179 lines
18 KiB
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179 lines
18 KiB
XML
<document id="7EB58FB9AD0DA5CB4857C9D895118052" ID-DOI="10.5252/z2015n3a4" ID-ISSN="1638-9387" ID-Zenodo-Dep="5156547" ID-ZooBank="urn:lsid:zoobank.org:pub:EDB0C7A2-7B5E-4E39-91CE-B823BB38ACA0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628016323462" checkinUser="felipe" docAuthor="Roland, Houart & Virginie, Héros" docDate="2015" docId="75255D75FF8BCF28FE94EFA8FE0DFB9C" docLanguage="en" docName="Zoosystema.37.3.481-503.pdf" docOrigin="Zoosystema 37 (3)" docSource="http://dx.doi.org/10.5252/z2015n3a4" docStyle="DocumentStyle:0AF8C315773078909029C6FC3CC05C6C.3:Zoosystema.2015-2017.journal_article" docStyleId="0AF8C315773078909029C6FC3CC05C6C" docStyleName="Zoosystema.2015-2017.journal_article" docStyleVersion="3" docTitle="Timbellus de Gregorio 1885" docType="treatment" docVersion="4" lastPageNumber="486" masterDocId="891C250DFF8ECF2DFF91EF7FFFA2FFD2" masterDocTitle="New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean" masterLastPageNumber="503" masterPageNumber="481" pageNumber="486" updateTime="1699033106268" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="2113E5BCC56BD080C44FD1ECE9321670">
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<mods:title id="E390A88C3A54301EFE70C0607251D1C5">New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean</mods:title>
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<mods:name id="766806D6E5E35049E1D8673931A7F9EA" type="personal">
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<mods:namePart id="BEF6F49D8395BE1BBB896D80E64057CB">Roland, Houart</mods:namePart>
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<mods:namePart id="7F7EB7D6C8AD270BDC992C64B277C066">Virginie, Héros</mods:namePart>
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<mods:typeOfResource id="4B27FD2AB7C790CC2C6B4398CB5F4056">text</mods:typeOfResource>
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<mods:title id="AF4452B9CAF6913AC6CA01FAF59D2589">Zoosystema</mods:title>
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</mods:titleInfo>
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<mods:part id="B5406B6CE19E05CD90F4A9431EF972F1">
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<mods:date id="C5F421BF5D28C8D8E1183689B67CBD2B">2015</mods:date>
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<mods:detail id="18F87CC1E94AB4CB4223077F1CE03DB2" type="pubDate">
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<mods:number id="76836B50C731A0400DC1AB160FAF8BF2">2015-09-30</mods:number>
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<mods:number id="6B9671710DC3FD84EECC26652CB0F57F">37</mods:number>
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<mods:url id="B6961A4D3C0E43B148F07CCF5D2512B1">http://dx.doi.org/10.5252/z2015n3a4</mods:url>
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<mods:classification id="095A96AFDCD60C0347DDDABB9051D5E2">journal article</mods:classification>
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<mods:identifier id="562BD4491DF3A43672D4165FB16DDB76" type="DOI">10.5252/z2015n3a4</mods:identifier>
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<mods:identifier id="EE5358E4DEDF7268AF23D8E890095AE5" type="ISSN">1638-9387</mods:identifier>
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<treatment id="75255D75FF8BCF28FE94EFA8FE0DFB9C" ID-DOI="http://doi.org/10.5281/zenodo.5157135" ID-Zenodo-Dep="5157135" LSID="urn:lsid:plazi:treatment:75255D75FF8BCF28FE94EFA8FE0DFB9C" httpUri="http://treatment.plazi.org/id/75255D75FF8BCF28FE94EFA8FE0DFB9C" lastPageNumber="486" pageId="5" pageNumber="486">
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<subSubSection id="B596BFE8FF8BCF28FE94EFA8FD24FF20" box="[261,646,215,242]" pageId="5" pageNumber="486" type="nomenclature">
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<paragraph id="FD33EC63FF8BCF28FE94EFA8FD24FF20" blockId="5.[261,646,215,242]" box="[261,646,215,242]" pageId="5" pageNumber="486">
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<heading id="A67B5B0FFF8BCF28FE94EFA8FD24FF20" box="[261,646,215,242]" centered="true" fontSize="9" level="2" pageId="5" pageNumber="486" reason="2">
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Genus
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<taxonomicName id="3A8C97E0FF8BCF28FEC3EFA8FD24FF20" ID-CoL="7PJK5" authority="de Gregorio, 1885" authorityName="de Gregorio" authorityYear="1885" box="[338,646,215,242]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="486" phylum="Mollusca" rank="genus">
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<emphasis id="CFF83071FF8BCF28FEC3EFA8FE1BFF23" box="[338,441,215,241]" italics="true" pageId="5" pageNumber="486">Timbellus</emphasis>
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de
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<bibRefCitation id="991D9192FF8BCF28FE71EFA8FD24FF20" author="GREGORIO A. & DE" box="[480,646,215,242]" pageId="5" pageNumber="486" pagination="36 - 288" refId="ref15128" refString="GREGORIO A. DE, 1885. - Studi su talune conchiglie mediterranee viventi e fossili con una rivista del gen. Vulsella e del gen. Ficula. Bulletin de la Societa Malacalogico Italiana 10: 36 - 288." type="journal article" year="1885">Gregorio, 1885</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="B596BFE8FF8BCF28FF15EE6EFDD1FE91" pageId="5" pageNumber="486" type="materials_examined">
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<paragraph id="FD33EC63FF8BCF28FF15EE6EFDD1FE91" blockId="5.[132,775,272,323]" pageId="5" pageNumber="486">
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<materialsCitation id="4DE4E63EFF8BCF28FF15EE6EFDCDFE91" country="Italy" location="Middle Miocene" pageId="5" pageNumber="486" specimenCount="1" typeStatus="holotype">
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<typeStatus id="223752C1FF8BCF28FF15EE6EFF17FEF4" box="[132,181,273,296]" pageId="5" pageNumber="486">TYPE</typeStatus>
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SPECIES. —
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<taxonomicName id="3A8C97E0FF8BCF28FEBCEE6EFDF6FEFA" authority="Bellardi, 1872" authorityName="Bellardi" authorityYear="1872" box="[301,596,272,296]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="486" phylum="Mollusca" rank="species" species="latifolius">
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<emphasis id="CFF83071FF8BCF28FEBCEE6EFE60FEFA" box="[301,450,272,296]" italics="true" pageId="5" pageNumber="486">Murex latifolius</emphasis>
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Bellardi, 1872
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</taxonomicName>
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,
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<location id="F853BAB8FF8BCF28FDCFEE6FFCA1FEFA" LSID="urn:lsid:plazi:treatment:75255D75FF8BCF28FE94EFA8FE0DFB9C:F853BAB8FF8BCF28FDCFEE6FFCA1FEFA" box="[606,771,272,296]" country="Italy" name="Middle Miocene" pageId="5" pageNumber="486">Middle Miocene</location>
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,
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<collectingCountry id="859BACF3FF8BCF28FF15EE54FF0DFE91" box="[132,175,299,323]" name="Italy" pageId="5" pageNumber="486">Italy</collectingCountry>
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(by subsequent designation;
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<bibRefCitation id="991D9192FF8BCF28FE5DEE54FDC5FE91" author="VOKES E. H." box="[460,615,299,323]" pageId="5" pageNumber="486" pagination="1 - 41" refId="ref15904" refString="VOKES E. H. 1964. - Supraspecific groups in the subfamilies Muricinae & Tritonaliinae (Gastropoda: Muricidae). Malacologia 2 (1): 1 - 41." type="journal article" year="1964">Vokes 1964: 14</bibRefCitation>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="B596BFE8FF8BCF28FF15EE08FE0DFB9C" pageId="5" pageNumber="486" type="discussion">
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<paragraph id="FD33EC63FF8BCF28FF15EE08FF4AFE5D" blockId="5.[130,777,375,1102]" box="[132,232,375,401]" pageId="5" pageNumber="486">REMARKS</paragraph>
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<paragraph id="FD33EC63FF8BCF28FF15EEE9FEEAFD3D" blockId="5.[130,777,375,1102]" pageId="5" pageNumber="486">
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The genus
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<taxonomicName id="3A8C97E0FF8BCF28FF6FEEE9FEC5FE62" authorityName="de Gregorio" authorityYear="1885" box="[254,359,406,432]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="486" phylum="Mollusca" rank="genus">
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<emphasis id="CFF83071FF8BCF28FF6FEEE9FEC5FE62" box="[254,359,406,432]" italics="true" pageId="5" pageNumber="486">Timbellus</emphasis>
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</taxonomicName>
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was reinstated by
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<bibRefCitation id="991D9192FF8BCF28FDA4EEE9FCA5FE63" author="MERLE D. & GARRIGUES B. & POINTIER J. P." box="[565,775,406,433]" pageId="5" pageNumber="486" refId="ref15629" refString="MERLE D., GARRIGUES B. & POINTIER J. P. 2011. - Fossil and Recent Muricidae of the World - Part Muricinae. Conchbooks, Hackenheim, 648 p." type="book" year="2011">
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Merle
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<emphasis id="CFF83071FF8BCF28FDEFEEE8FD16FE62" box="[638,692,406,432]" italics="true" pageId="5" pageNumber="486">et al.</emphasis>
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(2011)
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</bibRefCitation>
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for the species generally classified in
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<taxonomicName id="3A8C97E0FF8BCF28FD96EEC8FDD6FE02" box="[519,628,439,464]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="486" phylum="Mollusca" rank="genus" sensu="stricto">
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<emphasis id="CFF83071FF8BCF28FD96EEC8FDD6FE02" box="[519,628,439,464]" italics="true" pageId="5" pageNumber="486">Pterynotus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D4CB8D0AFF8BCF28FDE6EEC8FD36FE03" box="[631,660,439,465]" pageId="5" pageNumber="486" sensu="stricto">
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<emphasis id="CFF83071FF8BCF28FDE6EEC8FD2EFE02" box="[631,652,439,464]" italics="true" pageId="5" pageNumber="486">s.s</emphasis>
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.
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</taxonomicNameLabel>
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Swainson, 1833 but usually with a less scabrous shell morphology and with three major axial varices “appearing early during the ontogeny”. This distinction from the
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<taxonomicName id="3A8C97E0FF8BCF28FDBBED68FD3AFDE2" box="[554,664,535,560]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="486" phylum="Mollusca" rank="genus" sensu="stricto">
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<emphasis id="CFF83071FF8BCF28FDBBED68FD3AFDE2" box="[554,664,535,560]" italics="true" pageId="5" pageNumber="486">Pterynotus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D4CB8D0AFF8BCF28FD0EED68FD1FFDE3" box="[671,701,535,561]" pageId="5" pageNumber="486" sensu="stricto">
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<emphasis id="CFF83071FF8BCF28FD0EED68FD14FDE2" box="[671,694,535,560]" italics="true" pageId="5" pageNumber="486">s.s</emphasis>
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.
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</taxonomicNameLabel>
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group was noted by
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<bibRefCitation id="991D9192FF8BCF28FEB4ED48FE5CFD82" author="BARCO A. & CLAREMONT M. & REID D. G. & HOUART R. & BOUCHET P. & WILLIAMS S. T. & CRUAUD C. & COULOUX A. & OLIVERIO M." box="[293,510,566,593]" pageId="5" pageNumber="486" pagination="1025 - 1039" refId="ref14906" refString="BARCO A., CLAREMONT M., REID D. G., HOUART R., BOUCHET P., WILLIAMS S. T., CRUAUD C., COULOUX A. & OLIVERIO M. 2010. - A molecular phylogenetic framework for the Muricidae, a diverse family of carnivorous gastropods. Molecular Phylogenetics and Evolution 56: 1025 - 1039." type="journal article" year="2010">
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Barco
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<emphasis id="CFF83071FF8BCF28FEFFED48FE04FD82" box="[366,422,566,592]" italics="true" pageId="5" pageNumber="486">et al.</emphasis>
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(2010)
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</bibRefCitation>
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, with
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<taxonomicName id="3A8C97E0FF8BCF28FDD6ED48FE83FDA2" authority="Houart, 1988" authorityName="Houart" authorityYear="1988" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="486" phylum="Mollusca" rank="species" species="fulgens">
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<emphasis id="CFF83071FF8BCF28FDD6ED48FCAAFD82" box="[583,776,566,592]" italics="true" pageId="5" pageNumber="486">Pterynotus fulgens</emphasis>
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Houart,
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<quantity id="3A744186FF8BCF28FF76ED29FE83FDA2" box="[231,289,598,624]" metricMagnitude="1" metricUnit="m" metricValue="5.04952" pageId="5" pageNumber="486" unit="in" value="1988.0">1988</quantity>
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</taxonomicName>
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in their DNA analysis even falling outside the Muricinae Rafinesque, 1815. The classification used here follows
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<bibRefCitation id="991D9192FF8BCF28FE9CEDE9FE76FD62" author="BARCO A. & CLAREMONT M. & REID D. G. & HOUART R. & BOUCHET P. & WILLIAMS S. T. & CRUAUD C. & COULOUX A. & OLIVERIO M." box="[269,468,662,688]" pageId="5" pageNumber="486" pagination="1025 - 1039" refId="ref14906" refString="BARCO A., CLAREMONT M., REID D. G., HOUART R., BOUCHET P., WILLIAMS S. T., CRUAUD C., COULOUX A. & OLIVERIO M. 2010. - A molecular phylogenetic framework for the Muricidae, a diverse family of carnivorous gastropods. Molecular Phylogenetics and Evolution 56: 1025 - 1039." type="journal article" year="2010">
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Barco
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<emphasis id="CFF83071FF8BCF28FEC0EDE9FE26FD62" box="[337,388,662,688]" italics="true" pageId="5" pageNumber="486">et al.</emphasis>
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(2010)
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</bibRefCitation>
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pending availability of more material for further DNA research and the publication of additional results.
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</paragraph>
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<paragraph id="FD33EC63FF8BCF28FF0AED8AFE0DFB9C" blockId="5.[130,777,375,1102]" pageId="5" pageNumber="486">
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On the other hand, the classification of
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<bibRefCitation id="991D9192FF8BCF28FDD1ED8AFCA5FCDD" author="MERLE D. & GARRIGUES B. & POINTIER J. P." box="[576,775,757,783]" pageId="5" pageNumber="486" refId="ref15629" refString="MERLE D., GARRIGUES B. & POINTIER J. P. 2011. - Fossil and Recent Muricidae of the World - Part Muricinae. Conchbooks, Hackenheim, 648 p." type="book" year="2011">
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Merle
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<emphasis id="CFF83071FF8BCF28FD14ED89FD15FCDD" box="[645,695,757,783]" italics="true" pageId="5" pageNumber="486">et al.</emphasis>
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(2011)
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</bibRefCitation>
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mixes the species with a trivaricate axial sculpture starting already on the first teleoconch whorl, such as
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<taxonomicName id="3A8C97E0FF8BCF28FD0AEC4AFF41FCBD" baseAuthorityName="Bellardi" baseAuthorityYear="1872" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="486" phylum="Mollusca" rank="species" species="latifolius">
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<emphasis id="CFF83071FF8BCF28FD0AEC4AFF41FCBD" italics="true" pageId="5" pageNumber="486">Timbellus latifolius</emphasis>
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</taxonomicName>
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, the
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<typeStatus id="223752C1FF8BCF28FEB0EC29FEF3FCA2" box="[289,337,854,880]" pageId="5" pageNumber="486">type</typeStatus>
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species, and some Recent species like
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<taxonomicName id="3A8C97E0FF8BCF28FF17EC09FE3CFC5D" authority="(Houart, 1988)" baseAuthorityName="Houart" baseAuthorityYear="1988" box="[134,414,885,911]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="486" phylum="Mollusca" rank="species" species="fulgens">
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<emphasis id="CFF83071FF8BCF28FF17EC09FF48FC5D" box="[134,234,885,911]" italics="true" pageId="5" pageNumber="486">T. fulgens</emphasis>
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(Houart, 1988)
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</taxonomicName>
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. In
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<taxonomicName id="3A8C97E0FF8BCF28FE54EC0AFD2FFC5D" authority="Roland & Virginie, 2015" authorityName="Roland & Virginie" authorityYear="2015" box="[453,653,885,911]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="486" phylum="Mollusca" rank="species" species="goniodes" status="sp. nov.">
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<emphasis id="CFF83071FF8BCF28FE54EC0AFD2FFC5D" box="[453,653,885,911]" italics="true" pageId="5" pageNumber="486">Timbellus goniodes</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D4CB8D0AFF8BCF28FD03EC0AFD6DFC5D" box="[658,719,885,911]" pageId="5" pageNumber="486" rank="species">n. sp.</taxonomicNameLabel>
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, here described, and a few other species such as
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<taxonomicName id="3A8C97E0FF8BCF28FDC5ECE9FF5FFC1D" authority="(Brazier, 1878)" baseAuthorityName="Brazier" baseAuthorityYear="1878" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="486" phylum="Mollusca" rank="species" species="bednalli">
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<emphasis id="CFF83071FF8BCF28FDC5ECE9FD6AFC7D" box="[596,712,917,943]" italics="true" pageId="5" pageNumber="486">T. bednalli</emphasis>
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(Brazier, 1878)
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</taxonomicName>
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,
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<taxonomicName id="3A8C97E0FF8BCF28FE96ECCAFDC0FC1D" authority="(Lamarck, 1822)" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[263,610,949,975]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="486" phylum="Mollusca" rank="species" species="phyllopterus">
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<emphasis id="CFF83071FF8BCF28FE96ECCAFE00FC1D" box="[263,418,949,975]" italics="true" pageId="5" pageNumber="486">T. phyllopterus</emphasis>
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(Lamarck, 1822)
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</taxonomicName>
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and
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<taxonomicName id="3A8C97E0FF8BCF28FD09ECCAFEDDFC3D" authority="(Roding, 1798)" baseAuthorityName="Roding" baseAuthorityYear="1798" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="486" phylum="Mollusca" rank="species" species="alatus">
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<emphasis id="CFF83071FF8BCF28FD09ECCAFF61FC3C" italics="true" pageId="5" pageNumber="486">Pterynotus alatus</emphasis>
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(Röding, 1798)
|
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</taxonomicName>
|
|
, the
|
|
<typeStatus id="223752C1FF8BCF28FE29ECAAFE4AFC3D" box="[440,488,981,1007]" pageId="5" pageNumber="486">type</typeStatus>
|
|
species of
|
|
<taxonomicName id="3A8C97E0FF8BCF28FDF4ECAAFD76FC3C" box="[613,724,981,1006]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="486" phylum="Mollusca" rank="genus">
|
|
<emphasis id="CFF83071FF8BCF28FDF4ECAAFD76FC3C" box="[613,724,981,1006]" italics="true" pageId="5" pageNumber="486">Pterynotus</emphasis>
|
|
</taxonomicName>
|
|
, the trivaricate sculpture appears only on the third or fourth teleoconch whorl. Their generic placement awaits confirmation by genetic analysis.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
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</document> |