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<document id="5EFCCD85F31D1B7F746A46605D8ED47E" ID-CLB-Dataset="8752" ID-DOI="10.11646/zootaxa.4899.1.16" ID-GBIF-Dataset="dde87a19-79a2-44a3-8533-b8970944cb99" ID-ISSN="1175-5326" ID-Zenodo-Dep="4456831" ID-ZooBank="27E67BBB-DFD0-4A96-8269-9E1CB6153B83" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609326131882" checkinUser="plazi" docAuthor="Salgado, Alexandre &amp; Ruiz, Gustavo R. S." docDate="2020" docId="03403F11FFBEFFA3538BFF73082DF8EE" docLanguage="en" docName="zootaxa.4899.1.16.pdf" docOrigin="Zootaxa 4899 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Marma wesolowskae Salgado &amp; Ruiz 2020, sp. nov." docType="treatment" docVersion="8" lastPageNumber="322" masterDocId="FF794769FF9EFF80531CFFE40C1AFF88" masterDocTitle="Taxonomic revision of Marma Simon, 1902 (Araneae: Salticidae: Euophryini)" masterLastPageNumber="353" masterPageNumber="287" pageNumber="319" updateTime="1698848557116" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="BD8C093643E245D8919ECAAFA3284D91">Taxonomic revision of Marma Simon, 1902 (Araneae: Salticidae: Euophryini)</mods:title>
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<mods:namePart id="26612D6770FB4A03A1ADCD61275E5E8E">Salgado, Alexandre</mods:namePart>
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<mods:affiliation id="DEB888FB937D9D3E55C75B429CD58E7F">Programa de Pós-Graduação em Zoologia, Universidade Federal do Pará / Museu Paraense Emílio Goeldi, Belém, PA, Brazil Instituto de Ciências Biológicas, Universidade Federal do Pará, Rua Augusto Corrêa, 01, CEP 66075 - 110, Belém, PA, Brazil</mods:affiliation>
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<mods:namePart id="6702D1BD6E1AD29D41372707EC24F89B">Ruiz, Gustavo R. S.</mods:namePart>
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<mods:affiliation id="4ED1CD1F3F10C5EDD5E1DEB847778E2E">Instituto de Ciências Biológicas, Universidade Federal do Pará, Rua Augusto Corrêa, 01, CEP 66075 - 110, Belém, PA, Brazil</mods:affiliation>
<mods:nameIdentifier id="383E4A8E8D78B4DFE4CF51069452E070" type="email">gustavoruiz86@hotmail.com</mods:nameIdentifier>
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<treatment id="03403F11FFBEFFA3538BFF73082DF8EE" ID-DOI="http://doi.org/10.5281/zenodo.4456881" ID-GBIF-Taxon="177010979" ID-Zenodo-Dep="4456881" LSID="urn:lsid:plazi:treatment:03403F11FFBEFFA3538BFF73082DF8EE" httpUri="http://treatment.plazi.org/id/03403F11FFBEFFA3538BFF73082DF8EE" lastPageId="35" lastPageNumber="322" pageId="32" pageNumber="319">
<subSubSection id="C3F3DD8CFFBEFFA0538BFF730DE2FF39" box="[151,504,151,178]" pageId="32" pageNumber="319" type="nomenclature">
<paragraph id="8B568E07FFBEFFA0538BFF730DE2FF39" blockId="32.[151,504,151,214]" box="[151,504,151,178]" pageId="32" pageNumber="319">
<heading id="D01E396BFFBEFFA0538BFF730DE2FF39" bold="true" box="[151,504,151,178]" fontSize="11" level="1" pageId="32" pageNumber="319" reason="1">
<emphasis id="B99D5215FFBEFFA0538BFF730DE2FF39" bold="true" box="[151,504,151,178]" pageId="32" pageNumber="319">
<taxonomicName id="4CE9F584FFBEFFA0538BFF730D89FF3A" ID-CoL="98Z86" authority="Salgado &amp; Ruiz, 2020" authorityName="Salgado &amp; Ruiz" authorityYear="2020" box="[151,403,151,178]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="319" phylum="Arthropoda" rank="species" species="wesolowskae" status="sp. nov.">
<emphasis id="B99D5215FFBEFFA0538BFF730D89FF3A" bold="true" box="[151,403,151,178]" italics="true" pageId="32" pageNumber="319">Marma wesolowskae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2AEEF6EFFBEFFA05286FF730DE2FF39" box="[410,504,151,177]" pageId="32" pageNumber="319" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3F3DD8CFFBEFFA0538BFF5F0DC7FF5E" box="[151,477,187,214]" pageId="32" pageNumber="319" type="description">
<paragraph id="8B568E07FFBEFFA0538BFF5F0DC7FF5E" blockId="32.[151,504,151,214]" box="[151,477,187,214]" pageId="32" pageNumber="319">
<figureCitation id="13D29282FFBEFFA0538BFF5F0D09FF5D" box="[151,275,187,213]" captionStart="FIGURE 1" captionStartId="3.[151,250,1910,1934]" captionTargetBox="[157,1430,309,1887]" captionTargetId="figure-90@3.[157,1430,309,1887]" captionTargetPageId="3" captionText="FIGURE 1. Marma Simon, 1902. AE live specimens in different habitats [A M. nigritarsis from Belém/Pará/Brazil, on urban construction (Photo credit: César Favacho); B M. linae sp. nov. from São Geraldo do Araguaia/Pará/Brazil, on tree trunk; C M. rosea from São Geraldo do Araguaia/Pará/Brazil, on a wall of an urban construction; D M. nigritarsis from Belém/Pará/Brazil, on a substrate with moss; E M. wesolowskae sp. nov. from Buíque/Pernambuco/Brazil, on rocks]." figureDoi="http://doi.org/10.5281/zenodo.4456835" httpUri="https://zenodo.org/record/4456835/files/figure.png" pageId="32" pageNumber="319">Figures 1E</figureCitation>
,
<figureCitation id="13D29282FFBEFFA05203FF5F0D72FF5D" box="[287,360,187,213]" captionStart-0="FIGURE 29" captionStart-1="FIGURE 30" captionStart-2="FIGURE 31" captionStart-3="FIGURE 32" captionStart-4="FIGURE 33" captionStartId-0="32.[151,250,1980,2004]" captionStartId-1="33.[151,250,1960,1984]" captionStartId-2="34.[151,250,1720,1744]" captionStartId-3="35.[151,250,1365,1389]" captionStartId-4="36.[151,250,1784,1808]" captionTargetBox-0="[151,1435,340,1957]" captionTargetBox-1="[151,1435,1173,1937]" captionTargetBox-2="[151,1435,489,1697]" captionTargetBox-3="[152,1405,191,1324]" captionTargetBox-4="[151,1435,181,1761]" captionTargetId-0="figure-60@32.[151,1435,340,1957]" captionTargetId-1="figure-480@33.[151,1435,1173,1937]" captionTargetId-2="figure-168@34.[151,1435,489,1697]" captionTargetId-3="figure-17@35.[151,1435,181,1341]" captionTargetId-4="figure-24@36.[151,1435,181,1761]" captionTargetPageId-0="32" captionTargetPageId-1="33" captionTargetPageId-2="34" captionTargetPageId-3="35" captionTargetPageId-4="36" captionText-0="FIGURE 29. Marma wesolowskae sp. nov. live specimens from Buíque/Pernambuco/Brazil on sandrock. AC male (AB dorsolateral, C dorsal); DF female (DE dorsolateral, F dorsal)." captionText-1="FIGURE 30. Male holotype of Marma wesolowskae sp. nov. (MPEG 34355). AB habitus (A dorsal, B ventral); CD left palp (C ventral; D retrolateral)." captionText-2="FIGURE 31. Female paratypes of Marma wesolowskae sp. nov. AB habitus, MPEG 34354 (A dorsal, B ventral); CD epi- gyne/vulva, MPEG 37131 (C ventral, D dorsal). Abbreviations: BG—Bennetts gland; cd—copulatory duct; co—copulatory opening; fd—fertilization duct; PS—Primary spermatheca; SS—secondary spermatheca." captionText-3="FIGURE 32. Marma wesolowskae sp. nov. AE left male palp (A ventral; B retrolateral; C proventral; D dorsal; E cleared bulb, ventral); FG epigyne/vulva (F ventral; G cleared, ventral).Abbreviations: BG—Bennetts gland; cc—cymbial conductor; cdcopulatory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PED—process on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca." captionText-4="FIGURE 33. Marma wesolowskae sp. nov. AI left male palp [A ventral; B apical portion of palp, ventral; C tegular shoulder, ventral; D dissected embolic disc and embolus, dorsal; E patella and tibia, retrolateral; F tip of RTA, retrolateral; G detail of patella and tibia setae (blue rectangle in 33E); H detail of patella setae (red rectangle in 33E); I detail of smooth scale of tibia (yellow arrow in 33E]. J left spermatheca and copulatory duct, ventral. Abbreviations: cd—copulatory duct; e—embolus shaft; PED—process on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca. Black arrow in C shows depression of tegular shoulder; in D shows groove; white arrows in D show spikes of embolus shaft; in J shows end region of the primary spermatheca (broken fertilization duct)." figureDoi-0="http://doi.org/10.5281/zenodo.4456917" figureDoi-1="http://doi.org/10.5281/zenodo.4456919" figureDoi-2="http://doi.org/10.5281/zenodo.4456921" figureDoi-3="http://doi.org/10.5281/zenodo.4456923" figureDoi-4="http://doi.org/10.5281/zenodo.4456925" httpUri-0="https://zenodo.org/record/4456917/files/figure.png" httpUri-1="https://zenodo.org/record/4456919/files/figure.png" httpUri-2="https://zenodo.org/record/4456921/files/figure.png" httpUri-3="https://zenodo.org/record/4456923/files/figure.png" httpUri-4="https://zenodo.org/record/4456925/files/figure.png" pageId="32" pageNumber="319">2933</figureCitation>
,
<figureCitation id="13D29282FFBEFFA0526FFF5F0DBCFF5E" box="[371,422,187,214]" captionStart="FIGURE 57" captionStartId="62.[151,250,1843,1867]" captionTargetBox="[159,1422,193,1813]" captionTargetId="figure-24@62.[151,1435,181,1819]" captionTargetPageId="62" captionText="FIGURE 57. Left male palp, ventral [A Marma baeri Simon, 1902; B M. linae sp. nov.; C M. pipa sp. nov.; D M. wesolowskae sp. nov.; E M. nigritarsis (Simon, 1900); F M. sinuosa sp. nov., G M. argentina (Mello-Leit„o, 1941), H M. rosea (Mello- Leit„o, 1941), I M. spelunca sp. nov.]. Colors represent homologous structures." figureDoi="http://doi.org/10.5281/zenodo.4456979" httpUri="https://zenodo.org/record/4456979/files/figure.png" pageId="32" pageNumber="319">57D</figureCitation>
,
<figureCitation id="13D29282FFBEFFA052AEFF5F0DC7FF5E" box="[434,477,187,214]" captionStart="FIGURE 58" captionStartId="63.[151,250,1595,1619]" captionTargetBox="[156,1429,271,1549]" captionTargetId="figure-41@63.[151,1435,237,1571]" captionTargetPageId="63" captionText="FIGURE 58. Epigyne/vulva, AC, EK ventral, D dorsal [A Marma baeri Simon, 1902; B M. linae sp. nov.; C M. abaira sp. nov.; D M. femella (Caporiacco, 1955); E M. pipa sp. nov.; F M. wesolowskae sp. nov.; G M. nigritarsis (Simon, 1900); H M. sinuosa sp. nov.; I M. argentina (Mello-Leitão, 1941); J M. rosea (Mello-Leitão, 1941); K M. spelunca sp. nov.]. Colors represent homologous structures; arrow in D shows hypothetical position of secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456981" httpUri="https://zenodo.org/record/4456981/files/figure.png" pageId="32" pageNumber="319">58F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3F3DD8CFFBEFFA0538BFF1A088DFEB4" pageId="32" pageNumber="319" type="etymology">
<paragraph id="8B568E07FFBEFFA0538BFF1A088DFEB4" blockId="32.[151,1436,254,316]" pageId="32" pageNumber="319">
<emphasis id="B99D5215FFBEFFA0538BFF1A0D04FE90" bold="true" box="[151,286,254,280]" pageId="32" pageNumber="319">Etymology.</emphasis>
The epithet commemorates the career and life of Polish Professor Wanda Wesołowska. During her career, she has authored/co-authored the description of hundreds of jumping spider species.
</paragraph>
</subSubSection>
<caption id="DF96DE8FFFBEFFA0538BF8580F32F870" ID-DOI="http://doi.org/10.5281/zenodo.4456917" ID-Zenodo-Dep="4456917" httpUri="https://zenodo.org/record/4456917/files/figure.png" pageId="32" pageNumber="319" startId="32.[151,250,1980,2004]" targetBox="[151,1435,340,1957]" targetPageId="32">
<paragraph id="8B568E07FFBEFFA0538BF8580F32F870" blockId="32.[151,1436,1980,2040]" pageId="32" pageNumber="319">
<emphasis id="B99D5215FFBEFFA0538BF8580D39F85C" bold="true" box="[151,291,1980,2004]" pageId="32" pageNumber="319">FIGURE 29.</emphasis>
<taxonomicName id="4CE9F584FFBEFFA05230F8590E1EF85C" authority="Salgado &amp; Ruiz, 2020" authorityName="Salgado &amp; Ruiz" authorityYear="2020" box="[300,516,1980,2004]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="319" phylum="Arthropoda" rank="species" species="wesolowskae" status="sp. nov.">
<emphasis id="B99D5215FFBEFFA05230F8590E1EF85C" box="[300,516,1980,2004]" italics="true" pageId="32" pageNumber="319">Marma wesolowskae</emphasis>
</taxonomicName>
<emphasis id="B99D5215FFBEFFA05111F8580E7BF85C" bold="true" box="[525,609,1980,2004]" pageId="32" pageNumber="319">
<taxonomicNameLabel id="A2AEEF6EFFBEFFA05111F8580E7BF85C" box="[525,609,1980,2004]" pageId="32" pageNumber="319" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
live specimens from Buíque/Pernambuco/Brazil on sandrock. AC male (AB dorsolateral, C dorsal); DF female (DE dorsolateral, F dorsal).
</paragraph>
</caption>
<subSubSection id="C3F3DD8CFFBFFFA153DBFF720E37FE25" pageId="33" pageNumber="320" type="diagnosis">
<paragraph id="8B568E07FFBFFFA153DBFF720E37FE25" blockId="33.[151,1437,150,1149]" pageId="33" pageNumber="320">
<emphasis id="B99D5215FFBFFFA153DBFF720D58FF38" bold="true" box="[199,322,150,176]" pageId="33" pageNumber="320">Diagnosis.</emphasis>
Among all
<taxonomicName id="4CE9F584FFBFFFA152DAFF730E02FF38" authorityName="Simon" authorityYear="1902" box="[454,536,151,176]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="320" phylum="Arthropoda" rank="genus">
<emphasis id="B99D5215FFBFFFA152DAFF730E02FF38" box="[454,536,151,176]" italics="true" pageId="33" pageNumber="320">Marma</emphasis>
</taxonomicName>
species,
<taxonomicName id="4CE9F584FFBFFFA15162FF730F73FF39" authority="Salgado &amp; Ruiz, 2020" authorityName="Salgado &amp; Ruiz" authorityYear="2020" box="[638,873,151,177]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="320" phylum="Arthropoda" rank="species" species="wesolowskae" status="sp. nov.">
<emphasis id="B99D5215FFBFFFA15162FF730F73FF39" box="[638,873,151,177]" italics="true" pageId="33" pageNumber="320">Marma wesolowskae</emphasis>
</taxonomicName>
<emphasis id="B99D5215FFBFFFA1506DFF730FD0FF39" bold="true" box="[881,970,151,177]" pageId="33" pageNumber="320">
<taxonomicNameLabel id="A2AEEF6EFFBFFFA1506DFF730FD0FF39" box="[881,970,151,177]" pageId="33" pageNumber="320" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
shares most similarities with
<taxonomicName id="4CE9F584FFBFFFA1560BFF730CAFFF5C" baseAuthorityName="Simon" baseAuthorityYear="1900" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="320" phylum="Arthropoda" rank="species" species="nigritarsis">
<emphasis id="B99D5215FFBFFFA1560BFF730CAFFF5C" italics="true" pageId="33" pageNumber="320">M. nigritarsis</emphasis>
</taxonomicName>
and
<taxonomicName id="4CE9F584FFBFFFA153F5FF5F0D25FF5C" authority="Salgado &amp; Ruiz, 2020" authorityName="Salgado &amp; Ruiz" authorityYear="2020" box="[233,319,187,212]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="320" phylum="Arthropoda" rank="species" species="pipa" status="sp. nov.">
<emphasis id="B99D5215FFBFFFA153F5FF5F0D25FF5C" box="[233,319,187,212]" italics="true" pageId="33" pageNumber="320">M. pipa</emphasis>
</taxonomicName>
<emphasis id="B99D5215FFBFFFA15259FF5F0D84FF5D" bold="true" box="[325,414,187,213]" pageId="33" pageNumber="320">
<taxonomicNameLabel id="A2AEEF6EFFBFFFA15259FF5F0D84FF5D" box="[325,414,187,213]" pageId="33" pageNumber="320" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
The males of these species have an embolic disc with a straight prolateral border and curved retrolateral border (
<figureCitation id="13D29282FFBFFFA1526EFF3B0DC6FF71" box="[370,476,222,249]" captionStart="FIGURE 25" captionStartId="28.[151,250,1819,1843]" captionTargetBox="[163,1432,927,1783]" captionTargetId="figure-290@28.[151,1435,921,1796]" captionTargetPageId="28" captionText="FIGURE 25. Male holotype of Marma pipa sp. nov. (MPEG 34352). AB habitus (A dorsal, B ventral); CD left palp (C ventral; D retrolateral)." figureDoi="http://doi.org/10.5281/zenodo.4456909" httpUri="https://zenodo.org/record/4456909/files/figure.png" pageId="33" pageNumber="320">Figs 25C</figureCitation>
,
<figureCitation id="13D29282FFBFFFA152F4FF3B0E01FF71" box="[488,539,223,249]" captionStart="FIGURE 27" captionStartId="30.[151,250,1858,1882]" captionTargetBox="[154,1416,859,1817]" captionTargetId="figure-284@30.[151,1435,849,1835]" captionTargetPageId="30" captionText="FIGURE 27. Marma pipa sp. nov. AE left male palp (A ventral; B retrolateral; C proventral; D dorsal; E cleared bulb, ventral); FG epigyne/vulva (F ventral; G cleared, ventral). Abbreviations: BG—Bennetts gland; cc—cymbial conductor; cd—copu- latory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PED—process on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456913" httpUri="https://zenodo.org/record/4456913/files/figure.png" pageId="33" pageNumber="320">27A</figureCitation>
,
<figureCitation id="13D29282FFBFFFA1513AFF3B0E4DFF71" box="[550,599,223,249]" captionStart="FIGURE 30" captionStartId="33.[151,250,1960,1984]" captionTargetBox="[151,1435,1173,1937]" captionTargetId="figure-480@33.[151,1435,1173,1937]" captionTargetPageId="33" captionText="FIGURE 30. Male holotype of Marma wesolowskae sp. nov. (MPEG 34355). AB habitus (A dorsal, B ventral); CD left palp (C ventral; D retrolateral)." figureDoi="http://doi.org/10.5281/zenodo.4456919" httpUri="https://zenodo.org/record/4456919/files/figure.png" pageId="33" pageNumber="320">30C</figureCitation>
,
<figureCitation id="13D29282FFBFFFA15178FF3B0E8DFF71" box="[612,663,223,249]" captionStart="FIGURE 32" captionStartId="35.[151,250,1365,1389]" captionTargetBox="[152,1405,191,1324]" captionTargetId="figure-17@35.[151,1435,181,1341]" captionTargetPageId="35" captionText="FIGURE 32. Marma wesolowskae sp. nov. AE left male palp (A ventral; B retrolateral; C proventral; D dorsal; E cleared bulb, ventral); FG epigyne/vulva (F ventral; G cleared, ventral).Abbreviations: BG—Bennetts gland; cc—cymbial conductor; cdcopulatory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PED—process on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456923" httpUri="https://zenodo.org/record/4456923/files/figure.png" pageId="33" pageNumber="320">32A</figureCitation>
,
<figureCitation id="13D29282FFBFFFA151BEFF3B0EC9FF71" box="[674,723,223,249]" captionStart="FIGURE 36" captionStartId="39.[151,250,1890,1914]" captionTargetBox="[151,1435,345,1867]" captionTargetId="figure-106@39.[151,1435,345,1867]" captionTargetPageId="39" captionText="FIGURE 36. Marma nigritarsis (Simon, 1900), specimens from Belém/Pará/Brasil. AD male, MPEG 34357 (A habitus, dorsal; B ventral; C left palp, ventral; D retrolateral). EF female, MPEG 34358 (E habitus, dorsal; F ventral)." figureDoi="http://doi.org/10.5281/zenodo.4456931" httpUri="https://zenodo.org/record/4456931/files/figure.png" pageId="33" pageNumber="320">36C</figureCitation>
,
<figureCitation id="13D29282FFBFFFA151C3FF3B0F08FF71" box="[735,786,223,249]" captionStart="FIGURE 37" captionStartId="40.[151,250,1798,1822]" captionTargetBox="[159,1411,194,1759]" captionTargetId="figure-24@40.[151,1435,181,1773]" captionTargetPageId="40" captionText="FIGURE 37. Marma nigritarsis (Simon, 1900). AE, G specimen from Belém/Pará/Brazil. F holotype illustrated by Galiano (1962), reflected.AG left male palp (A ventral, B retrolateral, C proventral, D dorsal, EF retroventral; G cleared bulb, ventral). HI female holotype of M. trifidocarinata Caporiacco, 1947, epigyne/vulva (H ventral; I cleared, ventral).Abbreviations: BG— Bennetts gland; cc—cymbial conductor; cd—copulatory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PED—process on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456933" httpUri="https://zenodo.org/record/4456933/files/figure.png" pageId="33" pageNumber="320">37A</figureCitation>
,
<figureCitation id="13D29282FFBFFFA15002FF3A0F21FF71" box="[798,827,222,249]" captionStart="FIGURE 57" captionStartId="62.[151,250,1843,1867]" captionTargetBox="[159,1422,193,1813]" captionTargetId="figure-24@62.[151,1435,181,1819]" captionTargetPageId="62" captionText="FIGURE 57. Left male palp, ventral [A Marma baeri Simon, 1902; B M. linae sp. nov.; C M. pipa sp. nov.; D M. wesolowskae sp. nov.; E M. nigritarsis (Simon, 1900); F M. sinuosa sp. nov., G M. argentina (Mello-Leit„o, 1941), H M. rosea (Mello- Leit„o, 1941), I M. spelunca sp. nov.]. Colors represent homologous structures." figureDoi="http://doi.org/10.5281/zenodo.4456979" httpUri="https://zenodo.org/record/4456979/files/figure.png" pageId="33" pageNumber="320">57</figureCitation>
CE). Besides, in the females, the copulatory ducts do not touch each other and they encircle the primary spermathecae (
<figureCitation id="13D29282FFBFFFA1508EFEE70FF0FE95" box="[914,1002,259,285]" captionStart="FIGURE 26" captionStartId="29.[151,250,1842,1866]" captionTargetBox="[166,1421,723,1797]" captionTargetId="figure-286@29.[151,1435,705,1819]" captionTargetPageId="29" captionText="FIGURE 26. Female paratype of Marma pipa sp. nov (MPEG 34349). AB habitus (A dorsal, B ventral); CD cleared epigyne/ vulva (C ventral, D dorsal). Abbreviations: BG—Bennetts gland; cd—copulatory duct; co—copulatory opening; ets—endothe- cal spikes; fd—fertilization duct; PS—Primary spermatheca; SS—secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456911" httpUri="https://zenodo.org/record/4456911/files/figure.png" pageId="33" pageNumber="320">Figs 26</figureCitation>
CD, 27FG, 31CD, 32FG, 35HI, 37HI, 58EG). The males of
<taxonomicName id="4CE9F584FFBFFFA152F6FEC30EBBFEC9" authority="Salgado &amp; Ruiz, 2020" authorityName="Salgado &amp; Ruiz" authorityYear="2020" box="[490,673,295,321]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="320" phylum="Arthropoda" rank="species" species="wesolowskae" status="sp. nov.">
<emphasis id="B99D5215FFBFFFA152F6FEC30EBBFEC9" box="[490,673,295,321]" italics="true" pageId="33" pageNumber="320">M. wesolowskae</emphasis>
</taxonomicName>
<emphasis id="B99D5215FFBFFFA151B4FEC30F1BFEC9" bold="true" box="[680,769,295,321]" pageId="33" pageNumber="320">
<taxonomicNameLabel id="A2AEEF6EFFBFFFA151B4FEC30F1BFEC9" box="[680,769,295,321]" pageId="33" pageNumber="320" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differ from the males of these species by having a narrower embolic disc (
<figureCitation id="13D29282FFBFFFA1522BFEAF0DBBFEED" box="[311,417,331,357]" captionStart="FIGURE 30" captionStartId="33.[151,250,1960,1984]" captionTargetBox="[151,1435,1173,1937]" captionTargetId="figure-480@33.[151,1435,1173,1937]" captionTargetPageId="33" captionText="FIGURE 30. Male holotype of Marma wesolowskae sp. nov. (MPEG 34355). AB habitus (A dorsal, B ventral); CD left palp (C ventral; D retrolateral)." figureDoi="http://doi.org/10.5281/zenodo.4456919" httpUri="https://zenodo.org/record/4456919/files/figure.png" pageId="33" pageNumber="320">Figs 30C</figureCitation>
,
<figureCitation id="13D29282FFBFFFA152B2FEAF0DFBFEED" box="[430,481,331,357]" captionStart="FIGURE 32" captionStartId="35.[151,250,1365,1389]" captionTargetBox="[152,1405,191,1324]" captionTargetId="figure-17@35.[151,1435,181,1341]" captionTargetPageId="35" captionText="FIGURE 32. Marma wesolowskae sp. nov. AE left male palp (A ventral; B retrolateral; C proventral; D dorsal; E cleared bulb, ventral); FG epigyne/vulva (F ventral; G cleared, ventral).Abbreviations: BG—Bennetts gland; cc—cymbial conductor; cdcopulatory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PED—process on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456923" httpUri="https://zenodo.org/record/4456923/files/figure.png" pageId="33" pageNumber="320">32A</figureCitation>
,
<figureCitation id="13D29282FFBFFFA152F2FEAE0E10FEED" box="[494,522,330,357]" captionStart="FIGURE 57" captionStartId="62.[151,250,1843,1867]" captionTargetBox="[159,1422,193,1813]" captionTargetId="figure-24@62.[151,1435,181,1819]" captionTargetPageId="62" captionText="FIGURE 57. Left male palp, ventral [A Marma baeri Simon, 1902; B M. linae sp. nov.; C M. pipa sp. nov.; D M. wesolowskae sp. nov.; E M. nigritarsis (Simon, 1900); F M. sinuosa sp. nov., G M. argentina (Mello-Leit„o, 1941), H M. rosea (Mello- Leit„o, 1941), I M. spelunca sp. nov.]. Colors represent homologous structures." figureDoi="http://doi.org/10.5281/zenodo.4456979" httpUri="https://zenodo.org/record/4456979/files/figure.png" pageId="33" pageNumber="320">57</figureCitation>
CE), and the females differ by having the posterior border of the epigynal plate projected and the copulatory openings placed more posteriorly (posterior to the distal region of the primary spermathecae) (
<figureCitation id="13D29282FFBFFFA153EBFE770D51FE25" box="[247,331,403,429]" captionStart="FIGURE 31" captionStartId="34.[151,250,1720,1744]" captionTargetBox="[151,1435,489,1697]" captionTargetId="figure-168@34.[151,1435,489,1697]" captionTargetPageId="34" captionText="FIGURE 31. Female paratypes of Marma wesolowskae sp. nov. AB habitus, MPEG 34354 (A dorsal, B ventral); CD epi- gyne/vulva, MPEG 37131 (C ventral, D dorsal). Abbreviations: BG—Bennetts gland; cd—copulatory duct; co—copulatory opening; fd—fertilization duct; PS—Primary spermatheca; SS—secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456921" httpUri="https://zenodo.org/record/4456921/files/figure.png" pageId="33" pageNumber="320">Figs 31</figureCitation>
CD, 32G, 58EG).
</paragraph>
</subSubSection>
<subSubSection id="C3F3DD8CFFBFFFA253DBFE520D61F844" lastPageId="34" lastPageNumber="321" pageId="33" pageNumber="320" type="description">
<paragraph id="8B568E07FFBFFFA153DBFE520942FD91" blockId="33.[151,1437,150,1149]" pageId="33" pageNumber="320">
<emphasis id="B99D5215FFBFFFA153DBFE520EC7FE58" bold="true" box="[199,733,438,465]" pageId="33" pageNumber="320">
Description.
<emphasis id="B99D5215FFBFFFA1527DFE520D86FE59" bold="true" box="[353,412,438,465]" italics="true" pageId="33" pageNumber="320">Male</emphasis>
(
<typeStatus id="545230A5FFBFFFA152B0FE520E06FE58" box="[428,540,438,464]" pageId="33" pageNumber="320" type="holotype">Holotype</typeStatus>
, MPEG 34355).
</emphasis>
Total length: 3.38. Carapace 1.96 long, 1.31 wide, 0.97 high. Ocular quadrangle 0.78 long. Anterior eye row 1.18 wide, posterior 0.97 wide. Legs 3412. Length of legs: I 3.53 (1.13 + 1.32 + 1.08); II 3.17 (1.02 + 1.15 + 1.00); III 4.09 (1.39 + 1.41 + 1.29); IV 3.94 (1.25 + 1.30 + 1.39).
</paragraph>
<paragraph id="8B568E07FFBFFFA153DBFDC70DBEFD21" blockId="33.[151,1437,150,1149]" pageId="33" pageNumber="320">Leg macrosetae: Femur IIII d1-1-1, p1di, r1di; IV d1-1-1, p0, r1di. Patella III 0; IIIIV p0, r1. Tibia I p0 (or p0-1-0), r0, v1r-2-2; II p0-1-0, r0-1-0, v1r-1r-2; III p0-1-1-0, r1-1-1-0 (or r0-1-1-0), v1p-0-0-2; IV p0-1-1-0, r1-1- 1-0, v1p-0-0-2. Metatarsus I p1-1, r0-1, v1r-2 (or v2-2); II p1-1, r1di, v2-2; III d1p-0-0, p1-0-2, r1-0-2, v2-0-2; IV p1-1-2, r1-1-2, v1p-0-2.</paragraph>
<paragraph id="8B568E07FFBFFFA153DBFD570D6BFC09" blockId="33.[151,1437,150,1149]" pageId="33" pageNumber="320">
Color in alcohol (
<figureCitation id="13D29282FFBFFFA15291FD570DFEFD45" box="[397,484,691,717]" captionStart="FIGURE 30" captionStartId="33.[151,250,1960,1984]" captionTargetBox="[151,1435,1173,1937]" captionTargetId="figure-480@33.[151,1435,1173,1937]" captionTargetPageId="33" captionText="FIGURE 30. Male holotype of Marma wesolowskae sp. nov. (MPEG 34355). AB habitus (A dorsal, B ventral); CD left palp (C ventral; D retrolateral)." figureDoi="http://doi.org/10.5281/zenodo.4456919" httpUri="https://zenodo.org/record/4456919/files/figure.png" pageId="33" pageNumber="320">Figs 30</figureCitation>
AB): carapace covered entirely by scales (homogenous distribution); abdomen with thick dark brown stripe and pale lateral borders; legs: femur III with dark dorsal spot distally, III with distal and proximal regions with incomplete dark ring (distal ring: not colored ventrally; proximal ring: not colored dorsally), IV with dark distal ring; patella IIV with dark proximal spot; tibia III with proximal region with dark dorsal spot, IIIIV proximal and distal regions with dark dorsal spot; tarsus I with dark tip, II without dark marks, IIIIV with dark proximal ring.
</paragraph>
<paragraph id="8B568E07FFBFFFA153DBFC6F0CEDFB99" blockId="33.[151,1437,150,1149]" pageId="33" pageNumber="320">
Palp: RTA finger-shaped (
<figureCitation id="13D29282FFBFFFA152F0FC6F0E42FC2D" box="[492,600,907,933]" captionStart="FIGURE 30" captionStartId="33.[151,250,1960,1984]" captionTargetBox="[151,1435,1173,1937]" captionTargetId="figure-480@33.[151,1435,1173,1937]" captionTargetPageId="33" captionText="FIGURE 30. Male holotype of Marma wesolowskae sp. nov. (MPEG 34355). AB habitus (A dorsal, B ventral); CD left palp (C ventral; D retrolateral)." figureDoi="http://doi.org/10.5281/zenodo.4456919" httpUri="https://zenodo.org/record/4456919/files/figure.png" pageId="33" pageNumber="320">Figs 30D</figureCitation>
,
<figureCitation id="13D29282FFBFFFA15178FC6F0E8CFC2D" box="[612,662,907,933]" captionStart="FIGURE 32" captionStartId="35.[151,250,1365,1389]" captionTargetBox="[152,1405,191,1324]" captionTargetId="figure-17@35.[151,1435,181,1341]" captionTargetPageId="35" captionText="FIGURE 32. Marma wesolowskae sp. nov. AE left male palp (A ventral; B retrolateral; C proventral; D dorsal; E cleared bulb, ventral); FG epigyne/vulva (F ventral; G cleared, ventral).Abbreviations: BG—Bennetts gland; cc—cymbial conductor; cdcopulatory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PED—process on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456923" httpUri="https://zenodo.org/record/4456923/files/figure.png" pageId="33" pageNumber="320">32B</figureCitation>
,
<figureCitation id="13D29282FFBFFFA151BEFC6F0EC8FC2D" box="[674,722,907,933]" captionStart="FIGURE 33" captionStartId="36.[151,250,1784,1808]" captionTargetBox="[151,1435,181,1761]" captionTargetId="figure-24@36.[151,1435,181,1761]" captionTargetPageId="36" captionText="FIGURE 33. Marma wesolowskae sp. nov. AI left male palp [A ventral; B apical portion of palp, ventral; C tegular shoulder, ventral; D dissected embolic disc and embolus, dorsal; E patella and tibia, retrolateral; F tip of RTA, retrolateral; G detail of patella and tibia setae (blue rectangle in 33E); H detail of patella setae (red rectangle in 33E); I detail of smooth scale of tibia (yellow arrow in 33E]. J left spermatheca and copulatory duct, ventral. Abbreviations: cd—copulatory duct; e—embolus shaft; PED—process on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca. Black arrow in C shows depression of tegular shoulder; in D shows groove; white arrows in D show spikes of embolus shaft; in J shows end region of the primary spermatheca (broken fertilization duct)." figureDoi="http://doi.org/10.5281/zenodo.4456925" httpUri="https://zenodo.org/record/4456925/files/figure.png" pageId="33" pageNumber="320">33E</figureCitation>
); embolic disc narrow, with straight prolateral edge and curved retrolateral edge (
<figureCitation id="13D29282FFBFFFA1527DFC4B0DD6FC41" box="[353,460,943,969]" captionStart="FIGURE 30" captionStartId="33.[151,250,1960,1984]" captionTargetBox="[151,1435,1173,1937]" captionTargetId="figure-480@33.[151,1435,1173,1937]" captionTargetPageId="33" captionText="FIGURE 30. Male holotype of Marma wesolowskae sp. nov. (MPEG 34355). AB habitus (A dorsal, B ventral); CD left palp (C ventral; D retrolateral)." figureDoi="http://doi.org/10.5281/zenodo.4456919" httpUri="https://zenodo.org/record/4456919/files/figure.png" pageId="33" pageNumber="320">Figs 30C</figureCitation>
,
<figureCitation id="13D29282FFBFFFA152C6FC4B0E17FC41" box="[474,525,943,969]" captionStart="FIGURE 32" captionStartId="35.[151,250,1365,1389]" captionTargetBox="[152,1405,191,1324]" captionTargetId="figure-17@35.[151,1435,181,1341]" captionTargetPageId="35" captionText="FIGURE 32. Marma wesolowskae sp. nov. AE left male palp (A ventral; B retrolateral; C proventral; D dorsal; E cleared bulb, ventral); FG epigyne/vulva (F ventral; G cleared, ventral).Abbreviations: BG—Bennetts gland; cc—cymbial conductor; cdcopulatory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PED—process on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456923" httpUri="https://zenodo.org/record/4456923/files/figure.png" pageId="33" pageNumber="320">32A</figureCitation>
,
<figureCitation id="13D29282FFBFFFA15106FC4B0E55FC41" box="[538,591,943,969]" captionStart="FIGURE 33" captionStartId="36.[151,250,1784,1808]" captionTargetBox="[151,1435,181,1761]" captionTargetId="figure-24@36.[151,1435,181,1761]" captionTargetPageId="36" captionText="FIGURE 33. Marma wesolowskae sp. nov. AI left male palp [A ventral; B apical portion of palp, ventral; C tegular shoulder, ventral; D dissected embolic disc and embolus, dorsal; E patella and tibia, retrolateral; F tip of RTA, retrolateral; G detail of patella and tibia setae (blue rectangle in 33E); H detail of patella setae (red rectangle in 33E); I detail of smooth scale of tibia (yellow arrow in 33E]. J left spermatheca and copulatory duct, ventral. Abbreviations: cd—copulatory duct; e—embolus shaft; PED—process on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca. Black arrow in C shows depression of tegular shoulder; in D shows groove; white arrows in D show spikes of embolus shaft; in J shows end region of the primary spermatheca (broken fertilization duct)." figureDoi="http://doi.org/10.5281/zenodo.4456925" httpUri="https://zenodo.org/record/4456925/files/figure.png" pageId="33" pageNumber="320">33A</figureCitation>
); PED slightly longer than exposed portion of embolic disc and emerging from the prodistal part of embolic disc (
<figureCitation id="13D29282FFBFFFA15143FC370EF6FC65" box="[607,748,979,1005]" captionStart="FIGURE 32" captionStartId="35.[151,250,1365,1389]" captionTargetBox="[152,1405,191,1324]" captionTargetId="figure-17@35.[151,1435,181,1341]" captionTargetPageId="35" captionText="FIGURE 32. Marma wesolowskae sp. nov. AE left male palp (A ventral; B retrolateral; C proventral; D dorsal; E cleared bulb, ventral); FG epigyne/vulva (F ventral; G cleared, ventral).Abbreviations: BG—Bennetts gland; cc—cymbial conductor; cdcopulatory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PED—process on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456923" httpUri="https://zenodo.org/record/4456923/files/figure.png" pageId="33" pageNumber="320">Figs 32A, E</figureCitation>
,
<figureCitation id="13D29282FFBFFFA151E6FC370F35FC65" box="[762,815,979,1005]" captionStart="FIGURE 33" captionStartId="36.[151,250,1784,1808]" captionTargetBox="[151,1435,181,1761]" captionTargetId="figure-24@36.[151,1435,181,1761]" captionTargetPageId="36" captionText="FIGURE 33. Marma wesolowskae sp. nov. AI left male palp [A ventral; B apical portion of palp, ventral; C tegular shoulder, ventral; D dissected embolic disc and embolus, dorsal; E patella and tibia, retrolateral; F tip of RTA, retrolateral; G detail of patella and tibia setae (blue rectangle in 33E); H detail of patella setae (red rectangle in 33E); I detail of smooth scale of tibia (yellow arrow in 33E]. J left spermatheca and copulatory duct, ventral. Abbreviations: cd—copulatory duct; e—embolus shaft; PED—process on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca. Black arrow in C shows depression of tegular shoulder; in D shows groove; white arrows in D show spikes of embolus shaft; in J shows end region of the primary spermatheca (broken fertilization duct)." figureDoi="http://doi.org/10.5281/zenodo.4456925" httpUri="https://zenodo.org/record/4456925/files/figure.png" pageId="33" pageNumber="320">33A</figureCitation>
); tip of embolus ending beyond tip of PED (
<figureCitation id="13D29282FFBFFFA15630FC37098DFC65" box="[1324,1431,979,1005]" captionStart="FIGURE 30" captionStartId="33.[151,250,1960,1984]" captionTargetBox="[151,1435,1173,1937]" captionTargetId="figure-480@33.[151,1435,1173,1937]" captionTargetPageId="33" captionText="FIGURE 30. Male holotype of Marma wesolowskae sp. nov. (MPEG 34355). AB habitus (A dorsal, B ventral); CD left palp (C ventral; D retrolateral)." figureDoi="http://doi.org/10.5281/zenodo.4456919" httpUri="https://zenodo.org/record/4456919/files/figure.png" pageId="33" pageNumber="320">Figs 30C</figureCitation>
,
<figureCitation id="13D29282FFBFFFA1538BFC130CF0FB99" box="[151,234,1015,1041]" captionStart="FIGURE 32" captionStartId="35.[151,250,1365,1389]" captionTargetBox="[152,1405,191,1324]" captionTargetId="figure-17@35.[151,1435,181,1341]" captionTargetPageId="35" captionText="FIGURE 32. Marma wesolowskae sp. nov. AE left male palp (A ventral; B retrolateral; C proventral; D dorsal; E cleared bulb, ventral); FG epigyne/vulva (F ventral; G cleared, ventral).Abbreviations: BG—Bennetts gland; cc—cymbial conductor; cdcopulatory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PED—process on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456923" httpUri="https://zenodo.org/record/4456923/files/figure.png" pageId="33" pageNumber="320">32A, E</figureCitation>
).
</paragraph>
<paragraph id="8B568E07FFBFFFA153DBFBFE08ABFBF5" blockId="33.[151,1437,150,1149]" pageId="33" pageNumber="320">
<emphasis id="B99D5215FFBFFFA153DBFBFE0E43FBBC" bold="true" box="[199,601,1050,1077]" pageId="33" pageNumber="320">
<emphasis id="B99D5215FFBFFFA153DBFBFE0D04FBBD" bold="true" box="[199,286,1050,1077]" italics="true" pageId="33" pageNumber="320">Female</emphasis>
(
<typeStatus id="545230A5FFBFFFA15230FBFE0D86FBBC" box="[300,412,1050,1076]" pageId="33" pageNumber="320" type="paratype">Paratype</typeStatus>
, MPEG 34354).
</emphasis>
Total length: 4.01. Carapace 1.90 long, 1.43 wide, 1.11 high. Ocular quadrangle 0.97 long. Anterior eye row 1.27 wide, posterior 1.07 wide. Legs 3421. Length of legs: I 2.97 (0.93 + 1.17 + 0.87); II 3.00 (0.95 + 1.16 + 0.89); III 4.17 (1.37 + 1.45 + 1.35); IV 4.09 (1.27 + 1.37 + 1.45).
</paragraph>
<caption id="DF96DE8FFFBFFFA1538BF84C0DB8F86D" ID-DOI="http://doi.org/10.5281/zenodo.4456919" ID-Zenodo-Dep="4456919" httpUri="https://zenodo.org/record/4456919/files/figure.png" pageId="33" pageNumber="320" startId="33.[151,250,1960,1984]" targetBox="[151,1435,1173,1937]" targetPageId="33">
<paragraph id="8B568E07FFBFFFA1538BF84C0DB8F86D" blockId="33.[151,1437,1960,2021]" pageId="33" pageNumber="320">
<emphasis id="B99D5215FFBFFFA1538BF84C0D3AF848" bold="true" box="[151,288,1960,1984]" pageId="33" pageNumber="320">FIGURE 30.</emphasis>
Male holotype of
<taxonomicName id="4CE9F584FFBFFFA152C5F84D0EB4F849" authority="Salgado &amp; Ruiz, 2020" authorityName="Salgado &amp; Ruiz" authorityYear="2020" box="[473,686,1961,1985]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="320" phylum="Arthropoda" rank="species" species="wesolowskae" status="sp. nov.">
<emphasis id="B99D5215FFBFFFA152C5F84D0EB4F849" box="[473,686,1961,1985]" italics="true" pageId="33" pageNumber="320">Marma wesolowskae</emphasis>
</taxonomicName>
<emphasis id="B99D5215FFBFFFA151A8F84D0F1EF849" bold="true" box="[692,772,1961,1985]" pageId="33" pageNumber="320">
<taxonomicNameLabel id="A2AEEF6EFFBFFFA151A8F84D0F1EF849" box="[692,772,1961,1985]" pageId="33" pageNumber="320" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(MPEG 34355). AB habitus (A dorsal, B ventral); CD left palp (C ventral; D retrolateral).
</paragraph>
</caption>
<paragraph id="8B568E07FFBCFFA253DBFF730D14FEC9" blockId="34.[151,1437,151,465]" pageId="34" pageNumber="321">Leg macrosetae: Femur I d1-1-1, p1di, r0; II d0 (d1-1-1), p0 (p1di), r0 (r1di); III d1-1-1, p1di, r1di; IV d1-1-1, p0, r0 (r1di). Patella III 0; IIIIV p0, r1. Tibia I p0-1-0, r0, v1r-2-2; II p1di (p1-1-1), r0, v0 (v1r-2-0); III p0-1-0- 0 (or p0-1-1-0), r1-1-1-0 (or r0-1-1-0), v1p-0-0-1-2; IV p1-1-1-0 (or p0-1-1-0), r1-1-1-0, v0-1p-0-2. Metatarsus I p1-1, r1di, v2-2; II p1di (p1-1), r1di, v2di (v2-2); III d1p-0-0, p1-0-2, r1-0-2, v2-2; IV p1-1-2, r1-1-2, v0-1p-2 (or v1p-1p-2).</paragraph>
<paragraph id="8B568E07FFBCFFA253DBFEAF0F43FE59" blockId="34.[151,1437,151,465]" pageId="34" pageNumber="321">
Color in alcohol (
<figureCitation id="13D29282FFBCFFA25296FEAF0DC4FEED" box="[394,478,331,357]" captionStart="FIGURE 31" captionStartId="34.[151,250,1720,1744]" captionTargetBox="[151,1435,489,1697]" captionTargetId="figure-168@34.[151,1435,489,1697]" captionTargetPageId="34" captionText="FIGURE 31. Female paratypes of Marma wesolowskae sp. nov. AB habitus, MPEG 34354 (A dorsal, B ventral); CD epi- gyne/vulva, MPEG 37131 (C ventral, D dorsal). Abbreviations: BG—Bennetts gland; cd—copulatory duct; co—copulatory opening; fd—fertilization duct; PS—Primary spermatheca; SS—secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456921" httpUri="https://zenodo.org/record/4456921/files/figure.png" pageId="34" pageNumber="321">Figs 31</figureCitation>
AB): carapace as in male; abdomen ventrally pale; legs: femur III with distal region with dark spot, III with dark distal ring and proximal region with dark proventral spot, IV with dark distal ring; patella III with dark proximal spot; tibia III with dark proximal ring, IIIIV with proximal and distal dark ring; tarsus III without dark marks, IIIIV with dark proximal ring.
</paragraph>
<caption id="DF96DE8FFFBCFFA2538BF95C0806F890" ID-DOI="http://doi.org/10.5281/zenodo.4456921" ID-Zenodo-Dep="4456921" httpUri="https://zenodo.org/record/4456921/files/figure.png" pageId="34" pageNumber="321" startId="34.[151,250,1720,1744]" targetBox="[151,1435,489,1697]" targetPageId="34">
<paragraph id="8B568E07FFBCFFA2538BF95C0806F890" blockId="34.[151,1437,1720,1816]" pageId="34" pageNumber="321">
<emphasis id="B99D5215FFBCFFA2538BF95C0D3BF958" bold="true" box="[151,289,1720,1744]" pageId="34" pageNumber="321">FIGURE 31.</emphasis>
Female paratypes of
<taxonomicName id="4CE9F584FFBCFFA2511CF95D0ECCF958" authority="Salgado &amp; Ruiz, 2020" authorityName="Salgado &amp; Ruiz" authorityYear="2020" box="[512,726,1720,1744]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="321" phylum="Arthropoda" rank="species" species="wesolowskae" status="sp. nov.">
<emphasis id="B99D5215FFBCFFA2511CF95D0ECCF958" box="[512,726,1720,1744]" italics="true" pageId="34" pageNumber="321">Marma wesolowskae</emphasis>
</taxonomicName>
<emphasis id="B99D5215FFBCFFA251C2F95C0F2AF958" bold="true" box="[734,816,1720,1744]" pageId="34" pageNumber="321">
<taxonomicNameLabel id="A2AEEF6EFFBCFFA251C2F95C0F2AF958" box="[734,816,1720,1744]" pageId="34" pageNumber="321" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
AB habitus, MPEG 34354 (A dorsal, B ventral); CD epigyne/vulva, MPEG 37131 (C ventral, D dorsal). Abbreviations: BG—Bennetts gland; cd—copulatory duct; co—copulatory opening; fd—fertilization duct; PS—Primary spermatheca; SS—secondary spermatheca.
</paragraph>
</caption>
<paragraph id="8B568E07FFBCFFA253DBF8A20D61F844" blockId="34.[151,1436,1862,1996]" pageId="34" pageNumber="321">
Epigyne (
<figureCitation id="13D29282FFBCFFA2522FF8A20D9DF8E8" box="[307,391,1862,1888]" captionStart="FIGURE 31" captionStartId="34.[151,250,1720,1744]" captionTargetBox="[151,1435,489,1697]" captionTargetId="figure-168@34.[151,1435,489,1697]" captionTargetPageId="34" captionText="FIGURE 31. Female paratypes of Marma wesolowskae sp. nov. AB habitus, MPEG 34354 (A dorsal, B ventral); CD epi- gyne/vulva, MPEG 37131 (C ventral, D dorsal). Abbreviations: BG—Bennetts gland; cd—copulatory duct; co—copulatory opening; fd—fertilization duct; PS—Primary spermatheca; SS—secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456921" httpUri="https://zenodo.org/record/4456921/files/figure.png" pageId="34" pageNumber="321">Figs 31</figureCitation>
CD, 32FG, 33J): copulatory openings very close to each other and placed more posteriorly than end region of primary spermathecae; copulatory ducts long, encircling primary spermathecae; proximal copulatory duct shorter than distal section; primary spermathecae with homogenous diameter, narrow distal end and anteriorly projected.
</paragraph>
</subSubSection>
<caption id="DF96DE8FFFBDFFA3538BFAB10996FA51" ID-DOI="http://doi.org/10.5281/zenodo.4456923" ID-Zenodo-Dep="4456923" httpUri="https://zenodo.org/record/4456923/files/figure.png" pageId="35" pageNumber="322" startId="35.[151,250,1365,1389]" targetBox="[152,1405,191,1324]" targetPageId="35">
<paragraph id="8B568E07FFBDFFA3538BFAB10996FA51" blockId="35.[151,1436,1365,1497]" pageId="35" pageNumber="322">
<emphasis id="B99D5215FFBDFFA3538BFAB10D05FAE5" bold="true" box="[151,287,1365,1389]" pageId="35" pageNumber="322">FIGURE 32.</emphasis>
<taxonomicName id="4CE9F584FFBDFFA35238FAB20DE2FAE5" authority="Salgado &amp; Ruiz, 2020" authorityName="Salgado &amp; Ruiz" authorityYear="2020" box="[292,504,1365,1389]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="322" phylum="Arthropoda" rank="species" species="wesolowskae" status="sp. nov.">
<emphasis id="B99D5215FFBDFFA35238FAB20DE2FAE5" box="[292,504,1365,1389]" italics="true" pageId="35" pageNumber="322">Marma wesolowskae</emphasis>
</taxonomicName>
<emphasis id="B99D5215FFBDFFA352E2FAB10E57FAE5" bold="true" box="[510,589,1365,1389]" pageId="35" pageNumber="322">
<taxonomicNameLabel id="A2AEEF6EFFBDFFA352E2FAB10E57FAE5" box="[510,589,1365,1389]" pageId="35" pageNumber="322" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
AE left male palp (A ventral; B retrolateral; C proventral; D dorsal; E cleared bulb, ventral); FG epigyne/vulva (F ventral; G cleared, ventral). Abbreviations: BG—Bennetts gland; cc—cymbial conductor; cdcopulatory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PED—process on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca.
</paragraph>
</caption>
<subSubSection id="C3F3DD8CFFBDFFA353DBF9EC0E6FF896" pageId="35" pageNumber="322" type="materials_examined">
<paragraph id="8B568E07FFBDFFA353DBF9EC0F5FF9CE" blockId="35.[151,1437,1543,1894]" pageId="35" pageNumber="322">
<emphasis id="B99D5215FFBDFFA353DBF9EC0D6FF9A9" bold="true" box="[199,373,1543,1570]" pageId="35" pageNumber="322">Type material.</emphasis>
<materialsCitation id="3B81845AFFBDFFA35260F9E30F5FF9CE" ID-GBIF-Occurrence="3020125408" collectingDate="2019-07-21" collectionCode="MPEG" collectorName="A. Salgado" country="Brazil" latitude="-8.585777" location="Paraiso Selvagem" longLatPrecision="1" longitude="-37.241222" municipality="Buique" pageId="35" pageNumber="322" specimenCode="MPEG 34355" specimenCount="1" stateProvince="Pernambuco" typeStatus="holotype">
<typeStatus id="545230A5FFBDFFA35260F9E30DFEF9A9" box="[380,484,1543,1569]" pageId="35" pageNumber="322" type="holotype">Holotype</typeStatus>
♁:
<emphasis id="B99D5215FFBDFFA35113F9E30E9EF9A9" bold="true" box="[527,644,1543,1569]" pageId="35" pageNumber="322">
<collectingCountry id="F3FECE97FFBDFFA35113F9E30E65F9A9" box="[527,639,1543,1569]" name="Brazil" pageId="35" pageNumber="322">BRAZIL</collectingCountry>
:
</emphasis>
<emphasis id="B99D5215FFBDFFA35197F9E30F00F9A9" box="[651,794,1543,1569]" italics="true" pageId="35" pageNumber="322">
<collectingRegion id="492D40E5FFBDFFA35197F9E30F00F9A9" box="[651,794,1543,1569]" country="Brazil" name="Pernambuco" pageId="35" pageNumber="322">Pernambuco</collectingRegion>
</emphasis>
:
<collectingMunicipality id="6B32147DFFBDFFA35035F9EC0F67F9AA" box="[809,893,1544,1570]" pageId="35" pageNumber="322">Buíque</collectingMunicipality>
,
<location id="8E36D8DCFFBDFFA35094F9EC084AF9A9" LSID="urn:lsid:plazi:treatment:03403F11FFBEFFA3538BFF73082DF8EE:8E36D8DCFFBDFFA35094F9EC084AF9A9" box="[904,1104,1543,1570]" country="Brazil" latitude="-8.585777" longLatPrecision="1" longitude="-37.241222" municipality="Buique" name="Paraiso Selvagem" pageId="35" pageNumber="322" stateProvince="Pernambuco">Paraíso Selvagem</location>
camping area,
<geoCoordinate id="EEDDE8C0FFBDFFA357E0F9E30982F9AA" box="[1276,1432,1543,1570]" degrees="08" direction="south" minutes="35" orientation="latitude" pageId="35" pageNumber="322" precision="1" seconds="08.8" value="-8.585777">08°3508.8”S</geoCoordinate>
,
<geoCoordinate id="EEDDE8C0FFBDFFA3538BF9C80D26F9CE" box="[151,316,1580,1606]" degrees="37" direction="west" minutes="14" orientation="longitude" pageId="35" pageNumber="322" precision="1" seconds="28.4" value="-37.241222">37°1428.4”W</geoCoordinate>
, leg.
<collectorName id="261CEBD1FFBDFFA3526AF9C80DEFF9CD" box="[374,501,1579,1606]" pageId="35" pageNumber="322">A. Salgado</collectorName>
,
<date id="FF57A8C7FFBDFFA3511CF9C80E93F9CE" box="[512,649,1580,1606]" pageId="35" pageNumber="322" value="2019-07-21">
<collectingDate id="EF13512FFFBDFFA3511CF9C80E93F9CE" box="[512,649,1580,1606]" pageId="35" pageNumber="322" value="2019-07-21">21.VII.2019</collectingDate>
</date>
(
<specimenCode id="DB4F267CFFBDFFA3518BF9C80F23F9CE" box="[663,825,1579,1606]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="35" pageNumber="322">MPEG 34355</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8B568E07FFBDFFA353DBF9B40CE6F93A" blockId="35.[151,1437,1543,1894]" pageId="35" pageNumber="322">
<materialsCitation id="3B81845AFFBDFFA353DBF9B40901F9E2" ID-GBIF-Occurrence="3020125326" box="[199,1307,1615,1644]" collectingDate="2019-07-21" collectionCode="MPEG, IBSP" collectorName="A. Salgado" country="Brazil" latitude="-8.585777" location="Paraiso Selvagem" longLatPrecision="1" longitude="-37.241222" municipality="Buique" pageId="35" pageNumber="322" specimenCode="MPEG 37130, MPEG 34354, 37131, IBSP 267943" specimenCount="3" specimenCount-female="3" stateProvince="Pernambuco" typeStatus="paratype">
<typeStatus id="545230A5FFBDFFA353DBF9B40D22F9E2" box="[199,312,1616,1642]" pageId="35" pageNumber="322" type="paratype">Paratypes</typeStatus>
: Same data as holotype, 1♁ (
<specimenCode id="DB4F267CFFBDFFA35194F9B40F31F9E2" box="[648,811,1616,1642]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="35" pageNumber="322">MPEG 37130</specimenCode>
),
<specimenCount id="9DEF458EFFBDFFA3505CF9B40F79F9E2" box="[832,867,1616,1642]" pageId="35" pageNumber="322" type="female">3♀</specimenCount>
(
<specimenCode id="DB4F267CFFBDFFA3506FF9B4080FF9E2" box="[883,1045,1615,1642]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="35" pageNumber="322">MPEG 34354</specimenCode>
,
<specimenCode id="DB4F267CFFBDFFA3573EF9B40871F9E2" box="[1058,1131,1616,1642]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="35" pageNumber="322">37131</specimenCode>
,
<specimenCode id="DB4F267CFFBDFFA3576BF9B40909F9E2" box="[1143,1299,1615,1642]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="35" pageNumber="322">IBSP 267943</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B81845AFFBDFFA35634F9AB0CE2F93A" ID-GBIF-Occurrence="3020125312" collectingDate="2019-07-21" collectionCode="IBSP" collectorName="A. Salgado" country="Brazil" latitude="-8.570167" location="Pedra Furada" longLatPrecision="1" longitude="-36.82622" municipality="Venturosa" pageId="35" pageNumber="322" specimenCode="IBSP 267944" specimenCount="1" stateProvince="Pernambuco" typeStatus="paratype">
<emphasis id="B99D5215FFBDFFA35634F9AB0986F9E1" bold="true" box="[1320,1436,1615,1641]" pageId="35" pageNumber="322">
<collectingCountry id="F3FECE97FFBDFFA35634F9AB0982F9E1" box="[1320,1432,1615,1641]" name="Brazil" pageId="35" pageNumber="322">BRAZIL</collectingCountry>
:
</emphasis>
<emphasis id="B99D5215FFBDFFA3538BF9970D3CF905" box="[151,294,1651,1677]" italics="true" pageId="35" pageNumber="322">
<collectingRegion id="492D40E5FFBDFFA3538BF9970D3CF905" box="[151,294,1651,1677]" country="Brazil" name="Pernambuco" pageId="35" pageNumber="322">Pernambuco</collectingRegion>
</emphasis>
:
<collectingMunicipality id="6B32147DFFBDFFA35225F9900DB6F906" box="[313,428,1652,1678]" pageId="35" pageNumber="322">Venturosa</collectingMunicipality>
,
<location id="8E36D8DCFFBDFFA352A0F9970E42F905" LSID="urn:lsid:plazi:treatment:03403F11FFBEFFA3538BFF73082DF8EE:8E36D8DCFFBDFFA352A0F9970E42F905" box="[444,600,1651,1677]" country="Brazil" latitude="-8.570167" longLatPrecision="1" longitude="-36.82622" municipality="Venturosa" name="Pedra Furada" pageId="35" pageNumber="322" stateProvince="Pernambuco">Pedra Furada</location>
,
<geoCoordinate id="EEDDE8C0FFBDFFA3517BF9970F19F905" box="[615,771,1651,1677]" degrees="08" direction="south" minutes="34" orientation="latitude" pageId="35" pageNumber="322" precision="1" seconds="12.6" value="-8.570167">08°3412.6”S</geoCoordinate>
,
<geoCoordinate id="EEDDE8C0FFBDFFA35008F9970FA2F905" box="[788,952,1651,1677]" degrees="36" direction="west" minutes="49" orientation="longitude" pageId="35" pageNumber="322" precision="1" seconds="34.4" value="-36.82622">36°4934.4”W</geoCoordinate>
, leg.
<collectorName id="261CEBD1FFBDFFA350E7F9900865F905" box="[1019,1151,1651,1678]" pageId="35" pageNumber="322">A. Salgado</collectorName>
,
<date id="FF57A8C7FFBDFFA35793F9900900F906" box="[1167,1306,1652,1678]" pageId="35" pageNumber="322" value="2019-07-21">
<collectingDate id="EF13512FFFBDFFA35793F9900900F906" box="[1167,1306,1652,1678]" pageId="35" pageNumber="322" value="2019-07-21">21.VII.2019</collectingDate>
</date>
, 1♁ (
<specimenCode id="DB4F267CFFBDFFA3567CF9900CEAF93A" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="35" pageNumber="322">IBSP 267944</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B568E07FFBDFFA353DBF95F0E6FF896" blockId="35.[151,1437,1543,1894]" pageId="35" pageNumber="322">
<materialsCitation id="3B81845AFFBDFFA353DBF95F0F30F950" ID-GBIF-Occurrence="3020125446" box="[199,810,1723,1752]" collectingDate="2019-07-21" collectionCode="MPEG" collectorName="Same" country="Brazil" latitude="-8.585777" location="Paraiso Selvagem" longLatPrecision="1" longitude="-37.241222" municipality="Buique" pageId="35" pageNumber="322" specimenCount="1" stateProvince="Pernambuco">
<emphasis id="B99D5215FFBDFFA353DBF95F0DE5F95D" bold="true" box="[199,511,1723,1749]" pageId="35" pageNumber="322">Other material examined.</emphasis>
<collectorName id="261CEBD1FFBDFFA35114F9580E5CF95E" box="[520,582,1724,1750]" pageId="35" pageNumber="322">Same</collectorName>
data as holotype, 2
</materialsCitation>
<materialsCitation id="3B81845AFFBDFFA35058F9580838F95E" ID-GBIF-Occurrence="3020125420" box="[836,1058,1724,1750]" collectionCode="MPEG" country="Brazil" pageId="35" pageNumber="322" specimenCode="MPEG 37132" specimenCount="1" specimenCount-female="1" stateProvince="Pernambuco">
<specimenCount id="9DEF458EFFBDFFA35058F9580F7DF95E" box="[836,871,1724,1750]" pageId="35" pageNumber="322" type="female">1♀</specimenCount>
(
<specimenCode id="DB4F267CFFBDFFA3506BF9580803F95E" box="[887,1049,1724,1750]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="35" pageNumber="322">MPEG 37132</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="3B81845AFFBDFFA35733F9590916F95E" ID-GBIF-Occurrence="3020125308" box="[1071,1292,1724,1752]" collectionCode="MPEG" country="Brazil" pageId="35" pageNumber="322" specimenCode="MPEG 37133" specimenCount="1" stateProvince="Pernambuco">
1♁ (
<specimenCode id="DB4F267CFFBDFFA3577EF958091EF95E" box="[1122,1284,1724,1750]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="35" pageNumber="322">MPEG 37133</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="3B81845AFFBDFFA35605F9590CF3F972" ID-GBIF-Occurrence="3020125381" collectionCode="MPEG" country="Brazil" pageId="35" pageNumber="322" specimenCode="MPEG 37134" specimenCount="1" stateProvince="Pernambuco">
1♁ (
<specimenCode id="DB4F267CFFBDFFA35650F9580CFBF972" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="35" pageNumber="322">MPEG 37134</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="3B81845AFFBDFFA353E8F9040DD7F972" ID-GBIF-Occurrence="3020125344" box="[244,461,1759,1786]" collectionCode="MPEG" country="Brazil" pageId="35" pageNumber="322" specimenCode="MPEG 37135" specimenCount="1" specimenCount-female="1" stateProvince="Pernambuco">
<specimenCount id="9DEF458EFFBDFFA353E8F9040D0DF972" box="[244,279,1760,1786]" pageId="35" pageNumber="322" type="female">1♀</specimenCount>
(
<specimenCode id="DB4F267CFFBDFFA35238F9040DDFF972" box="[292,453,1759,1786]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="35" pageNumber="322">MPEG 37135</specimenCode>
)
</materialsCitation>
.
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:
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<collectingRegion id="492D40E5FFBDFFA3514DF93B0EFAF971" box="[593,736,1759,1785]" country="Brazil" name="Pernambuco" pageId="35" pageNumber="322">Pernambuco</collectingRegion>
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:
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<collectorName id="261CEBD1FFBDFFA3569DF9040CEEF895" pageId="35" pageNumber="322">A. Salgado</collectorName>
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<date id="FF57A8C7FFBDFFA353E3F8E00D91F896" box="[255,395,1796,1822]" pageId="35" pageNumber="322" value="2019-07-21">
<collectingDate id="EF13512FFFBDFFA353E3F8E00D91F896" box="[255,395,1796,1822]" pageId="35" pageNumber="322" value="2019-07-21">21.VII.2019</collectingDate>
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, 1♁ (
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</paragraph>
</subSubSection>
<subSubSection id="C3F3DD8CFFBDFFA353DBF8C3082DF8EE" pageId="35" pageNumber="322" type="distribution">
<paragraph id="8B568E07FFBDFFA353DBF8C3082FF8CA" blockId="35.[151,1437,1543,1894]" box="[199,1077,1831,1858]" pageId="35" pageNumber="322">
<emphasis id="B99D5215FFBDFFA353DBF8C30D7BF8C9" bold="true" box="[199,353,1831,1857]" pageId="35" pageNumber="322">Distribution.</emphasis>
Known only from the Brazilian state of
<collectingRegion id="492D40E5FFBDFFA3503BF8C30FA9F8C9" box="[807,947,1831,1857]" country="Brazil" name="Pernambuco" pageId="35" pageNumber="322">Pernambuco</collectingRegion>
(
<figureCitation id="13D29282FFBDFFA350DEF8C30832F8CA" box="[962,1064,1831,1858]" captionStart="FIGURE 59" captionStartId="64.[151,250,1837,1861]" captionTargetBox="[162,1435,181,1814]" captionTargetId="figure-24@64.[151,1435,181,1814]" captionTargetPageId="64" captionText="FIGURE 59. Maps of South America showing records of Marma species (A species previously described; B new species). Record from the state of Amazonas (Brazil) in Fig. 59A is taken from Zhang &amp; Maddison (2015)." figureDoi="http://doi.org/10.5281/zenodo.4456983" httpUri="https://zenodo.org/record/4456983/files/figure.png" pageId="35" pageNumber="322">Fig. 59B</figureCitation>
).
</paragraph>
<paragraph id="8B568E07FFBDFFA353DBF8A8082DF8EE" blockId="35.[151,1437,1543,1894]" box="[199,1079,1867,1894]" pageId="35" pageNumber="322">
<emphasis id="B99D5215FFBDFFA353DBF8A80D91F8ED" bold="true" box="[199,395,1867,1894]" pageId="35" pageNumber="322">Natural History.</emphasis>
The specimens were collected from sandstone (
<figureCitation id="13D29282FFBDFFA350B9F8AF0FE4F8EE" box="[933,1022,1867,1894]" captionStart="FIGURE 1" captionStartId="3.[151,250,1910,1934]" captionTargetBox="[157,1430,309,1887]" captionTargetId="figure-90@3.[157,1430,309,1887]" captionTargetPageId="3" captionText="FIGURE 1. Marma Simon, 1902. AE live specimens in different habitats [A M. nigritarsis from Belém/Pará/Brazil, on urban construction (Photo credit: César Favacho); B M. linae sp. nov. from São Geraldo do Araguaia/Pará/Brazil, on tree trunk; C M. rosea from São Geraldo do Araguaia/Pará/Brazil, on a wall of an urban construction; D M. nigritarsis from Belém/Pará/Brazil, on a substrate with moss; E M. wesolowskae sp. nov. from Buíque/Pernambuco/Brazil, on rocks]." figureDoi="http://doi.org/10.5281/zenodo.4456835" httpUri="https://zenodo.org/record/4456835/files/figure.png" pageId="35" pageNumber="322">Figs 1E</figureCitation>
,
<figureCitation id="13D29282FFBDFFA35716F8A80830F8EE" box="[1034,1066,1868,1894]" captionStart="FIGURE 29" captionStartId="32.[151,250,1980,2004]" captionTargetBox="[151,1435,340,1957]" captionTargetId="figure-60@32.[151,1435,340,1957]" captionTargetPageId="32" captionText="FIGURE 29. Marma wesolowskae sp. nov. live specimens from Buíque/Pernambuco/Brazil on sandrock. AC male (AB dorsolateral, C dorsal); DF female (DE dorsolateral, F dorsal)." figureDoi="http://doi.org/10.5281/zenodo.4456917" httpUri="https://zenodo.org/record/4456917/files/figure.png" pageId="35" pageNumber="322">29</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
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