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<document ID-DOI="http://dx.doi.org/10.3897/subtbiol.41.75613" ID-GBIF-Dataset="d409aeb2-e394-4143-a84d-77ce363a8b73" ID-GBIF-Taxon="191263880" ID-Pensoft-Pub="1314-2615-41-69" ID-Pensoft-UUID="5992EA8732135F93917415967CC3DDE0" ID-ZooBank="917780D7509B43D7B6499D1D4FF0EC54" ModsDocID="1314-2615-41-69" checkinTime="1639196403999" checkinUser="pensoft" docAuthor="Curcic 1, Srecko, Vesovic 1, Nikola, Vrbica 1, Maja, Popovic 1,2, Slađana, Radovanovic 3, Zeljko, Curcic 4, Nina B. &amp; Rađa 5, Tonci" docDate="2021" docId="075A8FD8A3B45743840D9C7DBB0D8B38" docLanguage="en" docName="SubterBiol 41: 69-85" docOrigin="Subterranean Biology 41" docPubDate="2021-12-10" docSource="http://dx.doi.org/10.3897/subtbiol.41.75613" docTitle="Leonhardia solaki Curcic, Rađa, Vesovic &amp; Vrbica 2021, sp. nov." docType="treatment" docVersion="3" id="5992EA8732135F93917415967CC3DDE0" lastPageNumber="69" masterDocId="5992EA8732135F93917415967CC3DDE0" masterDocTitle="A new species of Leonhardia Reitter, 1901 (Coleoptera, Leiodidae, Leptodirini) from Bosnia and Herzegovina, with a key to species of the genus" masterLastPageNumber="85" masterPageNumber="69" pageId="0" pageNumber="69" updateTime="1639202123423" updateUser="ExternalLinkService">
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<mods:title>A new species of Leonhardia Reitter, 1901 (Coleoptera, Leiodidae, Leptodirini) from Bosnia and Herzegovina, with a key to species of the genus</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Curcic 1, Srecko</mods:namePart>
<mods:affiliation>University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vesovic 1, Nikola</mods:namePart>
<mods:affiliation>University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vrbica 1, Maja</mods:namePart>
<mods:affiliation>University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Popovic 1,2, Slađana</mods:namePart>
<mods:affiliation>University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Radovanovic 3, Zeljko</mods:namePart>
<mods:affiliation>University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Curcic 4, Nina B.</mods:namePart>
<mods:affiliation>University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rađa 5, Tonci</mods:namePart>
<mods:affiliation>University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia</mods:affiliation>
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<mods:title>Subterranean Biology</mods:title>
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<mods:date>2021</mods:date>
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<mods:number>2021-12-10</mods:number>
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<mods:number>41</mods:number>
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<mods:start>69</mods:start>
<mods:end>85</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/subtbiol.41.75613</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/subtbiol.41.75613</mods:identifier>
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<treatment ID-GBIF-Taxon="191263880" LSID="urn:lsid:plazi:treatment:075A8FD8A3B45743840D9C7DBB0D8B38" httpUri="http://treatment.plazi.org/id/075A8FD8A3B45743840D9C7DBB0D8B38" lastPageNumber="69" pageId="0" pageNumber="69">
<subSubSection pageId="0" pageNumber="69" type="nomenclature">
<paragraph pageId="0" pageNumber="69">
<taxonomicName LSID="075A8FD8-A3B4-5743-840D-9C7DBB0D8B38" authority="Ćurčić, Rađa, Vesović &amp; Vrbica" authorityName="Curcic, Rađa, Vesovic &amp; Vrbica" authorityYear="2021" class="Insecta" family="Leiodidae" genus="Leonhardia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leonhardia solaki" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="solaki" status="sp. nov.">
Leonhardia solaki
<normalizedToken originalValue="Ćurčić">Curcic</normalizedToken>
,
<normalizedToken originalValue="Rađa">Rađa</normalizedToken>
,
<normalizedToken originalValue="Vesović">Vesovic</normalizedToken>
&amp; Vrbica
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="69">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="69" type="description">
<paragraph pageId="0" pageNumber="69">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">http://zoobank.org/32557822-9245-4CE3-9E5B-3E8ABF5DEF12 Figures 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)." figureDoi="10.3897/subtbiol.41.75613.figure3" httpUri="https://binary.pensoft.net/fig/621564" pageId="0" pageNumber="69">, 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="69" type="type material">
<paragraph pageId="0" pageNumber="69">Type material.</paragraph>
<paragraph pageId="0" pageNumber="69">
<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Holotype</emphasis>
: male (IZFB) labeled as follows: &quot;WESTERN BOSNIA AND HERZEGOVINA: town of
<normalizedToken originalValue="Glamoč">Glamoc</normalizedToken>
, village of Skucani,
<normalizedToken originalValue="Golubnjača">Golubnjaca</normalizedToken>
kod Skucana Pit, 1,005 m a.s.l.,
<geoCoordinate degrees="43" direction="north" minutes="58" orientation="latitude" precision="1" seconds="31.9" value="43.97553">43°58'31.9&quot;N</geoCoordinate>
, 16°54'43.6&quot; E, June, 27, 2018,
<normalizedToken originalValue="Tonći">Tonci</normalizedToken>
<normalizedToken originalValue="Rađa“">Rađa&quot;</normalizedToken>
(white label, printed)/&quot;Holotypus
<taxonomicName authorityName="Curcic, Rađa, Vesovic &amp; Vrbica" authorityYear="2021" class="Insecta" family="Leiodidae" genus="Leonhardia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leonhardia solaki" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="solaki">
<emphasis italics="true" pageId="0" pageNumber="69">Leonhardia solaki</emphasis>
</taxonomicName>
sp. nov.
<normalizedToken originalValue="Ćurčić">Curcic</normalizedToken>
,
<normalizedToken originalValue="Rađa">Rađa</normalizedToken>
,
<normalizedToken originalValue="Vesović">Vesovic</normalizedToken>
&amp; Vrbica det. 2021&quot; (red label, printed) (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2</figureCitation>
).
</paragraph>
<caption doi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="69">
<emphasis bold="true" pageId="0" pageNumber="69">Figure 2.</emphasis>
Bright-field images of morphological features of
<taxonomicName authorityName="Curcic, Rađa, Vesovic &amp; Vrbica" authorityYear="2021" class="Insecta" family="Leiodidae" genus="Leonhardia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leonhardia solaki" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="solaki">
<emphasis italics="true" pageId="0" pageNumber="69">Leonhardia solaki</emphasis>
</taxonomicName>
sp. nov. from the
<normalizedToken originalValue="Golubnjača">Golubnjaca</normalizedToken>
kod Skucana Pit, village of Skucani, close to the town of
<normalizedToken originalValue="Glamoč">Glamoc</normalizedToken>
, western Bosnia and Herzegovina
<emphasis bold="true" pageId="0" pageNumber="69">A</emphasis>
holotype male, habitus, dorsal aspect
<emphasis bold="true" pageId="0" pageNumber="69">B</emphasis>
holotype male, mesosternal carina, lateral aspect
<emphasis bold="true" pageId="0" pageNumber="69">C</emphasis>
holotype male, aedeagus, dorsal aspect
<emphasis bold="true" pageId="0" pageNumber="69">D</emphasis>
holotype male, aedeagus, lateral aspect
<emphasis bold="true" pageId="0" pageNumber="69">E</emphasis>
holotype male, left paramere apex, dorsal aspect
<emphasis bold="true" pageId="0" pageNumber="69">F</emphasis>
paratype female, left gonostylus, dorsal aspect
<emphasis bold="true" pageId="0" pageNumber="69">G</emphasis>
paratype female, spermatheca, lateral aspect
<emphasis bold="true" pageId="0" pageNumber="69">H</emphasis>
paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (
<emphasis bold="true" pageId="0" pageNumber="69">A</emphasis>
); 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="69">B-D, H</emphasis>
); 0.1 mm (
<emphasis bold="true" pageId="0" pageNumber="69">F, G</emphasis>
); 0.05 mm (
<emphasis bold="true" pageId="0" pageNumber="69">E</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="69">
<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Paratypes</emphasis>
(seven specimens). Same data as for holotype [two males and two females, IZFB, SSM]; same data as for holotype except for date [one male and two females, IZFB, June, 12, 2018]. All paratypes are labeled with white, printed locality labels and with red printed labels &quot;Paratypus
<taxonomicName authorityName="Curcic, Rađa, Vesovic &amp; Vrbica" authorityYear="2021" class="Insecta" family="Leiodidae" genus="Leonhardia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leonhardia solaki" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="solaki">
<emphasis italics="true" pageId="0" pageNumber="69">Leonhardia solaki</emphasis>
</taxonomicName>
sp. nov.
<normalizedToken originalValue="Ćurčić">Curcic</normalizedToken>
,
<normalizedToken originalValue="Rađa">Rađa</normalizedToken>
,
<normalizedToken originalValue="Vesović">Vesovic</normalizedToken>
&amp; Vrbica det. 2021&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="69" type="etymology">
<paragraph pageId="0" pageNumber="69">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="69">
The new species is named after
<normalizedToken originalValue="Siniša">Sinisa</normalizedToken>
<normalizedToken originalValue="Šolak">Solak</normalizedToken>
, a geographer and naturalist, who was our guide during cave investigations conducted near the town of
<normalizedToken originalValue="Glamoč">Glamoc</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="69" type="type locality">
<paragraph pageId="0" pageNumber="69">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="69">
Western Bosnia and Herzegovina, town of
<normalizedToken originalValue="Glamoč">Glamoc</normalizedToken>
, village of Skucani,
<normalizedToken originalValue="Golubnjača">Golubnjaca</normalizedToken>
kod Skucana Pit.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="69" type="diagnosis">
<paragraph pageId="0" pageNumber="69">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="69">
The new species is closely related to some other species of
<taxonomicName class="Insecta" family="Leiodidae" genus="Leonhardia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leonhardia" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="69">Leonhardia</emphasis>
</taxonomicName>
by its large size and by the presence of a high mesosternal carina. Those other species are
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. hilfi" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="hilfi">
<emphasis italics="true" pageId="0" pageNumber="69">L. hilfi</emphasis>
</taxonomicName>
,
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. reitteri" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="reitteri">
<emphasis italics="true" pageId="0" pageNumber="69">L. reitteri</emphasis>
</taxonomicName>
,
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. delminiumica" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="delminiumica">
<emphasis italics="true" pageId="0" pageNumber="69">L. delminiumica</emphasis>
</taxonomicName>
,
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. jajcensis" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="jajcensis">
<emphasis italics="true" pageId="0" pageNumber="69">L. jajcensis</emphasis>
</taxonomicName>
, and
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. sebesicensis" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="sebesicensis">
<emphasis italics="true" pageId="0" pageNumber="69">L. sebesicensis</emphasis>
</taxonomicName>
(Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map of Bosnia and Herzegovina showing locality records for all taxa of Leonhardia. The grey pattern indicates karst terrain L. reitteri reitteri (blue circles), L. reitteri mersa (blue cross), L. reitteri zariquieyi (blue rhombuses), L. delminiumica (green squares), L. jajcensis (lime deltoid), L. hilfi hilfi (orange star), L. hilfi robusta (orange sun), L. sebesicensis (pink pentagon), L. droveniki (red hexagon), L. solaki sp. nov. (yellow triangle). Scale bar: 50 km." figureDoi="10.3897/subtbiol.41.75613.figure1" httpUri="https://binary.pensoft.net/fig/621562" pageId="0" pageNumber="69">1</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)." figureDoi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69">4</figureCitation>
) (
<bibRefCitation author="Jeannel, R" journalOrPublisher="Archives de Zoologie experimentale et generale" pageId="0" pageNumber="69" pagination="1 - 436" refId="B14" refString="Jeannel, R, 1924. Monographie des Bathysciinae. Biospeologica L. Archives de Zoologie experimentale et generale 63: 1 - 436" title="Monographie des Bathysciinae. Biospeologica L." volume="63" year="1924">Jeannel 1924</bibRefCitation>
;
<bibRefCitation author="Nonveiller, G" journalOrPublisher="Revue francaise d'Entomologie, Nouvelle serie" pageId="0" pageNumber="69" pagination="165 - 168" refId="B19" refString="Nonveiller, G, Pavicevic, D, Rada, T, Vujcic-Karlo, S, 2002. A new species of the genus Leonhardia Reitter, 1901 from Herzegovina [Coleoptera, Leiodidae, Cholevinae, Leptodirini]. Revue francaise d'Entomologie, Nouvelle serie 24: 165 - 168" title="A new species of the genus Leonhardia Reitter, 1901 from Herzegovina [Coleoptera, Leiodidae, Cholevinae, Leptodirini]." volume="24" year="2002">Nonveiller et al. 2002</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.2298/ABS1402919C" author="Curcic, S" journalOrPublisher="Archives of Biological Sciences, Belgrade" pageId="0" pageNumber="69" pagination="919 - 933" refId="B6" refString="Curcic, S, Rađa, T, Mulaomerovic, J, Vrbica, M, Antic, D, Curcic, B, Rađa, B, Vesovic, N, 2014. Three new cave-dwelling leiodid beetles (Coleoptera: Leiodidae: Cholevinae: Leptodirini) from Bosnia and Herzegovina. Archives of Biological Sciences, Belgrade 66: 919 - 933, DOI: https://doi.org/10.2298/ABS1402919C" title="Three new cave-dwelling leiodid beetles (Coleoptera: Leiodidae: Cholevinae: Leptodirini) from Bosnia and Herzegovina." url="https://doi.org/10.2298/ABS1402919C" volume="66" year="2014">
<normalizedToken originalValue="Ćurčić">Curcic</normalizedToken>
et al. 2014
</bibRefCitation>
,
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4483.3.6" author="Curcic, S" journalOrPublisher="Zootaxa" pageId="0" pageNumber="69" pagination="523 - 548" refId="B7" refString="Curcic, S, Pavicevic, D, Vesovic, N, Mulaomerovic, J, Rađa, T, Antic, D, Bosco, F, Markovic, Đ, Petkovic, M, 2018. Seven new taxa of Leptodirini (Coleoptera: Leiodidae: Cholevinae) from the Balkan Peninsula. Zootaxa 4483: 523 - 548, DOI: https://doi.org/10.11646/zootaxa.4483.3.6" title="Seven new taxa of Leptodirini (Coleoptera: Leiodidae: Cholevinae) from the Balkan Peninsula." url="https://doi.org/10.11646/zootaxa.4483.3.6" volume="4483" year="2018">2018</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="69">
<taxonomicName authorityName="Curcic, Rađa, Vesovic &amp; Vrbica" authorityYear="2021" class="Insecta" family="Leiodidae" genus="Leonhardia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leonhardia solaki" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="solaki">
<emphasis italics="true" pageId="0" pageNumber="69">Leonhardia solaki</emphasis>
</taxonomicName>
sp. nov. differs from
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. hilfi" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="hilfi">
<emphasis italics="true" pageId="0" pageNumber="69">L. hilfi</emphasis>
</taxonomicName>
in regard to AL in males (antennae exceeding the middle of elytra vs. antennae reaching the middle of elytra); A10L/A10W (less than 2.375 vs. more than 2.50); pronotum form (bell-shaped, lateral margins well-rounded anteriorly vs. subquadrate, lateral margins obtuse anteriorly); PL/PW (pronotum wider than long vs. pronotum as long as wide); shape of the mesosternal carina (higher, almost right-angled, posterior margin concave vs. lower, obtuse-angled, posterior margin somewhat elevated); shape of elytra (inversely ovate vs. widely oval); shape of the median lobe in dorsal view (apically flattened vs. apically narrowed); length of the median lobe (barely longer than parameres vs. markedly longer than parameres); shape and size of the basal bulb in dorsal view (larger, rounded vs. smaller, not rounded); and shape of the aedeagus in lateral view (median lobe more curved, basal bulb larger vs. median lobe less curved, basal bulb smaller) (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2</figureCitation>
-
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)." figureDoi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69">4</figureCitation>
) (
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.7311" author="Reitter, E" journalOrPublisher="Berichte des Naturwissenschaftlich-medizinischen Vereins in Innsbruck" pageId="0" pageNumber="69" refId="B28" refString="Reitter, E, 1901. Ein neuer blinder Grotten-Silphide aus der Herzegowina. Wiener entomologische Zeitung 20: 128. https://doi.org/10.5962/bhl.part.7311" title="Ein neuer blinder Grotten-Silphide aus der Herzegowina. Wiener entomologische Zeitung 20: 128." url="https://doi.org/10.5962/bhl.part.7311" year="1901">Reitter 1901</bibRefCitation>
;
<bibRefCitation author="Jeannel, R" journalOrPublisher="Archives de Zoologie experimentale et generale" pageId="0" pageNumber="69" pagination="1 - 436" refId="B14" refString="Jeannel, R, 1924. Monographie des Bathysciinae. Biospeologica L. Archives de Zoologie experimentale et generale 63: 1 - 436" title="Monographie des Bathysciinae. Biospeologica L." volume="63" year="1924">Jeannel 1924</bibRefCitation>
;
<bibRefCitation author="Knirsch, E" journalOrPublisher="Casopis Ceskoslovenske Spolecnosti Entomologicke" pageId="0" pageNumber="69" pagination="101 - 122" refId="B15" refString="Knirsch, E, 1928. Novi zastupci slepych Silphidu ze zapadniho Srbska. Casopis Ceskoslovenske Spolecnosti Entomologicke 24: 101 - 122" title="Novi zastupci slepych Silphidu ze zapadniho Srbska." volume="24" year="1928">Knirsch 1928</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.3406/bsef.1999.16602" author="Perreau, M" journalOrPublisher="Bulletin de la Societe entomologique de France" pageId="0" pageNumber="69" pagination="399 - 406" refId="B20" refString="Perreau, M, 1999. Nouveaux genres et nouvelles especes de Leptodirini (Coleoptera, Leiodidae). Bulletin de la Societe entomologique de France 104: 399 - 406, DOI: https://doi.org/10.3406/bsef.1999.16602" title="Nouveaux genres et nouvelles especes de Leptodirini (Coleoptera, Leiodidae)." url="https://doi.org/10.3406/bsef.1999.16602" volume="104" year="1999">Perreau 1999</bibRefCitation>
;
<bibRefCitation author="Nonveiller, G" journalOrPublisher="Revue francaise d'Entomologie, Nouvelle serie" pageId="0" pageNumber="69" pagination="165 - 168" refId="B19" refString="Nonveiller, G, Pavicevic, D, Rada, T, Vujcic-Karlo, S, 2002. A new species of the genus Leonhardia Reitter, 1901 from Herzegovina [Coleoptera, Leiodidae, Cholevinae, Leptodirini]. Revue francaise d'Entomologie, Nouvelle serie 24: 165 - 168" title="A new species of the genus Leonhardia Reitter, 1901 from Herzegovina [Coleoptera, Leiodidae, Cholevinae, Leptodirini]." volume="24" year="2002">Nonveiller et al. 2002</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="69">
<taxonomicName authorityName="Curcic, Rađa, Vesovic &amp; Vrbica" authorityYear="2021" class="Insecta" family="Leiodidae" genus="Leonhardia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leonhardia solaki" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="solaki">
<emphasis italics="true" pageId="0" pageNumber="69">Leonhardia solaki</emphasis>
</taxonomicName>
sp. nov. is easily distinguished from
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. reitteri" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="reitteri">
<emphasis italics="true" pageId="0" pageNumber="69">L. reitteri</emphasis>
</taxonomicName>
, from which it differs with respect to AL (antennae exceeding the middle of elytra in males and reaching the middle of elytra in females vs. antennae barely reaching the middle of body); A2L/A3L+A4L (antennomere II shorter than the following two antennomeres combined vs. antennomere II as long as the following two antennomeres combined); A8L/A8W (M in males 2.06, in females 1.58 vs. 1.50 in both genders); A10L/A10W (R in females 1.54-1.78 vs. more than 2.00 in both genders); shape of the hind pronotal angles (obtuse vs. right or weakly acute); shape of the mesosternal carina (almost right-angled, anterior margin convex, posterior margin regularly concave vs. obtuse-angled, anterior margin obtuse, posterior margin deeply incised); shape of elytra (inversely ovate vs. oval); shape of the median lobe in dorsal (wider in apical half, apically flattened vs. thinner in apical half, apically narrowed) and lateral (more elongate vs. less elongate) views; length of the median lobe (barely longer than parameres vs. markedly longer than parameres); and position of parameral setae in dorsal view (inner pre-apical seta somewhat separated from two remaining setae vs. all setae equidistant) (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2</figureCitation>
-
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)." figureDoi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69">4</figureCitation>
) (
<bibRefCitation author="Breit, J" journalOrPublisher="Cladistics" pageId="0" pageNumber="69" refId="B2" refString="Breit, J, 1902. Ein neuer Hoehlenkaefer aus Bosnien. Societas entomologica 17: 89." title="Ein neuer Hoehlenkaefer aus Bosnien. Societas entomologica 17: 89." year="1902">Breit 1902</bibRefCitation>
;
<bibRefCitation author="Jeannel, R" journalOrPublisher="Archives de Zoologie experimentale et generale" pageId="0" pageNumber="69" pagination="1 - 436" refId="B14" refString="Jeannel, R, 1924. Monographie des Bathysciinae. Biospeologica L. Archives de Zoologie experimentale et generale 63: 1 - 436" title="Monographie des Bathysciinae. Biospeologica L." volume="63" year="1924">Jeannel 1924</bibRefCitation>
;
<bibRefCitation author="Knirsch, E" journalOrPublisher="Casopis Ceskoslovenske Spolecnosti Entomologicke" pageId="0" pageNumber="69" pagination="101 - 122" refId="B15" refString="Knirsch, E, 1928. Novi zastupci slepych Silphidu ze zapadniho Srbska. Casopis Ceskoslovenske Spolecnosti Entomologicke 24: 101 - 122" title="Novi zastupci slepych Silphidu ze zapadniho Srbska." volume="24" year="1928">Knirsch 1928</bibRefCitation>
;
<bibRefCitation author="Mueller, G" journalOrPublisher="Atti del Museo Civico di Storia Naturale di Trieste" pageId="0" pageNumber="69" pagination="107 - 117" refId="B16" refString="Mueller, G, 1937. Nuovi silfidi cavernicoli della Balcania e osservazioni su specie gia descritte. Atti del Museo Civico di Storia Naturale di Trieste 13: 107 - 117" title="Nuovi silfidi cavernicoli della Balcania e osservazioni su specie gia descritte." volume="13" year="1937">
<normalizedToken originalValue="Müller">Mueller</normalizedToken>
1937
</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.3406/bsef.1999.16602" author="Perreau, M" journalOrPublisher="Bulletin de la Societe entomologique de France" pageId="0" pageNumber="69" pagination="399 - 406" refId="B20" refString="Perreau, M, 1999. Nouveaux genres et nouvelles especes de Leptodirini (Coleoptera, Leiodidae). Bulletin de la Societe entomologique de France 104: 399 - 406, DOI: https://doi.org/10.3406/bsef.1999.16602" title="Nouveaux genres et nouvelles especes de Leptodirini (Coleoptera, Leiodidae)." url="https://doi.org/10.3406/bsef.1999.16602" volume="104" year="1999">Perreau 1999</bibRefCitation>
;
<bibRefCitation author="Nonveiller, G" journalOrPublisher="Revue francaise d'Entomologie, Nouvelle serie" pageId="0" pageNumber="69" pagination="165 - 168" refId="B19" refString="Nonveiller, G, Pavicevic, D, Rada, T, Vujcic-Karlo, S, 2002. A new species of the genus Leonhardia Reitter, 1901 from Herzegovina [Coleoptera, Leiodidae, Cholevinae, Leptodirini]. Revue francaise d'Entomologie, Nouvelle serie 24: 165 - 168" title="A new species of the genus Leonhardia Reitter, 1901 from Herzegovina [Coleoptera, Leiodidae, Cholevinae, Leptodirini]." volume="24" year="2002">Nonveiller et al. 2002</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="69">
<taxonomicName authorityName="Curcic, Rađa, Vesovic &amp; Vrbica" authorityYear="2021" class="Insecta" family="Leiodidae" genus="Leonhardia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leonhardia solaki" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="solaki">
<emphasis italics="true" pageId="0" pageNumber="69">Leonhardia solaki</emphasis>
</taxonomicName>
sp. nov. differs from
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. delminiumica" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="delminiumica">
<emphasis italics="true" pageId="0" pageNumber="69">L. delminiumica</emphasis>
</taxonomicName>
in regard to TL (R 3.40-3.65 mm in males, 3.55-3.93 mm in females vs. R 3.10-3.20 mm in males, 3.40 mm in a female); A8L/A8W (M 2.06 in males, 1.58 in females vs. antennomere VIII barely longer than broad); shape of the mesosternal carina (almost right-angled, posterior margin less concave vs. obtuse-angled, posterior margin more concave); shape of the median lobe in dorsal (apically flattened, with a rounded apex vs. apically narrowed, with a pointed apex) and lateral (more curved, wider vs. less curved, narrower) views; length of the median lobe (barely longer than parameres vs. markedly longer than parameres); size of the basal bulb in dorsal view (larger vs. smaller); and position of parameral setae in dorsal view (inner pre-apical seta closer to the two remaining setae vs. inner pre-apical seta farther away from the two remaining setae) (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2</figureCitation>
-
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)." figureDoi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69">4</figureCitation>
) (
<bibRefCitation author="Nonveiller, G" journalOrPublisher="Revue francaise d'Entomologie, Nouvelle serie" pageId="0" pageNumber="69" pagination="165 - 168" refId="B19" refString="Nonveiller, G, Pavicevic, D, Rada, T, Vujcic-Karlo, S, 2002. A new species of the genus Leonhardia Reitter, 1901 from Herzegovina [Coleoptera, Leiodidae, Cholevinae, Leptodirini]. Revue francaise d'Entomologie, Nouvelle serie 24: 165 - 168" title="A new species of the genus Leonhardia Reitter, 1901 from Herzegovina [Coleoptera, Leiodidae, Cholevinae, Leptodirini]." volume="24" year="2002">Nonveiller et al. 2002</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="69">
<taxonomicName authorityName="Curcic, Rađa, Vesovic &amp; Vrbica" authorityYear="2021" class="Insecta" family="Leiodidae" genus="Leonhardia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leonhardia solaki" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="solaki">
<emphasis italics="true" pageId="0" pageNumber="69">Leonhardia solaki</emphasis>
</taxonomicName>
sp. nov. is easily distinguished from
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. jajcensis" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="jajcensis">
<emphasis italics="true" pageId="0" pageNumber="69">L. jajcensis</emphasis>
</taxonomicName>
, from which it differs with respect to AL in females (antennae reaching the middle of elytra vs. antennae ending before the middle of elytra); shape of the hind pronotal angles (obtuse vs. almost right); shape of the mesosternal carina (higher, anterior margin more convex, posterior margin more concave vs. lower, anterior margin less convex, posterior margin less concave); form of the elytra (more elongate vs. less elongate); shape of the median lobe in dorsal (apically flattened vs. apically narrowed) and lateral (more thickened in its basal half, less convex ventrally vs. less thickened in its basal half, more convex ventrally) views; length of the median lobe (barely longer than parameres vs. markedly longer than parameres); size of the basal bulb in dorsal view (larger vs. smaller); and position of parameral setae in dorsal view (inner pre-apical seta farther away from the two remaining setae vs. inner pre-apical seta closer to the two remaining setae) (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2</figureCitation>
-
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)." figureDoi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69">4</figureCitation>
) (
<bibRefCitation DOI="https://doi.org/10.2298/ABS1402919C" author="Curcic, S" journalOrPublisher="Archives of Biological Sciences, Belgrade" pageId="0" pageNumber="69" pagination="919 - 933" refId="B6" refString="Curcic, S, Rađa, T, Mulaomerovic, J, Vrbica, M, Antic, D, Curcic, B, Rađa, B, Vesovic, N, 2014. Three new cave-dwelling leiodid beetles (Coleoptera: Leiodidae: Cholevinae: Leptodirini) from Bosnia and Herzegovina. Archives of Biological Sciences, Belgrade 66: 919 - 933, DOI: https://doi.org/10.2298/ABS1402919C" title="Three new cave-dwelling leiodid beetles (Coleoptera: Leiodidae: Cholevinae: Leptodirini) from Bosnia and Herzegovina." url="https://doi.org/10.2298/ABS1402919C" volume="66" year="2014">
<normalizedToken originalValue="Ćurčić">Curcic</normalizedToken>
et al. 2014
</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="69">
<taxonomicName authorityName="Curcic, Rađa, Vesovic &amp; Vrbica" authorityYear="2021" class="Insecta" family="Leiodidae" genus="Leonhardia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leonhardia solaki" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="solaki">
<emphasis italics="true" pageId="0" pageNumber="69">Leonhardia solaki</emphasis>
</taxonomicName>
sp. nov. differs from
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. sebesicensis" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="sebesicensis">
<emphasis italics="true" pageId="0" pageNumber="69">L. sebesicensis</emphasis>
</taxonomicName>
in regard to AL in females (antennae reaching the middle of elytra vs. antennae ending before the middle of elytra); shape of the hind pronotal angles (obtuse-angled in both genders vs. almost right-angled in males); shape of the pronotum and elytra (less elongate vs. more elongate); shape of the mesosternal carina (apically rounded vs. apically toothed); shape of the median lobe in dorsal (apically flattened vs. apically rounded) and lateral (more curved, narrower in apical half vs. less curved, wider in apical half) views; length of the median lobe (barely longer than parameres vs. markedly longer than parameres); and position of parameral setae in dorsal view (inner pre-apical seta somewhat separated from two remaining setae vs. all setae equidistant) (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2</figureCitation>
-
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)." figureDoi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69">4</figureCitation>
) (
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4483.3.6" author="Curcic, S" journalOrPublisher="Zootaxa" pageId="0" pageNumber="69" pagination="523 - 548" refId="B7" refString="Curcic, S, Pavicevic, D, Vesovic, N, Mulaomerovic, J, Rađa, T, Antic, D, Bosco, F, Markovic, Đ, Petkovic, M, 2018. Seven new taxa of Leptodirini (Coleoptera: Leiodidae: Cholevinae) from the Balkan Peninsula. Zootaxa 4483: 523 - 548, DOI: https://doi.org/10.11646/zootaxa.4483.3.6" title="Seven new taxa of Leptodirini (Coleoptera: Leiodidae: Cholevinae) from the Balkan Peninsula." url="https://doi.org/10.11646/zootaxa.4483.3.6" volume="4483" year="2018">
<normalizedToken originalValue="Ćurčić">Curcic</normalizedToken>
et al. 2018
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="69" type="description">
<paragraph pageId="0" pageNumber="69">Description.</paragraph>
<paragraph pageId="0" pageNumber="69">Medium-sized leptodirine. TL M 3.62 mm (3.55 mm in males, 3.69 mm in females), R 3.40-3.93 mm (3.40-3.65 mm in males, 3.55-3.93 mm in females).</paragraph>
<paragraph pageId="0" pageNumber="69">
<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Habitus</emphasis>
: Body shape pholeuonoid, colour reddish-brown (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2A</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)." figureDoi="10.3897/subtbiol.41.75613.figure3" httpUri="https://binary.pensoft.net/fig/621564" pageId="0" pageNumber="69">3A, B</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="69">
<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Integument</emphasis>
: Shiny, microsculptured (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)." figureDoi="10.3897/subtbiol.41.75613.figure3" httpUri="https://binary.pensoft.net/fig/621564" pageId="0" pageNumber="69">3D, G, L</figureCitation>
). Body covered with a number of densely distributed deep punctures and yellow pubescence of medium length (erect on head, recumbent on both pronotum and elytra).
</paragraph>
<caption doi="10.3897/subtbiol.41.75613.figure3" httpUri="https://binary.pensoft.net/fig/621564" pageId="0" pageNumber="69" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="69">
<emphasis bold="true" pageId="0" pageNumber="69">Figure 3.</emphasis>
SEM images of morphological features of
<taxonomicName authorityName="Curcic, Rađa, Vesovic &amp; Vrbica" authorityYear="2021" class="Insecta" family="Leiodidae" genus="Leonhardia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leonhardia solaki" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="solaki">
<emphasis italics="true" pageId="0" pageNumber="69">Leonhardia solaki</emphasis>
</taxonomicName>
sp. nov. from the
<normalizedToken originalValue="Golubnjača">Golubnjaca</normalizedToken>
kod Skucana Pit, village of Skucani, close to the town of
<normalizedToken originalValue="Glamoč">Glamoc</normalizedToken>
, western Bosnia and Herzegovina
<emphasis bold="true" pageId="0" pageNumber="69">A</emphasis>
paratype male, habitus, dorsal aspect
<emphasis bold="true" pageId="0" pageNumber="69">B</emphasis>
paratype male, habitus, lateral aspect
<emphasis bold="true" pageId="0" pageNumber="69">C</emphasis>
paratype male, head, dorsal aspect
<emphasis bold="true" pageId="0" pageNumber="69">D</emphasis>
paratype male, microsculpture of head, dorsal aspect
<emphasis bold="true" pageId="0" pageNumber="69">E</emphasis>
paratype male, right antenna, dorsal aspect
<emphasis bold="true" pageId="0" pageNumber="69">F</emphasis>
paratype male, pronotum, dorsal aspect
<emphasis bold="true" pageId="0" pageNumber="69">G</emphasis>
paratype male, microsculpture of pronotum, dorsal aspect
<emphasis bold="true" pageId="0" pageNumber="69">H</emphasis>
paratype male, mesosternal carina, lateral aspect
<emphasis bold="true" pageId="0" pageNumber="69">I</emphasis>
paratype male, mesoventrite, ventral aspect
<emphasis bold="true" pageId="0" pageNumber="69">J</emphasis>
paratype male, scutellum, dorsal aspect
<emphasis bold="true" pageId="0" pageNumber="69">K</emphasis>
paratype male, elytra, dorsal aspect
<emphasis bold="true" pageId="0" pageNumber="69">L</emphasis>
paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (
<emphasis bold="true" pageId="0" pageNumber="69">A, B</emphasis>
); 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="69">E, F, H, I, K</emphasis>
); 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="69">C, J</emphasis>
); 0.1 mm (
<emphasis bold="true" pageId="0" pageNumber="69">D, G, L</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="69">
<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Head</emphasis>
: Longer than wide (HL/HW M 1.11, R 1.06-1.19), anophthalmous, with no occipital carina (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2A</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)." figureDoi="10.3897/subtbiol.41.75613.figure3" httpUri="https://binary.pensoft.net/fig/621564" pageId="0" pageNumber="69">3C</figureCitation>
). Labrum slightly emarginate, with several long setae. Penultimate maxillary palpomere widened apically. Ultimate maxillary palpomere short, thin, gradually narrowing apically. Vertex with a longitudinal impression. Antennae inserted in middle third of head, slender, narrow proximally, slightly dilated distally, longer in males, AL M 2.15 mm, R 1.93-2.33 mm (2.165-2.33 mm in males, 1.93-2.165 mm in females), exceeding middle of elytra in males and reaching middle of elytra in females (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2A</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)." figureDoi="10.3897/subtbiol.41.75613.figure3" httpUri="https://binary.pensoft.net/fig/621564" pageId="0" pageNumber="69">3E</figureCitation>
). Antennomere I short and wide. A1L/A2L M 0.54, R 0.52-0.58. Antennomere II narrow, elongate, shorter than III and IV combined (A2L/A3+A4L M 0.86, R 0.79-0.90). Antennomeres III-VI of similar shape and size, narrow, among which V is the longest. Antennomeres VII and IX-XI widened (VII, IX, and X dilated distally), large, among which VII is the shortest in males, VIII the shortest in females, and XI the longest in both sexes. Antennomere VIII small, elongate in males (A8L/A8W M 2.06, R 1.82-2.40), wide in females (A8L/A8W M 1.58, R 1.33-1.90). A9L/A9W in males M 2.35 (R 2.25-2.50), in females M 1.68 (R 1.60-1.78). A10L/A10W in males M 2.27 (R 2.21-2.375), in females M 1.65 (R 1.54-1.78). Ultimate antennomere slender, narrowing apically, A11L/A11W in males M 3.10 (R 2.54-3.50), in females M 2.82 (R 2.67-3.00).
</paragraph>
<caption doi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="69">
<emphasis bold="true" pageId="0" pageNumber="69">Figure 4.</emphasis>
Different morphological features of
<taxonomicName class="Insecta" family="Leiodidae" genus="Leonhardia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leonhardia" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="69">Leonhardia</emphasis>
</taxonomicName>
[
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. droveniki" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="droveniki">
<emphasis italics="true" pageId="0" pageNumber="69">L. droveniki</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="69">A</emphasis>
)
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. hilfi" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="hilfi">
<emphasis italics="true" pageId="0" pageNumber="69">L. hilfi</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="69">B, G, L, Q</emphasis>
)
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. reitteri" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="reitteri">
<emphasis italics="true" pageId="0" pageNumber="69">L. reitteri</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="69">C, H, M, R</emphasis>
)
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. delminiumica" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="delminiumica">
<emphasis italics="true" pageId="0" pageNumber="69">L. delminiumica</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="69">D, I, N, S</emphasis>
)
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. jajcensis" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="jajcensis">
<emphasis italics="true" pageId="0" pageNumber="69">L. jajcensis</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="69">E, J, O, T</emphasis>
) and
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. sebesicensis" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="sebesicensis">
<emphasis italics="true" pageId="0" pageNumber="69">L. sebesicensis</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="69">F, K, P, U</emphasis>
)] [modified after
<bibRefCitation author="Nonveiller, G" journalOrPublisher="Revue francaise d'Entomologie, Nouvelle serie" pageId="0" pageNumber="69" pagination="165 - 168" refId="B19" refString="Nonveiller, G, Pavicevic, D, Rada, T, Vujcic-Karlo, S, 2002. A new species of the genus Leonhardia Reitter, 1901 from Herzegovina [Coleoptera, Leiodidae, Cholevinae, Leptodirini]. Revue francaise d'Entomologie, Nouvelle serie 24: 165 - 168" title="A new species of the genus Leonhardia Reitter, 1901 from Herzegovina [Coleoptera, Leiodidae, Cholevinae, Leptodirini]." volume="24" year="2002">Nonveiller et al. (2002)</bibRefCitation>
and
<bibRefCitation DOI="https://doi.org/10.2298/ABS1402919C" author="Curcic, S" journalOrPublisher="Archives of Biological Sciences, Belgrade" pageId="0" pageNumber="69" pagination="919 - 933" refId="B6" refString="Curcic, S, Rađa, T, Mulaomerovic, J, Vrbica, M, Antic, D, Curcic, B, Rađa, B, Vesovic, N, 2014. Three new cave-dwelling leiodid beetles (Coleoptera: Leiodidae: Cholevinae: Leptodirini) from Bosnia and Herzegovina. Archives of Biological Sciences, Belgrade 66: 919 - 933, DOI: https://doi.org/10.2298/ABS1402919C" title="Three new cave-dwelling leiodid beetles (Coleoptera: Leiodidae: Cholevinae: Leptodirini) from Bosnia and Herzegovina." url="https://doi.org/10.2298/ABS1402919C" volume="66" year="2014">
<normalizedToken originalValue="Ćurčić">Curcic</normalizedToken>
et al. (2014
</bibRefCitation>
,
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4483.3.6" author="Curcic, S" journalOrPublisher="Zootaxa" pageId="0" pageNumber="69" pagination="523 - 548" refId="B7" refString="Curcic, S, Pavicevic, D, Vesovic, N, Mulaomerovic, J, Rađa, T, Antic, D, Bosco, F, Markovic, Đ, Petkovic, M, 2018. Seven new taxa of Leptodirini (Coleoptera: Leiodidae: Cholevinae) from the Balkan Peninsula. Zootaxa 4483: 523 - 548, DOI: https://doi.org/10.11646/zootaxa.4483.3.6" title="Seven new taxa of Leptodirini (Coleoptera: Leiodidae: Cholevinae) from the Balkan Peninsula." url="https://doi.org/10.11646/zootaxa.4483.3.6" volume="4483" year="2018">2018</bibRefCitation>
)]
<emphasis bold="true" pageId="0" pageNumber="69">A-F</emphasis>
mesosternal carina, lateral aspect
<emphasis bold="true" pageId="0" pageNumber="69">G-K</emphasis>
aedeagus, dorsal aspect
<emphasis bold="true" pageId="0" pageNumber="69">L-P</emphasis>
aedeagus, lateral aspect
<emphasis bold="true" pageId="0" pageNumber="69">Q-U</emphasis>
left paramere apex, dorsal aspect. Scale bars: 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="69">G-P</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="69">
<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Prothorax</emphasis>
: Pronotum bell-shaped, transverse (PL/PW M 0.87, R 0.84-0.91), widest slightly after anterior third, markedly broader than head (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2A</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)." figureDoi="10.3897/subtbiol.41.75613.figure3" httpUri="https://binary.pensoft.net/fig/621564" pageId="0" pageNumber="69">3F</figureCitation>
). Lateral margins well-rounded anteriorly, somewhat concave posteriorly, sub-parallel basally. Pronotal base straight, markedly shorter than elytral base (PB/EB M 0.95, R 0.90-0.99). PB/AM M 1.40, R 1.18-1.55. Anterior margin somewhat convex medially. Fore pronotal angles small, rounded, obtuse. Hind pronotal angles obtuse, prominent, not protruding backwards. Pronotal disc moderately convex (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)." figureDoi="10.3897/subtbiol.41.75613.figure3" httpUri="https://binary.pensoft.net/fig/621564" pageId="0" pageNumber="69">3B</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="69">
<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Mesothorax</emphasis>
: Mesosternal carina high anteriorly, low posteriorly (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2B</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)." figureDoi="10.3897/subtbiol.41.75613.figure3" httpUri="https://binary.pensoft.net/fig/621564" pageId="0" pageNumber="69">3H</figureCitation>
). Anterior margin convex, posterior margin concave, with a few setae. Tooth obtuse, rounded. No mesoventral processus on mesoventrite (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)." figureDoi="10.3897/subtbiol.41.75613.figure3" httpUri="https://binary.pensoft.net/fig/621564" pageId="0" pageNumber="69">3I</figureCitation>
). Scutellum large, triangular (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2A</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)." figureDoi="10.3897/subtbiol.41.75613.figure3" httpUri="https://binary.pensoft.net/fig/621564" pageId="0" pageNumber="69">3J</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="69">
<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Metathorax</emphasis>
: Metasternum without carina.
</paragraph>
<paragraph pageId="0" pageNumber="69">
<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Elytra</emphasis>
: Wide, obovoid, slightly wider in females (EL/EW in males M 1.47, R 1.40-1.57; in females M 1.44, R 1.41-1.48), markedly wider than pronotum (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2A</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)." figureDoi="10.3897/subtbiol.41.75613.figure3" httpUri="https://binary.pensoft.net/fig/621564" pageId="0" pageNumber="69">3K</figureCitation>
). Maximum width slightly before middle. Lateral margins arcuate. Marginal furrows visible on nearly entire elytra. Shoulders weakly expressed, almost rounded. Elytral disc markedly convex, gently declining basally and steeply declining apically in lateral view (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)." figureDoi="10.3897/subtbiol.41.75613.figure3" httpUri="https://binary.pensoft.net/fig/621564" pageId="0" pageNumber="69">3B</figureCitation>
). Sutural striae developed. Elytral apex attenuated, rounded. Pygidium covered by elytra.
</paragraph>
<paragraph pageId="0" pageNumber="69">
<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Legs</emphasis>
: Elongate (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2A</figureCitation>
). Femora widened basally, constricted sub-apically. Tibiae thin, very gently curved, gradually widening distally, each with several spines. Each fore tibia with an outer distal brush-like structure. Male fore tarsi five-segmented, weakly dilated. Tarsal claws elongate, narrow, curved, pointed apically.
</paragraph>
<paragraph pageId="0" pageNumber="69">
<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Male genitalia</emphasis>
: Aedeagus elongate, straight, thin, sclerotized (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2C, D</figureCitation>
). Basal bulb small and rounded in dorsal view (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2C</figureCitation>
), moderately large in lateral view (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2D</figureCitation>
). Median lobe in dorsal view sub-parallel, rounded anteriorly, with a flattened apex, longer than parameres (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2C</figureCitation>
). Median lobe in lateral view curved, straight in basal half, sub-apically concave dorsally, almost straight ventrally in basal part, with an acute apex curved downwards (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2D</figureCitation>
). Parameres slender, narrow, arcuate, sub-apically curved towards exterior, each with a dilated rounded apex in dorsal view (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2C, E</figureCitation>
), while gently narrowing distally in lateral view (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2D</figureCitation>
). Paramere bearing three setae: two strong, long, in apical and pre-apical position, respectively, and one inner, thin, short, in pre-apical position (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2E</figureCitation>
). Two long parameral setae close-set, short one somewhat separated, positioned somewhat below level of other two parameral setae (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2E</figureCitation>
). Endophallus elongate, tubular. Copulatory piece having a Y-shaped phanera and two lateral basal sclerifications, in front of which a weakly chitinized structure resembling teeth is present. In addition, two lateral bands are present in apical half of median lobe (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2C</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="69">
<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Female genitalia</emphasis>
: Gonostyli slender, thin, gradually narrowing distally, straight, pointed apically (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2F</figureCitation>
). Each gonostylus carrying one apical seta, three inner setae, and one outer seta (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2F</figureCitation>
). Spermatheca elongate, chitinized, curved, constricted posteriorly in sub-apical part, spherical apically (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2G</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="69">
<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Female abdominal sternite VIII</emphasis>
: Large, transverse, with a thin anterior process, carrying pubescence (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="69" type="intraspecific variability">
<paragraph pageId="0" pageNumber="69">Intraspecific variability.</paragraph>
<paragraph pageId="0" pageNumber="69">A certain level of intraspecific variability is observed in the new species. Several morphological differences between males and females point to the occurrence of sexual dimorphism. To be specific: (i) males are on average slightly shorter than females; (ii) the antennae in males are longer than in females; (iii) antennomeres VIII-XI in males are more slender than in females; (iv) the head in males is larger than in females; (v) the pronotum in males is more elongate than in females; (vi) the elytra in males are more elongate than in females.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="69" type="distribution">
<paragraph pageId="0" pageNumber="69">Geographic distribution.</paragraph>
<paragraph pageId="0" pageNumber="69">
Thus far, the species is known only from its type locality - the
<normalizedToken originalValue="Golubnjača">Golubnjaca</normalizedToken>
kod Skucana Pit, situated in the village of Skucani, close to the town of
<normalizedToken originalValue="Glamoč">Glamoc</normalizedToken>
in western Bosnia and Herzegovina. This site represents the westernmost location of a
<taxonomicName class="Insecta" family="Leiodidae" genus="Leonhardia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leonhardia" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="69">Leonhardia</emphasis>
</taxonomicName>
species. We assume that the new species probably inhabits other subterranean sites in the surrounding areas of western Bosnia and Herzegovina, although one of us (TR) visited the
<normalizedToken originalValue="Ledenjača">Ledenjaca</normalizedToken>
Cave situated in the same village (Skucani), but found no specimens of the new species there.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="69" type="bionomy and habitat">
<paragraph pageId="0" pageNumber="69">Bionomy and habitat.</paragraph>
<paragraph pageId="0" pageNumber="69">
All specimens of
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. solaki" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="solaki">
<emphasis italics="true" pageId="0" pageNumber="69">L. solaki</emphasis>
</taxonomicName>
sp. nov. were collected deep in the
<normalizedToken originalValue="Golubnjača">Golubnjaca</normalizedToken>
kod Skucana Pit. They were recorded at this subterranean site at a depth of 30 m under conditions of permanent darkness and high humidity, along with the presence of trickling water. All individuals of
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. solaki" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="solaki">
<emphasis italics="true" pageId="0" pageNumber="69">L. solaki</emphasis>
</taxonomicName>
sp. nov. were gathered manually from the floor and walls in the innermost part of the pit. Aside from the new species,
<normalizedToken originalValue="Golubnjača">Golubnjaca</normalizedToken>
kod Skucana Pit is inhabited by another subterranean leiodid,
<taxonomicName authorityName="Maran" authorityYear="1943" class="Insecta" family="Leiodidae" genus="Parapropus" kingdom="Animalia" lsidName="Parapropus ganglbaueri subsp. obenbergeri" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="subSpecies" species="ganglbaueri" subSpecies="obenbergeri">
<emphasis italics="true" pageId="0" pageNumber="69">Parapropus ganglbaueri obenbergeri</emphasis>
</taxonomicName>
<normalizedToken originalValue="Mařan">Maran</normalizedToken>
, 1943, which is recorded in the same habitat where individuals of the new species were found, so these two taxa can be treated as sympatric. The same pit is also the type locality of the recently described moth fly
<taxonomicName authorityName="Wagner &amp; Rada" authorityYear="2020" class="Insecta" family="Psychodidae" genus="Psychoda" kingdom="Animalia" lsidName="Psychoda glamocensis" order="Diptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="glamocensis">
<emphasis italics="true" pageId="0" pageNumber="69">Psychoda glamocensis</emphasis>
</taxonomicName>
Wagner &amp; Rada, 2020 (
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4845.2.8" author="Wagner, R" journalOrPublisher="Zootaxa" pageId="0" pageNumber="69" pagination="275 - 282" refId="B32" refString="Wagner, R, Rada, T, 2020. Moth flies (Diptera, Psychodidae) living in the dark of caves in the Dinaric Karst. Zootaxa 4845: 275 - 282, DOI: https://doi.org/10.11646/zootaxa.4845.2.8" title="Moth flies (Diptera, Psychodidae) living in the dark of caves in the Dinaric Karst." url="https://doi.org/10.11646/zootaxa.4845.2.8" volume="4845" year="2020">Wagner and Rada 2020</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="69" type="key for identification of species and subspecies of leonhardia (figs 24)">
<paragraph pageId="0" pageNumber="69">
Key for identification of species and subspecies of
<taxonomicName class="Insecta" family="Leiodidae" genus="Leonhardia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leonhardia" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="69">Leonhardia</emphasis>
</taxonomicName>
(Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2</figureCitation>
-
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)." figureDoi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69">4</figureCitation>
)
</paragraph>
<paragraph pageId="0" pageNumber="69">
<table inLine="true" pageId="0" pageNumber="69">
<tr pageId="0" pageNumber="69">
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
Smaller body size (2.80-3.00 mm in length). Mesosternal carina low (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)." figureDoi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69">4A</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="69">
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. droveniki" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="droveniki">
<emphasis italics="true" pageId="0" pageNumber="69">L. droveniki</emphasis>
</taxonomicName>
Perreau, 1999
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="69">
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
Larger body size (3.10-4.00 mm in length). Mesosternal carina high (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2B</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)." figureDoi="10.3897/subtbiol.41.75613.figure3" httpUri="https://binary.pensoft.net/fig/621564" pageId="0" pageNumber="69">3H</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)." figureDoi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69">4B-F</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="69">2</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="69">
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
Anterior margin of mesosternal carina obtuse, posterior margin deeply incised (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)." figureDoi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69">4C</figureCitation>
). Median lobe in lateral view less elongate and more curved (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)." figureDoi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69">4M</figureCitation>
) (
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. reitteri" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="reitteri">
<emphasis italics="true" pageId="0" pageNumber="69">L. reitteri</emphasis>
</taxonomicName>
Breit, 1902)
</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="69">3</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="69">
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
Anterior margin of mesosternal carina convex, posterior margin regularly concave (Figs
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)." figureDoi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69">4B, D-F</figureCitation>
). Median lobe in lateral view more elongate and less curved (Figs
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)." figureDoi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69">4L, N-P</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="69">5</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="69">
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">3</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">Larger body size (3.50-3.70 mm in length). Pronotum less transverse. Lateral elytral margins more arcuate, elytra apically rounded</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="69">4</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="69">
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">Smaller body size (3.20 mm in length). Pronotum more transverse. Lateral elytral margins less arcuate, elytra apically pointed</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="69">
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. reitteri subsp. mersa" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="subSpecies" species="reitteri" subSpecies="mersa">
<emphasis italics="true" pageId="0" pageNumber="69">L. reitteri mersa</emphasis>
</taxonomicName>
Knirsch, 1928
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="69">
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">Pronotum narrowed basally, elytra narrower and less convex, pronotal depressions lacking or barely discernible</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="69">
<taxonomicName authority="reitteri" class="Insecta" genus="L." kingdom="Animalia" lsidName="L. reitteri subsp. reitteri" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="subSpecies" species="reitteri" subSpecies="reitteri">
<emphasis italics="true" pageId="0" pageNumber="69">L. reitteri reitteri</emphasis>
</taxonomicName>
Breit, 1902
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="69">
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">Pronotum not narrowed basally, elytra wider and more convex, two depressions present on pronotum</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="69">
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. reitteri subsp. zariquieyi" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="subSpecies" species="reitteri" subSpecies="zariquieyi">
<emphasis italics="true" pageId="0" pageNumber="69">L. reitteri zariquieyi</emphasis>
</taxonomicName>
<normalizedToken originalValue="Müller">Mueller</normalizedToken>
, 1937
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="69">
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
Mesosternal carina almost right-angled (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2A</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)." figureDoi="10.3897/subtbiol.41.75613.figure3" httpUri="https://binary.pensoft.net/fig/621564" pageId="0" pageNumber="69">3H</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)." figureDoi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69">4E</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="69">6</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="69">
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
Mesosternal carina obtuse-angled (Figs
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)." figureDoi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69">4B, D, F</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="69">7</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="69">
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
Hind pronotal angles almost right. Mesosternal carina lower, anterior margin less convex, posterior margin less concave (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)." figureDoi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69">4E</figureCitation>
). Median lobe in dorsal view apically narrowed, markedly longer than parameres (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)." figureDoi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69">4J</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="69">
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. jajcensis" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="jajcensis">
<emphasis italics="true" pageId="0" pageNumber="69">L. jajcensis</emphasis>
</taxonomicName>
S.
<normalizedToken originalValue="Ćurčić">Curcic</normalizedToken>
&amp;
<normalizedToken originalValue="Rađa">Rađa</normalizedToken>
, 2014
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="69">
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
Hind pronotal angles obtuse (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2A</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)." figureDoi="10.3897/subtbiol.41.75613.figure3" httpUri="https://binary.pensoft.net/fig/621564" pageId="0" pageNumber="69">3F</figureCitation>
). Mesosternal carina higher, anterior margin more convex, posterior margin more concave (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2B</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)." figureDoi="10.3897/subtbiol.41.75613.figure3" httpUri="https://binary.pensoft.net/fig/621564" pageId="0" pageNumber="69">3H</figureCitation>
). Median lobe in dorsal view apically flattened, barely longer than parameres (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)." figureDoi="10.3897/subtbiol.41.75613.figure2" httpUri="https://binary.pensoft.net/fig/621563" pageId="0" pageNumber="69">2C</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="69">
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. solaki" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="solaki">
<emphasis italics="true" pageId="0" pageNumber="69">L. solaki</emphasis>
</taxonomicName>
sp. nov.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="69">
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">7</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
Median lobe in dorsal view apically pointed (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)." figureDoi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69">4I</figureCitation>
). Inner pre-apical seta markedly separated from two remaining parameral setae (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)." figureDoi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69">4S</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="69">
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. delminiumica" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="delminiumica">
<emphasis italics="true" pageId="0" pageNumber="69">L. delminiumica</emphasis>
</taxonomicName>
Nonveiller,
<normalizedToken originalValue="Pavićević">Pavicevic</normalizedToken>
,
<normalizedToken originalValue="Rađa">Rađa</normalizedToken>
&amp;
<normalizedToken originalValue="Vujčić-Karlo">Vujcic-Karlo</normalizedToken>
, 2002
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="69">
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
Median lobe in dorsal view apically rounded (Figs
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)." figureDoi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69">4G, K</figureCitation>
). Parameral setae close-set (Figs
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)." figureDoi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69">4Q, U</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="69">8</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="69">
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">8</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
Mesosternal carina higher, apically toothed (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)." figureDoi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69">4F</figureCitation>
). Median lobe in dorsal view wider, apically rounded (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)." figureDoi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69">4K</figureCitation>
). Parameral setae equidistant (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)." figureDoi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69">4U</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="69">
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. sebesicensis" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="sebesicensis">
<emphasis italics="true" pageId="0" pageNumber="69">L. sebesicensis</emphasis>
</taxonomicName>
S.
<normalizedToken originalValue="Ćurčić">Curcic</normalizedToken>
,
<normalizedToken originalValue="Pavićević">Pavicevic</normalizedToken>
&amp;
<normalizedToken originalValue="Mulaomerović">Mulaomerovic</normalizedToken>
, 2018
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="69">
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
Mesosternal carina lower, apically rounded (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)." figureDoi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69">4B</figureCitation>
). Median lobe in dorsal view narrower, apically flattened (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)." figureDoi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69">4G</figureCitation>
). Inner pre-apical parameral seta somewhat separated from two remaining parameral setae (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)." figureDoi="10.3897/subtbiol.41.75613.figure4" httpUri="https://binary.pensoft.net/fig/621565" pageId="0" pageNumber="69">4Q</figureCitation>
) (
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. hilfi" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="hilfi">
<emphasis italics="true" pageId="0" pageNumber="69">L. hilfi</emphasis>
</taxonomicName>
Reitter, 1901)
</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="69">9</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="69">
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">9</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">Smaller body size (3.40-3.50 mm in length). Antennae shorter and narrower. Punctures smaller. Pubescence shorter</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="69">
<taxonomicName authority="hilfi" class="Insecta" genus="L." kingdom="Animalia" lsidName="L. hilfi subsp. hilfi" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="subSpecies" species="hilfi" subSpecies="hilfi">
<emphasis italics="true" pageId="0" pageNumber="69">L. hilfi hilfi</emphasis>
</taxonomicName>
Reitter, 1901
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="69">
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">Larger body size (3.80-4.00 mm in length). Antennae longer and wider. Punctures larger. Pubescence longer</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="69">
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. hilfi subsp. robusta" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="subSpecies" species="hilfi" subSpecies="robusta">
<emphasis italics="true" pageId="0" pageNumber="69">L. hilfi robusta</emphasis>
</taxonomicName>
Knirsch, 1928
</emphasis>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>