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<document id="9E1B54601BF1EFF8F1DF93D8DF83F0F0" ID-ISSN="1536-9307" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638232259732" checkinUser="felipe" docAuthor="Kovařík, František" docDate="2012" docId="03856D55156CFFCCFF01FBF06989AA64" docLanguage="en" docName="Euscorpius.141.1-22.pdf" docOrigin="Euscorpius 141" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Pandinus (Pandinurus) awashensis Kovarik 2012, sp. n." docType="treatment" docUuid="2D490D14-71B3-40CD-8D40-56667F6F2651" docUuidSource="ZooBank" docVersion="3" lastPageNumber="9" masterDocId="FFBC152D156BFFC6FFD8FFF26A0EAB3D" masterDocTitle="Review of the subgenus Pandinurus Fet, 1997 with descriptions of three new species (Scorpiones, Scorpionidae, Pandinus)" masterLastPageNumber="22" masterPageNumber="1" pageNumber="6" updateTime="1699434663504" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="C1AB924ABBFCBBEAC9A304B267774F31">Review of the subgenus Pandinurus Fet, 1997 with descriptions of three new species (Scorpiones, Scorpionidae, Pandinus)</mods:title>
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<mods:namePart id="7F85FF6DC04B8CC7A83D0A503617BC8E">Kovařík, František</mods:namePart>
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<mods:title id="E4C294C05D0DD75A0D9DEB420D42C96D">Euscorpius</mods:title>
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<mods:date id="A17402EAFCE56AA0F1110F5F12A5EC78">2012</mods:date>
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<treatment id="03856D55156CFFCCFF01FBF06989AA64" ID-DOI="http://doi.org/10.5281/zenodo.5739005" ID-Zenodo-Dep="5739005" LSID="urn:lsid:zoobank.org:act:2D490D14-71B3-40CD-8D40-56667F6F2651" httpUri="http://treatment.plazi.org/id/03856D55156CFFCCFF01FBF06989AA64" lastPageId="10" lastPageNumber="9" pageId="7" pageNumber="6">
<subSubSection id="C3368FC8156CFFC1FF01FBF06906AF27" box="[217,776,1026,1050]" pageId="7" pageNumber="6" type="nomenclature">
<paragraph id="8B93DC43156CFFC1FF01FBF06906AF27" blockId="7.[217,776,1026,1081]" box="[217,776,1026,1050]" pageId="7" pageNumber="6">
<heading id="D0DB6B2F156CFFC1FF01FBF06906AF27" box="[217,776,1026,1050]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="6" reason="4">
<taxonomicName id="4C2CA7C0156CFFC1FF01FBF068CFAF27" authority="Kovarik" authorityName="Kovarik" authorityYear="2012" box="[217,705,1026,1050]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="awashensis" status="sp. nov." subGenus="Pandinurus">
<emphasis id="B9580051156CFFC1FF01FBF0686EAF27" bold="true" box="[217,608,1026,1050]" pageId="7" pageNumber="6">
<emphasis id="B9580051156CFFC1FF01FBF06B4CAF27" bold="true" box="[217,322,1026,1050]" italics="true" pageId="7" pageNumber="6">Pandinus</emphasis>
(
<emphasis id="B9580051156CFFC1FE8AFBF06BDAAF27" bold="true" box="[338,468,1026,1050]" italics="true" pageId="7" pageNumber="6">Pandinurus</emphasis>
)
<emphasis id="B9580051156CFFC1FE3CFBF0686EAF27" bold="true" box="[484,608,1026,1050]" italics="true" pageId="7" pageNumber="6">awashensis</emphasis>
</emphasis>
Kovařík
</taxonomicName>
,
<emphasis id="B9580051156CFFC1FD14FBF16906AF27" bold="true" box="[716,776,1027,1050]" pageId="7" pageNumber="6">
<taxonomicNameLabel id="A26BBD2A156CFFC1FD14FBF16906AF27" box="[716,776,1027,1050]" pageId="7" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3368FC8156CFFC1FE85FBD06859AF04" box="[349,599,1058,1081]" pageId="7" pageNumber="6" type="description">
<paragraph id="8B93DC43156CFFC1FE85FBD06859AF04" blockId="7.[217,776,1026,1081]" box="[349,599,1058,1081]" pageId="7" pageNumber="6">
(
<figureCitation id="1317C0C6156CFFC1FEBDFBD06BD7AF04" box="[357,473,1058,1081]" captionStart-0="Figures 918" captionStart-1="Figures 1922" captionStart-2="Figures 2324" captionStart-3="Figures 2528" captionStartId-0="5.[192,278,1830,1853]" captionStartId-1="6.[192,278,1867,1890]" captionStartId-2="8.[192,278,1838,1861]" captionStartId-3="9.[192,278,1822,1845]" captionTargetBox-0="[212,1368,208,1788]" captionTargetBox-1="[221,1393,194,1842]" captionTargetBox-2="[192,1440,192,1808]" captionTargetBox-3="[232,1393,191,1791]" captionTargetId-0="figure-6@5.[208,1368,202,1800]" captionTargetId-1="figure-10@6.[221,1393,186,1848]" captionTargetId-2="figure-10@8.[192,1440,192,1808]" captionTargetId-3="figure-6@9.[232,1392,192,1790]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="8" captionTargetPageId-3="9" captionText-0="Figures 918: Pandinus (Pandinurus) awashensis sp. n. 913. Dorsal and ventral views, fifth metasomal segment and telson, carapace, and pectinal area, ♂ (110 mm) holotype. 1418. Dorsal and ventral views, fifth metasomal segment and telson, pectinal area, and carapace, ♀ (83 mm) allotype." captionText-1="Figures 1922: Pandinus (Pandinurus) awashensis sp. n. 1921. Chela dorsal, external and ventral/internal views, ♂ (110 mm) holotype. 22. Trichobothrial pattern, ♀ (83 mm) allotype." captionText-2="Figures 2324: Pandinus (Pandinurus) awashensis sp. n., ♀ (83 mm) allotype at the locality, Ethiopia, Awash, 09°00'34.5&quot;N 40°17'56.5&quot;E, 1012 m. a.s.l.." captionText-3="Figures 2528: Pandinus (Pandinurus) awashensis sp. n. 2526. Type locality, Ethiopia, Awash, Awash N. P., 08°52'35.15&quot;N 40°05'39.8&quot;E, 981 m a.s.l.. 26. Water-capture hole well that produced the dead male holotype. 2728. Locality of allotype and paratypes, Ethiopia, Awash, 09°00'34.5&quot;N 40°17'56.5&quot;E, 1012 m. a.s.l.. 28. Rock under which was captured the female allotype." pageId="7" pageNumber="6">Figs. 928</figureCitation>
,
<figureCitation id="1317C0C6156CFFC1FE3BFBD06826AF04" box="[483,552,1058,1081]" captionStart="Figures 5763" captionStartId="19.[192,278,1758,1781]" captionTargetBox="[220,1413,186,1722]" captionTargetId="figure-6@19.[212,1416,174,1728]" captionTargetPageId="19" captionText="Figures 5763: Tarsomere II and part of tarsomere I of 4th leg of subgenus Pandinurus. 57. P. lowei sp. n., ♂ (98 mm) holotype. 58. P. viatoris (Pocock, 1890), ♂ (105 mm), Democratic Republic of Congo, Parc National de Upemba, Kapelwa, 1780 m. a.s.l, ISNB. 59. P. somalilandus sp. n., ♀ (110 mm) holotype. 6061. P. awashensis sp. n.. 60. ♀ (83 mm) allotype. 61. ♂ (110 mm) holotype. 62. P. meidensis Karsch, 1879, ♂ (ca 140 mm), Somalia, Sar Uanle, about 20 km south of Chisimaio, 00°29'48&quot;S 42°25'30&quot;E, MZUF. 63. P. gregorii Pocock, 1896, ♀ (110 mm), Kenya, Maramtu Hill, Maramtu Village, Garissa, FKCP." pageId="7" pageNumber="6">6062</figureCitation>
,
<figureCitation id="1317C0C6156CFFC1FDECFBD0685EAF04" box="[564,592,1058,1081]" captionStart="Figure 64" captionStartId="20.[192,268,1768,1791]" captionTargetBox="[191,1441,191,1737]" captionTargetId="figure-10@20.[192,1440,192,1736]" captionTargetPageId="20" captionText="Figure 64: Distribution map of the subgenus Pandinurus." pageId="7" pageNumber="6">64</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3368FC8156CFFC1FF18FBAD685AAE94" pageId="7" pageNumber="6" type="materials_examined">
<paragraph id="8B93DC43156CFFC1FF18FBAD6B6BAF8E" blockId="7.[192,802,1119,1203]" pageId="7" pageNumber="6">
TYPE LOCALITY AND
<typeStatus id="549762E1156CFFC1FE0BFB916806AF4B" box="[467,520,1123,1142]" pageId="7" pageNumber="6">TYPE</typeStatus>
REPOSITORY.
<collectingCountry id="F33B9CD3156CFFC1FD18FBAD6913AF4B" box="[704,797,1119,1142]" name="Ethiopia" pageId="7" pageNumber="6">Ethiopia</collectingCountry>
, Awash, Awash N. P.,
<geoCoordinate id="EE18BA84156CFFC1FE6CFB8C685DAFA8" box="[436,595,1150,1173]" degrees="08" direction="north" minutes="52" orientation="latitude" pageId="7" pageNumber="6" precision="1" seconds="35.15" value="8.8764305">08°52'35.15&quot;N</geoCoordinate>
<geoCoordinate id="EE18BA84156CFFC1FD83FB8C68E3AFA8" box="[603,749,1150,1173]" degrees="40" direction="east" minutes="05" orientation="longitude" pageId="7" pageNumber="6" precision="1" seconds="39.8" value="40.094387">40°05'39.8&quot;E</geoCoordinate>
,
<quantity id="4CD471A6156CFFC1FD21FB8C6ADAAF8E" metricMagnitude="2" metricUnit="m" metricValue="9.809999999999999" pageId="7" pageNumber="6" unit="m" value="981.0">981 m</quantity>
a.s.l.;
<collectionCode id="ED3D4486156CFFC1FEC3FB6E6B6FAF8E" box="[283,353,1180,1203]" pageId="7" pageNumber="6">FKCP</collectionCode>
.
</paragraph>
<paragraph id="8B93DC43156CFFC1FF18FB28685AAE94" blockId="7.[192,802,1242,1449]" pageId="7" pageNumber="6">
<materialsCitation id="3B44D61E156CFFC1FF18FB28685FAE94" collectingDate="2011-07-19" collectingDateMax="2011-07-20" collectingDateMin="2011-07-19" collectionCode="FKCP" collectorName="Awash N. P. &amp; F. Kovarik &amp; P. Novak. All" country="Ethiopia" elevation="981" latitude="9.009583" location="Awash" longLatPrecision="1" longitude="40.299026" pageId="7" pageNumber="6" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="holotype">
TYPE MATERIAL.
<collectingCountry id="F33B9CD3156CFFC1FE40FB286BFBAFCC" box="[408,501,1242,1265]" name="Ethiopia" pageId="7" pageNumber="6">Ethiopia</collectingCountry>
,
<location id="8EF38A98156CFFC1FDD7FB286852AFCC" LSID="urn:lsid:plazi:treatment:03856D55156CFFCCFF01FBF06989AA64:8EF38A98156CFFC1FDD7FB286852AFCC" box="[527,604,1242,1265]" country="Ethiopia" latitude="9.009583" longLatPrecision="1" longitude="40.299026" name="Awash" pageId="7" pageNumber="6">Awash</location>
,
<collectorName id="26D9B995156CFFC1FDAEFB286913AFCC" box="[630,797,1242,1265]" pageId="7" pageNumber="6">Awash N. P.</collectorName>
,
<geoCoordinate id="EE18BA84156CFFC1FF18FB0A6B51AE32" box="[192,351,1272,1295]" degrees="08" direction="north" minutes="52" orientation="latitude" pageId="7" pageNumber="6" precision="1" seconds="35.15" value="8.8764305">08°52'35.15&quot;N</geoCoordinate>
<geoCoordinate id="EE18BA84156CFFC1FEB0FB0A6BF7AE32" box="[360,505,1272,1295]" degrees="40" direction="east" minutes="05" orientation="longitude" pageId="7" pageNumber="6" precision="1" seconds="39.8" value="40.094387">40°05'39.8&quot;E</geoCoordinate>
,
<elevation id="00013B70156CFFC1FDDFFB0A6887AE32" box="[519,649,1272,1295]" metricMagnitude="2" metricUnit="m" metricValue="9.809999999999999" pageId="7" pageNumber="6" unit="m" value="981.0">
<quantity id="4CD471A6156CFFC1FDDFFB0A6843AE32" box="[519,589,1272,1295]" metricMagnitude="2" metricUnit="m" metricValue="9.809999999999999" pageId="7" pageNumber="6" unit="m" value="981.0">981 m</quantity>
a.s.l.
</elevation>
,
<date id="FF92FA83156CFFC1FD41FB0A6910AE32" box="[665,798,1272,1295]" pageId="7" pageNumber="6" value="2011-07-20">
<collectingDate id="EFD6036B156CFFC1FD41FB0A6910AE32" box="[665,798,1272,1295]" pageId="7" pageNumber="6" value="2011-07-20">20.VII.2011</collectingDate>
</date>
,
<specimenCount id="9D2A17CA156CFFC1FF18FAE56AEFAE13" box="[192,225,1303,1326]" pageId="7" pageNumber="6" type="male">1♂</specimenCount>
(
<typeStatus id="549762E1156CFFC1FF2EFAE56B57AE13" box="[246,345,1303,1326]" pageId="7" pageNumber="6" type="holotype">holotype</typeStatus>
), leg.
<collectorName id="26D9B995156CFFC1FE7DFAE5682CAE13" box="[421,546,1303,1326]" pageId="7" pageNumber="6">F. Kovařík</collectorName>
; Awash,
<geoCoordinate id="EE18BA84156CFFC1FD48FAE5692CAE13" box="[656,802,1303,1326]" degrees="09" direction="north" minutes="00" orientation="latitude" pageId="7" pageNumber="6" precision="1" seconds="34.5" value="9.009583">09°00'34.5&quot;N</geoCoordinate>
<geoCoordinate id="EE18BA84156CFFC1FF18FAC46B5FAE70" box="[192,337,1334,1357]" degrees="40" direction="east" minutes="17" orientation="longitude" pageId="7" pageNumber="6" precision="1" seconds="56.5" value="40.299026">40°17'56.5&quot;E</geoCoordinate>
,
<elevation id="00013B70156CFFC1FE87FAC46BE2AE70" box="[351,492,1334,1357]" metricMagnitude="3" metricUnit="m" metricValue="1.012" pageId="7" pageNumber="6" unit="m" value="1012.0">
<quantity id="4CD471A6156CFFC1FE87FAC46BBCAE70" box="[351,434,1334,1357]" metricMagnitude="3" metricUnit="m" metricValue="1.012" pageId="7" pageNumber="6" unit="m" value="1012.0">1012 m</quantity>
a.s.l.
</elevation>
(
<figureCitation id="1317C0C6156CFFC1FE26FAC4688AAE70" box="[510,644,1334,1357]" captionStart="Figures 2528" captionStartId="9.[192,278,1822,1845]" captionTargetBox="[232,1393,191,1791]" captionTargetId="figure-6@9.[232,1392,192,1790]" captionTargetPageId="9" captionText="Figures 2528: Pandinus (Pandinurus) awashensis sp. n. 2526. Type locality, Ethiopia, Awash, Awash N. P., 08°52'35.15&quot;N 40°05'39.8&quot;E, 981 m a.s.l.. 26. Water-capture hole well that produced the dead male holotype. 2728. Locality of allotype and paratypes, Ethiopia, Awash, 09°00'34.5&quot;N 40°17'56.5&quot;E, 1012 m. a.s.l.. 28. Rock under which was captured the female allotype." pageId="7" pageNumber="6">Figs. 2728</figureCitation>
),
<date id="FF92FA83156CFFC1FD40FAC46913AE70" box="[664,797,1334,1357]" pageId="7" pageNumber="6" value="2011-07-19">
<collectingDate id="EFD6036B156CFFC1FD40FAC46913AE70" box="[664,797,1334,1357]" pageId="7" pageNumber="6" value="2011-07-19">19.VII.2011</collectingDate>
</date>
,
<specimenCount id="9D2A17CA156CFFC1FF18FAA66AEFAE56" box="[192,225,1364,1387]" pageId="7" pageNumber="6" type="female">1♀</specimenCount>
(
<typeStatus id="549762E1156CFFC1FF2DFAA66B40AE56" box="[245,334,1364,1387]" pageId="7" pageNumber="6" type="allotype">allotype</typeStatus>
) 5juvs. ca.
<quantity id="4CD471A6156CFFC1FE01FAA66826AE56" box="[473,552,1364,1387]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="6" unit="mm" value="35.0">35 mm</quantity>
long (
<typeStatus id="549762E1156CFFC1FDA0FAA668EDAE56" box="[632,739,1364,1387]" pageId="7" pageNumber="6" type="paratype">paratypes</typeStatus>
, 4 of them still alive), leg.
<collectorName id="26D9B995156CFFC1FE7DFA816817AEB7" box="[421,537,1395,1418]" pageId="7" pageNumber="6">F. Kovařík</collectorName>
and
<collectorName id="26D9B995156CFFC1FD96FA8168ECAEB7" box="[590,738,1395,1418]" pageId="7" pageNumber="6">P. Novák. All</collectorName>
types are in the author´s collection (
<collectionCode id="ED3D4486156CFFC1FDD8FA606847AE94" box="[512,585,1426,1449]" pageId="7" pageNumber="6">FKCP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3368FC8156CFFC1FF18FA3D6B7CAD5C" pageId="7" pageNumber="6" type="etymology">
<paragraph id="8B93DC43156CFFC1FF18FA3D6B7CAD5C" blockId="7.[192,802,1487,1633]" pageId="7" pageNumber="6">
ETYMOLOGY. Named after Awash National Park, the type locality. The male
<typeStatus id="549762E1156CFFC1FE65FA1C6814AD38" box="[445,538,1518,1541]" pageId="7" pageNumber="6" type="holotype">holotype</typeStatus>
was with the help of the parks manager removed dead from a water-capture hole (
<figureCitation id="1317C0C6156CFFC1FF10F9D96B10AD7F" box="[200,286,1579,1602]" captionStart="Figures 2528" captionStartId="9.[192,278,1822,1845]" captionTargetBox="[232,1393,191,1791]" captionTargetId="figure-6@9.[232,1392,192,1790]" captionTargetPageId="9" captionText="Figures 2528: Pandinus (Pandinurus) awashensis sp. n. 2526. Type locality, Ethiopia, Awash, Awash N. P., 08°52'35.15&quot;N 40°05'39.8&quot;E, 981 m a.s.l.. 26. Water-capture hole well that produced the dead male holotype. 2728. Locality of allotype and paratypes, Ethiopia, Awash, 09°00'34.5&quot;N 40°17'56.5&quot;E, 1012 m. a.s.l.. 28. Rock under which was captured the female allotype." pageId="7" pageNumber="6">Fig. 26</figureCitation>
). Other specimens (
<typeStatus id="549762E1156CFFC1FDDBF9D96863AD7F" box="[515,621,1579,1602]" pageId="7" pageNumber="6" type="paratype">paratypes</typeStatus>
) were collected outside the park.
</paragraph>
</subSubSection>
<subSubSection id="C3368FC8156CFFC1FF18F9756EF0AAEE" pageId="7" pageNumber="6" type="diagnosis">
<paragraph id="8B93DC43156CFFC1FF18F9756EF0AAEE" blockId="7.[192,802,1671,1909]" lastBlockId="7.[830,1441,199,467]" pageId="7" pageNumber="6">
DIAGNOSIS. Total length
<quantity id="4CD471A6156CFFC1FE17F975685DADA3" box="[463,595,1671,1694]" metricMagnitude="-2" metricUnit="m" metricValue="9.65" metricValueMax="11.0" metricValueMin="8.3" pageId="7" pageNumber="6" unit="mm" value="96.5" valueMax="110.0" valueMin="83.0">83110 mm</quantity>
. Movable finger of pedipalp without noticeable sexual dimorphism. Color uniformly reddish black, only legs, telson and chela reddish brown, slightly lighter-colored than body. Chelicerae brown, reticulate, with black fingers and anterior margins. External trichobothria on patella number 15 (5
<emphasis id="B9580051156CFFC1FF18F8CD6AD7AC6B" box="[192,217,1855,1878]" italics="true" pageId="7" pageNumber="6">eb</emphasis>
, 3
<emphasis id="B9580051156CFFC1FEDEF8CD6B27AC6B" box="[262,297,1855,1878]" italics="true" pageId="7" pageNumber="6">esb</emphasis>
, 2
<emphasis id="B9580051156CFFC1FE8EF8CD6B7BAC6B" box="[342,373,1855,1878]" italics="true" pageId="7" pageNumber="6">em</emphasis>
, 1
<emphasis id="B9580051156CFFC1FE79F8CD6BB1AC6B" box="[417,447,1855,1878]" italics="true" pageId="7" pageNumber="6">est</emphasis>
, 3
<emphasis id="B9580051156CFFC1FE33F8CD6BF0AC6B" box="[491,510,1855,1878]" italics="true" pageId="7" pageNumber="6">et</emphasis>
); ventral trichobothria on chela number 9 to 11. Pectinal teeth number 1822. Spination formula of tarsomere II = 6/5: 7/5: 7-8/5: 7- 8/5. Tarsomere II legs with 3 spines on inclined anteroventral surface. Tarsomere I legs IIII with spina but leg IV with seta distal prosuperior. All body sparsely hirsute, chela hirsute densely. Dorsal surface of manus with evenly sized conspicuous granules. External surface of chela smooth, with several granules in anterior part and with two granulated carinae. Chela of pedipalp length to width ratio ca.
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both sexes.
</paragraph>
</subSubSection>
<subSubSection id="C3368FC8156CFFCCFCE6FE0A6989AA64" lastPageId="10" lastPageNumber="9" pageId="7" pageNumber="6" type="description">
<paragraph id="8B93DC43156CFFC1FCE6FE0A69F8A842" blockId="7.[830,1448,504,1907]" pageId="7" pageNumber="6">
DESCRIPTION. The adults are
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long. The habitus is shown in
<figureCitation id="1317C0C6156CFFC1FBC5FDE56F01A913" box="[1053,1295,535,558]" captionStart="Figures 918" captionStartId="5.[192,278,1830,1853]" captionTargetBox="[212,1368,208,1788]" captionTargetId="figure-6@5.[208,1368,202,1800]" captionTargetPageId="5" captionText="Figures 918: Pandinus (Pandinurus) awashensis sp. n. 913. Dorsal and ventral views, fifth metasomal segment and telson, carapace, and pectinal area, ♂ (110 mm) holotype. 1418. Dorsal and ventral views, fifth metasomal segment and telson, pectinal area, and carapace, ♀ (83 mm) allotype." pageId="7" pageNumber="6">Figs. 910 and 1415</figureCitation>
. For position and distribution of trichobothria of pedipalps see
<figureCitation id="1317C0C6156CFFC1FA96FDC76F93A971" box="[1358,1437,565,588]" captionStart="Figures 1922" captionStartId="6.[192,278,1867,1890]" captionTargetBox="[221,1393,194,1842]" captionTargetId="figure-10@6.[221,1393,186,1848]" captionTargetPageId="6" captionText="Figures 1922: Pandinus (Pandinurus) awashensis sp. n. 1921. Chela dorsal, external and ventral/internal views, ♂ (110 mm) holotype. 22. Trichobothrial pattern, ♀ (83 mm) allotype." pageId="7" pageNumber="6">Fig. 22</figureCitation>
. External trichobothria on patella number 15 (5
<emphasis id="B9580051156CFFC1FAE2FDA66F5DA956" box="[1338,1363,596,619]" italics="true" pageId="7" pageNumber="6">eb</emphasis>
, 3
<emphasis id="B9580051156CFFC1FAAEFDA66F97A956" box="[1398,1433,596,619]" italics="true" pageId="7" pageNumber="6">esb</emphasis>
, 2
<emphasis id="B9580051156CFFC1FC8CFD81697DA9B7" box="[852,883,627,650]" italics="true" pageId="7" pageNumber="6">em</emphasis>
, 1
<emphasis id="B9580051156CFFC1FC40FD8169B8A9B7" box="[920,950,627,650]" italics="true" pageId="7" pageNumber="6">est</emphasis>
, 3
<emphasis id="B9580051156CFFC1FC02FD8169E3A9B7" box="[986,1005,627,650]" italics="true" pageId="7" pageNumber="6">et</emphasis>
); ventral trichobothria on chela number 9 to 11. The male has a slightly larger telson than the female (
<figureCitation id="1317C0C6156CFFC1FC42FD4269F9A9FA" box="[922,1015,688,711]" captionStart="Figures 918" captionStartId="5.[192,278,1830,1853]" captionTargetBox="[212,1368,208,1788]" captionTargetId="figure-6@5.[208,1368,202,1800]" captionTargetPageId="5" captionText="Figures 918: Pandinus (Pandinurus) awashensis sp. n. 913. Dorsal and ventral views, fifth metasomal segment and telson, carapace, and pectinal area, ♂ (110 mm) holotype. 1418. Dorsal and ventral views, fifth metasomal segment and telson, pectinal area, and carapace, ♀ (83 mm) allotype." pageId="7" pageNumber="6">Figs. 11</figureCitation>
versus 16). Movable fingers of pedipalps do not show any noticeable sexual dimorphism. COLORATION (
<figureCitation id="1317C0C6156CFFC1FC07FD1F6E57A839" box="[991,1113,749,772]" captionStart="Figures 918" captionStartId="5.[192,278,1830,1853]" captionTargetBox="[212,1368,208,1788]" captionTargetId="figure-6@5.[208,1368,202,1800]" captionTargetPageId="5" captionText="Figures 918: Pandinus (Pandinurus) awashensis sp. n. 913. Dorsal and ventral views, fifth metasomal segment and telson, carapace, and pectinal area, ♂ (110 mm) holotype. 1418. Dorsal and ventral views, fifth metasomal segment and telson, pectinal area, and carapace, ♀ (83 mm) allotype." pageId="7" pageNumber="6">Figs. 918</figureCitation>
). The color is uniformly reddish black, only the legs, telson and chela are reddish brown, slightly lighter-colored than the body. The chelicerae are brown, reticulate, with black fingers and anterior margins.
</paragraph>
<paragraph id="8B93DC43156CFFC1FCE6FC756E7AAFAE" blockId="7.[830,1448,504,1907]" pageId="7" pageNumber="6">
CARAPACE. The carapace (
<figureCitation id="1317C0C6156CFFC1FB87FC756F0BA8A3" box="[1119,1285,903,926]" captionStart="Figures 918" captionStartId="5.[192,278,1830,1853]" captionTargetBox="[212,1368,208,1788]" captionTargetId="figure-6@5.[208,1368,202,1800]" captionTargetPageId="5" captionText="Figures 918: Pandinus (Pandinurus) awashensis sp. n. 913. Dorsal and ventral views, fifth metasomal segment and telson, carapace, and pectinal area, ♂ (110 mm) holotype. 1418. Dorsal and ventral views, fifth metasomal segment and telson, pectinal area, and carapace, ♀ (83 mm) allotype." pageId="7" pageNumber="6">Figs. 12 and 18</figureCitation>
) lacks carinae but has a deep sagittal furrow with a forked furrow on each side in the posterior part. The surface is smooth, with sparse granules and punctures. The anteromedial margin of the carapace is strongly concave. Present are a pair of median eyes and three lateral eyes, one of them smaller. The distance ratio of the pair of median eyes from the anterior or the posterior margin of the carapace is, respectively, 0.55 or 0.45.
</paragraph>
<paragraph id="8B93DC43156CFFC1FCE6FB696E80AEB5" blockId="7.[830,1448,504,1907]" pageId="7" pageNumber="6">
MESOSOMA. The tergites are tuberculate and finely granulated and bear an incomplete, smooth sagittal carina and shallow, symmetrical furrows. The sternites are smooth, lack carinae and bear two pronounced furrows. The pectinal teeth number
<quantity id="4CD471A6156CFFC1FBB9FAE76ECFAE11" box="[1121,1217,1301,1324]" metricMagnitude="-1" metricUnit="m" metricValue="4.953" metricValueMax="5.08" metricValueMin="4.826" pageId="7" pageNumber="6" unit="in" value="19.5" valueMax="20.0" valueMin="19.0">1920 in</quantity>
males,
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females and
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juveniles. The pectines have three marginal lamellae and four middle lamellae. The fulcra are long and bear numerous white setae.
</paragraph>
<paragraph id="8B93DC43156CFFC1FCE6FA6269C0ADC5" blockId="7.[830,1448,504,1907]" pageId="7" pageNumber="6">
METASOMA AND TELSON. The first throught fourth segments bear eight carinae. Most of the carinae are smooth. The dorsal carinae are composed of three to seven teeth of similar size. The fifth segment bears five to seven carinae of which the lateral carinae may be incomplete. All carinae on the fifth metasomal segment are composed of granules. The surface between the carinae is smooth, with solitary granules. The telson is elongate, slightly larger in the male (
<figureCitation id="1317C0C6156CFFC1FB16F9776F12ADA1" box="[1230,1308,1669,1692]" captionStart="Figures 918" captionStartId="5.[192,278,1830,1853]" captionTargetBox="[212,1368,208,1788]" captionTargetId="figure-6@5.[208,1368,202,1800]" captionTargetPageId="5" captionText="Figures 918: Pandinus (Pandinurus) awashensis sp. n. 913. Dorsal and ventral views, fifth metasomal segment and telson, carapace, and pectinal area, ♂ (110 mm) holotype. 1418. Dorsal and ventral views, fifth metasomal segment and telson, pectinal area, and carapace, ♀ (83 mm) allotype." pageId="7" pageNumber="6">Fig. 11</figureCitation>
) than in the female (
<figureCitation id="1317C0C6156CFFC1FC42F95669E1AD86" box="[922,1007,1700,1723]" captionStart="Figures 918" captionStartId="5.[192,278,1830,1853]" captionTargetBox="[212,1368,208,1788]" captionTargetId="figure-6@5.[208,1368,202,1800]" captionTargetPageId="5" captionText="Figures 918: Pandinus (Pandinurus) awashensis sp. n. 913. Dorsal and ventral views, fifth metasomal segment and telson, carapace, and pectinal area, ♂ (110 mm) holotype. 1418. Dorsal and ventral views, fifth metasomal segment and telson, pectinal area, and carapace, ♀ (83 mm) allotype." pageId="7" pageNumber="6">Fig. 16</figureCitation>
). The surface of the telson is smooth, hirsute, with several granules which compose incomplete carinae.
</paragraph>
<paragraph id="8B93DC43156CFFCCFCE6F8F26890ABC0" blockId="7.[830,1448,504,1907]" lastBlockId="10.[192,802,199,1602]" lastPageId="10" lastPageNumber="9" pageId="7" pageNumber="6">
LEGS. The legs are smooth, hirsute, without carinae and granules. The spination formula of tarsomere II = 6/5: 7/5: 7-8/5: 7-8/5. Tarsomere II bears 3 spines on the inclined anteroventral surface (
<figureCitation id="1317C0C6156CFFC1FB49F8AE6EEEAC4E" box="[1169,1248,1884,1907]" captionStart="Figures 5763" captionStartId="19.[192,278,1758,1781]" captionTargetBox="[220,1413,186,1722]" captionTargetId="figure-6@19.[212,1416,174,1728]" captionTargetPageId="19" captionText="Figures 5763: Tarsomere II and part of tarsomere I of 4th leg of subgenus Pandinurus. 57. P. lowei sp. n., ♂ (98 mm) holotype. 58. P. viatoris (Pocock, 1890), ♂ (105 mm), Democratic Republic of Congo, Parc National de Upemba, Kapelwa, 1780 m. a.s.l, ISNB. 59. P. somalilandus sp. n., ♀ (110 mm) holotype. 6061. P. awashensis sp. n.. 60. ♀ (83 mm) allotype. 61. ♂ (110 mm) holotype. 62. P. meidensis Karsch, 1879, ♂ (ca 140 mm), Somalia, Sar Uanle, about 20 km south of Chisimaio, 00°29'48&quot;S 42°25'30&quot;E, MZUF. 63. P. gregorii Pocock, 1896, ♀ (110 mm), Kenya, Maramtu Hill, Maramtu Village, Garissa, FKCP." pageId="7" pageNumber="6">Fig. 60</figureCitation>
). Tarsomere I of legs IIII bears a spina, but on leg IV it bears a seta distal prosuperior (see red arrow on
<figureCitation id="1317C0C61561FFCCFD9AFF14689FABC0" box="[578,657,230,253]" captionStart="Figures 5763" captionStartId="19.[192,278,1758,1781]" captionTargetBox="[220,1413,186,1722]" captionTargetId="figure-6@19.[212,1416,174,1728]" captionTargetPageId="19" captionText="Figures 5763: Tarsomere II and part of tarsomere I of 4th leg of subgenus Pandinurus. 57. P. lowei sp. n., ♂ (98 mm) holotype. 58. P. viatoris (Pocock, 1890), ♂ (105 mm), Democratic Republic of Congo, Parc National de Upemba, Kapelwa, 1780 m. a.s.l, ISNB. 59. P. somalilandus sp. n., ♀ (110 mm) holotype. 6061. P. awashensis sp. n.. 60. ♀ (83 mm) allotype. 61. ♂ (110 mm) holotype. 62. P. meidensis Karsch, 1879, ♂ (ca 140 mm), Somalia, Sar Uanle, about 20 km south of Chisimaio, 00°29'48&quot;S 42°25'30&quot;E, MZUF. 63. P. gregorii Pocock, 1896, ♀ (110 mm), Kenya, Maramtu Hill, Maramtu Village, Garissa, FKCP." pageId="10" pageNumber="9">Fig. 61</figureCitation>
).
</paragraph>
<caption id="DF538CCB1563FFCEFF18F8DC6BDAAC60" pageId="8" pageNumber="7" startId="8.[192,278,1838,1861]" targetBox="[192,1440,192,1808]" targetPageId="8">
<paragraph id="8B93DC431563FFCEFF18F8DC6BDAAC60" blockId="8.[192,1440,1838,1885]" pageId="8" pageNumber="7">
<emphasis id="B95800511563FFCEFF18F8DC6903AC79" bold="true" box="[192,781,1838,1861]" pageId="8" pageNumber="7">
Figures 2324:
<taxonomicName id="4C2CA7C01563FFCEFEB7F8DC68DEAC79" authority="Kovařík, 2012" authorityName="Kovařík" authorityYear="2012" box="[367,720,1838,1860]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="awashensis" status="sp. nov." subGenus="Pandinurus">
<emphasis id="B95800511563FFCEFEB7F8DC6BC0AC79" bold="true" box="[367,462,1838,1860]" italics="true" pageId="8" pageNumber="7">Pandinus</emphasis>
(
<emphasis id="B95800511563FFCEFE04F8DC685FAC79" bold="true" box="[476,593,1838,1860]" italics="true" pageId="8" pageNumber="7">Pandinurus</emphasis>
)
<emphasis id="B95800511563FFCEFDB8F8DC68DEAC79" bold="true" box="[608,720,1838,1860]" italics="true" pageId="8" pageNumber="7">awashensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26BBD2A1563FFCEFD0FF8DD6903AC79" box="[727,781,1839,1860]" pageId="8" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, ♀ (83 mm) allotype at the locality, Ethiopia, Awash, 09°00'34.5&quot;N 40°17'56.5&quot;E, 1012 m. a.s.l..
</paragraph>
</caption>
<caption id="DF538CCB1562FFCFFF18F8EC6B55AC42" pageId="9" pageNumber="8" startId="9.[192,278,1822,1845]" targetBox="[232,1393,191,1791]" targetPageId="9">
<paragraph id="8B93DC431562FFCFFF18F8EC6B55AC42" blockId="9.[192,1440,1822,1919]" pageId="9" pageNumber="8">
<emphasis id="B95800511562FFCFFF18F8EC699BAC09" bold="true" box="[192,917,1822,1845]" pageId="9" pageNumber="8">
Figures 2528:
<taxonomicName id="4C2CA7C01562FFCFFE5BF8EC68F6AC09" authority="Kovařík, 2012" authorityName="Kovařík" authorityYear="2012" box="[387,760,1822,1844]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="awashensis" status="sp. nov." subGenus="Pandinurus">
<emphasis id="B95800511562FFCFFE5BF8EC6BECAC09" bold="true" box="[387,482,1822,1844]" italics="true" pageId="9" pageNumber="8">Pandinus</emphasis>
(
<emphasis id="B95800511562FFCFFE22F8EC6861AC09" bold="true" box="[506,623,1822,1844]" italics="true" pageId="9" pageNumber="8">Pandinurus</emphasis>
)
<emphasis id="B95800511562FFCFFD50F8EC68F6AC09" bold="true" box="[648,760,1822,1844]" italics="true" pageId="9" pageNumber="8">awashensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26BBD2A1562FFCFFCD0F8ED6947AC09" box="[776,841,1823,1844]" pageId="9" pageNumber="8" rank="species">sp. n.</taxonomicNameLabel>
2526
</emphasis>
. Type locality, Ethiopia, Awash, Awash N. P., 08°52'35.15&quot;N 40°05'39.8&quot;E, 981 m a.s.l..
<emphasis id="B95800511562FFCFFDB8F8CA6876AC70" bold="true" box="[608,632,1848,1869]" pageId="9" pageNumber="8">26</emphasis>
. Water-capture hole well that produced the dead male holotype.
<emphasis id="B95800511562FFCFFB35F8CA6F27AC70" bold="true" box="[1261,1321,1848,1869]" pageId="9" pageNumber="8">2728</emphasis>
. Locality of allotype and paratypes, Ethiopia, Awash, 09°00'34.5&quot;N 40°17'56.5&quot;E, 1012 m. a.s.l..
<emphasis id="B95800511562FFCFFBCAF8A36E24AC5B" bold="true" box="[1042,1066,1873,1894]" pageId="9" pageNumber="8">28</emphasis>
. Rock under which was captured the female allotype.
</paragraph>
</caption>
<paragraph id="8B93DC431561FFCCFF18FEF66828A883" blockId="10.[192,802,199,1602]" pageId="10" pageNumber="9">
PEDIPALPS (
<figureCitation id="1317C0C61561FFCCFE99FEF66BCBAA26" box="[321,453,260,283]" captionStart="Figures 1922" captionStartId="6.[192,278,1867,1890]" captionTargetBox="[221,1393,194,1842]" captionTargetId="figure-10@6.[221,1393,186,1848]" captionTargetPageId="6" captionText="Figures 1922: Pandinus (Pandinurus) awashensis sp. n. 1921. Chela dorsal, external and ventral/internal views, ♂ (110 mm) holotype. 22. Trichobothrial pattern, ♀ (83 mm) allotype." pageId="10" pageNumber="9">Figs. 1922</figureCitation>
). The pedipalps are hirsute. The femur and patella bear several large granules. The femur bears four carinae composed of several large, round granules, only the externoventral carina is smooth. The patella bears five complete carinae with granules or smooth. Several granules are only on the external surface of the patella. The chela is lobate and bears only two smooth ventral carinae. The dorsal surface of the chela bears granules that are neither conical nor pointed and whose summits may be confluent. The external surface of the chela is smooth, with several conical granules in anterior part and without carinae. The chela has a lobe. The dentate margins of the movable and fixed fingers are armed with two parallel rows of denticles extending the entire length of the finger, with larger granules which indicate five or six subrows on the movable finger and the fixed finger. The dorsal surface of the manus bears evenly sized conspicuous granules. The internal surface of the chela is smooth, with several granules in anterior part and with two granulated carinae. The dorsal and external surfaces of the chela bear four carinae; the total number of carinae on the is nine, but some of them are incomplete.
</paragraph>
<paragraph id="8B93DC431561FFCCFF18FC346BE6AE32" blockId="10.[192,802,199,1602]" pageId="10" pageNumber="9">
MEASUREMENTS IN MM.
<emphasis id="B95800511561FFCCFE38FC34688CA8E0" box="[480,642,966,989]" italics="true" pageId="10" pageNumber="9">
Male
<typeStatus id="549762E11561FFCCFDFEFC34688CA8E0" box="[550,642,966,989]" pageId="10" pageNumber="9" type="holotype">holotype</typeStatus>
</emphasis>
. Total length 110; carapace length 13.8, width 15.1; metasoma and telson length 55.1; first metasomal segment length 6.4, width 6.1, depth 5.1; second metasomal segment length 7.4, width 5.5, depth 4.8; third metasomal segment length 8.3, width 5.0; fourth metasomal segment length 9.3, width 4.5; fifth metasomal segment length 12.0, width 4.1; telson length 11.7; telson width 5.3; pedipalp femur length 11.1, width 4.9; pedipalp patella length 10.8, width 5.4; chela length 23.4; manus width 11.1; movable finger length 14.8.
</paragraph>
<paragraph id="8B93DC431561FFCCFF18FAE568C1AD7F" blockId="10.[192,802,199,1602]" pageId="10" pageNumber="9">
<emphasis id="B95800511561FFCCFF18FAE56B7EAE13" box="[192,368,1303,1326]" italics="true" pageId="10" pageNumber="9">
Female
<typeStatus id="549762E11561FFCCFEC2FAE56B7EAE13" box="[282,368,1303,1326]" pageId="10" pageNumber="9" type="allotype">allotype</typeStatus>
</emphasis>
. Total length 83; carapace length 12.9, width 12.9; metasoma and telson length 39.5; first metasomal segment length 4.8, width 4.5, depth 4.1; second metasomal segment length 5.5, width 4.1, depth 3.9; third metasomal segment length 5.9, width 3.8; fourth metasomal segment length 6.6, width 3.3; fifth metasomal segment length 8.7, width 3.2; telson length 8.0; telson width 3.2; pedipalp femur length 8.5, width 4.0; pedipalp patella length 9.0, width 4.5; chela length 19; manus width 9.2; movable finger length 11.3.
</paragraph>
<paragraph id="8B93DC431561FFCCFF18F99A6989AA64" blockId="10.[192,802,1640,1909]" lastBlockId="10.[830,1440,199,345]" pageId="10" pageNumber="9">
AFFINITIES. The described features distinguish
<taxonomicName id="4C2CA7C01561FFCCFD65F99A6B34ADA3" authority="Kovařík, 2012" authorityName="Kovařík" authorityYear="2012" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="awashensis" status="sp. nov.">
<emphasis id="B95800511561FFCCFD65F99A6B34ADA3" italics="true" pageId="10" pageNumber="9">Pandinus awashensis</emphasis>
</taxonomicName>
<emphasis id="B95800511561FFCCFE99F9756B70ADA3" bold="true" box="[321,382,1671,1694]" pageId="10" pageNumber="9">
<taxonomicNameLabel id="A26BBD2A1561FFCCFE99F9756B70ADA3" box="[321,382,1671,1694]" pageId="10" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
from all other species of the subgenus
<taxonomicName id="4C2CA7C01561FFCCFF18F9546B32AD80" authorityName="Fet" authorityYear="1997" box="[192,316,1702,1725]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="subGenus" subGenus="Pandinurus">
<emphasis id="B95800511561FFCCFF18F9546B32AD80" box="[192,316,1702,1725]" italics="true" pageId="10" pageNumber="9">Pandinurus</emphasis>
</taxonomicName>
. The shape and granulations of the chela distinguish
<taxonomicName id="4C2CA7C01561FFCCFEE7F9366BD4ADE6" authority="Kovařík, 2012" authorityName="Kovařík" authorityYear="2012" box="[319,474,1732,1755]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="awashensis" status="sp. nov.">
<emphasis id="B95800511561FFCCFEE7F9366BD4ADE6" box="[319,474,1732,1755]" italics="true" pageId="10" pageNumber="9">P. awashensis</emphasis>
</taxonomicName>
<emphasis id="B95800511561FFCCFE3BF936682CADE6" bold="true" box="[483,546,1732,1755]" pageId="10" pageNumber="9">
<taxonomicNameLabel id="A26BBD2A1561FFCCFE3BF936682CADE6" box="[483,546,1732,1755]" pageId="10" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
from
<taxonomicName id="4C2CA7C01561FFCCFDB2F93668E3ADE6" baseAuthorityName="Birula" baseAuthorityYear="1928" box="[618,749,1732,1755]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="magrettii">
<emphasis id="B95800511561FFCCFDB2F93668E3ADE6" box="[618,749,1732,1755]" italics="true" pageId="10" pageNumber="9">P. magrettii</emphasis>
</taxonomicName>
(see key) with which it shares sexual dimorphism (the male has a larger telson than the female but movable fingers of pedipalps do not show any noticeable sexual dimorphism). This distinguishes the new species from the other two species in the region,
<taxonomicName id="4C2CA7C01561FFCCFD80F8AC68C0AC48" baseAuthorityName="Hirst" baseAuthorityYear="1911" box="[600,718,1886,1909]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="exitialis">
<emphasis id="B95800511561FFCCFD80F8AC68C0AC48" box="[600,718,1886,1909]" italics="true" pageId="10" pageNumber="9">P. exitialis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2CA7C01561FFCCFCD3F8AC69A7ABE3" authorityName="Hirst" authorityYear="1911" box="[779,937,199,1909]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="sudanicus">
<emphasis id="B95800511561FFCCFCD3F8AC69A7ABE3" box="[779,937,199,1909]" italics="true" pageId="10" pageNumber="9">P. sudanicus</emphasis>
</taxonomicName>
, whose males have a more pronounced tooth on the movable finger of pedipalp. It appears that the most similar species is
<taxonomicName id="4C2CA7C01561FFCCFBEFFEF66EECAA26" authority="Kovařík, 2012" authorityName="Kovařík" authorityYear="2012" box="[1079,1250,260,283]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="somalilandus" status="sp. nov.">
<emphasis id="B95800511561FFCCFBEFFEF66EECAA26" box="[1079,1250,260,283]" italics="true" pageId="10" pageNumber="9">P. somalilandus</emphasis>
</taxonomicName>
<emphasis id="B95800511561FFCCFB31FEF66F2BAA26" bold="true" box="[1257,1317,260,283]" pageId="10" pageNumber="9">
<taxonomicNameLabel id="A26BBD2A1561FFCCFB31FEF66F2BAA26" box="[1257,1317,260,283]" pageId="10" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, which can be distinguished by the characters given in the key below.
</paragraph>
</subSubSection>
</treatment>
</document>