lateral view not flattened subapically; straight, coplanar (
|
||
<figureCitation id="DDD5F3F5EC175A6E6A2F1E1EFECBFDDF" box="[263,353,583,610]" captionStart="FIGURE 6" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@16.[203,1385,181,1920]" captionTargetPageId="16" captionText="FIGURE 6. Liogenys forcipata Frey, 1970 male views: A) dorsal, B) head and pronotum, C) lateral, D) base of metatibia, white arrow indicates basal apophysis and metatrochanter, E) pygidium, and parameres in views: F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561276" httpUri="https://zenodo.org/record/4561276/files/figure.png" pageId="18" pageNumber="19">Fig. 6G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="1191BFF8EC155A6C6BBF1BCEFA83F869" ID-DOI="http://doi.org/10.5281/zenodo.4561276" ID-Zenodo-Dep="4561276" httpUri="https://zenodo.org/record/4561276/files/figure.png" pageId="16" pageNumber="17" startId="16.[151,250,1943,1967]" targetBox="[203,1385,181,1920]" targetPageId="16">
|
||
<paragraph id="4551EF70EC155A6C6BBF1BCEFA83F869" blockId="16.[151,1436,1943,2004]" pageId="16" pageNumber="17">
|
||
<emphasis id="779A3362EC155A6C6BBF1BCEFEB8F812" bold="true" box="[151,274,1943,1967]" pageId="16" pageNumber="17">FIGURE 6.</emphasis>
|
||
<taxonomicName id="82EE94F3EC155A6C6A301BC1FDEDF80D" authority="Frey, 1970" authorityName="Frey" authorityYear="1970" box="[280,583,1944,1968]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="forcipata">
|
||
<emphasis id="779A3362EC155A6C6A301BC1FE7FF80D" box="[280,469,1944,1968]" italics="true" pageId="16" pageNumber="17">Liogenys forcipata</emphasis>
|
||
<bibRefCitation id="217F9281EC155A6C6AF31BC1FDEDF80D" author="Frey, G." box="[475,583,1944,1968]" pageId="16" pageNumber="17" pagination="281 - 287" refId="ref36196" refString="Frey, G. (1970) Beitrag zur kenntnis der Sudamerikanischen Melolonthiden (Col.). Entomologische Arbeiten aus dem Museum G. Frey, 21, 281 - 287." type="journal article" year="1970">Frey, 1970</bibRefCitation>
|
||
</taxonomicName>
|
||
male views: A) dorsal, B) head and pronotum, C) lateral, D) base of metatibia, white arrow indicates basal apophysis and metatrochanter, E) pygidium, and parameres in views: F) dorsal and G) lateral.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="1191BFF8EC145A6D6BBF1BCEFC4BF869" ID-DOI="http://doi.org/10.5281/zenodo.4561280" ID-Zenodo-Dep="4561280" httpUri="https://zenodo.org/record/4561280/files/figure.png" pageId="17" pageNumber="18" startId="17.[151,250,1943,1967]" targetBox="[203,1385,181,1920]" targetPageId="17">
|
||
<paragraph id="4551EF70EC145A6D6BBF1BCEFC4BF869" blockId="17.[151,1437,1943,2004]" pageId="17" pageNumber="18">
|
||
<emphasis id="779A3362EC145A6D6BBF1BCEFEBFF80D" bold="true" box="[151,277,1943,1968]" pageId="17" pageNumber="18">FIGURE 7.</emphasis>
|
||
<taxonomicName id="82EE94F3EC145A6D6A351BC1FDFFF80D" authority="Moser, 1921" authorityName="Moser" authorityYear="1921" box="[285,597,1944,1968]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="tarsalis">
|
||
<emphasis id="779A3362EC145A6D6A351BC1FE66F80D" box="[285,460,1944,1968]" italics="true" pageId="17" pageNumber="18">Liogenys tarsalis</emphasis>
|
||
<bibRefCitation id="217F9281EC145A6D6AFC1BC1FDFFF80D" author="Moser, J." box="[468,597,1944,1968]" pageId="17" pageNumber="18" pagination="133 - 182" refId="ref36844" refString="Moser, J. (1921) Neue Melolonthiden von Mittel- und S ʾ d-Amerika. Stettiner Entomologische Zeitung, 82, 133 - 182. https: // doi. org / 10.1002 / mmnd. 192119210312" type="journal article" year="1921">Moser, 1921</bibRefCitation>
|
||
</taxonomicName>
|
||
female views: A) dorsal, C) head and pronotum, E) pygidium; compared with
|
||
<taxonomicName id="82EE94F3EC145A6D6EA01BC1FECCF869" authority="Frey, 1970" authorityName="Frey" authorityYear="1970" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="forcipata">
|
||
<emphasis id="779A3362EC145A6D6EA01BC1FF59F869" italics="true" pageId="17" pageNumber="18">L. forcipata</emphasis>
|
||
<bibRefCitation id="217F9281EC145A6D6BD11BE5FECCF869" author="Frey, G." box="[249,358,1980,2004]" pageId="17" pageNumber="18" pagination="281 - 287" refId="ref36196" refString="Frey, G. (1970) Beitrag zur kenntnis der Sudamerikanischen Melolonthiden (Col.). Entomologische Arbeiten aus dem Museum G. Frey, 21, 281 - 287." type="journal article" year="1970">Frey, 1970</bibRefCitation>
|
||
</taxonomicName>
|
||
female views: B) dorsal, D) head and pronotum, F) pygidium.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="0DF4BCFBEC175A6E6BEF1E32FC45FD14" pageId="18" pageNumber="19" type="materials_examined">
|
||
<paragraph id="4551EF70EC175A6E6BEF1E32FCC8FD38" blockId="18.[151,1437,151,1113]" box="[199,866,618,645]" pageId="18" pageNumber="19">
|
||
<materialsCitation id="F586E52DEC175A6E6BEF1E32FCF4FD38" ID-GBIF-Occurrence="3047558389" box="[199,862,618,645]" country="Argentina" location="Pronunciamento" pageId="18" pageNumber="19" specimenCount="1" stateProvince="Entre Rios" typeStatus="holotype">
|
||
<emphasis id="779A3362EC175A6E6BEF1E32FECAFD39" bold="true" box="[199,352,618,645]" pageId="18" pageNumber="19">
|
||
<typeStatus id="9A5551D2EC175A6E6BEF1E32FEA8FD38" box="[199,258,619,645]" pageId="18" pageNumber="19">Type</typeStatus>
|
||
locality
|
||
</emphasis>
|
||
.
|
||
<collectingCountry id="3DF9AFE0EC175A6E6A441E32FDB8FD38" box="[364,530,619,645]" name="Argentina" pageId="18" pageNumber="19">ARGENTINA</collectingCountry>
|
||
,
|
||
<collectingRegion id="872A2192EC175A6E69361E32FD3DFD38" box="[542,663,619,645]" country="Argentina" name="Entre Rios" pageId="18" pageNumber="19">Entre Ríos</collectingRegion>
|
||
,
|
||
<location id="4031B9ABEC175A6E698B1E32FCF4FD38" LSID="urn:lsid:plazi:treatment:CD475E66EC0A5A6E6BBF1E7AFE1BFBE4:4031B9ABEC175A6E698B1E32FCF4FD38" box="[675,862,619,645]" country="Argentina" name="Pronunciamento" pageId="18" pageNumber="19" stateProvince="Entre Rios">Pronunciamento</location>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="4551EF70EC175A6E6BEF1ED7FC45FD14" blockId="18.[151,1437,151,1113]" box="[199,1007,654,681]" pageId="18" pageNumber="19">
|
||
<emphasis id="779A3362EC175A6E6BEF1ED7FE54FD15" bold="true" box="[199,510,654,680]" pageId="18" pageNumber="19">Geographical distribution</emphasis>
|
||
.
|
||
<materialsCitation id="F586E52DEC175A6E69241ED6FC43FD14" ID-GBIF-Occurrence="3047558478" box="[524,1001,654,681]" country="Argentina" location="Entre Rios" pageId="18" pageNumber="19" specimenCount="1" stateProvince="Entre Rios">
|
||
<collectingCountry id="3DF9AFE0EC175A6E69241ED6FD04FD14" box="[524,686,655,681]" name="Argentina" pageId="18" pageNumber="19">ARGENTINA</collectingCountry>
|
||
(
|
||
<collectingRegion id="872A2192EC175A6E69941ED6FC9DFD14" box="[700,823,655,681]" country="Argentina" name="Entre Rios" pageId="18" pageNumber="19">Entre Ríos</collectingRegion>
|
||
,
|
||
<emphasis id="779A3362EC175A6E686B1ED6FC74FD15" bold="true" box="[835,990,654,681]" pageId="18" pageNumber="19">
|
||
<collectingRegion id="872A2192EC175A6E686B1ED6FC74FD15" box="[835,990,654,681]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="18" pageNumber="19">Buenos Aires</collectingRegion>
|
||
</emphasis>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DF4BCFBEC175A6E6BEF1EEAFE1BFBE4" pageId="18" pageNumber="19" type="discussion">
|
||
<paragraph id="4551EF70EC175A6E6BEF1EEAFE1BFBE4" blockId="18.[151,1437,151,1113]" pageId="18" pageNumber="19">
|
||
<emphasis id="779A3362EC175A6E6BEF1EEAFE91FD70" bold="true" box="[199,315,691,717]" pageId="18" pageNumber="19">Remarks.</emphasis>
|
||
<taxonomicName id="82EE94F3EC175A6E6A6D1EEAFDB3FD70" authorityName="Frey" authorityYear="1970" box="[325,537,691,717]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="forcipata">
|
||
<emphasis id="779A3362EC175A6E6A6D1EEAFDB3FD70" box="[325,537,691,717]" italics="true" pageId="18" pageNumber="19">Liogenys forcipata</emphasis>
|
||
</taxonomicName>
|
||
resembles
|
||
<taxonomicName id="82EE94F3EC175A6E69B41EEAFC03FD70" authority="Moser, 1921" authorityName="Moser" authorityYear="1921" box="[668,937,691,717]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="tarsalis">
|
||
<emphasis id="779A3362EC175A6E69B41EEAFCBAFD70" box="[668,784,691,717]" italics="true" pageId="18" pageNumber="19">L. tarsalis</emphasis>
|
||
<bibRefCitation id="217F9281EC175A6E68321EEAFC03FD70" author="Moser, J." box="[794,937,691,717]" pageId="18" pageNumber="19" pagination="133 - 182" refId="ref36844" refString="Moser, J. (1921) Neue Melolonthiden von Mittel- und S ʾ d-Amerika. Stettiner Entomologische Zeitung, 82, 133 - 182. https: // doi. org / 10.1002 / mmnd. 192119210312" type="journal article" year="1921">Moser, 1921</bibRefCitation>
|
||
</taxonomicName>
|
||
(Fig. 75,
|
||
<bibRefCitation id="217F9281EC175A6E6F361EEAFAA7FD70" author="Cherman, M. A. & Mise, K. M. & Moron, M. A. & Vaz-de-Mello, F. Z. & Almeida, L. M." box="[1054,1293,691,717]" pageId="18" pageNumber="19" pagination="1 - 120" refId="ref35643" refString="Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. & Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031" type="journal article" year="2017">
|
||
Cherman
|
||
<emphasis id="779A3362EC175A6E6FA71EEAFB63FD70" box="[1167,1225,691,717]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
) in the nonuniform color of the elytra and somewhat in the shape of the clypeus, which is strongly emarginate. It differs from
|
||
<taxonomicName id="82EE94F3EC175A6E6BBF1EA2FEA3FCA8" authorityName="Moser" authorityYear="1921" box="[151,265,763,789]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="tarsalis">
|
||
<emphasis id="779A3362EC175A6E6BBF1EA2FEA3FCA8" box="[151,265,763,789]" italics="true" pageId="18" pageNumber="19">L. tarsalis</emphasis>
|
||
</taxonomicName>
|
||
(in parenthesis) in the body size slightly wider on posterior third (uniform width (
|
||
<figureCitation id="DDD5F3F5EC175A6E6F811EA2FAA9FCAB" box="[1193,1283,763,790]" captionStart="FIGURE 6" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@16.[203,1385,181,1920]" captionTargetPageId="16" captionText="FIGURE 6. Liogenys forcipata Frey, 1970 male views: A) dorsal, B) head and pronotum, C) lateral, D) base of metatibia, white arrow indicates basal apophysis and metatrochanter, E) pygidium, and parameres in views: F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561276" httpUri="https://zenodo.org/record/4561276/files/figure.png" pageId="18" pageNumber="19">Fig. 6A</figureCitation>
|
||
)); the clypeal emargination slightly wider, with teeth shorter; the clypeal lateral margin more convex and produced (
|
||
<figureCitation id="DDD5F3F5EC175A6E6FD51F46FAF4FC84" box="[1277,1374,799,825]" captionStart="FIGURE 6" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@16.[203,1385,181,1920]" captionTargetPageId="16" captionText="FIGURE 6. Liogenys forcipata Frey, 1970 male views: A) dorsal, B) head and pronotum, C) lateral, D) base of metatibia, white arrow indicates basal apophysis and metatrochanter, E) pygidium, and parameres in views: F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561276" httpUri="https://zenodo.org/record/4561276/files/figure.png" pageId="18" pageNumber="19">Figs. 6C</figureCitation>
|
||
,
|
||
<figureCitation id="DDD5F3F5EC175A6E6E411F46FA25FC84" box="[1385,1423,799,825]" captionStart="FIGURE 7" captionStartId="17.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@17.[203,1385,181,1920]" captionTargetPageId="17" captionText="FIGURE 7. Liogenys tarsalis Moser, 1921 female views: A) dorsal, C) head and pronotum, E) pygidium; compared with L. forcipata Frey, 1970 female views: B) dorsal, D) head and pronotum, F) pygidium." figureDoi="http://doi.org/10.5281/zenodo.4561280" httpUri="https://zenodo.org/record/4561280/files/figure.png" pageId="18" pageNumber="19">7D</figureCitation>
|
||
); the pronotum globose (flatter); the metacoxae glabrous (scaly throughout); the pygidium wider (
|
||
<figureCitation id="DDD5F3F5EC175A6E6FF91F1AFA98FCE0" box="[1233,1330,835,861]" captionStart="FIGURE 6" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@16.[203,1385,181,1920]" captionTargetPageId="16" captionText="FIGURE 6. Liogenys forcipata Frey, 1970 male views: A) dorsal, B) head and pronotum, C) lateral, D) base of metatibia, white arrow indicates basal apophysis and metatrochanter, E) pygidium, and parameres in views: F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561276" httpUri="https://zenodo.org/record/4561276/files/figure.png" pageId="18" pageNumber="19">Figs. 6E</figureCitation>
|
||
,
|
||
<figureCitation id="DDD5F3F5EC175A6E6E161F1AFAF5FCE0" box="[1342,1375,835,861]" captionStart="FIGURE 7" captionStartId="17.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@17.[203,1385,181,1920]" captionTargetPageId="17" captionText="FIGURE 7. Liogenys tarsalis Moser, 1921 female views: A) dorsal, C) head and pronotum, E) pygidium; compared with L. forcipata Frey, 1970 female views: B) dorsal, D) head and pronotum, F) pygidium." figureDoi="http://doi.org/10.5281/zenodo.4561280" httpUri="https://zenodo.org/record/4561280/files/figure.png" pageId="18" pageNumber="19">7F</figureCitation>
|
||
); and in males the tarsi strongly enlarged but less than in
|
||
<taxonomicName id="82EE94F3EC175A6E69F01F3EFCE3FC3C" authorityName="Moser" authorityYear="1921" box="[728,841,871,897]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="tarsalis">
|
||
<emphasis id="779A3362EC175A6E69F01F3EFCE3FC3C" box="[728,841,871,897]" italics="true" pageId="18" pageNumber="19">L. tarsalis</emphasis>
|
||
</taxonomicName>
|
||
, and the metatarsi less enlarged than the protarsi and mesotarsi (tarsi strongly enlarged in all legs).
|
||
<taxonomicName id="82EE94F3EC175A6E69B71FD2FCD8FC18" authorityName="Frey" authorityYear="1970" box="[671,882,907,933]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="forcipata">
|
||
<emphasis id="779A3362EC175A6E69B71FD2FCD8FC18" box="[671,882,907,933]" italics="true" pageId="18" pageNumber="19">Liogenys forcipata</emphasis>
|
||
</taxonomicName>
|
||
does not occur in the
|
||
<collectingRegion id="872A2192EC175A6E6F471FD2FA8EFC18" box="[1135,1316,907,933]" country="Argentina" name="Chaco" pageId="18" pageNumber="19">Chaco Province</collectingRegion>
|
||
, but in the Espinal boundaring ecorregion (
|
||
<figureCitation id="DDD5F3F5EC175A6E6AD41FF6FDF2FC74" box="[508,600,943,969]" captionStart="FIGURE 23" captionStartId="51.[151,250,1451,1475]" captionTargetBox="[171,1416,192,1415]" captionTargetId="figure-20@51.[151,1435,181,1427]" captionTargetPageId="51" captionText="FIGURE 23. Geographical distribution of four known Liogenys species occurring in the Chacoan province and its boundaries. Shaped points: previously known species; colors: Neotropical provinces (Morrone 2014) and Monte province from the South American Transition Zone (Roig-Juñent et al. 2018)." figureDoi="http://doi.org/10.5281/zenodo.4561324" httpUri="https://zenodo.org/record/4561324/files/figure.png" pageId="18" pageNumber="19">Figs. 23</figureCitation>
|
||
,
|
||
<figureCitation id="DDD5F3F5EC175A6E694B1FF6FD29FC74" box="[611,643,943,969]" captionStart="FIGURE 24" captionStartId="52.[151,250,1459,1483]" captionTargetBox="[171,1416,196,1422]" captionTargetId="figure-21@52.[151,1435,181,1434]" captionTargetPageId="52" captionText="FIGURE 24. Geographical distribution of four known Liogenys species occurring in the Chacoan province and its boundaries. Shaped points: previously known species; colors: Neotropical provinces (Morrone 2014) and Monte province from the South American Transition Zone (Roig-Juñent et al. 2018)." figureDoi="http://doi.org/10.5281/zenodo.4561326" httpUri="https://zenodo.org/record/4561326/files/figure.png" pageId="18" pageNumber="19">24</figureCitation>
|
||
). Nonetheless, we opted to include this species due to its similarity with
|
||
<taxonomicName id="82EE94F3EC175A6E6BBF1F8AFEA0FC50" authorityName="Moser" authorityYear="1921" box="[151,266,979,1005]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="tarsalis">
|
||
<emphasis id="779A3362EC175A6E6BBF1F8AFEA0FC50" box="[151,266,979,1005]" italics="true" pageId="18" pageNumber="19">L. tarsalis</emphasis>
|
||
</taxonomicName>
|
||
, especially the females (
|
||
<figureCitation id="DDD5F3F5EC175A6E690A1F8AFD16FC50" box="[546,700,979,1005]" captionStart="FIGURE 7" captionStartId="17.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@17.[203,1385,181,1920]" captionTargetPageId="17" captionText="FIGURE 7. Liogenys tarsalis Moser, 1921 female views: A) dorsal, C) head and pronotum, E) pygidium; compared with L. forcipata Frey, 1970 female views: B) dorsal, D) head and pronotum, F) pygidium." figureDoi="http://doi.org/10.5281/zenodo.4561280" httpUri="https://zenodo.org/record/4561280/files/figure.png" pageId="18" pageNumber="19">Fig. 7B, D, F</figureCitation>
|
||
), and with
|
||
<taxonomicName id="82EE94F3EC175A6E68141F8AFBF0FC50" authority="Cherman" authorityName="Cherman" authorityYear="2021" box="[828,1114,979,1005]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="neoforcipata" status="sp. nov.">
|
||
<emphasis id="779A3362EC175A6E68141F8AFC42FC50" box="[828,1000,979,1005]" italics="true" pageId="18" pageNumber="19">L. neoforcipata</emphasis>
|
||
Cherman
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="779A3362EC175A6E6F4E1F8AFB5AFC51" bold="true" box="[1126,1264,978,1005]" pageId="18" pageNumber="19">
|
||
<taxonomicNameLabel id="6CA98E19EC175A6E6F4E1F8AFB5AFC51" box="[1126,1264,978,1005]" pageId="18" pageNumber="19" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="DDD5F3F5EC175A6E6E291F8AFAFEFC50" box="[1281,1364,979,1005]" captionStart="FIGURE 13" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@29.[203,1385,181,1920]" captionTargetPageId="29" captionText="FIGURE 13. Liogenys neoforcipata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) base of metatibia, white arrow indicates basal apophysis and metatrochanter, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561296" httpUri="https://zenodo.org/record/4561296/files/figure.png" pageId="18" pageNumber="19">Fig. 13</figureCitation>
|
||
), both species present in the
|
||
<collectingRegion id="872A2192EC175A6E6AA41FAEFE7DFBAC" box="[396,471,1015,1041]" country="Argentina" name="Chaco" pageId="18" pageNumber="19">Chaco</collectingRegion>
|
||
.
|
||
<taxonomicName id="82EE94F3EC175A6E6AC91FAEFD19FBAC" authorityName="Frey" authorityYear="1970" box="[481,691,1015,1041]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="forcipata">
|
||
<emphasis id="779A3362EC175A6E6AC91FAEFD19FBAC" box="[481,691,1015,1041]" italics="true" pageId="18" pageNumber="19">Liogenys forcipata</emphasis>
|
||
</taxonomicName>
|
||
can be found in sympatry with
|
||
<taxonomicName id="82EE94F3EC175A6E6F3D1FAEFB3CFBAC" authorityName="Moser" authorityYear="1918" box="[1045,1174,1015,1041]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="latitarsis">
|
||
<emphasis id="779A3362EC175A6E6F3D1FAEFB3CFBAC" box="[1045,1174,1015,1041]" italics="true" pageId="18" pageNumber="19">L. latitarsis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="82EE94F3EC175A6E6F8C1FAEFF7BFB88" authority="Blanchard, 1851" authorityName="Blanchard" authorityYear="1851" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="obscura">
|
||
<emphasis id="779A3362EC175A6E6F8C1FAEFAB1FBAC" box="[1188,1307,1015,1041]" italics="true" pageId="18" pageNumber="19">L. obscura</emphasis>
|
||
<bibRefCitation id="217F9281EC175A6E6E0A1FAEFF7BFB88" author="Blanchard, E." pageId="18" pageNumber="19" pagination="129 - 240" refId="ref35099" refString="Blanchard, E. (1851) s. n. In: Museum D'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Ordre des Coleopteres. Tome I. Part 2. Gide et Baudry, Paris, pp. 129 - 240." type="book chapter" year="1851">Blanchard, 1851</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="82EE94F3EC175A6E6BF61842FDB8FB88" authority="Blanchard, 1851" authorityName="Blanchard" authorityYear="1851" box="[222,530,1050,1077]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pallens">
|
||
<emphasis id="779A3362EC175A6E6BF61842FEE4FB88" box="[222,334,1051,1077]" italics="true" pageId="18" pageNumber="19">L. pallens</emphasis>
|
||
<bibRefCitation id="217F9281EC175A6E6A7E1842FDB8FB88" author="Blanchard, E." box="[342,530,1050,1077]" pageId="18" pageNumber="19" pagination="129 - 240" refId="ref35099" refString="Blanchard, E. (1851) s. n. In: Museum D'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Ordre des Coleopteres. Tome I. Part 2. Gide et Baudry, Paris, pp. 129 - 240." type="book chapter" year="1851">Blanchard, 1851</bibRefCitation>
|
||
</taxonomicName>
|
||
, and sometimes with
|
||
<taxonomicName id="82EE94F3EC175A6E68211842FC0EFB88" authorityName="Moser" authorityYear="1921" box="[777,932,1051,1077]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="cribricollis">
|
||
<emphasis id="779A3362EC175A6E68211842FC0EFB88" box="[777,932,1051,1077]" italics="true" pageId="18" pageNumber="19">L. cribricollis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="82EE94F3EC175A6E68F51842FA98FB88" authority="Fairmaire, 1892" authorityName="Fairmaire" authorityYear="1892" box="[989,1330,1051,1077]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="opacicollis">
|
||
<emphasis id="779A3362EC175A6E68F51842FBDCFB88" box="[989,1142,1051,1077]" italics="true" pageId="18" pageNumber="19">L. opacicollis</emphasis>
|
||
<bibRefCitation id="217F9281EC175A6E6F561842FA98FB88" author="Fairmaire, M. L." box="[1150,1330,1051,1077]" pageId="18" pageNumber="19" pagination="242 - 253" refId="ref36073" refString="Fairmaire, M. L. (1892) Descriptions de quelques coleopteres Argentins. Annales de la Societe Entomologique de Belgique, 36, 242 - 253." type="journal article" year="1892">
|
||
Fairmaire,
|
||
<quantity id="82164295EC175A6E6FD21842FA98FB88" box="[1274,1330,1051,1077]" metricMagnitude="1" metricUnit="m" metricValue="4.80568" pageId="18" pageNumber="19" unit="in" value="1892.0">1892</quantity>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
in relicts of xerophytic vegetation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |