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<document id="C8E81A72518CBE2CE4A12594F0A4F8B1" ID-DOI="10.5281/zenodo.281839" ID-GBIF-Dataset="da06f1c1-190d-4ada-b1f0-8f847291210d" ID-ISSN="1175-5326" ID-Zenodo-Dep="281839" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460501162585" checkinUser="plazi" docAuthor="Iwan, Dariusz &amp; Kamiński, Marcin Jan" docDate="2012" docId="03851206FFC3935FFF60F95F6D51CEA5" docLanguage="en" docName="zt03399p034.pdf" docOrigin="Zootaxa 3399" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Lechius steineri Iwan 1995" docType="treatment" docVersion="11" lastPageNumber="32" masterDocId="FFBC6A7EFFC59356FFF7FF816E04C915" masterDocTitle="Revision of the Malagasy genus Lechius Iwan, 1995 (Coleoptera: Tenebrionidae: Pedinini)" masterLastPageNumber="34" masterPageNumber="23" pageNumber="29" updateTime="1698311846854" updateUser="plazi">
<mods:mods id="0FAF6CE4AA62DD66CF73D78D3A55A459" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5D98124DF63D374FDF4E5B0E1B3F21E0">
<mods:title id="E11B07DDDA31325C618DFB8242334B38">Revision of the Malagasy genus Lechius Iwan, 1995 (Coleoptera: Tenebrionidae: Pedinini)</mods:title>
</mods:titleInfo>
<mods:name id="8012C9FE191F9DF3B1F5E818755A8205" type="personal">
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</mods:role>
<mods:namePart id="F53A771403E8E89838FABB49A1FA4F2E">Iwan, Dariusz</mods:namePart>
</mods:name>
<mods:name id="B20A260E8CD5DCE1A19EE1E7E8B370CC" type="personal">
<mods:role id="FE77CE826441FD1C01293555DE1801C8">
<mods:roleTerm id="EA5FAFAC7A1AC9871D818DF0C909A395">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6983B9833D1C30600810DC54FBCDDE52">Kamiński, Marcin Jan</mods:namePart>
</mods:name>
<mods:typeOfResource id="11AE538C8ADC418FD11D99F3AB86C30C">text</mods:typeOfResource>
<mods:relatedItem id="02A2F29576DA9C0FF9FEF9ACEFC67857" type="host">
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<mods:title id="BFAE72C653A5D0607778BD80F1ECEA4B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="78F3482B60A1A93AF6E6AAADE62BF3DA">
<mods:date id="00B597590FE5B23450D39008F19DDC91">2012</mods:date>
<mods:detail id="F1EA5D0AC7B7AE9B2C750A2C04D4DE78" type="volume">
<mods:number id="716539B5C1A15F191BE2B578C4168C9A">3399</mods:number>
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<mods:start id="6855D2881BDC5831D0B8ED640210252D">23</mods:start>
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<mods:classification id="E11C0140FADF74FE5A7EF6A8C993F8D8">journal article</mods:classification>
<mods:identifier id="4D42E303485424EADD493D99507214CB" type="DOI">10.5281/zenodo.281839</mods:identifier>
<mods:identifier id="6E4010B8DE7D5F11495D20D6D8FE6F7A" type="GBIF-Dataset">da06f1c1-190d-4ada-b1f0-8f847291210d</mods:identifier>
<mods:identifier id="73B67A2B4BC835AD126CA16C3B725141" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="BFF25238B749979A127B542BB83DB7FD" type="Zenodo-Dep">281839</mods:identifier>
</mods:mods>
<treatment id="03851206FFC3935FFF60F95F6D51CEA5" ID-DOI="http://doi.org/10.5281/zenodo.6170867" ID-GBIF-Taxon="119521399" ID-Zenodo-Dep="6170867" LSID="urn:lsid:plazi:treatment:03851206FFC3935FFF60F95F6D51CEA5" httpUri="http://treatment.plazi.org/id/03851206FFC3935FFF60F95F6D51CEA5" lastPageId="9" lastPageNumber="32" pageId="6" pageNumber="29">
<subSubSection id="C336F09BFFC39350FF60F95F6C10CE0C" pageId="6" pageNumber="29" type="nomenclature">
<paragraph id="8B93A310FFC39350FF60F95F6FE3CFED" blockId="6.[151,532,1758,1817]" box="[151,487,1758,1784]" pageId="6" pageNumber="29">
<heading id="D0DB147CFFC39350FF60F95F6FE3CFED" bold="true" box="[151,487,1758,1784]" fontSize="11" level="1" pageId="6" pageNumber="29" reason="1">
<taxonomicName id="4C2CD893FFC39350FF60F95F6FE3CFED" authority="Iwan, 1995" authorityName="Iwan" authorityYear="1995" box="[151,487,1758,1784]" class="Insecta" family="Tenebrionidae" genus="Lechius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="steineri">
<emphasis id="B9587F02FFC39350FF60F95F6FE3CFED" bold="true" box="[151,487,1758,1784]" pageId="6" pageNumber="29">
<emphasis id="B9587F02FFC39350FF60F95F6F5CCFED" bold="true" box="[151,344,1758,1784]" italics="true" pageId="6" pageNumber="29">Lechius steineri</emphasis>
<bibRefCitation id="EFBDDEE1FFC39350FEA8F95F6FE3CFED" author="Iwan" box="[351,487,1758,1784]" pageId="6" pageNumber="29" refString="Iwan, D. (1995) A Madagascan genus Lechius gen. nov. (Coleoptera: Tenebrionidae: Platynotini). Genus, 6 (3 - 4), 401 - 414." type="journal article" year="1995">Iwan, 1995</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B93A310FFC39350FF60F8806C10CE0C" blockId="6.[151,532,1758,1817]" box="[151,532,1793,1817]" pageId="6" pageNumber="29">(Figs 5, 12, 24, 31, 33, 35, 36, 37)</paragraph>
</subSubSection>
<subSubSection id="C336F09BFFC39350FF60F8C86AE7CE4A" box="[151,1251,1865,1887]" pageId="6" pageNumber="29" type="reference_group">
<paragraph id="8B93A310FFC39350FF60F8C86AE7CE4A" blockId="6.[151,1251,1865,1887]" box="[151,1251,1865,1887]" pageId="6" pageNumber="29">
<treatmentCitationGroup id="AB3C843EFFC39350FF60F8C86AE7CE4A" box="[151,1251,1865,1887]" pageId="6" pageNumber="29">
<taxonomicName id="4C2CD893FFC39350FF60F8C86FDACE4A" authority="Iwan 1995: 408" authorityName="Iwan" authorityPageNumber="408" authorityYear="1995" box="[151,478,1865,1887]" class="Insecta" family="Tenebrionidae" genus="Lechius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="steineri">
<emphasis id="B9587F02FFC39350FF60F8C86F3CCE4A" box="[151,312,1865,1887]" italics="true" pageId="6" pageNumber="29">Lechius steineri</emphasis>
<treatmentCitation id="0A8D8501FFC39350FEC9F8CB6FDACE4A" author="Iwan" box="[318,478,1865,1887]" page="408" pageId="6" pageNumber="29" year="1995">
<bibRefCitation id="EFBDDEE1FFC39350FEC9F8CB6FA9CE4A" author="Iwan" box="[318,429,1865,1887]" pageId="6" pageNumber="29" refString="Iwan, D. (1995) A Madagascan genus Lechius gen. nov. (Coleoptera: Tenebrionidae: Platynotini). Genus, 6 (3 - 4), 401 - 414." type="journal article" year="1995">Iwan 1995</bibRefCitation>
: 408
</treatmentCitation>
</taxonomicName>
.—
<treatmentCitation id="0A8D8501FFC39350FE0BF8CB6C9ACE4A" author="Iwan" box="[508,670,1865,1887]" page="504" pageId="6" pageNumber="29" year="2001">
<bibRefCitation id="EFBDDEE1FFC39350FE0BF8CB6C68CE4A" author="Iwan" box="[508,620,1866,1887]" pageId="6" pageNumber="29" refString="Iwan, D. (2001) Systematics of the Madagascan genera Clastopus Fairmaire, 1898 and Lechius Iwan, 1995 (Coleoptera: Tenebrionidae: Platynotini). Annales Zoologici, 51 (4), 497 - 504." type="journal article" year="2001">Iwan 2001</bibRefCitation>
: 504
</treatmentCitation>
, 2002a: 74, 2002b: 274, 2010: 40;
<treatmentCitation id="0A8D8501FFC39350FC08F8C86ADBCE4A" author="Schawaller" box="[1023,1247,1865,1887]" page="279" pageId="6" pageNumber="29" year="2010">
<bibRefCitation id="EFBDDEE1FFC39350FC08F8C86AA8CE4A" author="Schawaller" box="[1023,1196,1865,1887]" pageId="6" pageNumber="29" refString="Schawaller, W. (2010) Check-list and bibliography of the Tenebrionidae (sensu stricto) from Madagascar (Coleoptera). Stuttgarter Beitrage zur Naturkunde A (NS), 3, 277 - 289." type="journal article" year="2010">Schawaller 2010</bibRefCitation>
: 279
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C336F09BFFC39351FF60F80D6DDAC8B9" lastPageId="7" lastPageNumber="30" pageId="6" pageNumber="29" type="materials_examined">
<paragraph id="8B93A310FFC39351FF60F80D6DDAC8B9" blockId="6.[151,1436,1932,2029]" lastBlockId="7.[151,1436,151,429]" lastPageId="7" lastPageNumber="30" pageId="6" pageNumber="29">
<emphasis id="B9587F02FFC39350FF60F80D6FE7CEB0" bold="true" box="[151,483,1932,1957]" pageId="6" pageNumber="29">
Material studied.
<typeStatus id="54971DB2FFC39350FE81F80C6FE7CEB0" box="[374,483,1933,1957]" pageId="6" pageNumber="29" type="holotype">Holotype</typeStatus>
</emphasis>
, male (
<collectionCode id="ED3D3BD5FFC39350FDB6F80D6C9ACEB1" LSID="urn:lsid:biocol.org:col:34988" box="[577,670,1932,1956]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="29">MNHN</collectionCode>
): &quot;
<collectingCountry id="F33BE380FFC39350FD49F80D6D48CEB1" box="[702,844,1932,1956]" name="Madagascar" pageId="6" pageNumber="29">Madagascar</collectingCountry>
: Province Toliara
<quantity id="4CD40EF5FFC39350FBD0F80D6A61CEB1" box="[1063,1125,1932,1956]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="29" unit="km" value="4.0">4 km</quantity>
N Bemanonga
<date id="FF9285D0FFC39350FAD3F80D6B9DCEB1" box="[1316,1433,1932,1956]" pageId="6" pageNumber="29" value="1929-03">29 March</date>
. 1990”, “Under leaf litter on loose sandy soil in partial shade and shrubs”, “W.E. Steiner, C. Kremen, V. Razafimahatratra collectors&quot;.
<typeStatus id="54971DB2FFC39350FE70F8546FFBCEF8" box="[391,511,2005,2029]" pageId="6" pageNumber="29" type="paratype">
<emphasis id="B9587F02FFC39350FE70F8546FFBCEF8" bold="true" box="[391,511,2005,2029]" pageId="6" pageNumber="29">Paratypes</emphasis>
</typeStatus>
,
<specimenCount id="9D2A6899FFC39350FDE4F8556C82CEF9" box="[531,646,2004,2028]" pageId="6" pageNumber="29" type="female">3 females</specimenCount>
(
<collectionCode id="ED3D3BD5FFC39350FD6BF8556CDBCEF9" box="[668,735,2004,2028]" pageId="6" pageNumber="29">MIIZ</collectionCode>
): “
<collectingCountry id="F33BE380FFC39350FCF4F8556D95CEF9" box="[771,913,2004,2028]" name="Madagascar" pageId="6" pageNumber="29">Madagascar</collectingCountry>
: Province Toliara, Andranomena,
<date id="FF9285D0FFC39351FADEF8556ED7C9BA" lastPageId="7" lastPageNumber="30" pageId="6" pageNumber="29" value="1990-03-29">29 March 1990</date>
”, ”W. E. Steiner, C. Kremen, V. Razafimahatratra, collectors”.
<specimenCount id="9D2A6899FFC29351FC5EFF166A1CC9BA" box="[937,1048,151,175]" pageId="7" pageNumber="30" type="female">2 females</specimenCount>
(
<collectionCode id="ED3D3BD5FFC29351FBDDFF166A69C9BA" box="[1066,1133,151,175]" pageId="7" pageNumber="30">MIIZ</collectionCode>
): “
<collectingCountry id="F33BE380FFC29351FB7BFF166B12C9BA" box="[1164,1302,151,175]" name="Madagascar" pageId="7" pageNumber="30">Madagascar</collectingCountry>
Sud-Ouest, LAC
<collectionCode id="ED3D3BD5FFC29351FF2EFF3D6F29C9C1" box="[217,301,188,212]" pageId="7" pageNumber="30">IOTRY</collectionCode>
<quantity id="4CD40EF5FFC29351FEC0FF3D6F68C9C1" box="[311,364,188,212]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="30" unit="m" value="40.0">40m</quantity>
, Morombe,
<date id="FF9285D0FFC29351FE0CFF3D6C42C9C1" box="[507,582,188,212]" pageId="7" pageNumber="30" value="1957-07">VII-57</date>
Andria R.”, “Muséum Paris”, “Institut Scientifique,
<collectingCountry id="F33BE380FFC29351FB58FF3D6B38C9C1" box="[1199,1340,188,212]" name="Madagascar" pageId="7" pageNumber="30">Madagascar</collectingCountry>
”.
<emphasis id="B9587F02FFC29351FAA3FF3A6EF8C9ED" bold="true" pageId="7" pageNumber="30">Other material</emphasis>
:
<specimenCount id="9D2A6899FFC29351FEFAFF5E6F62C9E2" box="[269,358,223,247]" pageId="7" pageNumber="30" type="male">4 males</specimenCount>
and
<specimenCount id="9D2A6899FFC29351FE55FF5E6C14C9E2" box="[418,528,223,247]" pageId="7" pageNumber="30" type="female">3 females</specimenCount>
(USNMNH): “
<collectingCountry id="F33BE380FFC29351FD35FF5E6D54C9E2" box="[706,848,223,247]" name="Madagascar" pageId="7" pageNumber="30">Madagascar</collectingCountry>
: Province Toliara, Andranomena,
<date id="FF9285D0FFC29351FB2FFF5E6B88C9E2" box="[1240,1420,223,247]" pageId="7" pageNumber="30" value="1990-03-29">29 March 1990</date>
”, ”W. E. Steiner, C. Kremen, V. Razafimahatratra, collectors”. Female (
<collectionCode id="ED3D3BD5FFC29351FC59FE856A0FC809" LSID="urn:lsid:biocol.org:col:34988" box="[942,1035,260,284]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="30">MNHN</collectionCode>
): “1957,
<collectingCountry id="F33BE380FFC29351FB85FE856B3DC809" box="[1138,1337,260,284]" name="Madagascar" pageId="7" pageNumber="30">MADAGASCAR</collectingCountry>
SUD, P. Renaud leg.”, “Museum Paris coll. P. Ardoin 1978”. Female (
<collectionCode id="ED3D3BD5FFC29351FCBDFEA66D9EC82A" box="[842,922,295,319]" pageId="7" pageNumber="30">CASC</collectionCode>
): “
<collectingCountry id="F33BE380FFC29351FC40FEA66A81C82A" box="[951,1157,295,319]" name="Madagascar" pageId="7" pageNumber="30">MADAGASCAR</collectingCountry>
: Toliara Prov. Sept Lacs, elev
<quantity id="4CD40EF5FFC29351FF38FECD6F00C871" box="[207,260,332,356]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="30" unit="m" value="80.0">80m</quantity>
,
<geoCoordinate id="EE18C5D7FFC29351FEE5FECD6F99C871" box="[274,413,332,356]" direction="south" orientation="latitude" pageId="7" pageNumber="30" precision="15" value="-23.528334">23°31`42``S</geoCoordinate>
<geoCoordinate id="EE18C5D7FFC29351FE50FECD6C2CC871" box="[423,552,332,356]" direction="east" orientation="longitude" pageId="7" pageNumber="30" precision="15" value="44.155556">44°9`20``E</geoCoordinate>
”,
<date id="FF9285D0FFC29351FDB4FECD6CE3C871" box="[579,743,332,356]" pageId="7" pageNumber="30" value="2002-03-08">8 March 2002</date>
” “California Acad. of Sciences, collectors: Frontier Project, gallery forest, leaf litter extraction, code:
<collectionCode id="ED3D3BD5FFC29351FD90FEEE6CA3C892" box="[615,679,367,391]" httpUri="http://grbio.org/cool/nsep-8f7n" name="Museum George Frey" pageId="7" pageNumber="30">MGF</collectionCode>
020”, “CASENT 8037308”.
<specimenCount id="9D2A6899FFC29351FC13FEEE6A3EC892" box="[996,1082,367,391]" pageId="7" pageNumber="30" type="male">4 males</specimenCount>
and
<specimenCount id="9D2A6899FFC29351FB86FEEE6AD9C892" box="[1137,1245,367,391]" pageId="7" pageNumber="30" type="female">3 females</specimenCount>
(
<collectionCode id="ED3D3BD5FFC29351FB1BFEEE6B42C892" box="[1260,1350,367,391]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="7" pageNumber="30">MRAC</collectionCode>
): “Coll. Mus. Tervuren
<collectingCountry id="F33BE380FFC29351FEB5FE156FCAC8B9" box="[322,462,404,428]" name="Madagascar" pageId="7" pageNumber="30">Madagascar</collectingCountry>
: L. Yotry, Tanandava,
<date id="FF9285D0FFC29351FD3EFE156D4CC8B9" box="[713,840,404,428]" pageId="7" pageNumber="30" value="1964-06-14">14.VI.1964</date>
G. Schmitz”.
</paragraph>
</subSubSection>
<caption id="DF53F398FFC29351FF60FAE16F68CC80" httpUri="https://zenodo.org/record/281842/files/figure.png" pageId="7" pageNumber="30" targetBox="[163,1417,497,1347]" targetPageId="7">
<paragraph id="8B93A310FFC29351FF60FAE16F68CC80" blockId="7.[151,1435,1376,1429]" pageId="7" pageNumber="30">
<emphasis id="B9587F02FFC29351FF60FAE16F49CC60" bold="true" box="[151,333,1376,1399]" pageId="7" pageNumber="30">FIGURES 20–24</emphasis>
. Body structure.
<taxonomicName id="4C2CD893FFC29351FE0BFAE06C79CC63" box="[508,637,1376,1398]" class="Insecta" family="Tenebrionidae" genus="Selinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="30" phylum="Arthropoda" rank="species" species="abacoides">
<emphasis id="B9587F02FFC29351FE0BFAE06C79CC63" box="[508,637,1376,1398]" italics="true" pageId="7" pageNumber="30">L. abacoides</emphasis>
</taxonomicName>
(20),
<taxonomicName id="4C2CD893FFC29351FD4CFAE06D41CC63" box="[699,837,1376,1398]" class="Insecta" family="Tenebrionidae" genus="Lechius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="30" phylum="Arthropoda" rank="species" species="longiclavis" status="sp. nov.">
<emphasis id="B9587F02FFC29351FD4CFAE06D41CC63" box="[699,837,1376,1398]" italics="true" pageId="7" pageNumber="30">L. longiclavis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26BC279FFC29351FCBBFAE06D98CC63" box="[844,924,1377,1398]" pageId="7" pageNumber="30" rank="species">
<emphasis id="B9587F02FFC29351FCBBFAE06D98CC63" bold="true" box="[844,924,1377,1398]" pageId="7" pageNumber="30">sp. nov.</emphasis>
</taxonomicNameLabel>
(21),
<taxonomicName id="4C2CD893FFC29351FC2DFAE06AAFCC63" box="[986,1195,1376,1398]" class="Insecta" family="Tenebrionidae" genus="Hovademulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="30" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B9587F02FFC29351FC2DFAE06AAFCC63" box="[986,1195,1376,1398]" italics="true" pageId="7" pageNumber="30">L. madagascariensis</emphasis>
</taxonomicName>
(22),
<taxonomicName id="4C2CD893FFC29351FB1EFAE06B41CC63" box="[1257,1349,1376,1398]" class="Insecta" family="Tenebrionidae" genus="Lechius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="30" phylum="Arthropoda" rank="species" species="aalbui" status="sp. nov.">
<emphasis id="B9587F02FFC29351FB1EFAE06B41CC63" box="[1257,1349,1376,1398]" italics="true" pageId="7" pageNumber="30">L. aalbui</emphasis>
</taxonomicName>
<emphasis id="B9587F02FFC29351FABCFAE06B9FCC63" bold="true" box="[1355,1435,1377,1398]" pageId="7" pageNumber="30">
<taxonomicNameLabel id="A26BC279FFC29351FABCFAE06B9FCC63" box="[1355,1435,1377,1398]" pageId="7" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(23),
<taxonomicName id="4C2CD893FFC29351FF3AFA016F30CC80" box="[205,308,1408,1429]" class="Insecta" family="Tenebrionidae" genus="Lechius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="30" phylum="Arthropoda" rank="species" species="steineri">
<emphasis id="B9587F02FFC29351FF3AFA016F30CC80" box="[205,308,1408,1429]" italics="true" pageId="7" pageNumber="30">L. steineri</emphasis>
</taxonomicName>
(24).
</paragraph>
</caption>
<subSubSection id="C336F09BFFC2935FFF3EFA646D31CE35" lastPageId="9" lastPageNumber="32" pageId="7" pageNumber="30" type="description">
<paragraph id="8B93A310FFC29351FF3EFA646CCCCE54" blockId="7.[151,1437,1509,2038]" pageId="7" pageNumber="30">
<emphasis id="B9587F02FFC29351FF3EFA646F72CCEB" bold="true" box="[201,374,1509,1534]" pageId="7" pageNumber="30">Redescription.</emphasis>
Measurements. Body—length = 12.0
<emphasis id="B9587F02FFC29351FCDEFA676D33CCEB" bold="true" box="[809,823,1510,1534]" pageId="7" pageNumber="30">–</emphasis>
13.0 mm; width ratio elytra/pronotum ca. 1.1; length ratio elytra/pronotum (pl) ca. 2.3. Clypeus—clypeal emargination width/depth ratio ca. 5.0. Antennae—length ratio antenna/pronotum (apl) ca. 0.9; ratio length of antenna/width of 3rd antennomere ca. 7.3; length ratio of antennomere 3rd/2nd ca. 3.0. Maxillary palp—ratio width of maxillary palp/length of 3rd antennomere ca. 1.0. Prono-tum—pronotum length (pl)/width ratio ca. 0.6; length ratio pronotum (pl)/anterior pronotal angles (apl) ca. 0.9. Elytra—elytra length/width ratio ca. 1.3. Scutellum—width ratio anterior margin of elytra/scutellum at base ca. 3.2. Metaventrite—length ratio cavity of hind coxa/metaventrite between the insertions of mid and hind coxae ca. 2.0. Abdomen—width ratio process of I abdominal ventrite/process of metaventrite ca. 2.0. Legs—male protibia length/width ratio ca. 4.5; female protibia length/width ratio ca. 4.5; both sexes profemur length/width ratio ca. 4.6. Tarsi—length ratio metatarsomere 1st/2nd ca. 2.8.
</paragraph>
<paragraph id="8B93A310FFC29351FF30F8CC6FB3CE70" blockId="7.[151,1437,1509,2038]" box="[199,439,1869,1893]" pageId="7" pageNumber="30">
Habitus as in
<figureCitation id="1317BF95FFC29351FE97F8CC6FB6CE70" box="[352,434,1869,1893]" captionStart="FIGURES 20 – 24" captionStartId="7.[151,264,1376,1399]" captionTargetBox="[163,1417,497,1347]" captionTargetId="figure@7.[151,1436,481,1355]" captionTargetPageId="7" captionText="FIGURES 20 – 24. Body structure. L. abacoides (20), L. longiclavis sp. nov. (21), L. madagascariensis (22), L. aalbui sp. nov. (23), L. steineri (24)." httpUri="https://zenodo.org/record/281842/files/figure.png" pageId="7" pageNumber="30">Fig. 24</figureCitation>
.
</paragraph>
<paragraph id="8B93A310FFC29351FF30F8F06A97CEB8" blockId="7.[151,1437,1509,2038]" pageId="7" pageNumber="30">Punctation on head dense and conspicuous (Fig. 5). Anterior margin of clypeus with shallow median emargination. Mentum slightly narrowing towards apex; submentum with elongated base (Fig. 33).</paragraph>
<paragraph id="8B93A310FFC29351FF30F8386DD4CEE0" blockId="7.[151,1437,1509,2038]" pageId="7" pageNumber="30">
Pronotal disc dull with conspicuous punctures (3
<emphasis id="B9587F02FFC29351FD0CF83B6D0DCEC7" bold="true" box="[763,777,1978,2002]" pageId="7" pageNumber="30">–</emphasis>
4 diameters apart). Lateral border of pronotum complete. Hypomeron with longitudinal depression near the lateral border (
<figureCitation id="1317BF95FFC29351FC98F85C6DC6CEE0" box="[879,962,2013,2037]" captionStart="FIGURES 8 – 16" captionStartId="4.[151,264,1090,1113]" captionTargetBox="[152,1431,197,1068]" captionTargetId="figure@4.[151,1436,195,1069]" captionTargetPageId="4" captionText="FIGURES 8 – 16. Chosen body elements: L. abacoides (8, 11, 14), L. longiclavis sp. nov. (9, 13), L. aalbui sp. nov. (10, 16), L. steineri (12), L. madagascariensis (15). Pronotum and its puncturation (8 – 10), pronotal hypomeron (11,12), elytral humeri (13), puncturation of elytral disc (14 – 16)." httpUri="https://zenodo.org/record/281840/files/figure.png" pageId="7" pageNumber="30">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="8B93A310FFCD935EFF60FBC16EEACD81" blockId="8.[151,1436,1088,1172]" pageId="8" pageNumber="31">
<emphasis id="B9587F02FFCD935EFF60FBC16F49CD40" bold="true" box="[151,333,1088,1111]" pageId="8" pageNumber="31">FIGURES 25–34</emphasis>
.
<taxonomicName id="4C2CD893FFCD935EFEADFBC06FDFCD43" box="[346,475,1088,1110]" class="Insecta" family="Tenebrionidae" genus="Selinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="abacoides">
<emphasis id="B9587F02FFCD935EFEADFBC06FDFCD43" box="[346,475,1088,1110]" italics="true" pageId="8" pageNumber="31">L. abacoides</emphasis>
</taxonomicName>
(25, 29, 32),
<taxonomicName id="4C2CD893FFCD935EFD93FBC06CEACD43" box="[612,750,1088,1110]" class="Insecta" family="Tenebrionidae" genus="Lechius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="longiclavis" status="sp. nov.">
<emphasis id="B9587F02FFCD935EFD93FBC06CEACD43" box="[612,750,1088,1110]" italics="true" pageId="8" pageNumber="31">L. longiclavis</emphasis>
</taxonomicName>
<emphasis id="B9587F02FFCD935EFD03FBC06D40CD43" bold="true" box="[756,836,1089,1110]" pageId="8" pageNumber="31">
<taxonomicNameLabel id="A26BC279FFCD935EFD03FBC06D40CD43" box="[756,836,1089,1110]" pageId="8" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(28, 34),
<taxonomicName id="4C2CD893FFCD935EFC5FFBC06A0BCD43" box="[936,1039,1089,1110]" class="Insecta" family="Tenebrionidae" genus="Lechius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="steineri">
<emphasis id="B9587F02FFCD935EFC5FFBC06A0BCD43" box="[936,1039,1089,1110]" italics="true" pageId="8" pageNumber="31">L. steineri</emphasis>
</taxonomicName>
(31, 33),
<taxonomicName id="4C2CD893FFCD935EFB85FBC06AC9CD43" box="[1138,1229,1088,1110]" class="Insecta" family="Tenebrionidae" genus="Lechius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="aalbui" status="sp. nov.">
<emphasis id="B9587F02FFCD935EFB85FBC06AC9CD43" box="[1138,1229,1088,1110]" italics="true" pageId="8" pageNumber="31">L. aalbui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26BC279FFCD935EFB23FBC06B20CD43" box="[1236,1316,1089,1110]" pageId="8" pageNumber="31" rank="species">
<emphasis id="B9587F02FFCD935EFB23FBC06B20CD43" bold="true" box="[1236,1316,1089,1110]" pageId="8" pageNumber="31">sp. nov.</emphasis>
</taxonomicNameLabel>
(27, 30),
<taxonomicName id="4C2CD893FFCD935EFA70FBC06F4ACD60" class="Insecta" family="Tenebrionidae" genus="Hovademulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B9587F02FFCD935EFA70FBC06F4ACD60" italics="true" pageId="8" pageNumber="31">L. madagascariensis</emphasis>
</taxonomicName>
(26). Aedeagal tegmen dorsal and ventral view (27
<emphasis id="B9587F02FFCD935EFC9AFBDE6D7DCD61" bold="true" box="[877,889,1119,1140]" pageId="8" pageNumber="31">–</emphasis>
27). Ovipositor (28). Spermatheca (29
<emphasis id="B9587F02FFCD935EFAFFFBDE6B11CD61" bold="true" box="[1288,1301,1119,1140]" pageId="8" pageNumber="31">–</emphasis>
31). Mentum (32
<emphasis id="B9587F02FFCD935EFF4FFBFF6EC1CD86" bold="true" box="[184,197,1150,1171]" pageId="8" pageNumber="31">–</emphasis>
34).
</paragraph>
<caption id="DF53F398FFCD935EFF60F8FF6DF1CE80" box="[151,1013,1918,1941]" httpUri="https://zenodo.org/record/281843/files/figure.png" pageId="8" pageNumber="31" targetBox="[158,1421,1231,1884]" targetPageId="8">
<paragraph id="8B93A310FFCD935EFF60F8FF6DF1CE80" blockId="8.[151,1013,1918,1941]" box="[151,1013,1918,1941]" pageId="8" pageNumber="31">
<emphasis id="B9587F02FFCD935EFF60F8FF6F57CE81" bold="true" box="[151,339,1918,1941]" pageId="8" pageNumber="31">FIGURES 35–37.</emphasis>
Ovipositor of
<taxonomicName id="4C2CD893FFCD935EFE10F8016C4ACE80" box="[487,590,1920,1941]" class="Insecta" family="Tenebrionidae" genus="Lechius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="steineri">
<emphasis id="B9587F02FFCD935EFE10F8016C4ACE80" box="[487,590,1920,1941]" italics="true" pageId="8" pageNumber="31">L. steineri</emphasis>
</taxonomicName>
: dorsal (35), ventral (36) and lateral (37).
</paragraph>
</caption>
<caption id="DF53F398FFCC935FFF60FA056C8ACCAF" httpUri="https://zenodo.org/record/281844/files/figure.png" pageId="9" pageNumber="32" targetBox="[417,1169,193,1390]" targetPageId="9">
<paragraph id="8B93A310FFCC935FFF60FA056C8ACCAF" blockId="9.[151,1437,1412,1466]" pageId="9" pageNumber="32">
<emphasis id="B9587F02FFCC935FFF60FA056F24CC8F" bold="true" box="[151,288,1412,1435]" pageId="9" pageNumber="32">FIGURE 38.</emphasis>
Distribution of the genus
<taxonomicName id="4C2CD893FFCC935FFDDDFA046C7DCC8E" box="[554,633,1413,1435]" class="Insecta" family="Tenebrionidae" genus="Lechius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B9587F02FFCC935FFDDDFA046C7DCC8E" box="[554,633,1413,1435]" italics="true" pageId="9" pageNumber="32">Lechius</emphasis>
</taxonomicName>
in Malagasy ecoregions (Species richness – number of species in a given area. Ecoregions were adopted after Olson et al. 2001).
</paragraph>
</caption>
<paragraph id="8B93A310FFCC935FFF30F98B6DE3CF53" blockId="9.[151,1436,1546,1969]" pageId="9" pageNumber="32">
Elytral row with punctures, most noticeable on the elytral slope. Interrows dull with punctures 4
<emphasis id="B9587F02FFCC935FFAFCF98B6B1DCF37" bold="true" box="[1291,1305,1546,1570]" pageId="9" pageNumber="32">–</emphasis>
5 diameters apart. Elytral base sinusoidal. Elytral humeri rounded. Scutellum triangular.
</paragraph>
<paragraph id="8B93A310FFCC935FFF30F9D46FD8CF85" blockId="9.[151,1436,1546,1969]" pageId="9" pageNumber="32">
Intercoxal process of prosternum straight with complete bordering (as in
<taxonomicName id="4C2CD893FFCC935FFBFDF9D76AA0CF78" box="[1034,1188,1621,1645]" class="Insecta" family="Tenebrionidae" genus="Lechius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="longiclavis" status="sp. nov.">
<emphasis id="B9587F02FFCC935FFBFDF9D76AA0CF78" box="[1034,1188,1621,1645]" italics="true" pageId="9" pageNumber="32">L. longiclavis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26BC279FFCC935FFB5AF9D46B0CCF78" box="[1197,1288,1621,1645]" pageId="9" pageNumber="32" rank="species">
<emphasis id="B9587F02FFCC935FFB5AF9D46B0CCF78" bold="true" box="[1197,1288,1621,1645]" pageId="9" pageNumber="32">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<figureCitation id="1317BF95FFCC935FFAEFF9D46B68CF78" box="[1304,1388,1621,1645]" captionStart="FIGURES 17 – 19" captionStartId="6.[151,264,1088,1111]" captionTargetBox="[154,1435,195,1063]" captionTargetId="figure@6.[151,1436,193,1067]" captionTargetPageId="6" captionText="FIGURES 17 – 19. Intercoxal process of prosternum of L. longiclavis sp. nov. (17). Male mesofemora of L. madagascariensis (18). Distribution of the species of Lechius (19)." httpUri="https://zenodo.org/record/281841/files/figure.png" pageId="9" pageNumber="32">Fig. 17</figureCitation>
). 5th abdominal ventrite bordered.
</paragraph>
<paragraph id="8B93A310FFCC935FFF30F91C6DD0CFA0" blockId="9.[151,1436,1546,1969]" box="[199,980,1693,1717]" pageId="9" pageNumber="32">
Male legs as in
<taxonomicName id="4C2CD893FFCC935FFE81F91F6C0BCFA0" box="[374,527,1693,1717]" class="Insecta" family="Tenebrionidae" genus="Lechius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="longiclavis" status="sp. nov.">
<emphasis id="B9587F02FFCC935FFE81F91F6C0BCFA0" box="[374,527,1693,1717]" italics="true" pageId="9" pageNumber="32">L. longiclavis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26BC279FFCC935FFDE1F91C6C6BCFA0" box="[534,623,1693,1717]" pageId="9" pageNumber="32" rank="species">
<emphasis id="B9587F02FFCC935FFDE1F91C6C6BCFA0" bold="true" box="[534,623,1693,1717]" pageId="9" pageNumber="32">sp. nov.</emphasis>
</taxonomicNameLabel>
(Fig. 6, 7). Female legs simple.
</paragraph>
<paragraph id="8B93A310FFCC935FFF30F9416F0FCFE8" blockId="9.[151,1436,1546,1969]" pageId="9" pageNumber="32">Apical part of aedeagal tegmen equal to 0.25 of the rest of tegmen. Parameres narrowing toward apex. Clavae elongated.</paragraph>
<paragraph id="8B93A310FFCC935FFF30F8896D31CE35" blockId="9.[151,1436,1546,1969]" box="[199,821,1800,1825]" pageId="9" pageNumber="32">
Ovipositor as in
<figureCitation id="1317BF95FFCC935FFE76F8896C05CE35" box="[385,513,1800,1825]" captionStart="FIGURES 35 – 37" captionStartId="8.[151,264,1918,1941]" captionTargetBox="[158,1421,1231,1884]" captionTargetId="figure@8.[151,1436,1221,1897]" captionTargetPageId="8" captionText="FIGURES 35 – 37. Ovipositor of L. steineri: dorsal (35), ventral (36) and lateral (37)." httpUri="https://zenodo.org/record/281843/files/figure.png" pageId="9" pageNumber="32">
Figs 35
<emphasis id="B9587F02FFCC935FFE23F8886FE6CE34" bold="true" box="[468,482,1801,1825]" pageId="9" pageNumber="32">–</emphasis>
37
</figureCitation>
. Spermatheca as in Fig. 31.
</paragraph>
</subSubSection>
<subSubSection id="C336F09BFFCC935FFF3EF8AD6DBDCE98" pageId="9" pageNumber="32" type="discussion">
<paragraph id="8B93A310FFCC935FFF3EF8AD6DBDCE98" blockId="9.[151,1436,1546,1969]" pageId="9" pageNumber="32">
<emphasis id="B9587F02FFCC935FFF3EF8AD6F55CE50" bold="true" box="[201,337,1836,1861]" pageId="9" pageNumber="32">Comments.</emphasis>
Following sampling periods (a), habitat descriptions (b) and altitudes (c) were recorded from specimen labels: (a) VII 1957,
<date id="FF9285D0FFCC935FFE04F8D16C71CE7D" box="[499,629,1872,1896]" pageId="9" pageNumber="32" value="1964-06-14">14 VI 1964</date>
,
<date id="FF9285D0FFCC935FFD76F8D16D05CE7D" box="[641,769,1872,1896]" pageId="9" pageNumber="32" value="1990-03-29">29 III 1990</date>
,
<date id="FF9285D0FFCC935FFCF9F8D16D86CE7D" box="[782,898,1872,1896]" pageId="9" pageNumber="32" value="2002-03-08">8 III 2002</date>
; (b) Gallery forest (leaf litter extraction), Under leaf litter on loose sandy soil in partial shade and shrubs; (c)
<quantity id="4CD40EF5FFCC935FFCB7F8F46D70CE98" box="[832,884,1909,1933]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="32" unit="m" value="40.0">40m</quantity>
,
<specimenCount id="9D2A6899FFCC935FFC77F8F46DBDCE98" box="[896,953,1909,1933]" pageId="9" pageNumber="32" type="generic">
<quantity id="4CD40EF5FFCC935FFC77F8F46DB1CE98" box="[896,949,1909,1933]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="32" unit="m" value="80.0">80m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="C336F09BFFCC935FFF3EF8196D51CEA5" box="[201,853,1944,1969]" pageId="9" pageNumber="32" type="distribution">
<paragraph id="8B93A310FFCC935FFF3EF8196D51CEA5" blockId="9.[151,1436,1546,1969]" box="[201,853,1944,1969]" pageId="9" pageNumber="32">
<emphasis id="B9587F02FFCC935FFF3EF8196F66CEA4" bold="true" box="[201,354,1944,1969]" pageId="9" pageNumber="32">Distribution.</emphasis>
Central and Southern
<collectingCountry id="F33BE380FFCC935FFDA9F8196CE1CEA5" box="[606,741,1944,1968]" name="Madagascar" pageId="9" pageNumber="32">Madagascar</collectingCountry>
(
<figureCitation id="1317BF95FFCC935FFD03F8196D4CCEA5" box="[756,840,1944,1968]" captionStart="FIGURES 17 – 19" captionStartId="6.[151,264,1088,1111]" captionTargetBox="[154,1435,195,1063]" captionTargetId="figure@6.[151,1436,193,1067]" captionTargetPageId="6" captionText="FIGURES 17 – 19. Intercoxal process of prosternum of L. longiclavis sp. nov. (17). Male mesofemora of L. madagascariensis (18). Distribution of the species of Lechius (19)." httpUri="https://zenodo.org/record/281841/files/figure.png" pageId="9" pageNumber="32">Fig. 19</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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