312 lines
37 KiB
XML
312 lines
37 KiB
XML
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<mods:title id="38D41B034CABD9E6AE30B09CF77378C2">Phylogeny, taxonomy and flower-breeding ecology of the Colocasiomyia cristata species group (Diptera: Drosophilidae), with descriptions of ten new species</mods:title>
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<mods:namePart id="7FC66B640AD8A6FF882E1FC070385B55">Takano, Kohei Takenaka</mods:namePart>
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<mods:namePart id="5CBF9F60BDB37D0FA03016AB2D7AA64F">Yafuso, Masako</mods:namePart>
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<mods:namePart id="01746AC60046DE9EE59807B60C8C66C7">Suwito, Awit</mods:namePart>
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<mods:namePart id="CD953889CA4CC5141833DDA9CD3807CE">Pungga, Runi Anak Sylvester</mods:namePart>
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<mods:namePart id="4219664739F392EF43C8084301B63DC1">Chong, Lucy</mods:namePart>
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<mods:namePart id="1C0746ED8513423D460211B9762A0A20">Toda, Masanori J.</mods:namePart>
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<mods:title id="0F27D198E0C7DC8D63C8D15318AA9F79">Zootaxa</mods:title>
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<mods:date id="CC5C7F087EB610D4E48BD8741E5FCDD3">2021</mods:date>
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<mods:number id="04AAC228DED34A56EAA6CC8D663D15C2">2021-12-08</mods:number>
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<mods:number id="1D0535206CDF59DBB40108B16E23A164">5079</mods:number>
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<treatment id="03D75C2D45702E23FF526551B7A1B27D" ID-DOI="http://doi.org/10.5281/zenodo.5786157" ID-GBIF-Taxon="191651672" ID-Zenodo-Dep="5786157" LSID="urn:lsid:plazi:treatment:03D75C2D45702E23FF526551B7A1B27D" httpUri="http://treatment.plazi.org/id/03D75C2D45702E23FF526551B7A1B27D" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27">
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<paragraph id="8BC1ED3B45702E25FF526551B118B652" blockId="26.[151,596,1363,1427]" box="[151,596,1363,1390]" pageId="26" pageNumber="27">
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<emphasis id="B90A312945702E25FF526551B118B652" bold="true" box="[151,596,1363,1390]" pageId="26" pageNumber="27">
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3)
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<taxonomicName id="4C7E96B845702E25FF726556B2A5B652" ID-CoL="9HTGB" authority="Toda" authorityName="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda" authorityYear="2021" box="[183,489,1363,1390]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="vieti" status="sp. nov.">
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<emphasis id="B90A312945702E25FF726556B2EFB652" bold="true" box="[183,419,1363,1390]" italics="true" pageId="26" pageNumber="27">Colocasiomyia vieti</emphasis>
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Toda
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2398C5245702E25FE336551B118B652" box="[502,596,1364,1390]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C364BEB045702E25FF52657DB22FB6AE" box="[151,355,1400,1427]" pageId="26" pageNumber="27" type="description">
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<paragraph id="8BC1ED3B45702E25FF52657DB22FB6AE" blockId="26.[151,596,1363,1427]" box="[151,355,1400,1427]" pageId="26" pageNumber="27">
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(
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<figureCitation id="1345F1BE45702E25FF5A657DB3BAB6AE" box="[159,246,1400,1426]" captionStart="FIGURE 9" captionStartId="13.[151,250,1780,1804]" captionTargetBox="[151,1435,181,1757]" captionTargetId="figure-18@13.[151,1435,181,1757]" captionTargetPageId="13" captionText="FIGURE 9. Male abdominal sternite VIs of Colocasiomyia colocasiae (Duda) (B in anteroventral view, C in lateral view), C. stamenicola (Carson & Okada) (D in anterior view) and all species of the C. cristata subgroup (A in ventral view, E‒R in anterior view): A, C. ecornuta sp. nov.; E, C. grandis sp. nov.; F, C. vieti sp. nov.; G, C. cristata de Meijere; H, C. matthewsi sp. nov.; I, C. sulawesiana Okada & Yafuso; J, C. kinabaluana sp. nov.; K, C. kotana sp. nov.; L, C. sabahana sp. nov.; M, C. sarawakana sp. nov.; N, C. pistilicola (Carson & Okada); O, C. xenalocasiae (Okada); P, C. diconica (Toda & Okada); Q, C. sumatrana sp. nov.; and R, C. leucocasiae sp. nov. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766420" httpUri="https://zenodo.org/record/5766420/files/figure.png" pageId="26" pageNumber="27">Figs 9F</figureCitation>
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,
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<figureCitation id="1345F1BE45702E25FEC4657DB27EB6AF" box="[257,306,1400,1427]" captionStart="FIGURE 15" captionStartId="21.[151,250,1936,1960]" captionTargetBox="[151,1435,860,1912]" captionTargetId="figure-242@21.[151,1435,860,1912]" captionTargetPageId="21" captionText="FIGURE 15. Habitus (upper, lateral view; lower, dorsal view) of species of the Colocasiomyia cristata subgroup.A, C. ecornuta sp. nov. ♂; B, C. grandis sp. nov. ♂; C, C. vieti sp. nov. ♂; D, C. cristata de Meijere ♂; E, C. sulawesiana Okada & Yafuso ♂; F, C. kinabaluana sp. nov. ♂; G, C. kotana sp. nov. ♂; and H, C. matthewsi sp. nov. ♂. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766434" httpUri="https://zenodo.org/record/5766434/files/figure.png" pageId="26" pageNumber="27">15C</figureCitation>
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,
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<figureCitation id="1345F1BE45702E25FEFB657DB217B6AE" box="[318,347,1400,1426]" captionStart="FIGURE 18" captionStartId="27.[151,250,1424,1448]" captionTargetBox="[151,1435,181,1400]" captionTargetId="figure-18@27.[151,1435,181,1400]" captionTargetPageId="27" captionText="FIGURE 18. Colocasiomyia vieti Toda, sp. nov. (♂ holotype, ♀ paratype, and puparium from Cuc Phuong, Nho Quan District, Vietnam). A, Periphallic organs (lateral view; inset: surstylus distal portion); B, phallic organs (lateral view); C, ditto (ventral view); D, female terminalia (lateral view); E, puparium (ventral habitus); F, caudal-most portion of puparium (ventral view); G, anterior spiracles of puparium (dorsal view); H, mouth hook; I, cephalopharyngeal skeleton. Scale bars: 0.1 mm in A–D and I, 1.0 mm in E." figureDoi="http://doi.org/10.5281/zenodo.5766442" httpUri="https://zenodo.org/record/5766442/files/figure.png" pageId="26" pageNumber="27">18</figureCitation>
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)
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</paragraph>
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<paragraph id="8BC1ED3B45702E25FF5265B9B1BDB6E8" blockId="26.[151,753,1468,1493]" box="[151,753,1468,1493]" pageId="26" pageNumber="27">
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<treatmentCitationGroup id="AB6ECA1545702E25FF5265B9B1BDB6E8" box="[151,753,1468,1493]" pageId="26" pageNumber="27">
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<taxonomicName id="4C7E96B845702E25FF5265B9B1A1B6E8" authority=": Sultana et al., 2006: 694" authorityName="Sultana" authorityPageNumber="694" authorityYear="2006" box="[151,749,1468,1493]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="undefined-2">
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<emphasis id="B90A312945702E25FF5265B9B263B6E8" box="[151,303,1468,1492]" italics="true" pageId="26" pageNumber="27">Colocasiomyia</emphasis>
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sp.2
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<treatmentCitation id="0ADFCB2A45702E25FEA065B9B1A1B6E8" author="Sultana, F. & Hu, Y. G. & Toda, M. J. & Takenaka, K. & Yafuso, M." box="[357,749,1468,1492]" page="694" pageId="26" pageNumber="27" year="2006">
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aff.
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<emphasis id="B90A312945702E25FE4B65B9B2A9B6E8" box="[398,485,1468,1492]" italics="true" pageId="26" pageNumber="27">diconica</emphasis>
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:
|
||
<bibRefCitation id="EFEF90CA45702E25FE3765B9B1A1B6E8" author="Sultana, F. & Hu, Y. G. & Toda, M. J. & Takenaka, K. & Yafuso, M." box="[498,749,1468,1492]" pageId="26" pageNumber="27" pagination="684 - 702" refId="ref33499" refString="Sultana, F., Hu, Y. G., Toda, M. J., Takenaka, K. & Yafuso, M. (2006) Phylogeny and classification of Colocasiomyia (Diptera, Drosophilidae), and its evolution of pollination mutualism with aroid plants. Systematic Entomology, 31, 684 - 702. https: // doi. org / 10.1111 / j. 1365 - 3113.2006.00344. x" type="journal article" year="2006">
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Sultana
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<emphasis id="B90A312945702E25FD8065B8B13DB6E8" box="[581,625,1468,1492]" italics="true" pageId="26" pageNumber="27">et al</emphasis>
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., 2006: 694
|
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</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C364BEB045702E25FF526607B602B521" box="[151,1358,1538,1565]" pageId="26" pageNumber="27" type="diagnosis">
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<paragraph id="8BC1ED3B45702E25FF526607B602B521" blockId="26.[151,1437,1538,2033]" box="[151,1358,1538,1565]" pageId="26" pageNumber="27">
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<emphasis id="B90A312945702E25FF526607B25EB520" bold="true" box="[151,274,1538,1564]" pageId="26" pageNumber="27">Diagnosis.</emphasis>
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Male abdominal sternite VI with a pair of long processes nearly straight and divergent (
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<figureCitation id="1345F1BE45702E25FB296606B60DB521" box="[1260,1345,1539,1565]" captionStart="FIGURE 9" captionStartId="13.[151,250,1780,1804]" captionTargetBox="[151,1435,181,1757]" captionTargetId="figure-18@13.[151,1435,181,1757]" captionTargetPageId="13" captionText="FIGURE 9. Male abdominal sternite VIs of Colocasiomyia colocasiae (Duda) (B in anteroventral view, C in lateral view), C. stamenicola (Carson & Okada) (D in anterior view) and all species of the C. cristata subgroup (A in ventral view, E‒R in anterior view): A, C. ecornuta sp. nov.; E, C. grandis sp. nov.; F, C. vieti sp. nov.; G, C. cristata de Meijere; H, C. matthewsi sp. nov.; I, C. sulawesiana Okada & Yafuso; J, C. kinabaluana sp. nov.; K, C. kotana sp. nov.; L, C. sabahana sp. nov.; M, C. sarawakana sp. nov.; N, C. pistilicola (Carson & Okada); O, C. xenalocasiae (Okada); P, C. diconica (Toda & Okada); Q, C. sumatrana sp. nov.; and R, C. leucocasiae sp. nov. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766420" httpUri="https://zenodo.org/record/5766420/files/figure.png" pageId="26" pageNumber="27">Fig. 9F</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C364BEB045702E24FF026623B0C1B5BD" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="description">
|
||
<paragraph id="8BC1ED3B45702E25FF026623B3BBB559" blockId="26.[151,1437,1538,2033]" pageId="26" pageNumber="27">
|
||
<emphasis id="B90A312945702E25FF026623B21DB57C" bold="true" box="[199,337,1574,1600]" pageId="26" pageNumber="27">Description</emphasis>
|
||
(
|
||
<specimenCount id="9D7826B245702E25FEA46622B23AB57D" box="[353,374,1575,1601]" pageId="26" pageNumber="27" type="male">♂</specimenCount>
|
||
and
|
||
<specimenCount id="9D7826B245702E25FE6E6622B28DB57D" box="[427,449,1575,1601]" pageId="26" pageNumber="27" type="female">♀</specimenCount>
|
||
; not repeating characters common to
|
||
<taxonomicName id="4C7E96B845702E25FCA46622B09BB57D" authority="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda, 2021" authorityName="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda" authorityYear="2021" box="[865,983,1575,1601]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
|
||
<emphasis id="B90A312945702E25FCA46622B038B57D" box="[865,884,1575,1601]" italics="true" pageId="26" pageNumber="27">C</emphasis>
|
||
.
|
||
<emphasis id="B90A312945702E25FC446622B09BB57D" box="[897,983,1575,1601]" italics="true" pageId="26" pageNumber="27">grandis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90A312945702E25FC186622B77AB57D" bold="true" box="[989,1078,1575,1601]" pageId="26" pageNumber="27">
|
||
<taxonomicNameLabel id="A2398C5245702E25FC186622B77AB57D" box="[989,1078,1575,1601]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
). Head. Supracervical setae 4–6 per side.
|
||
</paragraph>
|
||
<paragraph id="8BC1ED3B45702E25FF02666AB671B591" blockId="26.[151,1437,1538,2033]" pageId="26" pageNumber="27">Thorax. Anterior dorsocentral setae behind transverse suture. Prescutellar acrostichal setae absent; acrostichal setulae in 2 or 4 rows. Apical scutellar setae slightly more distant to each other than to basal scutellar seta.</paragraph>
|
||
<paragraph id="8BC1ED3B45702E25FF0266B2B67DB5ED" blockId="26.[151,1437,1538,2033]" box="[199,1329,1719,1745]" pageId="26" pageNumber="27">Legs. Patch covered with only minute pubescence absent on anterodorsal portion of fore tarsomere I.</paragraph>
|
||
<paragraph id="8BC1ED3B45702E25FF0266DEB012B425" blockId="26.[151,1437,1538,2033]" pageId="26" pageNumber="27">Abdomen. Male: sternites III and IV wider than long. Female: sternites III–V longer than wide; VI as wide as long, posteriorly dilated, medially notched on posterior margin.</paragraph>
|
||
<paragraph id="8BC1ED3B45702E25FF026726B041B4CD" blockId="26.[151,1437,1538,2033]" pageId="26" pageNumber="27">
|
||
Male terminalia. Epandrium pubescent except for narrow anterior to ventral margin and anteroventral elongation, roundish on posteroventral margin (
|
||
<figureCitation id="1345F1BE45702E25FD626742B043B45D" box="[679,783,1863,1889]" captionStart="FIGURE 18" captionStartId="27.[151,250,1424,1448]" captionTargetBox="[151,1435,181,1400]" captionTargetId="figure-18@27.[151,1435,181,1400]" captionTargetPageId="27" captionText="FIGURE 18. Colocasiomyia vieti Toda, sp. nov. (♂ holotype, ♀ paratype, and puparium from Cuc Phuong, Nho Quan District, Vietnam). A, Periphallic organs (lateral view; inset: surstylus distal portion); B, phallic organs (lateral view); C, ditto (ventral view); D, female terminalia (lateral view); E, puparium (ventral habitus); F, caudal-most portion of puparium (ventral view); G, anterior spiracles of puparium (dorsal view); H, mouth hook; I, cephalopharyngeal skeleton. Scale bars: 0.1 mm in A–D and I, 1.0 mm in E." figureDoi="http://doi.org/10.5281/zenodo.5766442" httpUri="https://zenodo.org/record/5766442/files/figure.png" pageId="26" pageNumber="27">Fig. 18A</figureCitation>
|
||
). Surstylus long, moderately broad plate with 5–6 recurved teeth apically and 1 trichoid seta ventrosubapically (
|
||
<figureCitation id="1345F1BE45702E25FD3F676EB02AB4B9" box="[762,870,1899,1925]" captionStart="FIGURE 18" captionStartId="27.[151,250,1424,1448]" captionTargetBox="[151,1435,181,1400]" captionTargetId="figure-18@27.[151,1435,181,1400]" captionTargetPageId="27" captionText="FIGURE 18. Colocasiomyia vieti Toda, sp. nov. (♂ holotype, ♀ paratype, and puparium from Cuc Phuong, Nho Quan District, Vietnam). A, Periphallic organs (lateral view; inset: surstylus distal portion); B, phallic organs (lateral view); C, ditto (ventral view); D, female terminalia (lateral view); E, puparium (ventral habitus); F, caudal-most portion of puparium (ventral view); G, anterior spiracles of puparium (dorsal view); H, mouth hook; I, cephalopharyngeal skeleton. Scale bars: 0.1 mm in A–D and I, 1.0 mm in E." figureDoi="http://doi.org/10.5281/zenodo.5766442" httpUri="https://zenodo.org/record/5766442/files/figure.png" pageId="26" pageNumber="27">Fig. 18A</figureCitation>
|
||
). Median piece of subepandrial sclerite medially and anteriorly bilobed into long lobes. Cercus pubescent except for anterior margin and apical portion, slightly constricted ventrosubapically, with 34–36 setae all over and small setae on apicoventral margin (
|
||
<figureCitation id="1345F1BE45702E25FB1767B6B677B4F1" box="[1234,1339,1971,1997]" captionStart="FIGURE 18" captionStartId="27.[151,250,1424,1448]" captionTargetBox="[151,1435,181,1400]" captionTargetId="figure-18@27.[151,1435,181,1400]" captionTargetPageId="27" captionText="FIGURE 18. Colocasiomyia vieti Toda, sp. nov. (♂ holotype, ♀ paratype, and puparium from Cuc Phuong, Nho Quan District, Vietnam). A, Periphallic organs (lateral view; inset: surstylus distal portion); B, phallic organs (lateral view); C, ditto (ventral view); D, female terminalia (lateral view); E, puparium (ventral habitus); F, caudal-most portion of puparium (ventral view); G, anterior spiracles of puparium (dorsal view); H, mouth hook; I, cephalopharyngeal skeleton. Scale bars: 0.1 mm in A–D and I, 1.0 mm in E." figureDoi="http://doi.org/10.5281/zenodo.5766442" httpUri="https://zenodo.org/record/5766442/files/figure.png" pageId="26" pageNumber="27">Fig. 18A</figureCitation>
|
||
). Phallal sheath apically broadly round in lateral view (
|
||
<figureCitation id="1345F1BE45702E25FD5F67D2B04CB4CD" box="[666,768,2007,2033]" captionStart="FIGURE 18" captionStartId="27.[151,250,1424,1448]" captionTargetBox="[151,1435,181,1400]" captionTargetId="figure-18@27.[151,1435,181,1400]" captionTargetPageId="27" captionText="FIGURE 18. Colocasiomyia vieti Toda, sp. nov. (♂ holotype, ♀ paratype, and puparium from Cuc Phuong, Nho Quan District, Vietnam). A, Periphallic organs (lateral view; inset: surstylus distal portion); B, phallic organs (lateral view); C, ditto (ventral view); D, female terminalia (lateral view); E, puparium (ventral habitus); F, caudal-most portion of puparium (ventral view); G, anterior spiracles of puparium (dorsal view); H, mouth hook; I, cephalopharyngeal skeleton. Scale bars: 0.1 mm in A–D and I, 1.0 mm in E." figureDoi="http://doi.org/10.5281/zenodo.5766442" httpUri="https://zenodo.org/record/5766442/files/figure.png" pageId="26" pageNumber="27">Fig. 18B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF01BDB345712E24FF526595B257B504" ID-DOI="http://doi.org/10.5281/zenodo.5766442" ID-Zenodo-Dep="5766442" httpUri="https://zenodo.org/record/5766442/files/figure.png" pageId="27" pageNumber="28" startId="27.[151,250,1424,1448]" targetBox="[151,1435,181,1400]" targetPageId="27">
|
||
<paragraph id="8BC1ED3B45712E24FF526595B257B504" blockId="27.[151,1437,1424,1592]" pageId="27" pageNumber="28">
|
||
<emphasis id="B90A312945712E24FF526595B253B694" bold="true" box="[151,287,1424,1448]" pageId="27" pageNumber="28">FIGURE 18.</emphasis>
|
||
<taxonomicName id="4C7E96B845712E24FEE06595B165B694" authority="Toda" authorityName="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda" authorityYear="2021" box="[293,553,1424,1448]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="vieti" status="sp. nov.">
|
||
<emphasis id="B90A312945712E24FEE06595B2A3B694" box="[293,495,1424,1448]" italics="true" pageId="27" pageNumber="28">Colocasiomyia vieti</emphasis>
|
||
Toda
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B90A312945712E24FDF76595B1CEB694" bold="true" box="[562,642,1424,1448]" pageId="27" pageNumber="28">
|
||
<taxonomicNameLabel id="A2398C5245712E24FDF76595B1CEB694" box="[562,642,1424,1448]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(♂ holotype, ♀ paratype, and puparium from Cuc Phuong, Nho Quan District, Vietnam). A, Periphallic organs (lateral view; inset: surstylus distal portion); B, phallic organs (lateral view); C, ditto (ventral view); D, female terminalia (lateral view); E, puparium (ventral habitus); F, caudal-most portion of puparium (ventral view); G, anterior spiracles of puparium (dorsal view); H, mouth hook; I, cephalopharyngeal skeleton. Scale bars: 0.1 mm in A–D and I, 1.0 mm in E.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC1ED3B45712E24FF026663B0C1B5BD" blockId="27.[151,1437,1638,2025]" box="[199,909,1638,1665]" pageId="27" pageNumber="28">
|
||
Female terminalia. Oviscapt with 16–18 ovisensilla (
|
||
<figureCitation id="1345F1BE45712E24FCDD6663B0CCB5BD" box="[792,896,1638,1665]" captionStart="FIGURE 18" captionStartId="27.[151,250,1424,1448]" captionTargetBox="[151,1435,181,1400]" captionTargetId="figure-18@27.[151,1435,181,1400]" captionTargetPageId="27" captionText="FIGURE 18. Colocasiomyia vieti Toda, sp. nov. (♂ holotype, ♀ paratype, and puparium from Cuc Phuong, Nho Quan District, Vietnam). A, Periphallic organs (lateral view; inset: surstylus distal portion); B, phallic organs (lateral view); C, ditto (ventral view); D, female terminalia (lateral view); E, puparium (ventral habitus); F, caudal-most portion of puparium (ventral view); G, anterior spiracles of puparium (dorsal view); H, mouth hook; I, cephalopharyngeal skeleton. Scale bars: 0.1 mm in A–D and I, 1.0 mm in E." figureDoi="http://doi.org/10.5281/zenodo.5766442" httpUri="https://zenodo.org/record/5766442/files/figure.png" pageId="27" pageNumber="28">Fig. 18D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364BEB045712E24FF02668FB20AB42C" pageId="27" pageNumber="28" type="materials_examined">
|
||
<paragraph id="8BC1ED3B45712E24FF02668FB20AB42C" blockId="27.[151,1437,1638,2025]" pageId="27" pageNumber="28">
|
||
Indices (range of
|
||
<typeStatus id="54C5539945712E24FE4A668FB2BDB598" box="[399,497,1674,1700]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
|
||
and
|
||
<specimenCount id="9D7826B245712E24FDEC668EB100B599" box="[553,588,1675,1701]" pageId="27" pageNumber="28" type="male">3♂</specimenCount>
|
||
and
|
||
<specimenCount id="9D7826B245712E24FD41668EB1EBB599" box="[644,679,1675,1701]" pageId="27" pageNumber="28" type="female">3♀</specimenCount>
|
||
<typeStatus id="54C5539945712E24FD6A668EB06CB599" box="[687,800,1675,1701]" pageId="27" pageNumber="28" type="paratype">paratypes</typeStatus>
|
||
): FW/HW = 0.54–0.57, ch/o = 0.43–0.56, prorb = 0.86– 1.03, rcorb = 0.44–0.52, vb = 0.34–0.47, dcl = 0.50–0.73, sctl = 0.65–0.83, sterno = 0.58–0.78, orbito = 0.64–0.89, dcp = 0.63–0.94, sctlp = 0.62–0.95, C = 2.08–2.39, 4c = 0.89–1.04, 4v = 1.46–1.70, 5x = 0.90–1.13, ac = 2.81–3.14, M = 0.36–0.47.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364BEB045712E24FF02671FB61AB441" pageId="27" pageNumber="28" type="description">
|
||
<paragraph id="8BC1ED3B45712E24FF02671FB61AB441" blockId="27.[151,1437,1638,2025]" pageId="27" pageNumber="28">
|
||
Puparium (3rd instar larva). Anterior spiracle with stalk ending in a whorl of 9–11 branches; stalk longer than longest branch; caudal segments elongate, ending in Y-shaped posterior spiracles (
|
||
<figureCitation id="1345F1BE45712E24FBE8673BB7F9B465" box="[1069,1205,1854,1881]" captionStart="FIGURE 18" captionStartId="27.[151,250,1424,1448]" captionTargetBox="[151,1435,181,1400]" captionTargetId="figure-18@27.[151,1435,181,1400]" captionTargetPageId="27" captionText="FIGURE 18. Colocasiomyia vieti Toda, sp. nov. (♂ holotype, ♀ paratype, and puparium from Cuc Phuong, Nho Quan District, Vietnam). A, Periphallic organs (lateral view; inset: surstylus distal portion); B, phallic organs (lateral view); C, ditto (ventral view); D, female terminalia (lateral view); E, puparium (ventral habitus); F, caudal-most portion of puparium (ventral view); G, anterior spiracles of puparium (dorsal view); H, mouth hook; I, cephalopharyngeal skeleton. Scale bars: 0.1 mm in A–D and I, 1.0 mm in E." figureDoi="http://doi.org/10.5281/zenodo.5766442" httpUri="https://zenodo.org/record/5766442/files/figure.png" pageId="27" pageNumber="28">Fig. 18E–G</figureCitation>
|
||
). Mouth hook: distal blade as long as basal portion, apically pointed, gently curved downward, with small, acute teeth (
|
||
<figureCitation id="1345F1BE45712E24FB246767B605B441" box="[1249,1353,1890,1917]" captionStart="FIGURE 18" captionStartId="27.[151,250,1424,1448]" captionTargetBox="[151,1435,181,1400]" captionTargetId="figure-18@27.[151,1435,181,1400]" captionTargetPageId="27" captionText="FIGURE 18. Colocasiomyia vieti Toda, sp. nov. (♂ holotype, ♀ paratype, and puparium from Cuc Phuong, Nho Quan District, Vietnam). A, Periphallic organs (lateral view; inset: surstylus distal portion); B, phallic organs (lateral view); C, ditto (ventral view); D, female terminalia (lateral view); E, puparium (ventral habitus); F, caudal-most portion of puparium (ventral view); G, anterior spiracles of puparium (dorsal view); H, mouth hook; I, cephalopharyngeal skeleton. Scale bars: 0.1 mm in A–D and I, 1.0 mm in E." figureDoi="http://doi.org/10.5281/zenodo.5766442" httpUri="https://zenodo.org/record/5766442/files/figure.png" pageId="27" pageNumber="28">Fig. 18H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364BEB045712E24FF026782B0EAB4D5" pageId="27" pageNumber="28" type="materials_examined">
|
||
<paragraph id="8BC1ED3B45712E24FF026782B3B5B4F8" blockId="27.[151,1437,1638,2025]" pageId="27" pageNumber="28">
|
||
<materialsCitation id="3B16E76645712E24FF026782B3B5B4F8" ID-GBIF-Occurrence="3418896659" collectingDate="2000-04-02" collectionCode="SEHU" country="Vietnam" location="Cuc Phuong" municipality="Nho Quan District" pageId="27" pageNumber="28" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<emphasis id="B90A312945712E24FF026782B277B49D" bold="true" box="[199,315,1927,1953]" pageId="27" pageNumber="28">
|
||
<typeStatus id="54C5539945712E24FF026782B27AB49D" box="[199,310,1927,1953]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCount id="9D7826B245712E24FE806782B216B49D" box="[325,346,1927,1953]" pageId="27" pageNumber="28" type="male">♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED6F75FE45712E24FEA96782B2F7B49D" box="[364,443,1927,1953]" pageId="27" pageNumber="28">SEHU</collectionCode>
|
||
), “
|
||
<location id="8EA1BBE045712E24FE1F6782B127B49C" LSID="urn:lsid:plazi:treatment:03D75C2D45702E23FF526551B7A1B27D:8EA1BBE045712E24FE1F6782B127B49C" box="[474,619,1926,1953]" country="Vietnam" municipality="Nho Quan District" name="Cuc Phuong" pageId="27" pageNumber="28">Cuc Phuong</location>
|
||
,
|
||
<collectingMunicipality id="6BA5774145712E24FDBF6783B01DB49C" box="[634,849,1926,1953]" pageId="27" pageNumber="28">Nho Quan District</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="F369ADAB45712E24FC9B6783B08CB49C" box="[862,960,1926,1952]" name="Vietnam" pageId="27" pageNumber="28">Vietnam</collectingCountry>
|
||
,
|
||
<date id="FFC0CBFB45712E24FC0B6783B774B49C" box="[974,1080,1926,1952]" pageId="27" pageNumber="28" value="2000-04-02">
|
||
<collectingDate id="EF84321345712E24FC0B6783B774B49C" box="[974,1080,1926,1952]" pageId="27" pageNumber="28" value="2000-04-02">2.iv.2000</collectingDate>
|
||
</date>
|
||
, ex
|
||
<taxonomicName id="4C7E96B845712E24FBAE6783B3A0B4F8" authority=", M. Yafuso" authorityName="M. Yafuso" class="Liliopsida" family="Araceae" genus="Alocasia" kingdom="Plantae" order="Alismatales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="atropurpurea">
|
||
<emphasis id="B90A312945712E24FBAE6783B620B49C" box="[1131,1388,1926,1952]" italics="true" pageId="27" pageNumber="28">Alocasia atropurpurea</emphasis>
|
||
, M. Yafuso
|
||
</taxonomicName>
|
||
”.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BC1ED3B45712E24FF0267CAB0EAB4D5" blockId="27.[151,1437,1638,2025]" box="[199,934,1998,2025]" pageId="27" pageNumber="28">
|
||
<materialsCitation id="3B16E76645712E24FF0267CAB0EEB4D5" ID-GBIF-Occurrence="3418896443" box="[199,930,1998,2025]" collectingDate="2000-04-02" collectionCode="SEHU" country="Vietnam" location="Cuc Phuong" municipality="Nho Quan District" pageId="27" pageNumber="28" specimenCount="6" specimenCount-female="3" specimenCount-male="3" typeStatus="paratype">
|
||
<emphasis id="B90A312945712E24FF0267CAB20AB4D5" bold="true" box="[199,326,1999,2025]" pageId="27" pageNumber="28">
|
||
<typeStatus id="54C5539945712E24FF0267CAB20DB4D5" box="[199,321,1999,2025]" pageId="27" pageNumber="28" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectingCountry id="F369ADAB45712E24FE8967CBB2E3B4D4" box="[332,431,1998,2024]" name="Vietnam" pageId="27" pageNumber="28">Vietnam</collectingCountry>
|
||
:
|
||
<specimenCount id="9D7826B245712E24FE7F67CAB293B4D5" box="[442,479,1999,2025]" pageId="27" pageNumber="28" type="male">3♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D7826B245712E24FE2E67CAB143B4D5" box="[491,527,1999,2025]" pageId="27" pageNumber="28" type="female">3♀</specimenCount>
|
||
, same data as the holotype (
|
||
<collectionCode id="ED6F75FE45712E24FC8E67CAB0D6B4D5" box="[843,922,1999,2025]" pageId="27" pageNumber="28">SEHU</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364BEB045762E23FF026093B281B38D" box="[199,461,150,177]" pageId="28" pageNumber="29" type="distribution">
|
||
<paragraph id="8BC1ED3B45762E23FF026093B281B38D" blockId="28.[151,1436,150,321]" box="[199,461,150,177]" pageId="28" pageNumber="29">
|
||
<emphasis id="B90A312945762E23FF026093B22DB38C" bold="true" box="[199,353,150,176]" pageId="28" pageNumber="29">Distribution.</emphasis>
|
||
<collectingCountry id="F369ADAB45762E23FEA26092B286B38D" box="[359,458,151,177]" name="Vietnam" pageId="28" pageNumber="29">Vietnam</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364BEB045762E23FF0260BEB21DB221" pageId="28" pageNumber="29" type="discussion">
|
||
<paragraph id="8BC1ED3B45762E23FF0260BEB21DB221" blockId="28.[151,1436,150,321]" pageId="28" pageNumber="29">
|
||
<emphasis id="B90A312945762E23FF0260BEB277B3E9" bold="true" box="[199,315,187,213]" pageId="28" pageNumber="29">Remarks.</emphasis>
|
||
This species resembles
|
||
<taxonomicName id="4C7E96B845762E23FD8960BEB188B3E9" authority="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda, 2021" authorityName="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda" authorityYear="2021" box="[588,708,187,213]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
|
||
<emphasis id="B90A312945762E23FD8960BEB113B3E9" box="[588,607,187,213]" italics="true" pageId="28" pageNumber="29">C</emphasis>
|
||
.
|
||
<emphasis id="B90A312945762E23FDAB60BEB188B3E9" box="[622,708,187,213]" italics="true" pageId="28" pageNumber="29">grandis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90A312945762E23FD0E60BEB06AB3E9" bold="true" box="[715,806,187,213]" pageId="28" pageNumber="29">
|
||
<taxonomicNameLabel id="A2398C5245762E23FD0E60BEB06AB3E9" box="[715,806,187,213]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in many characters, but can be distinguished from it by the diagnostic character and the absence of patch covered with only minute pubescence on anterodorsal portion of fore tarsomere I.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364BEB045762E23FF026122B7A1B27D" box="[199,1261,295,321]" pageId="28" pageNumber="29" type="etymology">
|
||
<paragraph id="8BC1ED3B45762E23FF026122B7A1B27D" blockId="28.[151,1436,150,321]" box="[199,1261,295,321]" pageId="28" pageNumber="29">
|
||
<emphasis id="B90A312945762E23FF026122B202B27D" bold="true" box="[199,334,295,321]" pageId="28" pageNumber="29">Etymology.</emphasis>
|
||
Patronym, dedicated to Dr. B. T. Viet who helped MY with field work in
|
||
<collectingCountry id="F369ADAB45762E23FB426122B7A6B27D" box="[1159,1258,295,321]" name="Vietnam" pageId="28" pageNumber="29">Vietnam</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |