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116 lines
15 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.179.21253" ID-GBIF-Dataset="144aa3b4-ee51-4c26-8436-aaa65ae3b22b" ID-PMC="PMC5740450" ID-Pensoft-Pub="1313-2970-179-75" ID-PubMed="29290697" ID-ZBK="36C4E2C8E07D4CC9A1D696B0FCE92CCF" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-179-75" ModsDocOrigin="ZooKeys 179" ModsDocTitle="A taxonomic revision of the subfamily Tillinae Leach sensu lato (Coleoptera, Cleridae) in the New World" checkinTime="1512688543562" checkinUser="pensoft" docAuthor="Burke, Alan & Zolnerowich, Gregory" docDate="2017" docId="28FEF2CA9BC990B3AF3B495943554F94" docLanguage="en" docName="ZooKeys 719: 75-157" docOrigin="ZooKeys 179" docSource="http://dx.doi.org/10.3897/zookeys.179.21253" docTitle="Monophylla pallipes Schaeffer 1908" docType="treatment" docVersion="5" lastPageNumber="131" masterDocId="8A60FFA894215C48ED67FF86FF8F3B5B" masterDocTitle="A taxonomic revision of the subfamily Tillinae Leach sensu lato (Coleoptera, Cleridae) in the New World" masterLastPageNumber="157" masterPageNumber="75" pageNumber="129" updateTime="1668165163296" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A taxonomic revision of the subfamily Tillinae Leach sensu lato (Coleoptera, Cleridae) in the New World</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Burke, Alan</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zolnerowich, Gregory</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>179</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>75</mods:start>
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<mods:end>157</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.179.21253</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.179.21253</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-179-75</mods:identifier>
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<mods:identifier type="ZBK">36C4E2C8E07D4CC9A1D696B0FCE92CCF</mods:identifier>
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<mods:identifier type="ZooBank">36C4E2C8E07D4CC9A1D696B0FCE92CCF</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="137663090" LSID="urn:lsid:plazi:treatment:28FEF2CA9BC990B3AF3B495943554F94" httpUri="http://treatment.plazi.org/id/28FEF2CA9BC990B3AF3B495943554F94" lastPageId="56" lastPageNumber="131" pageId="54" pageNumber="129">
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<subSubSection pageId="54" pageNumber="129" type="nomenclature">
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<paragraph pageId="54" pageNumber="129">
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<taxonomicName authority="Schaeffer, 1908" authorityName="Schaeffer" authorityYear="1908" class="Insecta" family="Cleridae" genus="Monophylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monophylla pallipes" order="Coleoptera" pageId="54" pageNumber="129" phylum="Arthropoda" rank="species" species="pallipes">Monophylla pallipes Schaeffer, 1908</taxonomicName>
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Figs 4C, 20C
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</paragraph>
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</subSubSection>
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<subSubSection pageId="54" pageNumber="129" type="type material">
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<paragraph pageId="54" pageNumber="129">Type material not examined.</paragraph>
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</subSubSection>
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<subSubSection pageId="54" pageNumber="129" type="type locality">
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<paragraph pageId="54" pageNumber="129">Type locality.</paragraph>
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<paragraph pageId="54" pageNumber="129">Brownsville, Texas. Type depository: United States National History Museum (USNM).</paragraph>
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<paragraph pageId="54" pageNumber="129">
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Holotype lost. Lectotype designated by
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<bibRefCitation author="Chapin, EA" journalOrPublisher="Smithsonian Miscellaneous Collection" pageId="75" pageNumber="150" pagination="1 - 12" title="Type material of the species of clerid beetles described by Charles Schaeffer." volume="11" year="1949">Chapin (1949)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="54" pageNumber="129" type="distribution">
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<paragraph pageId="54" pageNumber="129">Distribution.</paragraph>
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<paragraph pageId="54" pageNumber="129">USA: AZ, CA, TX; Mexico: Chiapas, Jalisco, Morelos, Quintana Roo, San Luis Potosi, Sinaloa, Tamaulipas, Yucatan; Central America: Costa Rica, Guatemala, Honduras; South America: Chile (introduced).</paragraph>
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</subSubSection>
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<subSubSection pageId="54" pageNumber="129" type="differential diagnosis">
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<paragraph pageId="54" pageNumber="129">Differential diagnosis.</paragraph>
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<paragraph pageId="54" pageNumber="129">
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<taxonomicName class="Insecta" family="Cleridae" genus="Monophylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monophylla pallipes" order="Coleoptera" pageId="54" pageNumber="129" phylum="Arthropoda" rank="species" species="pallipes">Monophylla pallipes</taxonomicName>
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is very similar to
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<taxonomicName lsidName="M. californica" pageId="54" pageNumber="129" rank="species" species="californica">M. californica</taxonomicName>
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. The two species are sympatric in distribution; therefore, they can be misidentified when collected simultaneously. Diagnostic characters are provided in the diagnosis of
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<taxonomicName lsidName="M. californica" pageId="54" pageNumber="129" rank="species" species="californica">M. californica</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="55" lastPageNumber="130" pageId="54" pageNumber="129" type="redescription">
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<paragraph pageId="54" pageNumber="129">Redescription.</paragraph>
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<paragraph pageId="54" pageNumber="129">Male. Form: Small to moderately large, slender individuals. Color: Head, antennae, pronotum, scutellum and legs dark testaceous to almost piceous; the anterior and posterior margins of the pronotum have a narrow ferrugineous to testaceous band; thorax ferrugineous to almost black; elytra testaceous to piceous, each elytron may have a pale to yellowish fascia on the median region of the elytral disc that initiates on the epipleural fold and does not reach the elytral suture; mouthparts fuscous, abdominal segments light testaceous to fuscous (Fig. 4C).</paragraph>
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<paragraph pageId="54" pageNumber="129">Head: Including eyes narrower than pronotum; eyes small, taller than wide, feebly bulging laterally, antennal notch in front of eye emargination; integument coarsely to slightly punctate; clothed with fine, pale, semirecumbent and semi-erect setae intermixed with some scattered erect, pale setae; antennae consisting of 10 antennomeres; second antennomeres robust, rather short, antennomeres 3-4 small, conspicuously compacted; antennomeres 5-9 serrate, serration and size gradually increasing toward distal end, last antennomeres noticeably enlarged, conspicuously compressed laterally, much longer than remaining antennomeres combined (Fig. 4C); last antennomere sexually dimorphic.</paragraph>
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<paragraph pageId="54" pageNumber="129">Thorax: Pronotum subparallel, widest at middle, constricted toward posterior margin; pronotal surface moderately to conspicuously punctate, rugose to rugulose; pronotal disc flat; clothed with fine, short, pale and dark, semirecumbent setae interspersed with some long and very long, erect, dark setae; anterior transverse depression feebly impressed, subbasal tumescence absent. Prosternum as long as wide; strongly punctate; punctation fine, shallow; surface feebly clothed. Mesoventrite: Surface moderately to coarsely punctate; punctations wide, deep; vested with fine, pale, semi-erect setae. Metepisterna visible throughout their length, not covered by elytra. Metaventrite: Convex; integument punctate laterally; punctation wide and shallow; moderately clothed with fine, pale, recumbent setae; longitudinal depression present, metaventral process absent. Scutellum wide, clothed with pale, fine, recumbent setae, compressed medially.</paragraph>
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<paragraph pageId="54" pageNumber="129">Elytra: Anterior margin slightly broader than pronotum; elongate; subparallel, humeri inconspicuously indicated, rounded; sides gradually expanding toward distal end, widest on middle third then narrowing toward apex; elytral apices subtriangular; inconspicuously dehiscent; elytral declivity gradual; surface clothed with fine, short, pale and dark, semi-erect and erect setae; conspicuously punctate, punctations small and shallow, irregularly arranged, punctations reaching the elytral apex; interstices smooth, narrow. Epipleural fold gradually narrowing toward apex.</paragraph>
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<paragraph pageId="55" pageNumber="130">
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<pageBreakToken pageId="55" pageNumber="130" start="start">Legs</pageBreakToken>
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: Femora rugulose to smooth; expanded behind middle; laterally compressed; clothed with some pale, fine, semi erect setae; surface feebly, shallowly punctate. Tibiae rugulose, slender, puncticulate; vestiture consisting of fine, pale, semi-erect setae mingled with some pale, semirecumbent setae.
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</paragraph>
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<paragraph pageId="55" pageNumber="130">Abdomen: Six visible ventrites. Ventrites 1-4 convex, smooth, shiny. First visible ventrite longer than second visible ventrite, feebly rugulose; ventrites 2-4 subquadrate; punctate; vested with fine, long, pale, recumbent setae; not compressed laterally; posterior margins truncate. Fifth visible ventrite convex; subquadrate; surface rugulose; puncticulate; vested with fine, pale, recumbent setae; lateral margins subparallel, arcuate; posterior margin broadly, shallowly emarginate. Sixth visible ventrite subquadrate; surface slightly to moderately rugulose, convex to almost flat; feebly compressed on the median-posterior region; clothed with some long, erect setae, vestiture more abundant on anterolateral margins; lateral margins oblique, slightly arcuate; posterior margin broadly, deeply emarginate, U-shaped emarginate, posterolateral angles round. Fifth tergite subquadrate, surface convex; posterior margin truncate. Sixth tergite subquadrate; finely rugulose; surface convex posterior median region compressed; clothed with long, fine, pale and dark recumbent setae; lateral margins oblique; posterior margin broadly, moderately deeply emarginate, U-shaped emargination, posterolateral angles rounded. Sixth tergite slightly extending beyond apical margin of sixth visible ventrite, covering sixth ventrite in dorsal view.</paragraph>
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<paragraph pageId="55" pageNumber="130">Aedeagus: Phallobasic apodeme present; phallus with copulatory piece swollen at apex; phallic plate armed with a row of denticles; intraspicular plate present, elongate; phallobasic apodeme conspicuously short, expanded distally; phallobase sinuate; parameres free; tegmen incomplete, partially covering the phallus; parameres pointed anteriorly; endophallic struts long, conspicuously robust distally (Fig. 20C).</paragraph>
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<paragraph pageId="55" pageNumber="130">
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Sexual dimorphism: Females of
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<taxonomicName lsidName="M. pallipes" pageId="55" pageNumber="130" rank="species" species="pallipes">M. pallipes</taxonomicName>
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differ from males based on the following characters: the tenth antennomere of females is shorter than in males; antennomeres 6-9 are larger and moderately serrate in females, but conspicuously reduced and compressed in males; and females have the last abdominal segment broadly rounded to feebly truncate, while males have this segment subquadrate in shape and emarginate posteriorly.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="56" lastPageNumber="131" pageId="55" pageNumber="130" type="materials_examined">
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<paragraph pageId="55" pageNumber="130">Material examined.</paragraph>
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<paragraph lastPageId="56" lastPageNumber="131" pageId="55" pageNumber="130">
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2 males, 2 females: Hidalgo Co., TX, IV-7-1950, D. J. and J. N. Knull; 2 males, 6 females: Brownsville, TX, V-25-1934, D. J. and J. N. Knull; 2 males, 5 females: Brownsville, TX, V-14, D. J. and J. N. Knull; 1 female: Brewster Co., TX, V-26-1948; 1 female: Cameron Co., TX, VI-4-1950, D. J. and J. N. Knull; 2 females: Corpus Christy, TX, III-30-1961, D. J. and J. N. Knull; l male: Gillespie Co., TX, IV-23, D. J. and J. N. Knull; 1 male, 2 females: Hidalgo Co., TX, III-20-1952, D. J. and J. N. Knull; 1 male, 1 female: Hidalgo Co., TX, III-29-1963, D. J. and J. N. Knull; 1 male: Hidalgo Co., TX, III-26-1953, D. J. and J. N. Knull; 1 male, 1 female: Starr Co., TX, IV-9-1963, D. J. and J. N. Knull; 2 males: Starr Co., TX, III-20-1952, D. J. and J. N. Knull; 1 female: Hidalgo Co., TX, 7-IV-1950, D. J. and J. N. Knull; 1 male, 1 female: Hidalgo Co., TX, V-23-1953, D. J. and J. N. Knull; 2 males: Jackson Co., TX, V-22, D. J. and J. N. Knull; 2 males, 1 female: Brownsville, TX, V-19, D. J. and J. N. Knull; 3 males, 2 females: Uvalde Co., TX, VI-13-1949, J.
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<pageBreakToken pageId="56" pageNumber="131" start="start">N</pageBreakToken>
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. Knull; 1 female: Santa Cruz Co., CA, Glenwood road, VI-16-1968, W. H. Tyson; 1 female: Gillespie Co., TX, VI-1, J. N. Knull; 1 male: Brownsville, TX, V-12, D. J. and J. N. Knull; 1 male: Brownsville, TX, V-5, D. J. and J. N. Knull; 1 female: Brownsville, TX, XI-19-1911, in pasture, Garden; 1 female: Cameron Co., TX, 2 mi E Los Indios, V-13-1978, N. M. Downie; 1 female: Cameron Co., TX, Sabal Palm Grove, IV-20-30-1986, D. H. Heffern; 2 males, 3 females: San Patricio Co., TX, Welder Wildlife Ref., VII-10-20-1981, R. H. Turnbow; 3 males: Cameron Co., TX, Sabal Palm Grove Sanctuary, III-16-1981, R. H. Turnbow; 2 males: Cameron Co., TX, Palm Groove Sanctuary, Brownsville, I-1977, F. T. Hovore; 1 female: Hidalgo Co., TX, Santa Ana National Refugee vic., Willow Lake, T. C. MacRae; 1 male: Starr Co., TX, Rio Grande City, X-10-1972, E. Giesbert; 1 female: San Patricio Co., TX, Welder Wildlife Refuge, V-10-12-1977, E. Giesbert; 1 male: TX, reared from mesquite logs, emerged X-10-1955, H. F. Howden; 1 male, 1 female: TX, Lake Corpus Christy State Park, VI-19-1971; G. H. Nelson; 1 male: Cameron Co., TX, Sabal Palm Grove Sanctuary, III-20-1982, R. Turnbow. MEXICO: 1 male: Sinaloa, Mexico, 3 km E El Marmol, VIII-8-1983, E. Giesbert; 1 male, 2 females: San Luis
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<normalizedToken originalValue="Potosí">Potosi</normalizedToken>
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, Mexico, 69.5 km N Tamazunchale, VI-5-1987, R. H. Turnbow; 2 males, 3 females: Jalisco, Mexico, 1.2 km S of La Cumbre, VII-19-2011, R. H. Turnbow; 2 males, 1 female: Tamaulipas, Mexico, 1-2 mi E Nuevo Morelos, VI-2-1982, R. H. Turnbow; 1 male: Guerrero, Mexico, 7.3 km NW Ixtapa, VII-17-1985, R. Turnbow; 2 males, 2 females: Quintana Roo, Mexico, highway 186, 17 km S jct. 307, V-30-1984, R. Turnbow; 1 female: Yucatan, Mexico, 2 km E Chichen Itza, V-26-1984, R. Turnbow; 1 female: Chiapas, Mexico, 4 mi NW of Pueblo Nuevo River Bajada, VII-15-1965, G. H. Nelson. CENTRAL AMERICA: 1 female: El Paraiso, Honduras, 31.5 km W Danli, VII-20-1995, R. Turnbow.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="56" pageNumber="131" type="remarks">
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<paragraph pageId="56" pageNumber="131">Remarks.</paragraph>
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<paragraph pageId="56" pageNumber="131">
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The holotype of
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<taxonomicName lsidName="M. pallipes" pageId="56" pageNumber="131" rank="species" species="pallipes">M. pallipes</taxonomicName>
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was lost and a lectotype was designated by
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<bibRefCitation author="Chapin, EA" journalOrPublisher="Smithsonian Miscellaneous Collection" pageId="75" pageNumber="150" pagination="1 - 12" title="Type material of the species of clerid beetles described by Charles Schaeffer." volume="11" year="1949">Chapin (1949)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |