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<document id="08E1C00F38FDDE04E87BA4879C40E811" ID-CLB-Dataset="32652" ID-DOI="10.11646/zootaxa.4294.2.3" ID-GBIF-Dataset="820a3d60-33c3-4934-9ce2-0ebef4de8532" ID-ISSN="1175-5326" ID-Zenodo-Dep="831790" ID-ZooBank="2D289A54-7600-4677-B487-84F91A4719CA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500408100684" checkinUser="plazi" docAuthor="Kočí, Tomáš, Veselská, Martina Kočová, Newman, William A., Buckeridge, John S. &amp; Sklenář, Jan" docDate="2017" docId="03960A54FF8199297EA5F9DB600F2495" docLanguage="en" docName="zootaxa.4294.2.3.pdf" docOrigin="Zootaxa 4294 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Archaeochionelasmus nekvasilovae Koci, Newman &amp; Buckeridge 2017, sp. nov." docType="treatment" docUuid="2F651BDE-7839-4995-A1F1-2A81D2B45D5C" docUuidSource="ZooBank" docVersion="8" lastPageNumber="192" masterDocId="FFAF722CFF8699227E32FF9D61422767" masterDocTitle="Archaeochionelasmus nekvasilovae gen. et sp. nov. (Cirripedia, Balanomorpha, Chionelasmatoidea) from the Bohemian Cretaceous Basin (Czech Republic): the first bona fide Cretaceous neobalanoform" masterLastPageNumber="196" masterPageNumber="181" pageNumber="188" updateTime="1698465761474" updateUser="ExternalLinkService">
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<mods:title id="20A7CC85F7C1B992BEA1A4A3C47A8061">Archaeochionelasmus nekvasilovae gen. et sp. nov. (Cirripedia, Balanomorpha, Chionelasmatoidea) from the Bohemian Cretaceous Basin (Czech Republic): the first bona fide Cretaceous neobalanoform</mods:title>
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<mods:namePart id="AC74B3938A00072184A8C1023CFC2620">Kočí, Tomáš</mods:namePart>
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<mods:namePart id="2CBDEC45F08E1D146CC389819DE7450B">Veselská, Martina Kočová</mods:namePart>
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<mods:namePart id="FE9B12D6FC29EC4C3ED4CAE13550EE16">Newman, William A.</mods:namePart>
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<mods:namePart id="8BE25E63A10E59E5B717521B64BCF616">Buckeridge, John S.</mods:namePart>
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<mods:namePart id="F1565039157908921D6C0648E5C2BABB">Sklenář, Jan</mods:namePart>
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<mods:title id="A5DED01C5927D6F5EDB03F8E07ACEC89">Zootaxa</mods:title>
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<mods:date id="558689227B5442BDA589F30600530E15">2017</mods:date>
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<treatment id="03960A54FF8199297EA5F9DB600F2495" ID-DOI="http://doi.org/10.5281/zenodo.6023946" ID-GBIF-Taxon="132550466" ID-Zenodo-Dep="6023946" LSID="urn:lsid:zoobank.org:act:2F651BDE-7839-4995-A1F1-2A81D2B45D5C" httpUri="http://treatment.plazi.org/id/03960A54FF8199297EA5F9DB600F2495" lastPageId="11" lastPageNumber="192" pageId="7" pageNumber="188">
<subSubSection id="C325E8C9FF8199257EA5F9DB603421E6" pageId="7" pageNumber="188" type="nomenclature">
<paragraph id="8B80BB42FF8199257EA5F9DB65742107" blockId="7.[151,1078,1605,1701]" box="[151,1078,1605,1632]" pageId="7" pageNumber="188">
<heading id="D0C80C2EFF8199257EA5F9DB65742107" bold="true" box="[151,1078,1605,1632]" fontSize="11" level="1" pageId="7" pageNumber="188" reason="1">
<materialsCitation id="3B57B11FFF8199257EA5F9DB65742107" ID-GBIF-Occurrence="1576158902" box="[151,1078,1605,1632]" location="Koci" pageId="7" pageNumber="188" specimenCount="1" stateProvince="Koci">
<emphasis id="B94B6750FF8199257EA5F9DB65742107" bold="true" box="[151,1078,1605,1632]" pageId="7" pageNumber="188">
<taxonomicName id="4C3FC0C1FF8199257E97F9D862892107" authority="Koci, Newman &amp; Buckeridge" authorityName="Koci, Newman &amp; Buckeridge" authorityYear="2017" box="[165,971,1605,1632]" class="Maxillopoda" family="Chionelasmatidae" genus="Archaeochionelasmus" kingdom="Animalia" order="Sessilia" pageId="7" pageNumber="188" phylum="Arthropoda" rank="species" species="nekvasilovae" status="sp. nov.">
<emphasis id="B94B6750FF8199257E97F9D863162138" bold="true" box="[165,596,1605,1631]" italics="true" pageId="7" pageNumber="188">Archaeochionelasmus nekvasilovae</emphasis>
<collectingRegion id="49FB75A0FF8199257C69F9DB63D52107" box="[603,663,1606,1632]" country="Japan" name="Kochi" pageId="7" pageNumber="188">Kočí</collectingRegion>
, Newman &amp; Buckeridge
</taxonomicName>
,
<taxonomicNameLabel id="A278DA2BFF8199257DEAF9DB65742107" box="[984,1078,1606,1632]" pageId="7" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</materialsCitation>
</heading>
</paragraph>
<paragraph id="8B80BB42FF8199257EA5F9F5603421E6" blockId="7.[151,1078,1605,1701]" box="[151,374,1640,1665]" pageId="7" pageNumber="188">Text-fig. 6; Pls. 1, 2</paragraph>
</subSubSection>
<subSubSection id="C325E8C9FF8199257EA5F91162A421C2" box="[151,998,1676,1701]" pageId="7" pageNumber="188" type="description">
<paragraph id="8B80BB42FF8199257EA5F91162A421C2" blockId="7.[151,1078,1605,1701]" box="[151,998,1676,1701]" pageId="7" pageNumber="188">urn:lsid:zoobank.org:pub: 177CC951-CEC1-425C-B693-46ADB83BD5D9</paragraph>
</subSubSection>
<subSubSection id="C325E8C9FF8199257EA5F949609A2189" box="[151,472,1748,1774]" pageId="7" pageNumber="188" type="diagnosis">
<paragraph id="8B80BB42FF8199257EA5F949609A2189" blockId="7.[151,1436,1748,1990]" box="[151,472,1748,1774]" pageId="7" pageNumber="188">
<emphasis id="B94B6750FF8199257EA5F9496053218A" bold="true" box="[151,273,1748,1773]" pageId="7" pageNumber="188">Diagnosis.</emphasis>
As for the genus.
</paragraph>
</subSubSection>
<subSubSection id="C325E8C9FF8199257EF5F96563E5203E" pageId="7" pageNumber="188" type="etymology">
<paragraph id="8B80BB42FF8199257EF5F96563E5203E" blockId="7.[151,1436,1748,1990]" pageId="7" pageNumber="188">
<emphasis id="B94B6750FF8199257EF5F965600C2076" bold="true" box="[199,334,1784,1809]" pageId="7" pageNumber="188">Etymology.</emphasis>
Named in honour of Dr Olga Nekvasilová (*1928), a prominent palaeontologist who contributed greatly to the knowledge of Mesozoic brachiopods as well as the Late Cretaceous hard-substrate communities. Dr Nekvasilová discovered this specimen in 1963.
</paragraph>
</subSubSection>
<subSubSection id="C325E8C9FF81992A7EF5F8F863152082" lastPageId="8" lastPageNumber="189" pageId="7" pageNumber="188" type="materials_examined">
<paragraph id="8B80BB42FF8199257EF5F8F862852019" blockId="7.[151,1436,1748,1990]" box="[199,967,1893,1918]" pageId="7" pageNumber="188">
<materialsCitation id="3B57B11FFF8199257EF5F8F862852019" ID-GBIF-Occurrence="1576158903" box="[199,967,1893,1918]" location="National Museum" pageId="7" pageNumber="188" specimenCount="1" stateProvince="Prague" typeStatus="holotype">
<typeStatus id="548405E0FF8199257EF5F8F860762019" box="[199,308,1893,1918]" pageId="7" pageNumber="188" type="holotype">
<emphasis id="B94B6750FF8199257EF5F8F860762019" bold="true" box="[199,308,1893,1918]" pageId="7" pageNumber="188">Holotype</emphasis>
</typeStatus>
. NM-O7615, collections of the
<location id="8EE0ED99FF8199257CA8F8F86225201A" LSID="urn:lsid:plazi:treatment:03960A54FF8199297EA5F9DB600F2495:8EE0ED99FF8199257CA8F8F86225201A" box="[666,871,1893,1918]" name="National Museum" pageId="7" pageNumber="188" stateProvince="Prague">National Museum</location>
,
<collectingRegion id="49FB75A0FF8199257D40F8FB62812019" box="[882,963,1894,1918]" country="Czech Republic" name="Praha" pageId="7" pageNumber="188">Prague</collectingRegion>
.
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</paragraph>
<paragraph id="8B80BB42FF8199257EF5F815651120A1" blockId="7.[151,1436,1748,1990]" pageId="7" pageNumber="188">
<emphasis id="B94B6750FF8199257EF5F815607420C6" bold="true" box="[199,310,1928,1953]" pageId="7" pageNumber="188">Material.</emphasis>
Three articulated parietal plates—carina, left carinal latus and left rostral latus. The form of the rostrum is readily reconstructed, but the nature of the operculum and IPs is unknown.
</paragraph>
<paragraph id="8B80BB42FF8E992A7EA5F8C863152082" blockId="8.[151,1436,1877,2021]" pageId="8" pageNumber="189">
<emphasis id="B94B6750FF8E992A7EA5F8C8607B200C" bold="true" box="[151,313,1877,1899]" pageId="8" pageNumber="189">
<collectionCode id="ED2E2387FF8E992A7EA5F8C861A8200C" box="[151,234,1877,1899]" pageId="8" pageNumber="189">PLATE</collectionCode>
1. AC
</emphasis>
General views of the specimen. Rostrolatera (RL) is oriented down and to the left in A and B; rightmost in
<emphasis id="B94B6750FF8E992A7B41F8C864C7200C" bold="true" box="[1395,1413,1877,1899]" pageId="8" pageNumber="189">C</emphasis>
is carina.
<emphasis id="B94B6750FF8E992A7ED4F8EE61BA20EE" bold="true" box="[230,248,1907,1929]" pageId="8" pageNumber="189">D</emphasis>
Detail of base of carinolatera (CL) with possible traces of original imbricating plates. Note the rough surface observable at CL base to left (see this feature also in Pl. 2G).
<emphasis id="B94B6750FF8E992A7D36F80F625720CF" bold="true" box="[772,789,1938,1960]" pageId="8" pageNumber="189">E</emphasis>
Detailed view of junction of CL and C plates at their basalmost portion.
<emphasis id="B94B6750FF8E992A7EC5F82C604420A0" bold="true" box="[247,262,1969,1991]" pageId="8" pageNumber="189">F</emphasis>
Detailed view of ala of C. AC are identical in scale, AC digital macrophotography, DE optical microphotography, F 3D microscope image.
</paragraph>
</subSubSection>
<subSubSection id="C325E8C9FF8F99297EA5F8B4600F2495" lastPageId="11" lastPageNumber="192" pageId="9" pageNumber="190" type="description">
<paragraph id="8B80BB42FF8F992B7EA5F8B465AB20DD" blockId="9.[151,1436,1833,1978]" pageId="9" pageNumber="190">
<emphasis id="B94B6750FF8F992B7EA5F8B4607D2058" bold="true" box="[151,319,1833,1855]" pageId="9" pageNumber="190">PLATE 2. AE</emphasis>
detailed views of rostrolatera (left)/carinolatera (right) junction.
<emphasis id="B94B6750FF8F992B7DD4F8B465542058" bold="true" box="[998,1046,1833,1855]" pageId="9" pageNumber="190">AB</emphasis>
Top-lateral views,
<emphasis id="B94B6750FF8F992B7AD3F8B465B12058" bold="true" box="[1249,1267,1833,1855]" pageId="9" pageNumber="190">C</emphasis>
Top view.
<emphasis id="B94B6750FF8F992B7B5EF8B464DE2058" bold="true" box="[1388,1436,1833,1855]" pageId="9" pageNumber="190">DE</emphasis>
Details in similar view using refracted light. Although these plates show partial recrystallization, some internal features are still observable in refracted light.
<emphasis id="B94B6750FF8F992B7F8CF8FB608F201B" bold="true" box="[446,461,1894,1916]" pageId="9" pageNumber="190">F</emphasis>
Detail of RL/CL junction showing overlapping margin of RL.
<emphasis id="B94B6750FF8F992B7A75F8FB6519201B" bold="true" box="[1095,1115,1894,1916]" pageId="9" pageNumber="190">G</emphasis>
Detail of the RL/CL junction at their basal portions with rough surface.
<emphasis id="B94B6750FF8F992B7C00F818630420FC" bold="true" box="[562,582,1925,1947]" pageId="9" pageNumber="190">H</emphasis>
Detail showing left radius of the RL plate. Compare delicate ornamentation of the plate surface with F. DE are identical in scale. A is a 3D microscope image, BH optical microphotography.
</paragraph>
<caption id="DF40EBCAFF8C99287EA5F9B165CE207F" httpUri="https://zenodo.org/record/831798/files/figure.png" pageId="10" pageNumber="191" targetBox="[170,1433,224,1502]" targetPageId="10">
<paragraph id="8B80BB42FF8C99287EA5F9B165CE207F" blockId="10.[151,1436,1580,1816]" pageId="10" pageNumber="191">
<emphasis id="B94B6750FF8C99287EA5F9B160572125" bold="true" box="[151,277,1580,1602]" pageId="10" pageNumber="191">FIGURE 6.</emphasis>
Reconstruction of
<emphasis id="B94B6750FF8C99287FEEF9B160AB2125" bold="true" box="[476,489,1580,1602]" pageId="10" pageNumber="191"></emphasis>
<taxonomicName id="4C3FC0C1FF8C99287FDBF9B165D02125" authority="Koci, Newman &amp; Buckeridge" authorityName="Koci, Newman &amp; Buckeridge" authorityYear="2017" box="[489,1170,1580,1602]" class="Maxillopoda" family="Chionelasmatidae" genus="Archaeochionelasmus" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="191" phylum="Arthropoda" rank="species" species="nekvasilovae" status="gen. et sp. nov.">
<emphasis id="B94B6750FF8C99287FDBF9B162112125" box="[489,851,1580,1602]" italics="true" pageId="10" pageNumber="191">Archaeochionelasmus nekvasilovae</emphasis>
Kočí, Newman &amp; Buckeridge
</taxonomicName>
,
<emphasis id="B94B6750FF8C99287AADF9B164262125" bold="true" box="[1183,1380,1580,1602]" pageId="10" pageNumber="191">
<taxonomicNameLabel id="A278DA2BFF8C99287AADF9B164012125" box="[1183,1347,1580,1602]" pageId="10" pageNumber="191" rank="species">gen. et sp. nov.</taxonomicNameLabel>
A.
</emphasis>
Wall based on the fossil, operculum low in accordance with the low profile and tight sutures of the wall that suggest it may have been an epibiont of large marine reptiles or cephalopods. The presence of imbricating plates is based on ribbons of sediment adhering to the base of the wall plates. Their arrangement is patterned after that found in B, the generalized form as typified by
<taxonomicName id="4C3FC0C1FF8C99287EA5F93B603A21DB" authorityName="Yamaguchi" authorityYear="1990" box="[151,376,1702,1724]" class="Maxillopoda" family="Chionelasmatidae" genus="Eochionelasmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="191" phylum="Arthropoda" rank="species" species="ohtai">
<emphasis id="B94B6750FF8C99287EA5F93B603A21DB" box="[151,376,1702,1724]" italics="true" pageId="10" pageNumber="191">Eochionelasmus ohtai</emphasis>
</taxonomicName>
Yamaguchi &amp; Newman, 1990. which is the basis for their arrangement in higher forms; cf.
<taxonomicName id="4C3FC0C1FF8C99287B13F93B60F621BC" authority="Buckeridge, 1996" authorityName="Buckeridge" authorityYear="1996" class="Maxillopoda" family="Waikalasmatidae" genus="Waikalasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="191" phylum="Arthropoda" rank="species" species="boucheti">
<emphasis id="B94B6750FF8C99287B13F93B61AC21BC" italics="true" pageId="10" pageNumber="191">Waikalasma boucheti</emphasis>
Buckeridge, 1996
</taxonomicName>
and
<taxonomicName id="4C3FC0C1FF8C99287FC3F95862FC21BC" authority="in Ross &amp; Newman (2001)" authorityName="in Ross &amp; Newman" authorityYear="2001" box="[497,958,1733,1755]" class="Maxillopoda" family="Catophragmidae" genus="Catophragmus" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="191" phylum="Arthropoda" rank="genus">
<emphasis id="B94B6750FF8C99287FC3F95863C521BC" box="[497,647,1733,1755]" italics="true" pageId="10" pageNumber="191">Catophragmus</emphasis>
in Ross &amp; Newman (2001)
</taxonomicName>
. Parietal plates R, RL, CL and C (rostrum, rostrolatus, carinolatus &amp; carina respectively) and opercular s and t (scutum and tergum respectively), with the relative positions of the paired and unpaired plates making up the imbricating whorls identified accordingly.
</paragraph>
</caption>
<paragraph id="8B80BB42FF8C99297EF5F8DF60D12622" blockId="10.[151,1437,1855,2030]" lastBlockId="11.[151,1437,151,1010]" lastPageId="11" lastPageNumber="192" pageId="10" pageNumber="191">
<emphasis id="B94B6750FF8C99287EF5F8DF601B203C" bold="true" box="[199,345,1858,1883]" pageId="10" pageNumber="191">Description.</emphasis>
Shell smooth, medium-low conical, comprising six compartmental plates—C, CL1, RL, R (rostrum reconstructed). Pseudo-radii narrow, running sub-parallel to the articular margins, slightly overlapping the edges of the parietes along both margins of the RL and the carinal margin of the CL1 (
<figureCitation id="1304A7C7FF8C99287A52F81065E720C1" box="[1120,1189,1933,1958]" captionStart="FIGURE 3" captionStartId="2.[151,250,1885,1907]" captionTargetBox="[188,1397,1016,1847]" captionTargetId="figure@2.[168,1419,994,1864]" captionTargetPageId="2" captionText="FIGURE 3. Comparison of (A) the reconstructed wall of Archaeochionelasmus nekvasilovae gen. et sp. nov. with (B) an illustration of a juvenile of Chionelasmus ohtai having the first whorl of basal imbricating plates (from a line drawing by D. S. Jones, with permission in litt. 2016), and (C) an adult of Chionelasmus ohtai; CL overlaps the ala of C, as is indicated on left side, whereas CL abuts rather than overlaps R (Yamaguchi &amp; Newman 1990, fig. 8 B), their articulation being strengthened and protected by the addition of the RL applied over the abutment of CL and R, dividing SRL into more or less equal halves." httpUri="https://zenodo.org/record/831796/files/figure.png" pageId="10" pageNumber="191">Fig. 3</figureCitation>
,
<figureCitation id="1304A7C7FF8C99287A80F810658020C1" box="[1202,1218,1933,1958]" captionStart="FIGURE 6" captionStartId="10.[151,250,1580,1602]" captionTargetBox="[170,1433,224,1502]" captionTargetId="figure@10.[151,1436,193,1558]" captionTargetPageId="10" captionText="FIGURE 6. Reconstruction of † Archaeochionelasmus nekvasilovae Kočí, Newman &amp; Buckeridge, gen. et sp. nov. A. Wall based on the fossil, operculum low in accordance with the low profile and tight sutures of the wall that suggest it may have been an epibiont of large marine reptiles or cephalopods. The presence of imbricating plates is based on ribbons of sediment adhering to the base of the wall plates. Their arrangement is patterned after that found in B, the generalized form as typified by Eochionelasmus ohtai Yamaguchi &amp; Newman, 1990. which is the basis for their arrangement in higher forms; cf. Waikalasma boucheti Buckeridge, 1996 and Catophragmus in Ross &amp; Newman (2001). Parietal plates R, RL, CL and C (rostrum, rostrolatus, carinolatus &amp; carina respectively) and opercular s and t (scutum and tergum respectively), with the relative positions of the paired and unpaired plates making up the imbricating whorls identified accordingly." httpUri="https://zenodo.org/record/831798/files/figure.png" pageId="10" pageNumber="191">6</figureCitation>
, 7), separated from them by a well-formed longitudinal groove; wider nearer summit, with width increasing by accretion of the fine transverse and longitudinal ridges; the transverse growth lines are not upturned (as in true radii that develop later in balanomorph evolution). Note that these pseudo-radii are not precursors of the true radii, which developed later in the Palaeogene. The longitudinal ridges represent successive growth increments, with new ridges and grooves gradually added at the base; RL and CL1 rhomboid, with summits parallel to the base; C semi-conic. RL—narrow:
<quantity id="4CC716A7FF8D99297EA5FE9461A72646" box="[151,229,265,290]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="192" unit="mm" value="16.0">16 mm</quantity>
basal length:
<quantity id="4CC716A7FF8D99297FB2FE97609A2646" box="[384,472,266,290]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="11" pageNumber="192" unit="mm" value="9.9">9.9 mm</quantity>
, length in upper portion
<quantity id="4CC716A7FF8D99297CDBFE9762002646" box="[745,834,266,290]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="11" pageNumber="192" unit="mm" value="2.4">2.4 mm</quantity>
; CL1—high 9
<quantity id="4CC716A7FF8D99297DD8FE9465692646" box="[1002,1067,265,290]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="192" unit="mm" value="5.0">5 mm</quantity>
, length in basis
<quantity id="4CC716A7FF8D99297AECFE97646C2646" box="[1246,1326,266,290]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="11" pageNumber="192" unit="mm" value="13.0">13 mm</quantity>
, length in upper portion
<quantity id="4CC716A7FF8D99297F07FEB360CC2622" box="[309,398,302,325]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="11" pageNumber="192" unit="mm" value="7.7">7.7 mm</quantity>
.
</paragraph>
<paragraph id="8B80BB42FF8D99297EF5FECC65E32579" blockId="11.[151,1437,151,1010]" pageId="11" pageNumber="192">Although no complete whorls of IPs are retained, the basal margin of the parietal plates (where not eroded or damaged) and the exposed ala (Pl.1, Fig. F) have variously arranged indentations indicating where tissue within extended out underneath the basal IPs and the wall. The delicate, sinuous, lateral calcareous ridges occurring near the base of the compartments (Pl. 1, Fig. D) are interpreted as sites that entrapped fine calcareous sediment along the zone where the imbrications abutted the compartments. There are also some fragments of shell near the base that could be the remains of IPs that survived the preparation of the specimen (Pl. 1, Fig. E).</paragraph>
<paragraph id="8B80BB42FF8D99297EF5FDB4600F2495" blockId="11.[151,1437,151,1010]" pageId="11" pageNumber="192">
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<emphasis id="B94B6750FF8D99297EF5FDB463152525" bold="true" box="[199,599,553,578]" pageId="11" pageNumber="192">
<location id="8EE0ED99FF8D99297EF5FDB4604E2525" LSID="urn:lsid:plazi:treatment:03960A54FF8199297EA5F9DB600F2495:8EE0ED99FF8D99297EF5FDB4604E2525" box="[199,268,553,578]" country="Austria" county="southern Sweden" municipality="The" name="Mode" pageId="11" pageNumber="192" stateProvince="Florida">Mode</location>
of life and palaeoecology.
</emphasis>
A reconstruction of
<taxonomicName id="4C3FC0C1FF8D99297D65FDB4650F2526" authority="Kočí &amp; Veselská &amp; Newman &amp; Buckeridge &amp; Sklenář, 2017" authorityName="Kočí &amp; Veselská &amp; Newman &amp; Buckeridge &amp; Sklenář" authorityYear="2017" box="[855,1101,553,577]" class="Maxillopoda" family="Chionelasmatidae" genus="Archaeochionelasmus" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="193" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94B6750FF8D99297D65FDB4650F2526" box="[855,1101,553,577]" italics="true" pageId="11" pageNumber="192">Archaeochionelasmus</emphasis>
</taxonomicName>
nov. gen. (
<figureCitation id="1304A7C7FF8D99297AEFFDB464642525" box="[1245,1318,553,578]" captionStart="FIGURE 6" captionStartId="10.[151,250,1580,1602]" captionTargetBox="[170,1433,224,1502]" captionTargetId="figure@10.[151,1436,193,1558]" captionTargetPageId="10" captionText="FIGURE 6. Reconstruction of † Archaeochionelasmus nekvasilovae Kočí, Newman &amp; Buckeridge, gen. et sp. nov. A. Wall based on the fossil, operculum low in accordance with the low profile and tight sutures of the wall that suggest it may have been an epibiont of large marine reptiles or cephalopods. The presence of imbricating plates is based on ribbons of sediment adhering to the base of the wall plates. Their arrangement is patterned after that found in B, the generalized form as typified by Eochionelasmus ohtai Yamaguchi &amp; Newman, 1990. which is the basis for their arrangement in higher forms; cf. Waikalasma boucheti Buckeridge, 1996 and Catophragmus in Ross &amp; Newman (2001). Parietal plates R, RL, CL and C (rostrum, rostrolatus, carinolatus &amp; carina respectively) and opercular s and t (scutum and tergum respectively), with the relative positions of the paired and unpaired plates making up the imbricating whorls identified accordingly." httpUri="https://zenodo.org/record/831798/files/figure.png" pageId="11" pageNumber="192">Fig. 6</figureCitation>
) shows a moderately low profile similar to that of the “turtle barnacle”
<taxonomicName id="4C3FC0C1FF8D99297D66FDD365A82501" authority="Ross &amp; Newman, 1967" authorityName="Ross &amp; Newman" authorityYear="1967" box="[852,1258,589,614]" class="Maxillopoda" family="Chelonibiidae" genus="Emersonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="192" phylum="Arthropoda" rank="genus">
<emphasis id="B94B6750FF8D99297D66FDD3629A2502" box="[852,984,590,613]" italics="true" pageId="11" pageNumber="192">Emersonius</emphasis>
<bibRefCitation id="EFAEC6B3FF8D99297DD3FDD365A82501" author="Ross" box="[993,1258,589,614]" pageId="11" pageNumber="192" refString="Ross, A. &amp; Newman, W. A. (1967) Eocene Balanidae of Florida, including a new genus and species with a unique plan of &quot; turtle-barnacle &quot; organization. American Museum Novitates, 2288, 1 - 22." type="journal article" year="1967">Ross &amp; Newman, 1967</bibRefCitation>
</taxonomicName>
recorded in the Eocene of
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(USA),
<taxonomicName id="4C3FC0C1FF8D99297FD4FDEC63D925EE" authorityName="Harzhauser &amp; Newman in Harzhauser, Newman &amp; Grunert" authorityYear="2011" box="[486,667,625,649]" class="Maxillopoda" family="Chelonibiidae" genus="Protochelonibia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="192" phylum="Arthropoda" rank="genus">
<emphasis id="B94B6750FF8D99297FD4FDEC63D925EE" box="[486,667,625,649]" italics="true" pageId="11" pageNumber="192">Protochelonibia</emphasis>
</taxonomicName>
from the
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of the
<location id="8EE0ED99FF8D99297A7CFDEC646425ED" LSID="urn:lsid:plazi:treatment:03960A54FF8199297EA5F9DB600F2495:8EE0ED99FF8D99297A7CFDEC646425ED" box="[1102,1318,625,650]" country="Austria" county="southern Sweden" municipality="The" name="Central Paratethys" pageId="11" pageNumber="192" stateProvince="Florida">Central Paratethys</location>
(
<collectingCountry id="F328FBD2FF8D99297B0CFDEC64D625ED" box="[1342,1428,625,650]" name="Austria" pageId="11" pageNumber="192">Austria</collectingCountry>
) (
<bibRefCitation id="EFAEC6B3FF8D99297EACFD0860E625C9" author="Harzhauser" box="[158,420,661,686]" pageId="11" pageNumber="192" refString="Harzhauser, M., Newman, W. A. &amp; Grunert, P. (2011) A new Early Miocene barnacle lineage and the roots of sea-turtle fouling Chelonibiidae (Cirripedia, Balanomorpha). Journal of Systematic Palaeontology, 9, 473 - 480. https: // doi. org / 10.1080 / 14772019.2010.528053" type="journal article" year="2011">
<location id="8EE0ED99FF8D99297EACFD08606225C9" LSID="urn:lsid:plazi:treatment:03960A54FF8199297EA5F9DB600F2495:8EE0ED99FF8D99297EACFD08606225C9" box="[158,288,661,686]" country="Austria" county="southern Sweden" municipality="The" name="Harzhauser" pageId="11" pageNumber="192" stateProvince="Florida">Harzhauser</location>
<emphasis id="B94B6750FF8D99297F1AFD0B602325CA" box="[296,353,661,685]" italics="true" pageId="11" pageNumber="192">et al.</emphasis>
2011
</bibRefCitation>
), extant
<taxonomicName id="4C3FC0C1FF8D99297C37FD08622D25C9" authority="Zardus et al. 2013" authorityName="Leach" authorityYear="1817" box="[517,879,661,686]" class="Maxillopoda" family="Chelonibiidae" genus="Chelonibia" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="192" phylum="Arthropoda" rank="genus">
<emphasis id="B94B6750FF8D99297C37FD0863C025CA" box="[517,642,661,685]" italics="true" pageId="11" pageNumber="192">Chelonibia</emphasis>
(Zardus
<emphasis id="B94B6750FF8D99297CDBFD0B626125CA" box="[745,803,661,685]" italics="true" pageId="11" pageNumber="192">et al.</emphasis>
2013)
</taxonomicName>
,
<taxonomicName id="4C3FC0C1FF8D99297D4EFD0862B025CA" authorityName="Gray" authorityYear="1825" box="[892,1010,661,685]" class="Maxillopoda" family="Platylepadidae" genus="Platylepas" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="192" phylum="Arthropoda" rank="genus">
<emphasis id="B94B6750FF8D99297D4EFD0862B025CA" box="[892,1010,661,685]" italics="true" pageId="11" pageNumber="192">Platylepas</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3FC0C1FF8D99297A19FD0864DA25C9" authority="Southward 2008" authorityName="Southward" authorityYear="2008" box="[1067,1432,661,686]" class="Maxillopoda" family="Platylepadidae" genus="Stomatolepas" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="192" phylum="Arthropoda" rank="genus">
<emphasis id="B94B6750FF8D99297A19FD08658325CA" box="[1067,1217,661,685]" italics="true" pageId="11" pageNumber="192">Stomatolepas</emphasis>
(
<bibRefCitation id="EFAEC6B3FF8D99297AE2FD0864D325C9" author="Southward" box="[1232,1425,661,686]" pageId="11" pageNumber="192" refString="Southward, A. J. (2008) Barnacles. Synopses of the British Fauna, New Series, 57, pp. 1 - 140, 4 pls." type="journal article" year="2008">Southward 2008</bibRefCitation>
)
</taxonomicName>
.
<collectingMunicipality id="6BE42138FF8D99297EA5FD24618125B5" box="[151,195,697,722]" pageId="11" pageNumber="192">The</collectingMunicipality>
form of
<taxonomicName id="4C3FC0C1FF8D99297F11FD24635B25B6" authority="Kočí &amp; Veselská &amp; Newman &amp; Buckeridge &amp; Sklenář, 2017" authorityName="Kočí &amp; Veselská &amp; Newman &amp; Buckeridge &amp; Sklenář" authorityYear="2017" box="[291,537,697,721]" class="Maxillopoda" family="Chionelasmatidae" genus="Archaeochionelasmus" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="193" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94B6750FF8D99297F11FD24635B25B6" box="[291,537,697,721]" italics="true" pageId="11" pageNumber="192">Archaeochionelasmus</emphasis>
</taxonomicName>
nov. gen. is consistent with that required for attachment to living substrates such as an ammonite or turtle, both of which are recorded in the
<collectionCode id="ED2E2387FF8D99297D76FD43623E2591" box="[836,892,734,758]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14402" name="Universitat Aut<75>noma de Barcelona" pageId="11" pageNumber="192">BCB</collectionCode>
(Ekrt
<emphasis id="B94B6750FF8D99297DF5FD4365432592" box="[967,1025,733,757]" italics="true" pageId="11" pageNumber="192">et al.</emphasis>
, 2001;
<bibRefCitation id="EFAEC6B3FF8D99297A6AFD43645C2591" author="Kear" box="[1112,1310,733,758]" pageId="11" pageNumber="192" refString="Kear, B. P., Ekrt, B., Prokop, J. &amp; Georgalis, G. L. (2014) Turonian marine amniotes from the Bohemian Cretaceous Basin, Czech Republic. Geological Magazine, 151, 183 - 198." type="journal article" year="2014">
Kear
<emphasis id="B94B6750FF8D99297AAAFD4365902592" box="[1176,1234,733,757]" italics="true" pageId="11" pageNumber="192">et al.</emphasis>
, 2014
</bibRefCitation>
). Although direct evidence is lacking, we suggest that
<taxonomicName id="4C3FC0C1FF8D99297C98FC9C62E2247E" authority="Kočí &amp; Veselská &amp; Newman &amp; Buckeridge &amp; Sklenář, 2017" authorityName="Kočí &amp; Veselská &amp; Newman &amp; Buckeridge &amp; Sklenář" authorityYear="2017" box="[682,928,769,793]" class="Maxillopoda" family="Chionelasmatidae" genus="Archaeochionelasmus" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="193" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94B6750FF8D99297C98FC9C62E2247E" box="[682,928,769,793]" italics="true" pageId="11" pageNumber="192">Archaeochionelasmus</emphasis>
</taxonomicName>
nov. gen. may have lived as an obligate commensal with ammonites and/or marine reptiles. Notwithstanding this, it is quite possible that it lived on hard, inorganic substrates in the littoral zone. However, further indirect support for commensal life is the absence of any other neobalanomorph plates, conspecific or otherwise, at this locality. Although negative evidence, it suggests there were no other barnacles present, compared to, for example, the variety of forms known to have lived on hard substrates along the rocky coast at Ivö Klack (
<collectingCounty id="62E1C3CEFF8D99297C80FC28623A24A9" box="[690,888,949,974]" pageId="11" pageNumber="192">southern Sweden</collectingCounty>
) during the Campanian (
<bibRefCitation id="EFAEC6B3FF8D99297AA8FC28647B24A9" author="Withers" box="[1178,1337,949,974]" pageId="11" pageNumber="192" refString="Withers, T. H. (1935) Catalogue of fossil Cirripedia in the Department of Geology II, Cretaceous. British Museum (Natural History), London, 534 pp." type="book" year="1935">Withers 1935</bibRefCitation>
;
<bibRefCitation id="EFAEC6B3FF8D99297B75FC2860002495" author="Gale" pageId="11" pageNumber="192" refString="Gale, A. S. &amp; Sorensen, A. M. (2015) Taxonomy and palaeoecology of thoracican cirripedes (Crustacea) from a Campanian rocky shoreline at Ivo Klack, southern Sweden. Cretaceous Research, 54, 212 - 242. https: // doi. org / 10.1016 / j. cretres. 2014.09.004" type="journal article" year="2015">Gale &amp; Sørensen 2015</bibRefCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>