155 lines
17 KiB
XML
155 lines
17 KiB
XML
<document id="D95E9E38A4ABC7A347D6AF91FFB19CC4" ID-CLB-Dataset="24546" ID-DOI="10.11646/zootaxa.4712.2.4" ID-GBIF-Dataset="cf61aa1d-1850-496a-92d4-2e5e5362ca24" ID-ISSN="1175-5326" ID-Zenodo-Dep="3587059" ID-ZooBank="6ECB5A18-2345-4259-BE51-7E77D7111BB7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1576824618479" checkinUser="plazi" docAuthor="Beaver, Roger A., Smith, Sarah M. & Sanguansub, Sunisa" docDate="2019" docId="03FA87EAFFFEFF925988010BFCF47774" docLanguage="en" docName="zootaxa.4712.2.4.pdf" docOrigin="Zootaxa 4712 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Procryphalus petioli Beaver & Smith & Sanguansub 2019, comb. n." docType="treatment" docVersion="9" lastPageNumber="248" masterDocId="FFC3FF92FFF5FF9E591F0627FFA3754B" masterDocTitle="A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae)" masterLastPageNumber="250" masterPageNumber="236" pageNumber="247" updateTime="1698775546065" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="6B4A7F36B7538F4CBE37D55AB5A1032A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="ADE2B1A51B07B4A1616AA16A28E3037D">A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae)</mods:title>
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<mods:namePart id="ED91ACF21298E3F8C26F3E91B657959D">Beaver, Roger A.</mods:namePart>
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<mods:roleTerm id="DCD1DEF83CCF3B7EC397BE2CD13572C0">Author</mods:roleTerm>
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<mods:namePart id="EDE1E398D60A19C8614675B9F849CC30">Smith, Sarah M.</mods:namePart>
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<mods:name id="546BA1458FD4A9C552A64EF2C5139610" type="personal">
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<mods:roleTerm id="681904EC455D74F408B068C34FC529D1">Author</mods:roleTerm>
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<mods:namePart id="A6A0FBEAF74F6665D92C39524C38A988">Sanguansub, Sunisa</mods:namePart>
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<mods:title id="30F913FCC582F2D538D8317E9F443C5D">Zootaxa</mods:title>
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<mods:part id="677EE2F123F1EE3B31318200088EDF2E">
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<mods:date id="5834CFCBA50F19519EE95CE349769FF4">2019</mods:date>
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<mods:number id="168BCBB4C285D63AE9321546547C294D">2019-12-19</mods:number>
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<mods:number id="474F30CD7241B0E201731C47E3EDD2D3">4712</mods:number>
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<mods:start id="BA241453EA9EFDCD92EB55D602BF9979">236</mods:start>
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<treatment id="03FA87EAFFFEFF925988010BFCF47774" ID-GBIF-Taxon="160798138" LSID="urn:lsid:plazi:treatment:03FA87EAFFFEFF925988010BFCF47774" httpUri="http://treatment.plazi.org/id/03FA87EAFFFEFF925988010BFCF47774" lastPageId="12" lastPageNumber="248" pageId="11" pageNumber="247">
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<subSubSection id="C3496577FFFEFF955988010BFDD1720C" box="[151,626,1836,1863]" pageId="11" pageNumber="247" type="nomenclature">
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<paragraph id="8BEC36FCFFFEFF955988010BFDD1720C" blockId="11.[151,626,1836,1863]" box="[151,626,1836,1863]" pageId="11" pageNumber="247">
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<heading id="D0A48190FFFEFF955988010BFDD1720C" bold="true" box="[151,626,1836,1863]" fontSize="11" level="1" pageId="11" pageNumber="247" reason="1">
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<emphasis id="B927EAEEFFFEFF955988010BFDD1720C" bold="true" box="[151,626,1836,1863]" pageId="11" pageNumber="247">
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<taxonomicName id="4C534D7FFFFEFF955988010BFE5C720D" authority="(Beaver)" authorityName="Beaver & Smith & Sanguansub" authorityYear="2019" baseAuthorityName="Beaver" baseAuthorityYear="1990" box="[151,511,1836,1863]" class="Insecta" family="Curculionidae" genus="Procryphalus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="247" phylum="Arthropoda" rank="species" species="petioli" status="comb. nov.">
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<emphasis id="B927EAEEFFFEFF955988010BFE2F720C" bold="true" box="[151,396,1836,1863]" italics="true" pageId="11" pageNumber="247">Procryphalus petioli</emphasis>
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(Beaver)
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</taxonomicName>
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<taxonomicNameLabel id="A2145795FFFEFF955B18010AFDD1720C" box="[519,626,1837,1863]" pageId="11" pageNumber="247" rank="species">comb. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3496577FFFEFF9259880156FCA374C0" lastPageId="12" lastPageNumber="248" pageId="11" pageNumber="247" type="reference_group">
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<paragraph id="8BEC36FCFFFEFF9559880156FD8B72C2" blockId="11.[151,552,1905,1929]" box="[151,552,1905,1929]" pageId="11" pageNumber="247">
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<treatmentCitationGroup id="AB4311D2FFFEFF9559880156FD8B72C2" box="[151,552,1905,1929]" pageId="11" pageNumber="247">
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<taxonomicName id="4C534D7FFFFEFF9559880156FD8772C2" ID-CoL="37VBW" authority="Beaver 1990: 281" authorityName="Beaver" authorityPageNumber="281" authorityYear="1990" box="[151,548,1905,1929]" class="Insecta" family="Curculionidae" genus="Dryocoetiops" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="247" phylum="Arthropoda" rank="species" species="petioli">
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<emphasis id="B927EAEEFFFEFF9559880156FEC472C2" box="[151,359,1905,1929]" italics="true" pageId="11" pageNumber="247">Dryocoetiops petioli</emphasis>
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<treatmentCitation id="0AF210EDFFFEFF9558720156FD8772C2" author="Beaver, R. A." box="[365,548,1905,1929]" page="281" pageId="11" pageNumber="247" year="1990">
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<bibRefCitation id="EFC24B0DFFFEFF9558720156FD8772C2" author="Beaver, R. A." box="[365,548,1905,1929]" pageId="11" pageNumber="247" pagination="279 - 284" refId="ref10784" refString="Beaver, R. A. (1990) New records and new species of bark and ambrosia beetles from Thailand (Coleoptera: Scolytidae and Platypodidae). Deutsche Entomologische Zeitschrift, Neue Folge, 37, 279 - 284. https: // doi. org / 10.1002 / mmnd. 19900370408" type="journal article" year="1990">Beaver 1990: 281</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BEC36FCFFFEFF9259880190FCA374C0" blockId="11.[151,1436,1975,2038]" lastBlockId="12.[151,1436,153,575]" lastPageId="12" lastPageNumber="248" pageId="11" pageNumber="247">
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<emphasis id="B927EAEEFFFEFF9559880190FEB9729A" bold="true" box="[151,282,1975,2001]" pageId="11" pageNumber="247">Taxonomy.</emphasis>
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We have examined the
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<typeStatus id="54E8885EFFFEFF955B39019FFDF57299" box="[550,598,1976,2002]" pageId="11" pageNumber="247">type</typeStatus>
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series (NHML, RAB) and further specimens collected by RAB at the
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<typeStatus id="54E8885EFFFEFF955C73019FFA3F7299" box="[1388,1436,1976,2002]" pageId="11" pageNumber="247">type</typeStatus>
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locality. The species was provisionally placed in the genus
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<taxonomicName id="4C534D7FFFFEFF955A2E01FBFC6B72BE" ID-CoL="8HHSH" authorityName="Schedl" authorityYear="1957" box="[817,968,2012,2037]" class="Insecta" family="Curculionidae" genus="Dryocoetiops" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="247" phylum="Arthropoda" rank="genus">
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<emphasis id="B927EAEEFFFEFF955A2E01FBFC6B72BE" box="[817,968,2012,2037]" italics="true" pageId="11" pageNumber="247">Dryocoetiops</emphasis>
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</taxonomicName>
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on the suggestion of S. L. Wood who ex- amined some specimens from the original collection (
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<bibRefCitation id="EFC24B0DFFF9FF925BE606BEFC2D75F8" author="Beaver, R. A." box="[761,910,153,179]" pageId="12" pageNumber="248" pagination="279 - 284" refId="ref10784" refString="Beaver, R. A. (1990) New records and new species of bark and ambrosia beetles from Thailand (Coleoptera: Scolytidae and Platypodidae). Deutsche Entomologische Zeitschrift, Neue Folge, 37, 279 - 284. https: // doi. org / 10.1002 / mmnd. 19900370408" type="journal article" year="1990">Beaver 1990</bibRefCitation>
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). However, dissections have shown that males are present, apparently indistinguishable from females using external morphology, and there is probably a 1: 1 sex ratio, unlike
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<taxonomicName id="4C534D7FFFF9FF92583A06C5FE1F75B0" ID-CoL="8HHSH" authorityName="Schedl" authorityYear="1957" box="[293,444,226,251]" class="Insecta" family="Curculionidae" genus="Dryocoetiops" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="248" phylum="Arthropoda" rank="genus">
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<emphasis id="B927EAEEFFF9FF92583A06C5FE1F75B0" box="[293,444,226,251]" italics="true" pageId="12" pageNumber="248">Dryocoetiops</emphasis>
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</taxonomicName>
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. B. Jordal (personal communication,
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<date id="FFED103CFFF9FF925A7E06C6FC5A75B0" box="[865,1017,224,251]" pageId="12" pageNumber="248" value="2015-01">January 2015</date>
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) has found that the male genitalia are of the cryphaline
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<typeStatus id="54E8885EFFF9FF9258400722FE307454" box="[351,403,261,287]" pageId="12" pageNumber="248">type</typeStatus>
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||
.
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||
<bibRefCitation id="EFC24B0DFFF9FF9258810722FD6D7454" author="Jordal, B. H. & Cognato, A. I." box="[414,718,261,287]" pageId="12" pageNumber="248" pagination="133" refId="ref11719" refString="Jordal, B. H. & Cognato, A. I. (2012) Molecular phylogeny of bark and ambrosia beetles reveals multiple origins of fungus farming during periods of global warming. BMC Evolutionary Biology, 12, 133. https: // doi. org / 10.1186 / 1471 - 2148 - 12 - 133" type="journal article" year="2012">Jordal and Cognato (2012)</bibRefCitation>
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, and
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||
<bibRefCitation id="EFC24B0DFFF9FF925A140722FC7D7454" author="Gohli, J. & Kirkendall, L. R. & Smith, S. M. & Cognato, A. I. & Hulcr, J. & Jordal, B. H." box="[779,990,261,287]" pageId="12" pageNumber="248" pagination="1258 - 1272" refId="ref11589" refString="Gohli, J., Kirkendall, L. R., Smith, S. M., Cognato, A. I., Hulcr, J. & Jordal, B. H. (2017) Biological factors contributing to bark and ambrosia beetle species diversification. Evolution, 71, 1258 - 1272. https: // doi. org / 10.1111 / evo. 13219" type="journal article" year="2017">
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Gohli
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<emphasis id="B927EAEEFFF9FF925A4C0721FC267454" box="[851,901,261,287]" italics="true" pageId="12" pageNumber="248">et al</emphasis>
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. (2017)
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</bibRefCitation>
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have shown, based on multiple genetic loci, that the species is closely related to two species of the cryphaline genus,
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<taxonomicName id="4C534D7FFFF9FF925AE7070EFA987408" ID-CoL="8HPSK" authority="Hopkins, 1915" authorityName="Hopkins" authorityYear="1915" box="[1016,1339,296,323]" class="Insecta" family="Curculionidae" genus="Procryphalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="248" phylum="Arthropoda" rank="genus">
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<emphasis id="B927EAEEFFF9FF925AE7070EFB2C7408" box="[1016,1167,297,323]" italics="true" pageId="12" pageNumber="248">Procryphalus</emphasis>
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<bibRefCitation id="EFC24B0DFFF9FF925D89070EFA987408" author="Hopkins, A. D." box="[1174,1339,296,323]" pageId="12" pageNumber="248" pagination="1 - 75" refId="ref11660" refString="Hopkins, A. D. (1915) Classification of the Cryphalinae, with descriptions of new genera and species. United States Department of Agriculture, Office of the Secretary, Report, 99, 1 - 75, pls. I-IV. https: // doi. org / 10.5962 / bhl. title. 65905" type="journal article" year="1915">Hopkins, 1915</bibRefCitation>
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</taxonomicName>
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(
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<taxonomicName id="4C534D7FFFF9FF925C54070DFEC37423" ID-CoL="4MPJT" authority="(Berger, 1916)" baseAuthorityName="Berger" baseAuthorityYear="1916" class="Insecta" family="Curculionidae" genus="Procryphalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="248" phylum="Arthropoda" rank="species" species="fraxini">
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<emphasis id="B927EAEEFFF9FF925C54070DFF16742C" italics="true" pageId="12" pageNumber="248">P. fraxini</emphasis>
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(Berger, 1916)
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</taxonomicName>
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,
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<taxonomicName id="4C534D7FFFF9FF9258740769FD6E742C" ID-CoL="4MPJV" authority="(Leconte, 1879))" authorityName="" baseAuthorityName="Leconte" baseAuthorityYear="1879" box="[363,717,333,359]" class="Insecta" family="Curculionidae" genus="Procryphalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="248" phylum="Arthropoda" rank="species" species="mucronatus">
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<emphasis id="B927EAEEFFF9FF9258740769FDAB742C" box="[363,520,334,359]" italics="true" pageId="12" pageNumber="248">P. mucronatus</emphasis>
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(Leconte, 1879))
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</taxonomicName>
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. The species is morphologically very distinct from these species, but it is provisionally transferred here to
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<taxonomicName id="4C534D7FFFF9FF925B7E0756FD5B74C0" ID-CoL="8HPSK" authorityName="Hopkins" authorityYear="1915" box="[609,760,369,395]" class="Insecta" family="Curculionidae" genus="Procryphalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="248" phylum="Arthropoda" rank="genus">
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<emphasis id="B927EAEEFFF9FF925B7E0756FD5B74C0" box="[609,760,369,395]" italics="true" pageId="12" pageNumber="248">Procryphalus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3496577FFF9FF9259D807B3FE7174E4" box="[199,466,404,431]" pageId="12" pageNumber="248" type="distribution">
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||
<paragraph id="8BEC36FCFFF9FF9259D807B3FE7174E4" blockId="12.[151,1436,153,575]" box="[199,466,404,431]" pageId="12" pageNumber="248">
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||
<emphasis id="B927EAEEFFF9FF9259D807B3FEC274E4" bold="true" box="[199,353,404,431]" pageId="12" pageNumber="248">Distribution.</emphasis>
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<collectingCountry id="F344766CFFF9FF92587807B2FE6D74E4" box="[359,462,405,431]" name="Thailand" pageId="12" pageNumber="248">Thailand</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3496577FFF9FF9259D8079FFCF47774" pageId="12" pageNumber="248" type="biology_ecology">
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||
<paragraph id="8BEC36FCFFF9FF9259D8079FFCF47774" blockId="12.[151,1436,153,575]" pageId="12" pageNumber="248">
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<emphasis id="B927EAEEFFF9FF9259D8079FFE847498" bold="true" box="[199,295,440,467]" pageId="12" pageNumber="248">Biology.</emphasis>
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Collected only from the petioles of fallen leaves of
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<taxonomicName id="4C534D7FFFF9FF925A6E079EFAEC749F" authority="Craib" authorityName="Craib" authorityYear="1922" box="[881,1359,441,468]" class="Magnoliopsida" family="Magnoliaceae" genus="Manglietia" higherTaxonomySource="CoL" kingdom="Plantae" order="Magnoliales" pageId="12" pageNumber="248" phylum="Tracheophyta" rank="species" species="garrettii">
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<emphasis id="B927EAEEFFF9FF925A6E079EFBEC7498" box="[881,1103,441,467]" italics="true" pageId="12" pageNumber="248">Manglietia garrettii</emphasis>
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Craib (Magnoliaceae)
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</taxonomicName>
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, incorrectly given as
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<taxonomicName id="4C534D7FFFF9FF92585F07F9FDD674BC" authority="in Beaver (1990)" authorityName="in Beaver" authorityYear="1990" box="[320,629,477,503]" class="Magnoliopsida" family="Magnoliaceae" genus="Manglietia" higherTaxonomySource="CoL" kingdom="Plantae" order="Magnoliales" pageId="12" pageNumber="248" phylum="Tracheophyta" rank="species" species="garelli">
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<emphasis id="B927EAEEFFF9FF92585F07F9FE0C74BC" box="[320,431,477,503]" italics="true" pageId="12" pageNumber="248">M. garelli</emphasis>
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in
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<bibRefCitation id="EFC24B0DFFF9FF9258CD07FAFDD674BC" author="Beaver, R. A." box="[466,629,477,503]" pageId="12" pageNumber="248" pagination="279 - 284" refId="ref10784" refString="Beaver, R. A. (1990) New records and new species of bark and ambrosia beetles from Thailand (Coleoptera: Scolytidae and Platypodidae). Deutsche Entomologische Zeitschrift, Neue Folge, 37, 279 - 284. https: // doi. org / 10.1002 / mmnd. 19900370408" type="journal article" year="1990">Beaver (1990)</bibRefCitation>
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</taxonomicName>
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. The beetles bored into the base of the petiole. The species appears to be monogamous. Immature stages were not found in the galleries, and it is possible that leafstalks are used primarily as a ‘refuge’ in the absence of other suitable breeding material.
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</paragraph>
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</subSubSection>
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||
</treatment>
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||
</document> |