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<document id="4ACD6B557C5CE063613967692982AF7A" ID-DOI="10.5281/zenodo.205759" ID-GBIF-Dataset="166f9754-caf3-479a-b4e5-d5dfdd382f31" ID-ISSN="1175-5326" ID-Zenodo-Dep="205759" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460246985593" checkinUser="plazi" docAuthor="Felippe-Bauer, Maria Luiza, Silva, Tiago Do Nascimento Da, Ribeiro, Enaly Silva &amp; Borkent, Art" docDate="2011" docId="03A487B0FFE29673BB8C0954FAA821B5" docLanguage="en" docName="zt02780p028.pdf" docOrigin="Zootaxa 2780" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Downeshelea lanei Felippe-Bauer &amp; Borkent, n. sp." docType="treatment" docVersion="9" lastPageNumber="25" masterDocId="FF9DFFC8FFE39676BB1B083EFF9D2344" masterDocTitle="A new species of Downeshelea Wirth &amp; Grogan and a redescription of the male of Downeshelea cebacoi (Lane &amp; Wirth) (Diptera: Ceratopogonidae)" masterLastPageNumber="28" masterPageNumber="20" pageNumber="21" updateTime="1698242004948" updateUser="plazi">
<mods:mods id="4B20EC6AD4305174DBCEBACEB0C9ACCE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E6AB7CA426241C99395EE92D3CF9D939">
<mods:title id="EE9235A45B33141A2E98E7ADB2912F66">A new species of Downeshelea Wirth &amp; Grogan and a redescription of the male of Downeshelea cebacoi (Lane &amp; Wirth) (Diptera: Ceratopogonidae)</mods:title>
</mods:titleInfo>
<mods:name id="1DCF0AD2B9255F4CD549A6C0E96DB16D" type="personal">
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<mods:namePart id="8EE3FD514DD88A17912C152730EC5323">Felippe-Bauer, Maria Luiza</mods:namePart>
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<mods:namePart id="B457579025C788BEE6BF5F0248C24A4F">Silva, Tiago Do Nascimento Da</mods:namePart>
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<mods:namePart id="F894E6C61E3733DDC1847F383FC763C2">Ribeiro, Enaly Silva</mods:namePart>
</mods:name>
<mods:name id="059CF9E0EB9FF136E9D9AB0DCB01813A" type="personal">
<mods:role id="F8012E1F4B9A5C08123C1FB68928C0F7">
<mods:roleTerm id="0583EA25908E9C5FB330B9448E65AEB1">Author</mods:roleTerm>
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<mods:namePart id="33264E6A869239C67CB8B3C9A35EB988">Borkent, Art</mods:namePart>
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<mods:title id="50A12512001CD0B22B597FE876E86263">Zootaxa</mods:title>
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<mods:part id="C7B41EAF8BD993AA69F7AFE5E059B9E4">
<mods:date id="13A0BB93CB835284AD7983D0E484245D">2011</mods:date>
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<mods:number id="BEB8578AD8A40D266BE2EE334F0C8C4B">2780</mods:number>
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<mods:classification id="ED577CE10227114FA104D3DEDA01FFF2">journal article</mods:classification>
<mods:identifier id="2A05F1C1820A64E903322E4CC476D8DA" type="DOI">10.5281/zenodo.205759</mods:identifier>
<mods:identifier id="59800F427FBA5F9FBE1B03510F695490" type="GBIF-Dataset">166f9754-caf3-479a-b4e5-d5dfdd382f31</mods:identifier>
<mods:identifier id="0E0DEB2459CBB690BB860F7F6E5F3609" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="64BB9B7F278195680F6194EB54D98D46" type="Zenodo-Dep">205759</mods:identifier>
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<treatment id="03A487B0FFE29673BB8C0954FAA821B5" ID-DOI="http://doi.org/10.5281/zenodo.6191815" ID-GBIF-Taxon="119390625" ID-Zenodo-Dep="6191815" LSID="urn:lsid:plazi:treatment:03A487B0FFE29673BB8C0954FAA821B5" httpUri="http://treatment.plazi.org/id/03A487B0FFE29673BB8C0954FAA821B5" lastPageId="5" lastPageNumber="25" pageId="1" pageNumber="21">
<subSubSection id="C317652DFFE29677BB8C0954FE9022E1" pageId="1" pageNumber="21" type="nomenclature">
<paragraph id="8BB236A6FFE29677BB8C0954FC8922C0" blockId="1.[151,788,362,421]" box="[151,788,362,388]" pageId="1" pageNumber="21">
<heading id="D0FA81CAFFE29677BB8C0954FC8922C0" bold="true" box="[151,788,362,388]" fontSize="11" level="1" pageId="1" pageNumber="21" reason="1">
<emphasis id="B979EAB4FFE29677BB8C0954FC8922C0" bold="true" box="[151,788,362,388]" pageId="1" pageNumber="21">
<taxonomicName id="4C0D4D25FFE29677BB8C0954FD5822C0" ID-CoL="8BVKB" authority="Felippe-Bauer &amp; Borkent" authorityName="Felippe-Bauer &amp; Borkent" box="[151,709,362,388]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="21" phylum="Arthropoda" rank="species" species="lanei" status="sp. nov.">
<emphasis id="B979EAB4FFE29677BB8C0954FEE622C0" bold="true" box="[151,379,362,388]" italics="true" pageId="1" pageNumber="21">Downeshelea lanei</emphasis>
Felippe-Bauer &amp; Borkent
</taxonomicName>
,
<taxonomicNameLabel id="A24A57CFFFE29677B9C90954FC8922C0" box="[722,788,362,388]" pageId="1" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB236A6FFE29677BB8C09B3FE9022E1" blockId="1.[151,788,362,421]" box="[151,269,397,421]" pageId="1" pageNumber="21">
<figureCitation id="13362A23FFE29677BB8C09B3FE9022E1" box="[151,269,397,421]" captionStart-0="FIGURES 1 – 2" captionStart-1="FIGURES 3 – 8" captionStart-2="FIGURES 9 – 12" captionStart-3="FIGURES 13 – 14" captionStartId-0="1.[151,264,1624,1647]" captionStartId-1="2.[151,264,1975,1998]" captionStartId-2="3.[151,264,1665,1688]" captionStartId-3="4.[151,264,2009,2032]" captionTargetBox-0="[253,1283,717,1573]" captionTargetBox-1="[251,1331,598,1932]" captionTargetBox-2="[216,1363,198,1631]" captionTargetBox-3="[476,1119,665,1969]" captionTargetId-0="figure@1.[245,1336,666,1607]" captionTargetId-1="figure@2.[236,1350,580,1952]" captionTargetId-2="figure@3.[212,1376,193,1644]" captionTargetId-3="figure@4.[456,1130,644,1988]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionText-0="FIGURES 1 – 2. Wing photographs of Downeshelea lanei n. sp. 1. Female; 2. Male." captionText-1="FIGURES 3 – 8. Downeshelea lanei n. sp., female. 3. Antenna; 4. Dorsal portion of head capsule, anterior view; 5. Palpus; 6, Spermathecae; 7. Femora and tibiae (left to right) of fore-, mid- and hindlegs; 8. Tarsi (left to right) of fore-, mid- and hindlegs." captionText-2="FIGURES 9 – 12. Downeshelea lanei n. sp., male. 9. Antennal flagellum; 10. Palpus; 11. Tarsi (top to bottom) of fore-, mid- and hindlegs; 12. Femora and tibiae (left to right) of fore-, mid- and hindlegs." captionText-3="FIGURES 13 – 14. Downeshelea lanei n. sp., male. 13. Terminalia, parameres removed; 14. Parameres." httpUri-0="https://zenodo.org/record/205760/files/figure.png" httpUri-1="https://zenodo.org/record/205761/files/figure.png" httpUri-2="https://zenodo.org/record/205762/files/figure.png" httpUri-3="https://zenodo.org/record/205763/files/figure.png" pageId="1" pageNumber="21">Figs. 1–14</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C317652DFFE29677BB8C09EEFC7022A2" box="[151,1005,464,486]" pageId="1" pageNumber="21" type="reference_group">
<paragraph id="8BB236A6FFE29677BB8C09EEFC7022A2" blockId="1.[151,1005,464,486]" box="[151,1005,464,486]" pageId="1" pageNumber="21">
<treatmentCitationGroup id="AB1D1188FFE29677BB8C09EEFC7022A2" box="[151,1005,464,486]" pageId="1" pageNumber="21">
<taxonomicName id="4C0D4D25FFE29677BB8C09EEFDEB22A1" ID-CoL="8CWDD" authority="Lane and Wirth 1964: 218" authorityName="Lane and Wirth" authorityPageNumber="218" authorityYear="1964" box="[151,630,464,486]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="21" phylum="Arthropoda" rank="species" species="cebacoi">
<emphasis id="B979EAB4FFE29677BB8C09EEFEC322A2" box="[151,350,464,486]" italics="true" pageId="1" pageNumber="21">Monohelea cebacoi</emphasis>
:
<treatmentCitation id="0AAC10B7FFE29677BA7009EFFDEB22A1" author="Lane" box="[363,630,464,486]" page="218" pageId="1" pageNumber="21" year="1964">
<bibRefCitation id="EF9C4B57FFE29677BA7009EFFDDB22A2" author="Lane" box="[363,582,464,486]" pageId="1" pageNumber="21" refString="Lane, J. &amp; Wirth, W. W. (1964) The biting midge genus Monohelea Kieffer in the Neotropical Region (Diptera, Ceratopogonidae). Studia Entomologica, 7, 209 - 236." type="journal article" year="1964">Lane and Wirth 1964</bibRefCitation>
: 218
</treatmentCitation>
</taxonomicName>
(in part; male, Brazilian specimens).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C317652DFFE29677BB8C0A31FD222136" pageId="1" pageNumber="21" type="diagnosis">
<paragraph id="8BB236A6FFE29677BB8C0A31FD222136" blockId="1.[151,1436,527,626]" pageId="1" pageNumber="21">
<emphasis id="B979EAB4FFE29677BB8C0A31FE8C216C" bold="true" box="[151,273,527,552]" pageId="1" pageNumber="21">Diagnosis.</emphasis>
Only species of
<taxonomicName id="4C0D4D25FFE29677BAD40A2EFDFB216C" box="[463,614,528,552]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAB4FFE29677BAD40A2EFDFB216C" box="[463,614,528,552]" italics="true" pageId="1" pageNumber="21">Downeshelea</emphasis>
</taxonomicName>
with dark spots on the apical portion of r3 and m1 (
<figureCitation id="13362A23FFE29677BFBC0A31FA8E2163" box="[1191,1299,527,551]" captionStart="FIGURES 1 – 2" captionStartId="1.[151,264,1624,1647]" captionTargetBox="[253,1283,717,1573]" captionTargetId="figure@1.[245,1336,666,1607]" captionTargetPageId="1" captionText="FIGURES 1 – 2. Wing photographs of Downeshelea lanei n. sp. 1. Female; 2. Male." httpUri="https://zenodo.org/record/205760/files/figure.png" pageId="1" pageNumber="21">Figs. 1, 2</figureCitation>
), dark areas on the subapical portion of the fore- and midfemora (
<figureCitation id="13362A23FFE29677B9F60A09FCAC210B" box="[749,817,567,591]" captionStart="FIGURES 3 – 8" captionStartId="2.[151,264,1975,1998]" captionTargetBox="[251,1331,598,1932]" captionTargetId="figure@2.[236,1350,580,1952]" captionTargetPageId="2" captionText="FIGURES 3 – 8. Downeshelea lanei n. sp., female. 3. Antenna; 4. Dorsal portion of head capsule, anterior view; 5. Palpus; 6, Spermathecae; 7. Femora and tibiae (left to right) of fore-, mid- and hindlegs; 8. Tarsi (left to right) of fore-, mid- and hindlegs." httpUri="https://zenodo.org/record/205761/files/figure.png" pageId="1" pageNumber="21">Fig. 7</figureCitation>
) and the mid portion of each paramere straight and the distal portion sharply directed medially (
<figureCitation id="13362A23FFE29677B9450A64FD2F2136" box="[606,690,602,626]" captionStart="FIGURES 13 – 14" captionStartId="4.[151,264,2009,2032]" captionTargetBox="[476,1119,665,1969]" captionTargetId="figure@4.[456,1130,644,1988]" captionTargetPageId="4" captionText="FIGURES 13 – 14. Downeshelea lanei n. sp., male. 13. Terminalia, parameres removed; 14. Parameres." httpUri="https://zenodo.org/record/205763/files/figure.png" pageId="1" pageNumber="21">Fig. 14</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF72662EFFE29677BB8C0E66FC74252A" box="[151,1001,1624,1647]" httpUri="https://zenodo.org/record/205760/files/figure.png" pageId="1" pageNumber="21" targetBox="[253,1283,717,1573]" targetPageId="1">
<paragraph id="8BB236A6FFE29677BB8C0E66FC74252A" blockId="1.[151,1001,1624,1647]" box="[151,1001,1624,1647]" pageId="1" pageNumber="21">
<emphasis id="B979EAB4FFE29677BB8C0E66FEA7252A" bold="true" box="[151,314,1624,1647]" pageId="1" pageNumber="21">FIGURES 1–2.</emphasis>
Wing photographs of
<taxonomicName id="4C0D4D25FFE29677B9070E66FD43252A" box="[540,734,1624,1646]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="21" phylum="Arthropoda" rank="species" species="lanei" status="sp. nov.">
<emphasis id="B979EAB4FFE29677B9070E66FD43252A" box="[540,734,1624,1646]" italics="true" pageId="1" pageNumber="21">Downeshelea lanei</emphasis>
</taxonomicName>
<emphasis id="B979EAB4FFE29677B9FF0E67FC83252A" bold="true" box="[740,798,1625,1646]" pageId="1" pageNumber="21">
<taxonomicNameLabel id="A24A57CFFFE29677B9FF0E67FC83252A" box="[740,798,1625,1646]" pageId="1" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
1. Female; 2. Male.
</paragraph>
</caption>
<subSubSection id="C317652DFFE29672BBDE0EA2FF7922B1" lastPageId="4" lastPageNumber="24" pageId="1" pageNumber="21" type="description">
<paragraph id="8BB236A6FFE29677BBDE0EA2FB18245F" blockId="1.[151,1436,1691,2012]" pageId="1" pageNumber="21">
<emphasis id="B979EAB4FFE29677BBDE0EA2FEBE25F0" bold="true" box="[197,291,1692,1716]" pageId="1" pageNumber="21">Female.</emphasis>
Head. Brown. Eyes (
<figureCitation id="13362A23FFE29677B9030EA5FDC125F7" box="[536,604,1691,1715]" captionStart="FIGURES 3 – 8" captionStartId="2.[151,264,1975,1998]" captionTargetBox="[251,1331,598,1932]" captionTargetId="figure@2.[236,1350,580,1952]" captionTargetPageId="2" captionText="FIGURES 3 – 8. Downeshelea lanei n. sp., female. 3. Antenna; 4. Dorsal portion of head capsule, anterior view; 5. Palpus; 6, Spermathecae; 7. Femora and tibiae (left to right) of fore-, mid- and hindlegs; 8. Tarsi (left to right) of fore-, mid- and hindlegs." httpUri="https://zenodo.org/record/205761/files/figure.png" pageId="1" pageNumber="21">Fig. 4</figureCitation>
) bare, narrowly contiguous. Antenna (
<figureCitation id="13362A23FFE29677BF080EA5FBCB25F7" box="[1043,1110,1691,1715]" captionStart="FIGURES 3 – 8" captionStartId="2.[151,264,1975,1998]" captionTargetBox="[251,1331,598,1932]" captionTargetId="figure@2.[236,1350,580,1952]" captionTargetPageId="2" captionText="FIGURES 3 – 8. Downeshelea lanei n. sp., female. 3. Antenna; 4. Dorsal portion of head capsule, anterior view; 5. Palpus; 6, Spermathecae; 7. Femora and tibiae (left to right) of fore-, mid- and hindlegs; 8. Tarsi (left to right) of fore-, mid- and hindlegs." httpUri="https://zenodo.org/record/205761/files/figure.png" pageId="1" pageNumber="21">Fig. 3</figureCitation>
) brown except on basal portion of flagellomeres; flagellomeres cylindrical, elongate; antennal ratio 0.97–1.15 (1.06; n=3); palpus (
<figureCitation id="13362A23FFE29677BE060E80FAC22592" box="[1309,1375,1726,1750]" captionStart="FIGURES 3 – 8" captionStartId="2.[151,264,1975,1998]" captionTargetBox="[251,1331,598,1932]" captionTargetId="figure@2.[236,1350,580,1952]" captionTargetPageId="2" captionText="FIGURES 3 – 8. Downeshelea lanei n. sp., female. 3. Antenna; 4. Dorsal portion of head capsule, anterior view; 5. Palpus; 6, Spermathecae; 7. Femora and tibiae (left to right) of fore-, mid- and hindlegs; 8. Tarsi (left to right) of fore-, mid- and hindlegs." httpUri="https://zenodo.org/record/205761/files/figure.png" pageId="1" pageNumber="21">Fig. 5</figureCitation>
) uniformly brown, longer than proboscis; segment 3 cylindrical, proximal 2/3 slightly swollen, with moderately broad, deep sensory pit at midlength; palpal ratio 2.5–2.7 (2.6; n=3). Mandible with 10–12 teeth.
</paragraph>
<paragraph id="8BB236A6FFE29674BBDE0F18FEA22154" blockId="1.[151,1436,1691,2012]" lastBlockId="2.[151,1437,151,528]" lastPageId="2" lastPageNumber="22" pageId="1" pageNumber="21">
Thorax. Brown, without definite pattern in slide mounted specimens. Wing (
<figureCitation id="13362A23FFE29677BF2F0F18FBEB247A" box="[1076,1142,1830,1854]" captionStart="FIGURES 1 – 2" captionStartId="1.[151,264,1624,1647]" captionTargetBox="[253,1283,717,1573]" captionTargetId="figure@1.[245,1336,666,1607]" captionTargetPageId="1" captionText="FIGURES 1 – 2. Wing photographs of Downeshelea lanei n. sp. 1. Female; 2. Male." httpUri="https://zenodo.org/record/205760/files/figure.png" pageId="1" pageNumber="21">Fig. 1</figureCitation>
) hyaline, with dark thick setae on costa, scarce apical microtrichia; two conspicuous black spots, one located on r-m extending posteriorly to medial fork and M2, the other in r3, extending posteriorly from end of 2nd radial cell to M1; four conspicuous grayish spots: 1st in apical portion of r3, 2nd near apex of M1 extending to wing margin in m1, 3rd sigmoid shaped near apex of M2 extending to wing margin in m2, 4th with markings on fork of CuA1 and CuA2 extending on these veins, extending into cua1 and anal cell; 2nd radial cell nearly twice as long as 1st; wing length 1.35–1.54 (1.43; n=3) mm; breadth 0.54–0.65 (0.59; n=3) mm; costal ratio 0.79–0.82 (0.80, n=3). Halter stem pale, knob brown. Legs with femora, tibiae (
<figureCitation id="13362A23FFE19674BBDC08DCFE9623BE" box="[199,267,226,250]" captionStart="FIGURES 3 – 8" captionStartId="2.[151,264,1975,1998]" captionTargetBox="[251,1331,598,1932]" captionTargetId="figure@2.[236,1350,580,1952]" captionTargetPageId="2" captionText="FIGURES 3 – 8. Downeshelea lanei n. sp., female. 3. Antenna; 4. Dorsal portion of head capsule, anterior view; 5. Palpus; 6, Spermathecae; 7. Femora and tibiae (left to right) of fore-, mid- and hindlegs; 8. Tarsi (left to right) of fore-, mid- and hindlegs." httpUri="https://zenodo.org/record/205761/files/figure.png" pageId="2" pageNumber="22">Fig. 7</figureCitation>
) yellowish brown, fore- and midfemur slightly darker on subapical portions, hind femur darker on subapical third, hind tibia darker on proximal half and apical portion; femorotibial joint areas yellowish; fore- and hind tibiae with apical spur, longer on foreleg; hind tibial comb with 6–7 large setae; lengths of trochanters, femora, tibiae of fore-, mid-, hind legs: 135-595-568, 108-730-676, 135-838-757 Μm (n=3). Tarsi (
<figureCitation id="13362A23FFE19674BF9A0974FB582226" box="[1153,1221,330,354]" captionStart="FIGURES 3 – 8" captionStartId="2.[151,264,1975,1998]" captionTargetBox="[251,1331,598,1932]" captionTargetId="figure@2.[236,1350,580,1952]" captionTargetPageId="2" captionText="FIGURES 3 – 8. Downeshelea lanei n. sp., female. 3. Antenna; 4. Dorsal portion of head capsule, anterior view; 5. Palpus; 6, Spermathecae; 7. Femora and tibiae (left to right) of fore-, mid- and hindlegs; 8. Tarsi (left to right) of fore-, mid- and hindlegs." httpUri="https://zenodo.org/record/205761/files/figure.png" pageId="2" pageNumber="22">Fig. 8</figureCitation>
) pale, pilose; palisade setae in single row on hind tarsomere 1; fore-, hind tarsomere 1 with one basal, one apical spine; mid tarsomere 1 with 2 basal, 2 apical, 1–3 other ventral spines; apical spines of tarsomeres 2–4 of fore-, mid-, hind legs: 2-2-2, 2-2-2, 1-1-2, basal spines absent; fore-, mid-, hind tarsal ratios: 2.3, 2.5, 2.5 (n=3); claws of fore-, midlegs paired, equal, about 0.7 length of their respective tarsomeres 5; hind leg with single claw, about 1.1 x as long as tarsomere 5.
</paragraph>
<caption id="DF72662EFFE19674BB8C0F89FA0A24A8" httpUri="https://zenodo.org/record/205761/files/figure.png" pageId="2" pageNumber="22" targetBox="[251,1331,598,1932]" targetPageId="2">
<paragraph id="8BB236A6FFE19674BB8C0F89FA0A24A8" blockId="2.[151,1436,1975,2028]" pageId="2" pageNumber="22">
<emphasis id="B979EAB4FFE19674BB8C0F89FEA62489" bold="true" box="[151,315,1975,1998]" pageId="2" pageNumber="22">FIGURES 3–8.</emphasis>
<taxonomicName id="4C0D4D25FFE19674BA590F89FD992489" box="[322,516,1975,1997]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="22" phylum="Arthropoda" rank="species" species="lanei" status="sp. nov.">
<emphasis id="B979EAB4FFE19674BA590F89FD992489" box="[322,516,1975,1997]" italics="true" pageId="2" pageNumber="22">Downeshelea lanei</emphasis>
</taxonomicName>
<emphasis id="B979EAB4FFE19674B9170F86FDDB2489" bold="true" box="[524,582,1976,1997]" pageId="2" pageNumber="22">
<taxonomicNameLabel id="A24A57CFFFE19674B9170F86FDDB2489" box="[524,582,1976,1997]" pageId="2" pageNumber="22" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female. 3. Antenna; 4. Dorsal portion of head capsule, anterior view; 5. Palpus; 6, Spermathecae; 7. Femora and tibiae (left to right) of fore-, mid- and hindlegs; 8. Tarsi (left to right) of fore-, mid- and hindlegs.
</paragraph>
</caption>
<caption id="DF72662EFFE09675BB8C0EBFFC3525F2" httpUri="https://zenodo.org/record/205762/files/figure.png" pageId="3" pageNumber="23" targetBox="[216,1363,198,1631]" targetPageId="3">
<paragraph id="8BB236A6FFE09675BB8C0EBFFC3525F2" blockId="3.[151,1436,1665,1718]" pageId="3" pageNumber="23">
<emphasis id="B979EAB4FFE09675BB8C0EBFFED525D3" bold="true" box="[151,328,1665,1688]" pageId="3" pageNumber="23">FIGURES 9–12.</emphasis>
<taxonomicName id="4C0D4D25FFE09675BA4B0EBFFD8E25D3" box="[336,531,1665,1687]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="23" phylum="Arthropoda" rank="species" species="lanei" status="sp. nov.">
<emphasis id="B979EAB4FFE09675BA4B0EBFFD8E25D3" box="[336,531,1665,1687]" italics="true" pageId="3" pageNumber="23">Downeshelea lanei</emphasis>
</taxonomicName>
<emphasis id="B979EAB4FFE09675B9010EBCFDC825D3" bold="true" box="[538,597,1666,1687]" pageId="3" pageNumber="23">
<taxonomicNameLabel id="A24A57CFFFE09675B9010EBCFDC825D3" box="[538,597,1666,1687]" pageId="3" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male. 9. Antennal flagellum; 10. Palpus; 11. Tarsi (top to bottom) of fore-, mid- and hindlegs; 12. Femora and tibiae (left to right) of fore-, mid- and hindlegs.
</paragraph>
</caption>
<paragraph id="8BB236A6FFE09675BBDE0EDCFD512465" blockId="3.[151,1436,1762,2005]" pageId="3" pageNumber="23">
Abdomen. Brown. Two slightly unequal subspherical spermathecae (
<figureCitation id="13362A23FFE09675B8DD0EDCFB9725BE" box="[966,1034,1762,1786]" captionStart="FIGURES 3 – 8" captionStartId="2.[151,264,1975,1998]" captionTargetBox="[251,1331,598,1932]" captionTargetId="figure@2.[236,1350,580,1952]" captionTargetPageId="2" captionText="FIGURES 3 – 8. Downeshelea lanei n. sp., female. 3. Antenna; 4. Dorsal portion of head capsule, anterior view; 5. Palpus; 6, Spermathecae; 7. Femora and tibiae (left to right) of fore-, mid- and hindlegs; 8. Tarsi (left to right) of fore-, mid- and hindlegs." httpUri="https://zenodo.org/record/205761/files/figure.png" pageId="3" pageNumber="23">Fig. 6</figureCitation>
), measuring 64 by 58 Μm and 50 by 48 Μm; small rudimentary 3rd present, 10 Μm long.
</paragraph>
<paragraph id="8BB236A6FFE09672BBDE0F10FF7922B1" blockId="3.[151,1436,1762,2005]" lastBlockId="4.[151,1437,151,611]" lastPageId="4" lastPageNumber="24" pageId="3" pageNumber="23">
<emphasis id="B979EAB4FFE09675BBDE0F10FE942402" bold="true" box="[197,265,1838,1862]" pageId="3" pageNumber="23">Male.</emphasis>
Similar to female with the usual sexual differences; antenna (
<figureCitation id="13362A23FFE09675B8A40F13FB9E2401" box="[959,1027,1837,1861]" captionStart="FIGURES 9 – 12" captionStartId="3.[151,264,1665,1688]" captionTargetBox="[216,1363,198,1631]" captionTargetId="figure@3.[212,1376,193,1644]" captionTargetPageId="3" captionText="FIGURES 9 – 12. Downeshelea lanei n. sp., male. 9. Antennal flagellum; 10. Palpus; 11. Tarsi (top to bottom) of fore-, mid- and hindlegs; 12. Femora and tibiae (left to right) of fore-, mid- and hindlegs." httpUri="https://zenodo.org/record/205762/files/figure.png" pageId="3" pageNumber="23">Fig. 9</figureCitation>
) with pedicel and flagellomeres 11– 13 brown; 2–9 somewhat barrel-shaped, 10 about
<date id="FFB31066FFE09675B9D40F6FFD77242D" box="[719,746,1873,1897]" pageId="3" pageNumber="23">3x</date>
longer than wide, 11–13 elongate; antennal ratio 0.92 (n=2); palpus as in
<figureCitation id="13362A23FFE09675BA380F4BFEE824C9" box="[291,373,1909,1933]" captionStart="FIGURES 9 – 12" captionStartId="3.[151,264,1665,1688]" captionTargetBox="[216,1363,198,1631]" captionTargetId="figure@3.[212,1376,193,1644]" captionTargetPageId="3" captionText="FIGURES 9 – 12. Downeshelea lanei n. sp., male. 9. Antennal flagellum; 10. Palpus; 11. Tarsi (top to bottom) of fore-, mid- and hindlegs; 12. Femora and tibiae (left to right) of fore-, mid- and hindlegs." httpUri="https://zenodo.org/record/205762/files/figure.png" pageId="3" pageNumber="23">Fig. 10</figureCitation>
, palpal ratio 2.8. Legs with femora, tibiae as in
<figureCitation id="13362A23FFE09675B88B0F4BFC7E24C9" box="[912,995,1909,1933]" captionStart="FIGURES 9 – 12" captionStartId="3.[151,264,1665,1688]" captionTargetBox="[216,1363,198,1631]" captionTargetId="figure@3.[212,1376,193,1644]" captionTargetPageId="3" captionText="FIGURES 9 – 12. Downeshelea lanei n. sp., male. 9. Antennal flagellum; 10. Palpus; 11. Tarsi (top to bottom) of fore-, mid- and hindlegs; 12. Femora and tibiae (left to right) of fore-, mid- and hindlegs." httpUri="https://zenodo.org/record/205762/files/figure.png" pageId="3" pageNumber="23">Fig. 12</figureCitation>
. Tarsi (
<figureCitation id="13362A23FFE09675BF2E0F4BFB1824C9" box="[1077,1157,1909,1933]" captionStart="FIGURES 9 – 12" captionStartId="3.[151,264,1665,1688]" captionTargetBox="[216,1363,198,1631]" captionTargetId="figure@3.[212,1376,193,1644]" captionTargetPageId="3" captionText="FIGURES 9 – 12. Downeshelea lanei n. sp., male. 9. Antennal flagellum; 10. Palpus; 11. Tarsi (top to bottom) of fore-, mid- and hindlegs; 12. Femora and tibiae (left to right) of fore-, mid- and hindlegs." httpUri="https://zenodo.org/record/205762/files/figure.png" pageId="3" pageNumber="23">Fig. 11</figureCitation>
) pale, pilose; fore-, hind tarsomere 1 each with one basal and one apical spine; midleg tarsomere 1 with 2 basal, 2 apical, 1–5 other ventral spines; apical spines of tarsomeres 2–4 of fore-, mid-, hind legs: 2-2-2, 2-2-2, 1-1-1, basal spines absent; fore-, mid-, hind tarsal ratios: 2.2, 2.5, 2.4. Wing (
<figureCitation id="13362A23FFE79672B9BA08A9FD7523EB" box="[673,744,151,175]" captionStart="FIGURES 1 – 2" captionStartId="1.[151,264,1624,1647]" captionTargetBox="[253,1283,717,1573]" captionTargetId="figure@1.[245,1336,666,1607]" captionTargetPageId="1" captionText="FIGURES 1 – 2. Wing photographs of Downeshelea lanei n. sp. 1. Female; 2. Male." httpUri="https://zenodo.org/record/205760/files/figure.png" pageId="4" pageNumber="24">Fig. 2</figureCitation>
) hyaline, pattern of dark spots as in female, narrower than female; wing length 1.32–1.35 (1.33, n=2) mm; breadth 0.43 (n=2) mm; costal ratio 0.70–0.75 (0.73, n=2). Terminalia (
<figureCitation id="13362A23FFE79672BBC508E1FEAF23B3" box="[222,306,223,247]" captionStart="FIGURES 13 – 14" captionStartId="4.[151,264,2009,2032]" captionTargetBox="[476,1119,665,1969]" captionTargetId="figure@4.[456,1130,644,1988]" captionTargetPageId="4" captionText="FIGURES 13 – 14. Downeshelea lanei n. sp., male. 13. Terminalia, parameres removed; 14. Parameres." httpUri="https://zenodo.org/record/205763/files/figure.png" pageId="4" pageNumber="24">Fig. 13</figureCitation>
): tergite 9 missing in
<typeStatus id="54B68804FFE79672B93708E1FD1323B3" box="[556,654,223,247]" pageId="4" pageNumber="24" type="holotype">holotype</typeStatus>
specimen, damaged in
<typeStatus id="54B68804FFE79672B88008DEFC6123BC" box="[923,1020,224,248]" pageId="4" pageNumber="24" type="paratype">paratype</typeStatus>
specimen; sternite 9 spiculate, basal portion not visible, posterior margin with large, long, greatly convex median extension bearing two long setae. Gonocoxite elongate, straight, twice as long as basal width; gonostylus nearly straight, 0.63 length of gonocoxite, proximal ½ moderately pilose. Parameres (
<figureCitation id="13362A23FFE79672B9980973FD482221" box="[643,725,333,357]" captionStart="FIGURES 13 – 14" captionStartId="4.[151,264,2009,2032]" captionTargetBox="[476,1119,665,1969]" captionTargetId="figure@4.[456,1130,644,1988]" captionTargetPageId="4" captionText="FIGURES 13 – 14. Downeshelea lanei n. sp., male. 13. Terminalia, parameres removed; 14. Parameres." httpUri="https://zenodo.org/record/205763/files/figure.png" pageId="4" pageNumber="24">Fig. 14</figureCitation>
) 1.4x longer than aedeagus, broadly fused basally; stem thickened on proximal portion, mid portion nearly straight, slender, distal portion a long, hyaline, very slender, straight, medially directed process. Aedeagus (
<figureCitation id="13362A23FFE79672B95409ABFD3F22E9" box="[591,674,405,429]" captionStart="FIGURES 13 – 14" captionStartId="4.[151,264,2009,2032]" captionTargetBox="[476,1119,665,1969]" captionTargetId="figure@4.[456,1130,644,1988]" captionTargetPageId="4" captionText="FIGURES 13 – 14. Downeshelea lanei n. sp., male. 13. Terminalia, parameres removed; 14. Parameres." httpUri="https://zenodo.org/record/205763/files/figure.png" pageId="4" pageNumber="24">Fig. 13</figureCitation>
) nearly subtriangular, uniformly sclerotized, basal arch extending 0.45 of total length, distal portion with moderately deep mesal excavation and two slender, pointed, serrate processes.
</paragraph>
</subSubSection>
<subSubSection id="C317652DFFE79673BBDE0A3EFB9522CD" lastPageId="5" lastPageNumber="25" pageId="4" pageNumber="24" type="distribution">
<paragraph id="8BB236A6FFE79672BBDE0A3EFB302127" blockId="4.[151,1437,151,611]" pageId="4" pageNumber="24">
<emphasis id="B979EAB4FFE79672BBDE0A3EFD89215D" bold="true" box="[197,532,512,537]" pageId="4" pageNumber="24">Distribution and bionomics.</emphasis>
<taxonomicName id="4C0D4D25FFE79672B9000A3FFD6F215D" box="[539,754,513,537]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="24" phylum="Arthropoda" rank="species" species="lanei">
<emphasis id="B979EAB4FFE79672B9000A3FFD6F215D" box="[539,754,513,537]" italics="true" pageId="4" pageNumber="24">Downeshelea lanei</emphasis>
</taxonomicName>
is known only from the northern Brazilian states of Amazonas and Pará. Some specimens were collected by Thomas H.G. Aitken in Floresta da Á rea de Pesquisas Ecológicas do
<collectingCountry id="F31A7636FFE79672BBA10A74FE962126" box="[186,267,586,610]" name="Guam" pageId="4" pageNumber="24">Guamá</collectingCountry>
(APEG Forest), with a light trap at ground level in a typical freshwater ecosystem.
</paragraph>
<caption id="DF72662EFFE79672BB8C0FE7FB3224AB" box="[151,1199,2009,2032]" httpUri="https://zenodo.org/record/205763/files/figure.png" pageId="4" pageNumber="24" targetBox="[476,1119,665,1969]" targetPageId="4">
<paragraph id="8BB236A6FFE79672BB8C0FE7FB3224AB" blockId="4.[151,1199,2009,2032]" box="[151,1199,2009,2032]" pageId="4" pageNumber="24">
<emphasis id="B979EAB4FFE79672BB8C0FE7FECE24AA" bold="true" box="[151,339,2009,2032]" pageId="4" pageNumber="24">FIGURES 13–14.</emphasis>
<taxonomicName id="4C0D4D25FFE79672BA410FE7FD8624AB" box="[346,539,2009,2031]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="24" phylum="Arthropoda" rank="species" species="lanei" status="sp. nov.">
<emphasis id="B979EAB4FFE79672BA410FE7FD8624AB" box="[346,539,2009,2031]" italics="true" pageId="4" pageNumber="24">Downeshelea lanei</emphasis>
</taxonomicName>
<emphasis id="B979EAB4FFE79672B93A0FE4FDC724AB" bold="true" box="[545,602,2010,2031]" pageId="4" pageNumber="24">
<taxonomicNameLabel id="A24A57CFFFE79672B93A0FE4FDC724AB" box="[545,602,2010,2031]" pageId="4" pageNumber="24" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male. 13. Terminalia, parameres removed; 14. Parameres.
</paragraph>
</caption>
<paragraph id="8BB236A6FFE69673BBDE08A6FB9522CD" blockId="5.[151,1436,151,753]" pageId="5" pageNumber="25">
<emphasis id="B979EAB4FFE69673BBDE08A6FE9223F4" bold="true" box="[197,271,152,176]" pageId="5" pageNumber="25">
<typeStatus id="54B68804FFE69673BBDE08A6FE9523F4" box="[197,264,152,176]" pageId="5" pageNumber="25">Types</typeStatus>
.
</emphasis>
<typeStatus id="54B68804FFE69673BA0508A9FE1B23EB" box="[286,390,151,175]" pageId="5" pageNumber="25" type="holotype">Holotype</typeStatus>
<specimenCount id="9D0BFD2FFFE69673BA8F08A6FE6C23EB" box="[404,497,151,176]" pageId="5" pageNumber="25" type="adult">3 adult</specimenCount>
, in drop of
<collectingCountry id="F31A7636FFE69673B99008A9FD4223EB" box="[651,735,151,175]" name="Canada" pageId="5" pageNumber="25">Canada</collectingCountry>
Balsam on five celluloid strips on pin, labeled &quot;
<typeStatus id="54B68804FFE69673BE2A08A9FA0123EB" box="[1329,1436,151,175]" pageId="5" pageNumber="25" type="holotype">Holotype</typeStatus>
<taxonomicName id="4C0D4D25FFE69673BB8C0882FEF32390" box="[151,366,188,212]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="25" phylum="Arthropoda" rank="species" species="lanei">
<emphasis id="B979EAB4FFE69673BB8C0882FEF32390" box="[151,366,188,212]" italics="true" pageId="5" pageNumber="25">Downeshelea lanei</emphasis>
</taxonomicName>
&quot;, &quot;
<taxonomicName id="4C0D4D25FFE69673BA880882FDED2390" box="[403,624,188,212]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="25" phylum="Arthropoda" rank="species" species="cebacoi">
<emphasis id="B979EAB4FFE69673BA880882FDED2390" box="[403,624,188,212]" italics="true" pageId="5" pageNumber="25">Monohelea cebacoi</emphasis>
</taxonomicName>
,
<typeStatus id="54B68804FFE69673B9650883FD7C2391" box="[638,737,189,213]" pageId="5" pageNumber="25" type="paratype">paratype</typeStatus>
, John Lane det.1961, drawn,
<collectingCountry id="F31A7636FFE69673BF340882FBEB2390" box="[1071,1142,188,212]" name="Brazil" pageId="5" pageNumber="25">Brazil</collectingCountry>
, Pará, Cachimbo,
<date id="FFB31066FFE69673BE5E0883FA052390" box="[1349,1432,188,212]" pageId="5" pageNumber="25" value="1956-01">I. 1956</date>
, S.J. Oliveira col. #14,943&quot; (FSPC). Allotype female, labeled &quot;Allotype
<taxonomicName id="4C0D4D25FFE69673B8D908DEFB0423BC" box="[962,1177,224,248]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="25" phylum="Arthropoda" rank="species" species="lanei">
<emphasis id="B979EAB4FFE69673B8D908DEFB0423BC" box="[962,1177,224,248]" italics="true" pageId="5" pageNumber="25">Downeshelea lanei</emphasis>
</taxonomicName>
&quot;, &quot;
<taxonomicName id="4C0D4D25FFE69673BFA408DEFF242258" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="25" phylum="Arthropoda" rank="species" species="cebacoi">
<emphasis id="B979EAB4FFE69673BFA408DEFF242258" italics="true" pageId="5" pageNumber="25">Downeshelea cebacoi</emphasis>
</taxonomicName>
,
<collectingCountry id="F31A7636FFE69673BBD3093AFE922258" box="[200,271,260,284]" name="Brazil" pageId="5" pageNumber="25">Brazil</collectingCountry>
, Amazon River, Floresta, Prainha,
<date id="FFB31066FFE69673B980093AFC802258" box="[667,797,260,284]" pageId="5" pageNumber="25" value="1969-09-16">16.IX.1969</date>
, HA Wright col., light trap&quot;, #458 (CCER).
<typeStatus id="54B68804FFE69673BE08093BFA1E2259" box="[1299,1411,261,285]" pageId="5" pageNumber="25" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="9D0BFD2FFFE69673BE95093AFF4D2204" pageId="5" pageNumber="25" type="male">1 male</specimenCount>
,
<specimenCount id="9D0BFD2FFFE69673BBC70916FED02204" box="[220,333,296,320]" pageId="5" pageNumber="25" type="female">2 females</specimenCount>
, as follows:
<specimenCount id="9D0BFD2FFFE69673BAC60916FDB12204" box="[477,556,296,320]" pageId="5" pageNumber="25" type="male">1 male</specimenCount>
,
<collectingCountry id="F31A7636FFE69673B9220917FD3C2205" box="[569,673,297,321]" name="Brazil" pageId="5" pageNumber="25">BRAZIL</collectingCountry>
, Pará, Belém, Floresta da Á rea de Pesquisas Ecológicas do
<collectingCountry id="F31A7636FFE69673BE500916FA012204" box="[1355,1436,296,320]" name="Guam" pageId="5" pageNumber="25">Guamá</collectingCountry>
(APEG Forest),
<date id="FFB31066FFE69673BA550973FE342221" box="[334,425,333,357]" pageId="5" pageNumber="25" value="1970-04">IV.1970</date>
, THG Aitken col., light trap., #459 (CCER);
<specimenCount id="9D0BFD2FFFE69673B8BC0973FB962221" box="[935,1035,333,357]" pageId="5" pageNumber="25" type="female">1 female</specimenCount>
, same data, except
<date id="FFB31066FFE69673BFFE0973FAD02221" box="[1253,1357,333,357]" pageId="5" pageNumber="25" value="1970-07">VII.1970</date>
, sticky trap, #460 (CCER);
<specimenCount id="9D0BFD2FFFE69673BA61094FFE4322CD" box="[378,478,369,393]" pageId="5" pageNumber="25" type="female">1 female</specimenCount>
, same data, except
<date id="FFB31066FFE69673B9AD094FFC9722CD" box="[694,778,369,393]" pageId="5" pageNumber="25" value="1970-02">II.1970</date>
, &quot;terra firme&quot; (CNCI).
</paragraph>
</subSubSection>
<subSubSection id="C317652DFFE69673BBDE09ABFAA821B5" pageId="5" pageNumber="25" type="etymology">
<paragraph id="8BB236A6FFE69673BBDE09ABFC592295" blockId="5.[151,1436,151,753]" pageId="5" pageNumber="25">
<emphasis id="B979EAB4FFE69673BBDE09ABFED622E9" bold="true" box="[197,331,405,429]" pageId="5" pageNumber="25">Etymology.</emphasis>
This species is named in honor of Dr. John Lane in recognition of his important contributions to the study of Brazilian
<taxonomicName id="4C0D4D25FFE69673BA940987FDD32295" box="[399,590,441,465]" class="Insecta" family="Ceratopogonidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="25" phylum="Arthropoda" rank="family">Ceratopogonidae</taxonomicName>
and other nematocerous
<taxonomicName id="4C0D4D25FFE69673B8730987FC5D2295" box="[872,960,441,465]" kingdom="Plantae" order="Diptera" pageId="5" pageNumber="25" rank="order">Diptera</taxonomicName>
.
</paragraph>
<paragraph id="8BB236A6FFE69673BBDE09E3FD6B21C1" blockId="5.[151,1436,151,753]" pageId="5" pageNumber="25">
<emphasis id="B979EAB4FFE69673BBDE09E3FE4F22B2" bold="true" box="[197,466,477,502]" pageId="5" pageNumber="25">Taxonomic discussion.</emphasis>
<taxonomicName id="4C0D4D25FFE69673BAC009E3FD2E22B1" box="[475,691,477,501]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="25" phylum="Arthropoda" rank="species" species="lanei">
<emphasis id="B979EAB4FFE69673BAC009E3FD2E22B1" box="[475,691,477,501]" italics="true" pageId="5" pageNumber="25">Downeshelea lanei</emphasis>
</taxonomicName>
most closely resembles
<taxonomicName id="4C0D4D25FFE69673B8D609E0FBD522B1" box="[973,1096,477,501]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="25" phylum="Arthropoda" rank="species" species="cebacoi">
<emphasis id="B979EAB4FFE69673B8D609E0FBD522B1" box="[973,1096,477,501]" italics="true" pageId="5" pageNumber="25">D. cebacoi</emphasis>
</taxonomicName>
especially in its similar wing pattern. It can be distinguished from that species by the darker areas on the subapical portions of the fore- and midfemora (fore-, midfemora entirely yellowish brown in
<taxonomicName id="4C0D4D25FFE69673B9E70A18FCEB2179" box="[764,886,549,573]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="25" phylum="Arthropoda" rank="species" species="cebacoi">
<emphasis id="B979EAB4FFE69673B9E70A18FCEB2179" box="[764,886,549,573]" italics="true" pageId="5" pageNumber="25">D. cebacoi</emphasis>
</taxonomicName>
) and the parameres with the mid portion of each stem nearly straight and the very slender distal portion medially directed (
<figureCitation id="13362A23FFE69673B8CF0A77FBBB2125" box="[980,1062,585,609]" captionStart="FIGURES 13 – 14" captionStartId="4.[151,264,2009,2032]" captionTargetBox="[476,1119,665,1969]" captionTargetId="figure@4.[456,1130,644,1988]" captionTargetPageId="4" captionText="FIGURES 13 – 14. Downeshelea lanei n. sp., male. 13. Terminalia, parameres removed; 14. Parameres." httpUri="https://zenodo.org/record/205763/files/figure.png" pageId="5" pageNumber="25">Fig. 14</figureCitation>
) (stem sinuous and distal portion a posteriorly directed process in
<taxonomicName id="4C0D4D25FFE69673B91F0A50FDE021C1" box="[516,637,621,645]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="25" phylum="Arthropoda" rank="species" species="cebacoi">
<emphasis id="B979EAB4FFE69673B91F0A50FDE021C1" box="[516,637,621,645]" italics="true" pageId="5" pageNumber="25">D. cebacoi</emphasis>
</taxonomicName>
(
<figureCitation id="13362A23FFE69673B9970A53FD4321C1" box="[652,734,621,645]" captionStart="FIGURES 22 – 24" captionStartId="7.[151,264,1770,1793]" captionTargetBox="[435,1151,290,1759]" captionTargetId="figure@7.[435,1152,277,1759]" captionTargetPageId="7" captionText="FIGURES 22 – 24. Downeshelea cebacoi (Lane &amp; Wirth), male. 22. Terminalia, aedeagus and parameres removed; 23. Parameres; 24. Aedeagus." httpUri="https://zenodo.org/record/205765/files/figure.png" pageId="5" pageNumber="25">Fig. 23</figureCitation>
)).
</paragraph>
<paragraph id="8BB236A6FFE69673BBDC0AACFAA821B5" blockId="5.[151,1436,151,753]" pageId="5" pageNumber="25">
A key to the Neotropical species of
<taxonomicName id="4C0D4D25FFE69673B9750AAFFC9821ED" box="[622,773,657,681]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAB4FFE69673B9750AAFFC9821ED" box="[622,773,657,681]" italics="true" pageId="5" pageNumber="25">Downeshelea</emphasis>
</taxonomicName>
was provided by
<bibRefCitation id="EF9C4B57FFE69673B8C30AACFB7E21ED" author="Lane" box="[984,1251,657,681]" pageId="5" pageNumber="25" refString="Lane, J. &amp; Wirth, W. W. (1964) The biting midge genus Monohelea Kieffer in the Neotropical Region (Diptera, Ceratopogonidae). Studia Entomologica, 7, 209 - 236." type="journal article" year="1964">Lane and Wirth (1964)</bibRefCitation>
, as members of
<taxonomicName id="4C0D4D25FFE69673BB8C0A8BFEEE2189" authority="Kieffer." authorityName="Kieffer." box="[151,371,693,717]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAB4FFE69673BB8C0A8BFE882189" box="[151,277,693,717]" italics="true" pageId="5" pageNumber="25">Monohelea</emphasis>
Kieffer.
</taxonomicName>
Males of
<taxonomicName id="4C0D4D25FFE69673BAFF0A8BFD272189" box="[484,698,693,717]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="25" phylum="Arthropoda" rank="species" species="lanei">
<emphasis id="B979EAB4FFE69673BAFF0A8BFD272189" box="[484,698,693,717]" italics="true" pageId="5" pageNumber="25">Downeshelea lanei</emphasis>
</taxonomicName>
key out to couplet
<quantity id="4CF59B43FFE69673B8890A88FC232189" box="[914,958,693,717]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="5" pageNumber="25" unit="in" value="7.0">7 in</quantity>
that key and can be distinguished from
<taxonomicName id="4C0D4D25FFE69673BE9A0A88FF7321B5" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="25" phylum="Arthropoda" rank="species" species="cebacoi">
<emphasis id="B979EAB4FFE69673BE9A0A88FF7321B5" italics="true" pageId="5" pageNumber="25">D. cebacoi</emphasis>
</taxonomicName>
by the shape of their parameres which have a straight stem and a medially directed distal portion.
</paragraph>
</subSubSection>
</treatment>
</document>
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