202 lines
27 KiB
XML
202 lines
27 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.463.8499" ID-PMC="PMC4294300" ID-Pensoft-Pub="1313-2970-463-21" ID-Pensoft-UUID="3E5CFFC6FFC91628FFF6E355951DFFF8" ID-PubMed="25589861" ID-Zenodo-Dep="578654" ID-ZooBank="71C1AE92B0CE4D26B1E5076E480180AD" ModsDocID="1313-2970-463-21" checkinTime="1451244962410" checkinUser="pensoft" docAuthor="Solodovnikov, Igor A., Zamotajlov, Alexandr S., Fominykh, Dmitriy D. & Titarenko, Andrey Y." docDate="2014" docId="56C62A04B98059CD8B5DCBF7850C80D9" docLanguage="en" docName="ZooKeys 463: 21-56" docOrigin="ZooKeys 463" docPubDate="2014-12-12" docSource="http://dx.doi.org/10.3897/zookeys.463.8499" docTitle="Carabus (Archiplectes) besleticus subsp. resheviensis Solodovnikov, Zamotajlov & Fominykh 2014, subsp. n." docType="treatment" docVersion="3" id="3E5CFFC6FFC91628FFF6E355951DFFF8" lastPageNumber="38" masterDocId="3E5CFFC6FFC91628FFF6E355951DFFF8" masterDocTitle="Contribution to the knowledge of the Carabus (Archiplectes) satyrus Kurnakov, 1962, species complex in Abkhazia (Coleoptera, Carabidae, Carabini)" masterLastPageNumber="56" masterPageNumber="21" pageNumber="33" updateTime="1668159733340" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Contribution to the knowledge of the Carabus (Archiplectes) satyrus Kurnakov, 1962, species complex in Abkhazia (Coleoptera, Carabidae, Carabini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Solodovnikov, Igor A.</mods:namePart>
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<mods:affiliation>Vitebsk State P. M. Masherov University, Moskovskyi prospect, 33, Vitebsk 210038, Belarus</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zamotajlov, Alexandr S.</mods:namePart>
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<mods:affiliation>Kuban State Agrarian University, Kalinin St. 13, Krasnodar 350044, Russia</mods:affiliation>
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<mods:nameIdentifier type="email">a_zamotajlov@mail.ru</mods:nameIdentifier>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fominykh, Dmitriy D.</mods:namePart>
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<mods:affiliation>Russian Entomological Society, Kuban Branch, Kalinin St. 13, Krasnodar 350044, Russia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Titarenko, Andrey Y.</mods:namePart>
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<mods:affiliation>Publicly Traded Company " Morpho Absoloni ", Bolotnikovskaya St. 5 / 3, Moscow 117556, Russia</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2014-12-12</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>463</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>21</mods:start>
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<mods:end>56</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.463.8499</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.463.8499</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-463-21</mods:identifier>
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<mods:identifier type="ZooBank">71C1AE92B0CE4D26B1E5076E480180AD</mods:identifier>
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<mods:identifier type="Pensoft-UUID">3E5CFFC6FFC91628FFF6E355951DFFF8</mods:identifier>
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<mods:identifier type="Zenodo-Dep">578654</mods:identifier>
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<treatment ID-GBIF-Taxon="182234819" LSID="urn:lsid:plazi:treatment:56C62A04B98059CD8B5DCBF7850C80D9" httpUri="http://treatment.plazi.org/id/56C62A04B98059CD8B5DCBF7850C80D9" lastPageId="17" lastPageNumber="38" pageId="12" pageNumber="33">
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<subSubSection pageId="12" pageNumber="33" type="nomenclature">
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<paragraph pageId="12" pageNumber="33">
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<taxonomicName LSID="56C62A04-B980-59CD-8B5D-CBF7850C80D9" authority="Solodovnikov, Zamotajlov & Fominykh" authorityName="Solodovnikov, Zamotajlov & Fominykh" authorityYear="2014" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. resheviensis" order="Coleoptera" pageId="12" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="besleticus" status="subsp. n." subGenus="Archiplectes" subSpecies="resheviensis">
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<pageBreakToken pageId="12" pageNumber="33" start="start">Carabus</pageBreakToken>
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(Archiplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh
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</taxonomicName>
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<taxonomicNameLabel pageId="12" pageNumber="33">subsp. n.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of morphometric characters in males of Carabus (Archiplectes) besleticus and Carabus (Archiplectes) satyrus subspecies constructed using discriminant analysis based on 8 parameters." pageId="12" pageNumber="33">Figs 1</figureCitation>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution of morphometric characters in females of Carabus (Archiplectes) besleticus and Carabus (Archiplectes) satyrus subspecies constructed using discriminant analysis based on 8 parameters." pageId="12" pageNumber="33">, 2</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distribution of morphometric characters in males of Carabus (Archiplectes) besleticus subspecies constructed using discriminant analysis based on 8 parameters." pageId="12" pageNumber="33">, 3</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution of morphometric characters in females of Carabus (Archiplectes) besleticus subspecies constructed using discriminant analysis based on 8 parameters." pageId="12" pageNumber="33">, 4</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distribution of morphometric characters in males of Carabus (Archiplectes) besleticus resheviensis (River Reshevie), Carabus (Archiplectes) pseudopshuensis (River Reshevie), and Carabus (Archiplectes) juenthneri juenthneri (Village Pskhu env.) constructed using discriminant analysis based on 8 parameters." pageId="12" pageNumber="33">, 5</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Distribution of morphometric characters in females of Carabus (Archiplectes) besleticus resheviensis (River Reshevie), Carabus (Archiplectes) pseudopshuensis (River Reshevie), and Carabus (Archiplectes) juenthneri juenthneri (Village Pskhu env.) constructed using discriminant analysis based on 8 parameters." pageId="12" pageNumber="33">, 6</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution map of Carabus (Archiplectes) satyrus, Carabus (Archiplectes) besleticus, and Carabus (Archiplectes) pseudopshuensis in Abkhazia." pageId="12" pageNumber="33">, 7</figureCitation>
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<figureCitation captionStart="Figures 28–33" captionStartId="F11" captionText="Figures 28 - 33. Fully inflated endophallus preparation and apical lamella of aedeagus of Carabus (Archiplectes) besleticus dsychvensis and Carabus (Archiplectes) besleticus resheviensis 28 - 30 Carabus (Archiplectes) besleticus dsychvensis 31 - 33 Carabus (Archiplectes) besleticus resheviensis." pageId="12" pageNumber="33">, 31-33</figureCitation>
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<figureCitation captionStart="Figures 66–73" captionStartId="F17" captionText="Figures 66 - 73. Dorsal habitus, Carabus (Archiplectes) besleticus resheviensis. 66 Abkhazia, N slopes of Bzybian Mt. Range, right bank of River Reshevie (holotype) 67 - 73 same locality (paratypes)." pageId="12" pageNumber="33">, 66-73</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="15" lastPageNumber="36" pageId="13" pageNumber="34" type="type material">
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<paragraph pageId="13" pageNumber="34">
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<pageBreakToken pageId="13" pageNumber="34" start="start">Type</pageBreakToken>
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material.
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</paragraph>
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<paragraph lastPageId="15" lastPageNumber="36" pageId="13" pageNumber="34">
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Holotype, a male: Abkhazia, N Slopes of Bzybian Mt. Range, right bank of River Reshevie (= Reshava) (left tributary of River Bzyb), 700-1000 m, 13.V-02.VII.2010, leg. D. Fominykh, I. Solodovnikov (ZISP). 577 paratypes: 34 males, 48 females, Abkhazia, N Slopes of Bzybian Mt. Range, right bank of River Reshevie (left tributary of River Bzyb), 700-1000 m, 13.V-02.VII.2010, leg. D. Fominykh, I. Solodovnikov (cZAM, cFDD, cSIA); 234 males, 261 females, same locality, 04.V-20.VII.2012, leg D.
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<pageBreakToken pageId="14" pageNumber="35" start="start">Fominykh</pageBreakToken>
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, A. Safronov (cZAM, cFDD, cSIA, cSAA, cBAS, cTAYu); 3 females, Abkhazia, Bzybian Mt. Range, right bank of River Reshevie (left tributary of River Bzyb), 670-720 m, 13-14.V.2010, leg. D. Fominykh, I. Solodovnikov (cPRYu); 5 males, 3 fe
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<pageBreakToken pageId="15" pageNumber="36" start="start">males</pageBreakToken>
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, Abkhazia, Sukhum Distr., left bank of River Bzyb, right bank of River Reshevie, beech, maple, fern forest site, 670-720 m, 13-14.V.2010, leg. I. Solodovnikov (cSIA); 5 males, 10 females, same locality, 13.V-02.VII.2010 leg. I. Solodovnikov, D. Fominykh (cSIA, cPIG, cKVM); 1 male, labeled "Cauc. occ. Abchasia Pskhu 1910,
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<taxonomicName class="Insecta" family="Carabidae" genus="Plectes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Plectes protensus" order="Coleoptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="protensus">
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<emphasis italics="true" pageId="15" pageNumber="36">Plectes protensus</emphasis>
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</taxonomicName>
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Schaum" (ZISP); 1 female, labeled "West Caucasus, Pskhu, VII.1913, Satunin" (ZISP). Holotype and 57 paratype specimens measured, 30 male genitalia preparations studied.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="15" pageNumber="36" type="description">
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<paragraph pageId="15" pageNumber="36">Description.</paragraph>
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<paragraph pageId="15" pageNumber="36">
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Large form, body size 34.5-43.0 mm, males 34.5-39.0 (37.6) mm, females 37.2-43.0 (40.6) mm long. Underside black, dorsum bright bronze, red-gold, green, violet, dark blue, black, with metallic lustre in males and matte in females. Appendages black, rarely the femora are red and basal antennomeres reddish-brown (approx. 5 % of individuals). Body usually monochromatic dorsally, less frequently bicolored (head and pronotum being usually greenish, the elytra bronze or reddish-bronze). Body massive and rather high (Figs
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<figureCitation captionStart="Figures 66–73" captionStartId="F17" captionText="Figures 66 - 73. Dorsal habitus, Carabus (Archiplectes) besleticus resheviensis. 66 Abkhazia, N slopes of Bzybian Mt. Range, right bank of River Reshevie (holotype) 67 - 73 same locality (paratypes)." pageId="15" pageNumber="36">66</figureCitation>
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-
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<figureCitation captionStart="Figures 66–73" captionStartId="F17" captionText="Figures 66 - 73. Dorsal habitus, Carabus (Archiplectes) besleticus resheviensis. 66 Abkhazia, N slopes of Bzybian Mt. Range, right bank of River Reshevie (holotype) 67 - 73 same locality (paratypes)." pageId="15" pageNumber="36">73</figureCitation>
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).
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</paragraph>
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<paragraph pageId="15" pageNumber="36">
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Head huge, somewhat inflated. Pronotum transverse to subquadrate, hind angles protruding backwards and in some individuals sidewards. PW/PL = 1,18-1,55 (1,37) in males and 1,29-1,67 (1,45) in females. Elytra faintly elongate, ovate; more elongate, sometimes with depression in the middle in females, EL/EW = 1,52-1,92 (1,72); generally more ovate, convex, and smoothed in males, EL/EW = 1,42-1,78 (1,59). Elytral sculpture precise to homodynamous and confused, with hardly any distinguishable primary interspaces in males. The main morphometric measurements are presented in Table
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<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphometric characteristic of males (n = 16) and females (n = 31) of Carabus (Archiplectes) satyrus and males (n = 83) and females (n = 91) of Carabus (Archiplectes) besleticus subspecies." httpUri="http://table.plazi.org/id/E3AA304075ED326595338A0A005BF64B" pageId="15" pageNumber="36" tableUuid="E3AA304075ED326595338A0A005BF64B">2</tableCitation>
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. Male genitalia practically identical to other taxa of
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<taxonomicName authorityName="Kurnakov" authorityYear="1972" baseAuthorityName="besleticus Kurnakov" baseAuthorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus" order="Coleoptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="besleticus" subGenus="Archiplectes">Carabus (Archiplectes) besleticus</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="16" lastPageNumber="37" pageId="15" pageNumber="36" type="differential diagnosis">
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<paragraph pageId="15" pageNumber="36">Differential diagnosis.</paragraph>
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<paragraph pageId="15" pageNumber="36">
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<taxonomicName authorityName="Solodovnikov, Zamotajlov & Fominykh" authorityYear="2014" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. resheviensis" order="Coleoptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="resheviensis">Carabus (Archiplectes) besleticus resheviensis</taxonomicName>
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resembles habitually a population of
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<taxonomicName authority="besleticus" authorityName="besleticus Kurnakov" authorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. besleticus" order="Coleoptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="besleticus">Carabus (Archiplectes) besleticus besleticus</taxonomicName>
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, inhabiting valley of River Basla (city of Sukhum vicinities), but differs in the following characters. Pronotum with a rather shallow median depression, being only poorly marked in males; disk impressed; hind angles huge, appreciably larger, lateral sides of pronotum strongly sinuated before hind angles; elytra broader in males, EL/EW = 1.59, while 1.75 in
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<taxonomicName authority="besleticus" authorityName="besleticus Kurnakov" authorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. besleticus" order="Coleoptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="besleticus">Carabus (Archiplectes) besleticus besleticus</taxonomicName>
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and 1.61-1.75 in
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<taxonomicName authorityName="Kurnakov" authorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. mtsaranus" order="Coleoptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="mtsaranus">Carabus (Archiplectes) besleticus mtsaranus</taxonomicName>
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. The new subspecies is the closest geographically to
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<taxonomicName authorityName="Kurnakov" authorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. mtsaranus" order="Coleoptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="mtsaranus">Carabus (Archiplectes) besleticus mtsaranus</taxonomicName>
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, populating southern slopes of Bzybian Mt. Range, but differs in having a larger body size, more ovate elytra, smooth elytral sculpture, more transverse pronotum with weaker protruding hind angles, and general dorsal coloration (PW/PL = 1.14-1.48 (1.35) in males and 1.26-1.56 (1.41) in females of
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<taxonomicName authorityName="Kurnakov" authorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. mtsaranus" order="Coleoptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="mtsaranus">Carabus (Archiplectes) besleticus mtsaranus</taxonomicName>
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); furthermore, the disk of the pronotum is more convex than in the new subspecies. The new subspecies differs from the recently described
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<taxonomicName authorityName="Retezar" authorityYear="2013" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. adzinbai" order="Coleoptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="adzinbai">Carabus (Archiplectes) besleticus adzinbai</taxonomicName>
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first of all in having a larger body size, more transverse and faintly cordate pronotum with more narrow base;
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<taxonomicName authorityName="Retezar" authorityYear="2013" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. adzinbai" order="Coleoptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="adzinbai">Carabus (Archiplectes) besleticus adzinbai</taxonomicName>
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possesses subquadrate pronotum with stronger protruding sidewards hind angles, PW/PL = 1.14-1.42 (1.25), PW/PB = 1.00-1.26 (1.14); elytra of
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<taxonomicName authorityName="Solodovnikov, Zamotajlov & Fominykh" authorityYear="2014" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. resheviensis" order="Coleoptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="resheviensis">Carabus (Archiplectes) besleticus resheviensis</taxonomicName>
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are more ovate and shorter both in males and females.
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</paragraph>
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<paragraph lastPageId="16" lastPageNumber="37" pageId="15" pageNumber="36">
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The bulk of studied individuals of
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<taxonomicName authorityName="Solodovnikov, Zamotajlov & Fominykh" authorityYear="2014" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. resheviensis" order="Coleoptera" pageId="16" pageNumber="37" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="resheviensis">
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<pageBreakToken pageId="16" pageNumber="37" start="start">Carabus</pageBreakToken>
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(Archiplectes) besleticus resheviensis
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</taxonomicName>
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(males) differ from the other subspecies, namely
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<taxonomicName authority="besleticus" authorityName="besleticus Kurnakov" authorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. besleticus" order="Coleoptera" pageId="16" pageNumber="37" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="besleticus">Carabus (Archiplectes) besleticus besleticus</taxonomicName>
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,
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<taxonomicName authorityName="Kurnakov" authorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. mtsaranus" order="Coleoptera" pageId="16" pageNumber="37" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="mtsaranus">Carabus (Archiplectes) besleticus mtsaranus</taxonomicName>
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,
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<taxonomicName authorityName="Kurnakov" authorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. duripshensis" order="Coleoptera" pageId="16" pageNumber="37" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="duripshensis">Carabus (Archiplectes) besleticus duripshensis</taxonomicName>
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, and
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<taxonomicName authorityName="Retezar" authorityYear="2013" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. adzinbai" order="Coleoptera" pageId="16" pageNumber="37" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="adzinbai">Carabus (Archiplectes) besleticus adzinbai</taxonomicName>
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in their elytral sculpture: tertiary interspaces are elevated almost as secondary ones, forming a smooth sculpture, primary interspaces interrupted more frequently. In the other subspecies tertiary interspaces are normally hardly raised. Morphometric characters are illustrated in Figures
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of morphometric characters in males of Carabus (Archiplectes) besleticus and Carabus (Archiplectes) satyrus subspecies constructed using discriminant analysis based on 8 parameters." pageId="16" pageNumber="37">1</figureCitation>
|
||
-
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Distribution of morphometric characters in females of Carabus (Archiplectes) besleticus resheviensis (River Reshevie), Carabus (Archiplectes) pseudopshuensis (River Reshevie), and Carabus (Archiplectes) juenthneri juenthneri (Village Pskhu env.) constructed using discriminant analysis based on 8 parameters." pageId="16" pageNumber="37">6</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="16" pageNumber="37" type="distribution">
|
||
<paragraph pageId="16" pageNumber="37">Distribution.</paragraph>
|
||
<paragraph pageId="16" pageNumber="37">
|
||
Populates boreal slopes of Bzybian Mt. Range on the right bank of River Reshevie (=Reshava) at 700-1000 m. Occurs sympatrically with
|
||
<taxonomicName authorityName="Zamotajlov" authorityYear="1991" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) pseudopshuensis" order="Coleoptera" pageId="16" pageNumber="37" phylum="Arthropoda" rank="species" species="pseudopshuensis" subGenus="Archiplectes">Carabus (Archiplectes) pseudopshuensis</taxonomicName>
|
||
, the population density of the latter being somewhat higher. Occurrence of this subspecies at the left bank of River Reshevie and downstream River Bzyb seems also possible, but collected material does not confirm this possibility at present.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="17" pageNumber="38" type="habitat">
|
||
<paragraph pageId="17" pageNumber="38">
|
||
<pageBreakToken pageId="17" pageNumber="38" start="start">Habitat</pageBreakToken>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="17" pageNumber="38">
|
||
Prefers slightly sloping beech and fir-beech forest sites rich in ground litter on karstic landforms. Activity of imago proceeds from April to July.
|
||
<taxonomicName authorityName="Zamotajlov" authorityYear="1991" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) pseudopshuensis" order="Coleoptera" pageId="17" pageNumber="38" phylum="Arthropoda" rank="species" species="pseudopshuensis" subGenus="Archiplectes">Carabus (Archiplectes) pseudopshuensis</taxonomicName>
|
||
,
|
||
<taxonomicName authority="apschuanus" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) apschuanus subsp. apschuanus" order="Coleoptera" pageId="17" pageNumber="38" phylum="Arthropoda" rank="subSpecies" species="apschuanus" subGenus="Tribax" subSpecies="apschuanus">Carabus (Tribax) apschuanus apschuanus</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) constantinovi subsp. otcharensis" order="Coleoptera" pageId="17" pageNumber="38" phylum="Arthropoda" rank="subSpecies" species="constantinovi" subGenus="Tribax" subSpecies="otcharensis">Carabus (Tribax) constantinovi otcharensis</taxonomicName>
|
||
,
|
||
<taxonomicName authority="circassicus" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) circassicus subsp. circassicus" order="Coleoptera" pageId="17" pageNumber="38" phylum="Arthropoda" rank="subSpecies" species="circassicus" subGenus="Tribax" subSpecies="circassicus">Carabus (Tribax) circassicus circassicus</taxonomicName>
|
||
(natio
|
||
<taxonomicName class="Insecta" family="Carabidae" form="abasinus" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="17" pageNumber="38" phylum="Arthropoda" rank="form">
|
||
<emphasis italics="true" pageId="17" pageNumber="38">abasinus</emphasis>
|
||
</taxonomicName>
|
||
),
|
||
<taxonomicName authorityName="Mandl" authorityYear="1955" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Microplectes) argonautarum subsp. reischitzi" order="Coleoptera" pageId="17" pageNumber="38" phylum="Arthropoda" rank="subSpecies" species="argonautarum" subGenus="Microplectes" subSpecies="reischitzi">Carabus (Microplectes) argonautarum reischitzi</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Motschulsky" authorityYear="1840" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Megodontus) septemcarinatus" order="Coleoptera" pageId="17" pageNumber="38" phylum="Arthropoda" rank="species" species="septemcarinatus" subGenus="Megodontus">Carabus (Megodontus) septemcarinatus</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Sphodristocarabus) armeniacus subsp. dvorschaki" order="Coleoptera" pageId="17" pageNumber="38" phylum="Arthropoda" rank="subSpecies" species="armeniacus" subGenus="Sphodristocarabus" subSpecies="dvorschaki">Carabus (Sphodristocarabus) armeniacus dvorschaki</taxonomicName>
|
||
, and
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Procerus) caucasicus subsp. colchicus" order="Coleoptera" pageId="17" pageNumber="38" phylum="Arthropoda" rank="subSpecies" species="caucasicus" subGenus="Procerus" subSpecies="colchicus">Carabus (Procerus) caucasicus colchicus</taxonomicName>
|
||
occur together with this subspecies.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="17" pageNumber="38" type="subspecific epithet">
|
||
<paragraph pageId="17" pageNumber="38">Subspecific epithet.</paragraph>
|
||
<paragraph pageId="17" pageNumber="38">The subspecific epithet refers to the name of the River Reshevie, the type locality of subspecies.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |