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110 lines
18 KiB
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<document id="EEE437A064892AF5D75A4CC2CC81F7A3" ID-ISSN="1536-9307" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638232259732" checkinUser="felipe" docAuthor="Kovařík, František" docDate="2012" docId="03856D55156FFFC2FEA8FBA96893AD84" docLanguage="en" docName="Euscorpius.141.1-22.pdf" docOrigin="Euscorpius 141" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Pandinus Thorell 1876" docType="treatment" docVersion="3" lastPageNumber="3" masterDocId="FFBC152D156BFFC6FFD8FFF26A0EAB3D" masterDocTitle="Review of the subgenus Pandinurus Fet, 1997 with descriptions of three new species (Scorpiones, Scorpionidae, Pandinus)" masterLastPageNumber="22" masterPageNumber="1" pageNumber="3" updateTime="1699434663504" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="3D53786986619D9D6216E25FEFBA52B6">Review of the subgenus Pandinurus Fet, 1997 with descriptions of three new species (Scorpiones, Scorpionidae, Pandinus)</mods:title>
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<mods:namePart id="BB6562A11DE670D8422BFB5C52F956E0">Kovařík, František</mods:namePart>
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<mods:title id="28583F8A0566A0AB21267C3A972D9F6C">Euscorpius</mods:title>
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<mods:date id="D7EB16263D709C1FA3B71FCE4E64F3AE">2012</mods:date>
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<mods:number id="04D9C724F35E5B64A6F67E8047281149">141</mods:number>
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<taxonomicName id="4C2CA7C0156FFFC2FEA8FBA9687CAF4E" ID-CoL="63JDT" authority="Thorell, 1876" authorityName="Thorell" authorityYear="1876" box="[368,626,1115,1139]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="B9580051156FFFC2FEA8FBA96BD7AF4E" bold="true" box="[368,473,1115,1139]" italics="true" pageId="4" pageNumber="3">Pandinus</emphasis>
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<bibRefCitation id="EFBDA1B2156FFFC2FE38FBAE687CAF4E" author="THORELL" box="[480,626,1116,1139]" pageId="4" pageNumber="3" pagination="1 - 15" refId="ref10606" refString="THORELL, T. 1876. On the classification of scorpions. Annals and Magazine of Natural History, 4 (17): 1 - 15." type="journal article" year="1876">Thorell, 1876</bibRefCitation>
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</paragraph>
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<paragraph id="8B93DC43156FFFC2FE68FB88683CAFAC" blockId="4.[368,626,1115,1169]" box="[432,562,1146,1169]" pageId="4" pageNumber="3">
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(
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<figureCitation id="1317C0C6156FFFC2FE6FFB886824AFAC" box="[439,554,1146,1169]" captionStart-0="Figures 1–8" captionStart-1="Figures 9–18" captionStart-10="Figures 57–63" captionStart-11="Figure 64" captionStart-2="Figures 19–22" captionStart-3="Figures 23–24" captionStart-4="Figures 25–28" captionStart-5="Figures 29–34" captionStart-6="Figures 35–37" captionStart-7="Figures 38–44" captionStart-8="Figures 45–50" captionStart-9="Figures 51–56" captionStartId-0="3.[192,278,1838,1861]" captionStartId-1="5.[192,278,1830,1853]" captionStartId-10="19.[192,278,1758,1781]" captionStartId-11="20.[192,268,1768,1791]" captionStartId-2="6.[192,278,1867,1890]" captionStartId-3="8.[192,278,1838,1861]" captionStartId-4="9.[192,278,1822,1845]" captionStartId-5="11.[192,278,1862,1885]" captionStartId-6="12.[192,278,1832,1855]" captionStartId-7="13.[192,278,1793,1816]" captionStartId-8="15.[192,278,1850,1873]" captionStartId-9="17.[192,278,1854,1877]" captionTargetBox-0="[192,1427,178,1800]" captionTargetBox-1="[212,1368,208,1788]" captionTargetBox-10="[220,1413,186,1722]" captionTargetBox-11="[191,1441,191,1737]" captionTargetBox-2="[221,1393,194,1842]" captionTargetBox-3="[192,1440,192,1808]" captionTargetBox-4="[232,1393,191,1791]" captionTargetBox-5="[234,1380,200,1832]" captionTargetBox-6="[215,1421,191,1801]" captionTargetBox-7="[246,1392,195,1752]" captionTargetBox-8="[239,1380,220,1815]" captionTargetBox-9="[236,1433,175,1821]" captionTargetId-0="figure-6@3.[192,1440,144,1808]" captionTargetId-1="figure-6@5.[208,1368,202,1800]" captionTargetId-10="figure-6@19.[212,1416,174,1728]" captionTargetId-11="figure-10@20.[192,1440,192,1736]" captionTargetId-2="figure-10@6.[221,1393,186,1848]" captionTargetId-3="figure-10@8.[192,1440,192,1808]" captionTargetId-4="figure-6@9.[232,1392,192,1790]" captionTargetId-5="figure-6@11.[224,1380,192,1832]" captionTargetId-6="figure-10@12.[216,1420,192,1800]" captionTargetId-7="figure-6@13.[236,1392,192,1763]" captionTargetId-8="figure-6@15.[228,1383,216,1820]" captionTargetId-9="figure-6@17.[236,1440,170,1824]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-10="19" captionTargetPageId-11="20" captionTargetPageId-2="6" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="11" captionTargetPageId-6="12" captionTargetPageId-7="13" captionTargetPageId-8="15" captionTargetPageId-9="17" captionText-0="Figures 1–8: Pandinus (Pandinurus) magrettii Borelli, 1901. 1–2. Dorsal view and label, ♀(ca. 110 mm), Eritrea, Dembelas, VII.1902, det. Borelli, MCSN. 3–8. Chela, dorsal and ventral views, tarsomere of 4th leg, carapace, fifth metasomal segment and telson, ♂(98 mm), Eritrea, FKCP." captionText-1="Figures 9–18: Pandinus (Pandinurus) awashensis sp. n. 9–13. Dorsal and ventral views, fifth metasomal segment and telson, carapace, and pectinal area, ♂ (110 mm) holotype. 14–18. Dorsal and ventral views, fifth metasomal segment and telson, pectinal area, and carapace, ♀ (83 mm) allotype." captionText-10="Figures 57–63: Tarsomere II and part of tarsomere I of 4th leg of subgenus Pandinurus. 57. P. lowei sp. n., ♂ (98 mm) holotype. 58. P. viatoris (Pocock, 1890), ♂ (105 mm), Democratic Republic of Congo, Parc National de Upemba, Kapelwa, 1780 m. a.s.l, ISNB. 59. P. somalilandus sp. n., ♀ (110 mm) holotype. 60–61. P. awashensis sp. n.. 60. ♀ (83 mm) allotype. 61. ♂ (110 mm) holotype. 62. P. meidensis Karsch, 1879, ♂ (ca 140 mm), Somalia, Sar Uanle, about 20 km south of Chisimaio, 00°29'48"S 42°25'30"E, MZUF. 63. P. gregorii Pocock, 1896, ♀ (110 mm), Kenya, Maramtu Hill, Maramtu Village, Garissa, FKCP." captionText-11="Figure 64: Distribution map of the subgenus Pandinurus." captionText-2="Figures 19–22: Pandinus (Pandinurus) awashensis sp. n. 19–21. Chela dorsal, external and ventral/internal views, ♂ (110 mm) holotype. 22. Trichobothrial pattern, ♀ (83 mm) allotype." captionText-3="Figures 23–24: Pandinus (Pandinurus) awashensis sp. n., ♀ (83 mm) allotype at the locality, Ethiopia, Awash, 09°00'34.5"N 40°17'56.5"E, 1012 m. a.s.l.." captionText-4="Figures 25–28: Pandinus (Pandinurus) awashensis sp. n. 25–26. Type locality, Ethiopia, Awash, Awash N. P., 08°52'35.15"N 40°05'39.8"E, 981 m a.s.l.. 26. Water-capture hole well that produced the dead male holotype. 27–28. Locality of allotype and paratypes, Ethiopia, Awash, 09°00'34.5"N 40°17'56.5"E, 1012 m. a.s.l.. 28. Rock under which was captured the female allotype." captionText-5="Figures 29–34: Pandinus (Pandinurus) somalilandus sp. n. Dorsal and ventral views, metasoma and telson lateral view, trichobothrial pattern, carapace, and pectinal area, ♀ (110 mm) holotype." captionText-6="Figures 35–37: Pandinus (Pandinurus) somalilandus sp. n. 35. Locality of paratype, Somaliland, near Sheikh, foothills of Goolis Mts., 09°59.881'N 45°09.762'E, 896 m a.s.l.. 36.♀ (110 mm) holotype at the type locality. 37. Type locality, Somaliland, 25 km N of Sheikh, 10°02.001'N 45°09.589'E, 763 m a.s.l." captionText-7="Figures 38–44: 38–39: Pandinus (Pandinurus) viatoris (Pocock, 1890). 38. Dorsal view, ♂(120 mm), Zambia, 92 km NW of Mpika, FKCP. 39. Chela, ♀(95 mm), Zambia, 92 km NW of Mpika, FKCP. 40–41: Pandinus (Pandinurus) sudanicus Hirst, 1911, stat. n., dorsal view and chela, ♂(98 mm), South Sudan, Kordofan Province, Lagowa; FKCP. 42–44: Pandinus (Pandinurus) exitialis (Pocock, 1888). 42. Type label, BMNH. 43. Chela, ♂(120 mm), Somalia, Afgoi, FKCP. 44. Chela, ♀(110 mm), Somalia, Afgoi, FKCP." captionText-8="Figures 45–50: Pandinus (Pandinurus) lowei sp. n. 45–47. Dorsal and ventral views, and carapace with chelicerae, ♂ (98 mm) holotype. 48–50. Dorsal and ventral views, and carapace with chelicerae, ♀ (95 mm) allotype." captionText-9="Figures 51–56: Pandinus (Pandinurus) lowei sp. n. 51–53. Trichobothrial pattern, metasoma and telson lateral view, and pectinal area, ♂ (98 mm) holotype. 54–56. Chela, metasoma and telson lateral view, and pectinal area, ♀ (95 mm) allotype." pageId="4" pageNumber="3">Figs. 1–64</figureCitation>
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)
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</paragraph>
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<subSubSection id="C3368FC8156FFFC2FF18FB4A6B3CAE74" pageId="4" pageNumber="3" type="reference_group">
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<treatmentCitationGroup id="AB3CFB6D156FFFC2FF18FB4A6B3CAE74" pageId="4" pageNumber="3">
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<treatmentCitation id="0A8DFA52156FFFC2FF18FB4A6804AFF2" author="THORELL" box="[192,522,1208,1231]" page="12" pageId="4" pageNumber="3" year="1876">
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<taxonomicName id="4C2CA7C0156FFFC2FF18FB4A6804AFF2" ID-CoL="63JDT" authority="Thorell, 1876: 12" authorityName="Thorell" authorityPageNumber="12" authorityYear="1876" box="[192,522,1208,1231]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="B9580051156FFFC2FF18FB4A6B2BAFF2" box="[192,293,1208,1231]" italics="true" pageId="4" pageNumber="3">Pandinus</emphasis>
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<bibRefCitation id="EFBDA1B2156FFFC2FEEFFB4A6804AFF2" author="THORELL" box="[311,522,1208,1231]" pageId="4" pageNumber="3" pagination="1 - 15" refId="ref10606" refString="THORELL, T. 1876. On the classification of scorpions. Annals and Magazine of Natural History, 4 (17): 1 - 15." type="journal article" year="1876">Thorell, 1876: 12</bibRefCitation>
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;
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<treatmentCitation id="0A8DFA52156FFFC2FDF9FB4A6913AFF2" author="KRAEPELIN" box="[545,797,1208,1231]" page="116" pageId="4" pageNumber="3" year="1899">
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<bibRefCitation id="EFBDA1B2156FFFC2FDF9FB4A6913AFF2" author="KRAEPELIN" box="[545,797,1208,1231]" pageId="4" pageNumber="3" refId="ref10409" refString="KRAEPELIN, K. 1899. Scorpiones und Pedipalpi. In F. DAHL (ed.), Das Tierreich. Herausgegeben von der Deutschen Zoologischen Gesellschaft. Berlin: R. Friedlander und Sohn Verlag, 8. Lieferung. 265 pp." type="book" year="1899">Kraepelin, 1899: 116</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0A8DFA52156FFFC2FF18FB246B98AFD0" author="VACHON" box="[192,406,1238,1261]" page="953" pageId="4" pageNumber="3" year="1974">
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<bibRefCitation id="EFBDA1B2156FFFC2FF18FB246B98AFD0" author="VACHON" box="[192,406,1238,1261]" pageId="4" pageNumber="3" pagination="857 - 958" refId="ref10692" refString="VACHON, M. 1974. Etude des caracteres utilises pour classer les familles et les genres de Scorpions. Bulletin du Museum National d'Histoire Naturelle Paris, 140: 857 - 958." type="journal article" year="1974">Vachon, 1974: 953</bibRefCitation>
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</treatmentCitation>
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||
, figs. 113–118;
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<treatmentCitation id="0A8DFA52156FFFC2FD95FB246913AFD0" author="SISSOM" box="[589,797,1238,1261]" page="136" pageId="4" pageNumber="3" year="1990">
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<bibRefCitation id="EFBDA1B2156FFFC2FD95FB246913AFD0" author="SISSOM" box="[589,797,1238,1261]" pageId="4" pageNumber="3" pagination="64 - 160" refId="ref10560" refString="SISSOM, W. D. 1990. Systematics, biogeography and paleontology. Pp. 64 - 160. In: POLIS G. A. (ed.): The biology of Scorpions. Stanford: Stanford University press, 587 pp." type="book chapter" year="1990">Sissom, 1990: 136</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0A8DFA52156FFFC2FF18FB076B60AE31" author="FET" box="[192,366,1269,1292]" page="248" pageId="4" pageNumber="3" year="1997">
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<bibRefCitation id="EFBDA1B2156FFFC2FF18FB076B60AE31" author="FET" box="[192,366,1269,1292]" pageId="4" pageNumber="3" pagination="245 - 250" refId="ref9876" refString="FET, V. 1997. Notes on the taxonomy of some old world scorpions (Scorpiones: Buthidae, Chactidae, Ischnuridae, Scorpionidae). Journal of Arachnology, 25: 245 - 250." type="journal article" year="1997">Fet, 1997: 248</bibRefCitation>
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;
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<treatmentCitation id="0A8DFA52156FFFC2FE5AFB07683EAE31" author="FET" box="[386,560,1269,1292]" page="465" pageId="4" pageNumber="3" year="2000">
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<bibRefCitation id="EFBDA1B2156FFFC2FE5AFB07683EAE31" author="FET" box="[386,560,1269,1292]" pageId="4" pageNumber="3" refId="ref9913" refString="FET, V., W. D. SISSOM, G. LOWE & M. E. BRAUN- WALDER. 2000. Catalog of the Scorpions of the World (1758 - 1998). Tne New York Entomological Society, New York, 689 pp." type="book" year="2000">Fet, 2000: 465</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0A8DFA52156FFFC2FD9CFB076913AE31" author="PRENDINI" box="[580,797,1269,1292]" page="44" pageId="4" pageNumber="3" year="2000">
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<bibRefCitation id="EFBDA1B2156FFFC2FD9CFB076913AE31" author="PRENDINI" box="[580,797,1269,1292]" pageId="4" pageNumber="3" pagination="1 - 78" refId="ref10527" refString="PRENDINI, L. 2000. Phylogeny and classification of the superfamily Scorpionoidea Latreille 1802 (Chelicerata, Scorpiones): An exemplar approach. Cladistics, 16: 1 - 78." type="journal article" year="2000">Prendini, 2000: 44</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0A8DFA52156FFFC2FF18FAE66B9AAE16" author="KOVARIK" box="[192,404,1300,1323]" page="50" pageId="4" pageNumber="3" year="2009">
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<bibRefCitation id="EFBDA1B2156FFFC2FF18FAE66B9AAE16" author="KOVARIK" box="[192,404,1300,1323]" pageId="4" pageNumber="3" refId="ref10153" refString="KOVARIK, F. 2009. Illustrated catalog of scorpions. Part I. Introductory remarks; keys to families and genera; subfamily Scorpioninae with keys to Heterometrus and Pandinus species. Clairon Production, Prague, 170 pp." type="book" year="2009">Kovařík, 2009: 50</bibRefCitation>
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, figs. 284–420; Kovařík, 2011: 1, figs. 1–42.
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</paragraph>
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<subSubSection id="C3368FC8156FFFC2FF18FA8268EFAEBA" box="[192,737,1392,1415]" pageId="4" pageNumber="3" type="materials_examined">
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<paragraph id="8B93DC43156FFFC2FF18FA8268EFAEBA" blockId="4.[192,737,1392,1415]" box="[192,737,1392,1415]" pageId="4" pageNumber="3">
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<typeStatus id="549762E1156FFFC2FF18FA826AF6AEBB" box="[192,248,1392,1415]" pageId="4" pageNumber="3">TYPE</typeStatus>
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SPECIES.
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<taxonomicName id="4C2CA7C0156FFFC2FE86FA8268D3AEBA" authority="C. L. Koch, 1841" authorityName="C. L. Koch" authorityYear="1841" box="[350,733,1392,1415]" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="imperator">
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<emphasis id="B9580051156FFFC2FE86FA826814AEBA" box="[350,538,1392,1415]" italics="true" pageId="4" pageNumber="3">Buthus imperator</emphasis>
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C. L. Koch, 1841
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3368FC8156FFFC2FF18FA5F6893AD84" pageId="4" pageNumber="3" type="diagnosis">
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<paragraph id="8B93DC43156FFFC2FF18FA5F6893AD84" blockId="4.[192,802,1453,1721]" pageId="4" pageNumber="3">
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DIAGNOSIS. Total length
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<quantity id="4CD471A6156FFFC2FE06FA5F6866AEF9" box="[478,616,1453,1476]" metricMagnitude="-1" metricUnit="m" metricValue="1.35" metricValueMax="2.2" metricValueMin="0.5" pageId="4" pageNumber="3" unit="mm" value="135.0" valueMax="220.0" valueMin="50.0">50–220 mm</quantity>
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. Pedipalp femur with 3 trichobothria, only one of them on internal surface. Pedipalp patella with 13–16 external and numerous (usually about 30) ventral trichobothria. Retrolateral pedal spurs absent. Lateroapical margins of tarsi produced into rounded lobes. Metasomal segments I–IV with paired ventral submedian carinae. Stridulatory organ located on opposing surfaces of pedipalp coxa and first leg. Telson without subaculear tubercle.
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</paragraph>
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</treatment>
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