144 lines
18 KiB
XML
144 lines
18 KiB
XML
<document id="FBDD91BC71846CE6C27F427CBA71942D" ID-DOI="10.5281/zenodo.203245" ID-GBIF-Dataset="8484b438-d520-4985-bccf-f914612a5798" ID-ISSN="1175-5326" ID-Zenodo-Dep="203245" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460235874530" checkinUser="plazi" docAuthor="Sites, Robert W. & Zettel, Herbert" docDate="2011" docId="513487E73669FF9EFF4BD198FD5B8257" docLanguage="en" docName="zt02760p017.pdf" docOrigin="Zootaxa 2760" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Diaphorocoris Montandon 1897" docType="treatment" docVersion="11" lastPageNumber="9" masterDocId="AD0DFF9F3661FF96FFDCD10FFFA98557" masterDocTitle="Waterfall-inhabiting Naucoridae (Hemiptera: Heteroptera) of southern India and Sri Lanka: Pogonocaudina Sites and Zettel, n. gen., and a review of Diaphorocoris with descriptions of two new species" masterLastPageNumber="17" masterPageNumber="1" pageNumber="9" updateTime="1698239580061" updateUser="plazi">
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<mods:title id="FCA4E209280C729696AFF954BDE7D876">Waterfall-inhabiting Naucoridae (Hemiptera: Heteroptera) of southern India and Sri Lanka: Pogonocaudina Sites and Zettel, n. gen., and a review of Diaphorocoris with descriptions of two new species</mods:title>
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<mods:namePart id="B876FC429429CF50EE3D50CEC18369D1">Sites, Robert W.</mods:namePart>
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<mods:namePart id="E8774FC07F950063C2095DD4E9FE54D8">Zettel, Herbert</mods:namePart>
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<mods:title id="9AC51884C40E1F240189CDEB606730F9">Zootaxa</mods:title>
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<mods:date id="169A1C02588194B33803D39452CA5E33">2011</mods:date>
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<mods:detail id="A0477213D465BD35ADA3ABBE0F37A11B" type="volume">
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<mods:number id="A0C703C69A428977645999894FF92F9E">2760</mods:number>
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<mods:identifier id="8F58C3AD61E8E27C6FAD5240CF7425F3" type="DOI">10.5281/zenodo.203245</mods:identifier>
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<mods:identifier id="12390146E3A9F9667795B857058FC2B8" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="91806E459C391C8A4C6D0802E92F7230" type="Zenodo-Dep">203245</mods:identifier>
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<treatment id="513487E73669FF9EFF4BD198FD5B8257" ID-DOI="http://doi.org/10.5281/zenodo.5670826" ID-GBIF-Taxon="119388190" ID-Zenodo-Dep="5670826" LSID="urn:lsid:plazi:treatment:513487E73669FF9EFF4BD198FD5B8257" httpUri="http://treatment.plazi.org/id/513487E73669FF9EFF4BD198FD5B8257" lastPageNumber="9" pageId="8" pageNumber="9">
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<subSubSection id="9187657A3669FF9EFF4BD198FF62815B" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph id="D92236F13669FF9EFF4BD198FDD685E6" blockId="8.[151,639,151,177]" box="[151,639,151,177]" pageId="8" pageNumber="9">
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<heading id="826A819D3669FF9EFF4BD198FDD685E6" bold="true" box="[151,639,151,177]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
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<emphasis id="EBE9EAE33669FF9EFF4BD198FDD685E6" bold="true" box="[151,639,151,177]" pageId="8" pageNumber="9">
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Genus
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<taxonomicName id="1E9D4D723669FF9EFF32D198FDD685E6" ID-CoL="449J" authority="Montandon, 1897" authorityName="Montandon" authorityYear="1897" box="[238,639,151,177]" class="Insecta" family="Naucoridae" genus="Diaphorocoris" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="1" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="EBE9EAE33669FF9EFF32D198FE3585E6" bold="true" box="[238,412,151,177]" italics="true" pageId="8" pageNumber="9">Diaphorocoris</emphasis>
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Montandon, 1897
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="D92236F13669FF9EFF4BD1D4FF62815B" blockId="8.[151,1437,217,1793]" pageId="8" pageNumber="9">
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<emphasis id="EBE9EAE33669FF9EFF4BD1D4FE4285A3" bold="true" box="[151,491,219,244]" pageId="8" pageNumber="9">Supplemental redescription.</emphasis>
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Head angled ventrad ca. 30o with respect to pronotum; compound eyes divergent, mesal margins concave, lateral margins convex in posterior half, concave in anterior half, convergent anteriorly; lateral hyperoche widening anteriorly; apex of head deflexed, oriented posteroventrally; anteroventrally impunctate, with line of elongate setae between eye and base of labrum; rostrum short, segment three (first visible anteriorly) partially concealed behind well developed labrum, segment four narrower, reaching to near middle of prothoracic coxae; maxillary plates small, with brush of long hairs distally; antennal segments pale, one and two short and inconspicuous, three widest in distal 1–2; antennae set behind strong proepisternal flange and brushes of long setae ventrally on head and proepimeron. Pronotum with lateral margins smooth, convex; posterolateral corners rounded. Hemelytra with well developed embolium and clavus, short clavate hairs in linear series, costal fracture beyond embolium. Embolium with marginal row of spines, yellowish anterior band gradually narrowing posteriorly and becoming obsolete before costal fracture, with notch-like mesal incursion of white or light yellow into dark brown at level of scutellar apex. Prosternal midline with wedge-shaped carina between coxae, lateral surface of carina yellow. Forelegs with femur broad; tibia narrow; tarsus two-segmented in male, one-segmented in female; paired, articulated claws in both sexes. Middle and hind coxae with ventromesal series of stout hairs. Middle and hind legs with femora flattened, slightly curved dorsad, with row of spines along anteroventral margin and stronger spines along posteroventral margin; tibiae with numerous stout spines. Middle tibia gradually widening apically; male with dense pad of hairs beginning at proximal 1/4 of ventral surface and widening distally, pad reduced on female; scattered weak posterior spines, multiple rows of heavy anterior spines, weak fringe of posterodorsal swimming hairs continuing onto tarsi.
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<collectingCountry id="A18A76613669FF9EFD4ED26BFD63862B" box="[658,714,868,892]" name="India" pageId="8" pageNumber="9">Hind</collectingCountry>
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leg with tibia elongate, scattered posterior spines, multiple rows of anterior spines, fringe of posterodorsal swimming hairs on tibia continuing onto tarsi; tarsomere one produced beneath base of tarsomere two. Mid- and hind pretarsi with claws elongate, evenly curved, symmetrical, each with small basal tooth. Hindwing with well-developed veins C, R, M, Cu, m-Cu, pCu, pCu-A, 1A, 2A; cells R, M; anal fold.
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</paragraph>
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</subSubSection>
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<subSubSection id="9187657A3669FF9EFF1BD518FAD08023" pageId="8" pageNumber="9" type="diagnosis">
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<paragraph id="D92236F13669FF9EFF1BD518FAD08023" blockId="8.[151,1437,217,1793]" pageId="8" pageNumber="9">
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<emphasis id="EBE9EAE33669FF9EFF1BD518FEE88167" bold="true" box="[199,321,1047,1072]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
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These naucorids are elongate and flattened, with eyes elevated above the level of the pronotum, and with a wide, triangular lateral hyperoche of the head. They have most of the features common to members of
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<taxonomicName id="1E9D4D723669FF9EFAB5D532FEAE812F" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="subFamily" subFamily="Laccocorinae">Laccocorinae</taxonomicName>
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, including the front of the head being folded posteroventrally; the labrum is well developed, transverse, and rounded apically; and the dense midventral abdominal band of hairs, which is present on only three other genera. The middle tibia has a sexually dimorphic pad of hairs that widens distally, which is well developed on the male and reduced on the female. The front tarsi are two-segmented in males, unlike
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<taxonomicName id="1E9D4D723669FF9EFB9CD5C3FB7481B3" box="[1088,1245,1228,1252]" class="Insecta" family="Naucoridae" genus="Namtokocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="EBE9EAE33669FF9EFB9CD5C3FB7481B3" box="[1088,1245,1228,1252]" italics="true" pageId="8" pageNumber="9">Namtokocoris</emphasis>
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</taxonomicName>
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and
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<emphasis id="EBE9EAE33669FF9EFACFD5C2FF66805F" italics="true" pageId="8" pageNumber="9">
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<taxonomicName id="1E9D4D723669FF9EFACFD5C2FF65805F" class="Insecta" family="Naucoridae" genus="Pogonocaudina" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">Pogonocaudina</taxonomicName>
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,
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</emphasis>
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which have single segmented front tarsi in males. The front pretarsal claws are paired in both sexes, unlike
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<taxonomicName id="1E9D4D723669FF9EFF4BD41BFE9D807B" box="[151,308,1300,1324]" class="Insecta" family="Naucoridae" genus="Namtokocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="EBE9EAE33669FF9EFF4BD41BFE9D807B" box="[151,308,1300,1324]" italics="true" pageId="8" pageNumber="9">Namtokocoris</emphasis>
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</taxonomicName>
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, which has a single, unmovable claw. The scutellum is very large, elevated, and with lateral tumescences. The hemelytra have linear series of short, stout hairs, a feature which has been overlooked by each previous taxonomic treatment of the genus, including by
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<bibRefCitation id="BD0C4B003669FF9EFD73D453FCFC8023" author="Distant" box="[687,853,1372,1396]" pageId="8" pageNumber="17" refString="Distant, W. L. (1906) The fauna of British India, including Ceylon and Burma. Rhynchota. - Vol. III. (Heteroptera-Homoptera). Taylor & Francis, London, 503 pp." type="book" year="1906">Distant (1906)</bibRefCitation>
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. This attribute is in common with
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<taxonomicName id="1E9D4D723669FF9EFB09D453FADB8023" box="[1237,1394,1372,1396]" class="Insecta" family="Naucoridae" genus="Namtokocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="EBE9EAE33669FF9EFB09D453FADB8023" box="[1237,1394,1372,1396]" italics="true" pageId="8" pageNumber="9">Namtokocoris</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="9187657A3669FF9EFF1BD470FD5B8257" pageId="8" pageNumber="9" type="discussion">
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<paragraph id="D92236F13669FF9EFF1BD470FC15831B" blockId="8.[151,1437,217,1793]" pageId="8" pageNumber="9">
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<emphasis id="EBE9EAE33669FF9EFF1BD470FEE580CF" bold="true" box="[199,332,1407,1432]" pageId="8" pageNumber="9">Discussion.</emphasis>
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Montandon erected this genus to contain his new species
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<taxonomicName id="1E9D4D723669FF9EFC08D48EFBE180CF" box="[980,1096,1409,1432]" class="Insecta" family="Naucoridae" genus="Diaphorocoris" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="notatus">
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<emphasis id="EBE9EAE33669FF9EFC08D48EFBE180CF" box="[980,1096,1409,1432]" italics="true" pageId="8" pageNumber="9">D. notatus</emphasis>
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</taxonomicName>
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in 1897. However, later in the same year, after examining specimens from
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<collectingCountry id="A18A76613669FF9EFD52D4ABFD5480EB" box="[654,765,1444,1468]" name="Sri Lanka" pageId="8" pageNumber="9">Sri Lanka</collectingCountry>
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given to him by Kirkaldy, he realized that this species had already been described by
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<bibRefCitation id="BD0C4B003669FF9EFE16D4C7FDF680B7" author="Kirby" box="[458,607,1480,1504]" pageId="8" pageNumber="17" refString="Kirby, W. F. (1891) Catalogue of the described Hemiptera: Heteroptera and Homoptera of Ceylon, based on the collections formed (chiefly at Pundaloya) by Mr. E. Ernest Green. Journal of the Linnean Society, Zoology, 24, 72 - 176." type="journal article" year="1891">Kirby (1891)</bibRefCitation>
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under the genus
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<taxonomicName id="1E9D4D723669FF9EFCF4D4C6FC2680B7" box="[808,911,1481,1504]" class="Insecta" family="Naucoridae" genus="Naucoris" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="EBE9EAE33669FF9EFCF4D4C6FC2680B7" box="[808,911,1481,1504]" italics="true" pageId="8" pageNumber="9">Naucoris</emphasis>
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</taxonomicName>
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as
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<taxonomicName id="1E9D4D723669FF9EFC64D4C6FACA80B7" authority="(Montandon 1897b)" baseAuthorityName="Montandon" baseAuthorityYear="1897" box="[952,1379,1480,1504]" class="Insecta" family="Naucoridae" genus="Naucoris" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="punctatissima">
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<emphasis id="EBE9EAE33669FF9EFC64D4C6FBDE80B7" box="[952,1143,1481,1504]" italics="true" pageId="8" pageNumber="9">N. punctatissima</emphasis>
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(
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<bibRefCitation id="BD0C4B003669FF9EFB5BD4C7FAF380B7" author="Montandon" box="[1159,1370,1480,1504]" pageId="8" pageNumber="17" refString="Montandon, A. L. (1897 b) Hemiptera Cryptocerata. Fam. Naucoridae. - Sous fam. Laccocorinae. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 47, 435 - 454." type="journal article" year="1897" yearSuffix="b">Montandon 1897b</bibRefCitation>
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)
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</taxonomicName>
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and, thus, synonymized
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<taxonomicName id="1E9D4D723669FF9EFEA9D4E2FE448353" box="[373,493,1517,1540]" class="Insecta" family="Naucoridae" genus="Diaphorocoris" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="notatus">
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<emphasis id="EBE9EAE33669FF9EFEA9D4E2FE448353" box="[373,493,1517,1540]" italics="true" pageId="8" pageNumber="9">D. notatus</emphasis>
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</taxonomicName>
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as
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<taxonomicName id="1E9D4D723669FF9EFDCBD4E2FD7E8353" box="[535,727,1517,1540]" class="Insecta" family="Naucoridae" genus="Naucoris" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="punctatissima">
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<emphasis id="EBE9EAE33669FF9EFDCBD4E2FD7E8353" box="[535,727,1517,1540]" italics="true" pageId="8" pageNumber="9">D. punctatissima</emphasis>
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</taxonomicName>
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. When Kirby described his species, he recognized the uncertainty of this generic placement and commented that this species was narrower than other species of
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<taxonomicName id="1E9D4D723669FF9EFAD9D71EFAC5837F" box="[1285,1388,1553,1576]" class="Insecta" family="Naucoridae" genus="Naucoris" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="EBE9EAE33669FF9EFAD9D71EFAC5837F" box="[1285,1388,1553,1576]" italics="true" pageId="8" pageNumber="9">Naucoris</emphasis>
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</taxonomicName>
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and even placed a question mark after the genus in the new species heading.
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</paragraph>
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<paragraph id="D92236F13669FF9EFF1BD757FD5B8257" blockId="8.[151,1437,217,1793]" pageId="8" pageNumber="9">
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Little was known about the biology or ecology of this genus. In the original description of the
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<typeStatus id="062688533669FF9EFAD7D756FA928326" box="[1291,1339,1625,1649]" pageId="8" pageNumber="9">type</typeStatus>
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species,
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<bibRefCitation id="BD0C4B003669FF9EFF4BD773FE8083C3" author="Kirby" box="[151,297,1660,1684]" pageId="8" pageNumber="17" refString="Kirby, W. F. (1891) Catalogue of the described Hemiptera: Heteroptera and Homoptera of Ceylon, based on the collections formed (chiefly at Pundaloya) by Mr. E. Ernest Green. Journal of the Linnean Society, Zoology, 24, 72 - 176." type="journal article" year="1891">Kirby (1891)</bibRefCitation>
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indicated that the
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<typeStatus id="062688533669FF9EFE2AD772FD8F83C2" box="[502,550,1661,1685]" pageId="8" pageNumber="9">type</typeStatus>
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species was found on "wet rocks by running water." In a treatment of the fauna of
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<collectingCountry id="A18A76613669FF9EFF6BD7AFFEBA83EF" box="[183,275,1696,1720]" name="Malaysia" pageId="8" pageNumber="9">Malesia</collectingCountry>
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, this genus was listed as occurring in a quiet pool with silt in a disturbed forest in northern
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<collectingCountry id="A18A76613669FF9EFAE5D7AFFA3583EF" box="[1337,1436,1696,1720]" name="Thailand" pageId="8" pageNumber="9">Thailand</collectingCountry>
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(
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<bibRefCitation id="BD0C4B003669FF9EFF42D7CBFECE838B" author="Chen" box="[158,359,1732,1756]" pageId="8" pageNumber="17" refString="Chen, P. - p., Nieser, N. & Zettel, H. (2005) The aquatic and semi-aquatic bugs Heteroptera: Nepomorpha & Gerromorpha of Malesia. Brill, Leiden, 546 p." type="book" year="2005">Chen et al. 2005</bibRefCitation>
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); however, this actually was in reference to
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<taxonomicName id="1E9D4D723669FF9EFCB3D7CBFBA5838B" box="[879,1036,1732,1756]" class="Insecta" family="Naucoridae" genus="Namtokocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="EBE9EAE33669FF9EFCB3D7CBFBA5838B" box="[879,1036,1732,1756]" italics="true" pageId="8" pageNumber="9">Namtokocoris</emphasis>
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</taxonomicName>
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(Nieser, pers. comm.), which was described two years later (Sites & Vitheepradit 2007).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |