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<document id="28BAC34D47E749862937425713DD57B8" ID-DOI="10.11646/zootaxa.5343.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="8324250" ID-ZooBank="20BF9AB4-F86D-42EA-AEAA-7C03EBA44EB6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1694069790509" checkinUser="plazi" docAuthor="Yamasaki, Takeshi, Hashimoto, Yoshiaki, Endo, Tomoji, Hyodo, Fujio, Itioka, Takao, Mohamed, Maryati &amp; Meleng, Paulus" docDate="2023" docId="03FA87E7FFBAFFD4FF7D3CC3FEBA8E13" docLanguage="en" docName="zootaxa.5343.1.3.pdf" docOrigin="Zootaxa 5343 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5343.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Utivarachna ichneumon Deeleman-Reinhold 2001" docType="treatment" docVersion="2" lastPageNumber="70" masterDocId="FFC3FF9FFFB7FFDBFFEA3E15FFEC8816" masterDocTitle="Taxonomic study of Bornean species of Utivarachna Kishida, 1940 (Araneae: Trachelidae), with the description of a new species" masterLastPageNumber="73" masterPageNumber="55" pageNumber="68" updateTime="1694071302809" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4778A8DD7A929CA8C5630DBD16BC1D80">Taxonomic study of Bornean species of Utivarachna Kishida, 1940 (Araneae: Trachelidae), with the description of a new species</mods:title>
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<emphasis id="B927EAE3FFBAFFD6FF7D3CC3FE538AE7" bold="true" box="[151,447,726,753]" italics="true" pageId="13" pageNumber="68">Utivarachna ichneumon</emphasis>
<bibRefCitation id="EFC24B00FFBAFFD6FE2C3CC3FCE48AE6" author="Deeleman-Reinhold, C. L." box="[454,776,726,752]" pageId="13" pageNumber="68" refId="ref8882" refString="Deeleman-Reinhold, C. L. (2001) Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]). Brill, Leiden, 591 pp. https: // doi. org / 10.1163 / 9789004475588" type="book" year="2001">Deeleman-Reinhold, 2001</bibRefCitation>
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<figureCitation id="13682A74FFBAFFD6FF7D3CEEFEF98B03" box="[151,277,763,789]" captionStart-0="FIGURE 20" captionStart-1="FIGURE 21" captionStart-2="FIGURE 22" captionStart-3="FIGURE 23" captionStartId-0="13.[151,250,1878,1902]" captionStartId-1="14.[151,250,686,710]" captionStartId-2="15.[151,250,836,860]" captionStartId-3="15.[151,250,1229,1253]" captionTargetBox-0="[151,1436,987,1854]" captionTargetBox-1="[151,1435,181,662]" captionTargetBox-2="[153,1419,181,806]" captionTargetBox-3="[151,1435,921,1205]" captionTargetId-0="figure-223@13.[151,1436,987,1854]" captionTargetId-1="figure-17@14.[151,1435,181,662]" captionTargetId-2="figure-17@15.[151,1436,181,812]" captionTargetId-3="figure-54@15.[151,1435,921,1205]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionTargetPageId-3="15" captionText-0="FIGURE 20. Utivarachna ichneumon Deeleman-Reinhold, 2001, male, holotype (RNMH.ARA.15223). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view; E label. Scale bars: AC, 1 mm; D, 0.50 mm." captionText-1="FIGURE 21. Utivarachna ichneumon, male, holotype (RNMH.ARA.15223). A left palp, prolateral view; B same, ventral view; C same, retrolateral view; D same, dorsal view. Scale bars: 0.50 mm. E—embolus, Sd—sperm duct, St—subtegulum." captionText-2="FIGURE 22. Utivarachna ichneumon Deeleman-Reinhold, 2001, female (DV53). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view. Scale bars AD: 1 mm." captionText-3="FIGURE 23. Utivarachna ichneumon, female (DV53). A epigyne, ventral view; B internal organs, ventral view; C same, dorsal view. Scale bars: 0.50 mm. Br—bursa, Fd—fertilization duct, S—spermatheca." figureDoi-0="http://doi.org/10.5281/zenodo.8324304" figureDoi-1="http://doi.org/10.5281/zenodo.8324308" figureDoi-2="http://doi.org/10.5281/zenodo.8324310" figureDoi-3="http://doi.org/10.5281/zenodo.8324314" httpUri-0="https://zenodo.org/record/8324304/files/figure.png" httpUri-1="https://zenodo.org/record/8324308/files/figure.png" httpUri-2="https://zenodo.org/record/8324310/files/figure.png" httpUri-3="https://zenodo.org/record/8324314/files/figure.png" pageId="13" pageNumber="68">Figs 2023</figureCitation>
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<emphasis id="B927EAE3FFBAFFD6FF7D3D2AFE618B41" box="[151,397,831,855]" italics="true" pageId="13" pageNumber="68">Utivarachna ichneumon</emphasis>
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, figs 581585.
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<emphasis id="B927EAE3FFBAFFD6FF7D3D90FE268B89" bold="true" box="[151,458,901,927]" pageId="13" pageNumber="68">Type material examined.</emphasis>
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<typeStatus id="54E88853FFBAFFD6FE3D3D90FDA88B89" box="[471,580,901,927]" pageId="13" pageNumber="68">Holotype</typeStatus>
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♁ (
<collectionCode id="ED42AE34FFBAFFD6FD903D90FD3F8B89" box="[634,723,901,927]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="13" pageNumber="68" type="Museum">RMNH</collectionCode>
.ARA. 15223;
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),
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F. C.
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, E.
<collectingRegion id="4997F813FFBAFFD6FAC23D90FA838B89" box="[1320,1391,901,927]" country="Malaysia" name="Sabah" pageId="13" pageNumber="68">Sabah</collectingRegion>
, 6 16.
<collectionCode id="ED42AE34FFBAFFD6FF523DBCFF238BD5" box="[184,207,937,963]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="13" pageNumber="68" type="Museum">V</collectionCode>
.[1991],
<collectorName id="26A65327FFBAFFD6FEC03DBCFE318BD5" box="[298,477,937,963]" pageId="13" pageNumber="68">Coll. Deeleman</collectorName>
, leg. C.L. &amp; P.
<collectionCode id="ED42AE34FFBAFFD6FD693DBFFD748BD2" box="[643,664,938,964]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="13" pageNumber="68">R</collectionCode>
. Deeleman.
</materialsCitation>
</paragraph>
<caption id="DF2C6679FFBAFFD6FF7D3943FEB38FA0" ID-DOI="http://doi.org/10.5281/zenodo.8324304" ID-Zenodo-Dep="8324304" httpUri="https://zenodo.org/record/8324304/files/figure.png" pageId="13" pageNumber="68" startId="13.[151,250,1878,1902]" targetBox="[151,1436,987,1854]" targetPageId="13" targetType="figure">
<paragraph id="8BEC36F1FFBAFFD6FF7D3943FEB38FA0" blockId="13.[151,1436,1878,1974]" pageId="13" pageNumber="68">
<emphasis id="B927EAE3FFBAFFD6FF7D3943FEC88F78" bold="true" box="[151,292,1878,1902]" pageId="13" pageNumber="68">FIGURE 20.</emphasis>
<taxonomicName id="4C534D72FFBAFFD6FEC43943FCA88F78" authority="Deeleman-Reinhold, 2001" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[302,836,1878,1902]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="68" phylum="Arthropoda" rank="species" species="ichneumon">
<emphasis id="B927EAE3FFBAFFD6FEC43943FDC48F78" box="[302,552,1878,1902]" italics="true" pageId="13" pageNumber="68">Utivarachna ichneumon</emphasis>
<bibRefCitation id="EFC24B00FFBAFFD6FDD93943FCA88F78" author="Deeleman-Reinhold, C. L." box="[563,836,1878,1902]" pageId="13" pageNumber="68" refId="ref8882" refString="Deeleman-Reinhold, C. L. (2001) Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]). Brill, Leiden, 591 pp. https: // doi. org / 10.1163 / 9789004475588" type="book" year="2001">Deeleman-Reinhold, 2001</bibRefCitation>
</taxonomicName>
, male, holotype (RNMH.ARA.15223).
<emphasis id="B927EAE3FFBAFFD6FB0D3943FB158F78" bold="true" box="[1255,1273,1878,1902]" pageId="13" pageNumber="68">A</emphasis>
habitus, dorsal view;
<emphasis id="B927EAE3FFBAFFD6FF3F396FFF0A8F84" bold="true" box="[213,230,1914,1938]" pageId="13" pageNumber="68">B</emphasis>
same, ventral view;
<emphasis id="B927EAE3FFBAFFD6FE5D396FFE258F84" bold="true" box="[439,457,1914,1938]" pageId="13" pageNumber="68">C</emphasis>
cephalic part and chelicerae, frontal view;
<emphasis id="B927EAE3FFBAFFD6FC91396FFC618F84" bold="true" box="[891,909,1914,1938]" pageId="13" pageNumber="68">D</emphasis>
leg I, left, prolateral view;
<emphasis id="B927EAE3FFBAFFD6FB75396FFB5C8F84" bold="true" box="[1183,1200,1914,1938]" pageId="13" pageNumber="68">E</emphasis>
label. Scale bars: AC, 1 mm; D, 0.50 mm.
</paragraph>
</caption>
<caption id="DF2C6679FFB9FFD5FF7D3CBBFAAE8AFC" ID-DOI="http://doi.org/10.5281/zenodo.8324308" ID-Zenodo-Dep="8324308" httpUri="https://zenodo.org/record/8324308/files/figure.png" pageId="14" pageNumber="69" startId="14.[151,250,686,710]" targetBox="[151,1435,181,662]" targetPageId="14" targetType="figure">
<paragraph id="8BEC36F1FFB9FFD5FF7D3CBBFAAE8AFC" blockId="14.[151,1436,686,746]" pageId="14" pageNumber="69">
<emphasis id="B927EAE3FFB9FFD5FF7D3CBBFEF38AD0" bold="true" box="[151,287,686,710]" pageId="14" pageNumber="69">FIGURE 21.</emphasis>
<taxonomicName id="4C534D72FFB9FFD5FECE3CBBFDF48AD0" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[292,536,686,710]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="69" phylum="Arthropoda" rank="species" species="ichneumon">
<emphasis id="B927EAE3FFB9FFD5FECE3CBBFDF48AD0" box="[292,536,686,710]" italics="true" pageId="14" pageNumber="69">Utivarachna ichneumon</emphasis>
</taxonomicName>
, male, holotype (RNMH.ARA.15223).
<emphasis id="B927EAE3FFB9FFD5FC433CBBFC578AD0" bold="true" box="[937,955,686,710]" pageId="14" pageNumber="69">A</emphasis>
left palp, prolateral view;
<emphasis id="B927EAE3FFB9FFD5FB283CBBFB3F8AD0" bold="true" box="[1218,1235,686,710]" pageId="14" pageNumber="69">B</emphasis>
same, ventral view;
<emphasis id="B927EAE3FFB9FFD5FF7D3CC7FF458AFC" bold="true" box="[151,169,722,746]" pageId="14" pageNumber="69">C</emphasis>
same, retrolateral view;
<emphasis id="B927EAE3FFB9FFD5FE4F3CC7FE5B8AFC" bold="true" box="[421,439,722,746]" pageId="14" pageNumber="69">D</emphasis>
same, dorsal view. Scale bars: 0.50 mm. E—embolus, Sd—sperm duct, St—subtegulum.
</paragraph>
</caption>
<paragraph id="8BEC36F1FFB9FFD5FF2D3D29FAF38B89" blockId="14.[151,1437,828,2043]" pageId="14" pageNumber="69">
<emphasis id="B927EAE3FFB9FFD5FF2D3D29FE698B40" bold="true" box="[199,389,828,854]" pageId="14" pageNumber="69">Other material.</emphasis>
<materialsCitation id="3B3B3CACFFB9FFD5FE7A3D28FD878B6D" collectingDate="2004-09-10" collectingDateMax="2005-02-22" collectingDateMin="2004-09-10" collectionCode="T" collectorName="Endo" location="Danum Valley" pageId="14" pageNumber="69" specimenCode="DV1, DV10" specimenCount="1" specimenCount-female="1" stateProvince="Sabah">
<specimenCount id="9D55FD78FFB9FFD5FE7A3D28FE528B41" box="[400,446,829,855]" pageId="14" pageNumber="69" type="female">1 ♀</specimenCount>
(
<specimenCode id="DBF59E8AFFB9FFD5FE3A3D29FDE88B41" box="[464,516,828,855]" collectionCode="DV" pageId="14" pageNumber="69">DV1</specimenCode>
_20040910-E5),
<location id="8E8C602AFFB9FFD5FD203D28FC9C8B40" LSID="urn:lsid:plazi:treatment:03FA87E7FFBAFFD4FF7D3CC3FEBA8E13:8E8C602AFFB9FFD5FD203D28FC9C8B40" box="[714,880,828,855]" name="Danum Valley" pageId="14" pageNumber="69" stateProvince="Sabah">Danum Valley</location>
,
<collectingRegion id="4997F813FFB9FFD5FC943D29FC298B40" box="[894,965,828,854]" country="Malaysia" name="Sabah" pageId="14" pageNumber="69">Sabah</collectingRegion>
,
<date id="FFED1031FFB9FFD5FC3E3D28FBBA8B41" box="[980,1110,829,855]" pageId="14" pageNumber="69" value="2004-09-10">
<collectingDate id="EFA9E9D9FFB9FFD5FC3E3D28FBBA8B41" box="[980,1110,829,855]" pageId="14" pageNumber="69" value="2004-09-10">10.IX.2004</collectingDate>
</date>
,
<collectionCode id="ED42AE34FFB9FFD5FB8F3D28FB9B8B41" box="[1125,1143,829,855]" name="Tavera, Department of Geology and Geophysics" pageId="14" pageNumber="69">T</collectionCode>
.
<collectorName id="26A65327FFB9FFD5FB6C3D29FB2D8B40" box="[1158,1217,828,854]" pageId="14" pageNumber="69">Endo</collectorName>
leg.; 1 ♁ (
<specimenCode id="DBF59E8AFFB9FFD5FAA23D28FA618B41" box="[1352,1421,829,855]" collectionCode="DV" pageId="14" pageNumber="69">DV10</specimenCode>
_ 20050222-I15), same locality,
<date id="FFED1031FFB9FFD5FE193D75FD878B6D" box="[499,619,864,891]" pageId="14" pageNumber="69" value="2005-02-22">
<collectingDate id="EFA9E9D9FFB9FFD5FE193D75FD878B6D" box="[499,619,864,891]" pageId="14" pageNumber="69" value="2005-02-22">22.II.2005</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B3B3CACFFB9FFD5FD923D74FA888B6C" box="[632,1380,864,891]" collectingDate="2005-02-20" collectingDateMax="2005-02-22" collectingDateMin="2004-09-10" collectionCode="R" collectorName="Oie" location="Danum Valley" pageId="14" pageNumber="69" specimenCode="DV7" specimenCount="1" specimenCount-female="1" stateProvince="Sabah">
<specimenCount id="9D55FD78FFB9FFD5FD923D74FD4F8B6D" box="[632,675,865,891]" pageId="14" pageNumber="69" type="female">1 ♀</specimenCount>
(
<specimenCode id="DBF59E8AFFB9FFD5FD593D75FD078B6D" box="[691,747,864,891]" collectionCode="DV" pageId="14" pageNumber="69">DV7</specimenCode>
_20050220-O16), same locality,
<date id="FFED1031FFB9FFD5FB8A3D75FB3A8B6D" box="[1120,1238,864,891]" pageId="14" pageNumber="69" value="2005-02-20">
<collectingDate id="EFA9E9D9FFB9FFD5FB8A3D75FB3A8B6D" box="[1120,1238,864,891]" pageId="14" pageNumber="69" value="2005-02-20">20.II.2005</collectingDate>
</date>
, leg.
<collectionCode id="ED42AE34FFB9FFD5FAFC3D74FAC08B6D" box="[1302,1324,865,891]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="14" pageNumber="69">R</collectionCode>
.
<collectorName id="26A65327FFB9FFD5FAD23D75FA888B6C" box="[1336,1380,864,890]" pageId="14" pageNumber="69">Oie</collectorName>
</materialsCitation>
;
<materialsCitation id="3B3B3CACFFB9FFD5FA9B3D74FAF68B89" collectingDate="2008-01-10" collectingDateMax="2005-02-22" collectingDateMin="2004-09-10" collectionCode="T, MNHAH" collectorName="Endo. All" location="Danum Valley" pageId="14" pageNumber="69" specimenCode="DV53" specimenCount="1" specimenCount-female="1" stateProvince="Sabah">
<specimenCount id="9D55FD78FFB9FFD5FA9B3D74FA708B6D" box="[1393,1436,865,891]" pageId="14" pageNumber="69" type="female">1 ♀</specimenCount>
(
<specimenCode id="DBF59E8AFFB9FFD5FF753D91FF088B89" box="[159,228,900,927]" collectionCode="DV" pageId="14" pageNumber="69">DV53</specimenCode>
_20080110-E7), same locality,
<date id="FFED1031FFB9FFD5FDA83D90FD5C8B89" box="[578,688,901,927]" pageId="14" pageNumber="69" value="2008-01-10">
<collectingDate id="EFA9E9D9FFB9FFD5FDA83D90FD5C8B89" box="[578,688,901,927]" pageId="14" pageNumber="69" value="2008-01-10">10.I.2008</collectingDate>
</date>
, leg.
<collectionCode id="ED42AE34FFB9FFD5FD013D90FD118B89" box="[747,765,901,927]" name="Tavera, Department of Geology and Geophysics" pageId="14" pageNumber="69">T</collectionCode>
.
<collectorName id="26A65327FFB9FFD5FCE23D91FC988B88" box="[776,884,900,926]" pageId="14" pageNumber="69">Endo. All</collectorName>
specimens are deposited at
<collectionCode id="ED42AE34FFB9FFD5FB473D90FAF68B89" box="[1197,1306,901,927]" pageId="14" pageNumber="69">MNHAH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C349657AFFB9FFD5FF2D3DBDFD508BD5" box="[199,700,936,963]" pageId="14" pageNumber="69" type="diagnosis">
<paragraph id="8BEC36F1FFB9FFD5FF2D3DBDFD508BD5" blockId="14.[151,1437,828,2043]" box="[199,700,936,963]" pageId="14" pageNumber="69">
<emphasis id="B927EAE3FFB9FFD5FF2D3DBDFEAE8BD4" bold="true" box="[199,322,936,962]" pageId="14" pageNumber="69">Diagnosis.</emphasis>
See Diagnosis of
<taxonomicName id="4C534D72FFB9FFD5FDE73DBCFD598BD4" authorityName="Kishida" authorityYear="1940" box="[525,693,936,962]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="69" phylum="Arthropoda" rank="species" species="fukasawana">
<emphasis id="B927EAE3FFB9FFD5FDE73DBCFD598BD4" box="[525,693,936,962]" italics="true" pageId="14" pageNumber="69">U. fukasawana</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C349657AFFB9FFD5FF2D3DD8FC678FEC" pageId="14" pageNumber="69" type="description">
<paragraph id="8BEC36F1FFB9FFD5FF2D3DD8FD3A8CF5" blockId="14.[151,1437,828,2043]" pageId="14" pageNumber="69">
<emphasis id="B927EAE3FFB9FFD5FF2D3DD8FE9A8BF1" bold="true" box="[199,374,973,999]" pageId="14" pageNumber="69">Measurements</emphasis>
(
<typeStatus id="54E88853FFB9FFD5FE6E3DD9FE048BF0" box="[388,488,972,998]" pageId="14" pageNumber="69">holotype</typeStatus>
♁/
<specimenCount id="9D55FD78FFB9FFD5FDE73DDBFDCC8BFF" box="[525,544,974,1001]" pageId="14" pageNumber="69" type="female"></specimenCount>
). Carapace length 3.95/3.16; cephalic width 1.50/1.37; thoracic width 1.93/1.80. Diameter of eyes: AME 0.18/0.17; ALE 0.16/0.15; PME 0.13/0.13; PLE 0.18/0.17. Interdistances between eyes: AMEAME 0.03/0.03; AMEALE 0.07/0.07; ALEALE 0.56/0.51; PMEPME 0.21/0.20; PMEPLE 0.33/0.27; PLEPLE 1.15/1.00; ALEPLE 0.18/0.16; AMEPME 0.06/0.06. MOA: length 0.35/0.32; anterior width 0.40/0.37; posterior width 0.50/0.46. Clypeus height 0.32/0.36. Abdomen length 3.25/2.86; width 1.36/1.83. Legs: I 12.44 (3.35, 1.13, 3.33, 3.50, 1.13)/9.75 (2.73, 0.93, 2.46, 2.53, 1.10); II 10.00 (2.83, 0.82, 2.53, 2.66, 1.16)/8.31 (2.36, 0.73, 2.06, 2.16, 1.00); III 6.35 (1.77, 0.70, 1.42, 1.75, 0.71)/5.51 (1.52, 0.61, 1.20, 1.53, 0.65); IV 8.51 (2.37, 0.68, 2.00, 2.66, 0.80)/7.54 (2.07, 0.61, 1.75, 2.30, 0.81).
</paragraph>
<paragraph id="8BEC36F1FFB9FFD5FF2D3AF8FD828DAD" blockId="14.[151,1437,828,2043]" pageId="14" pageNumber="69">
<emphasis id="B927EAE3FFB9FFD5FF2D3AF8FEE88D11" bold="true" box="[199,260,1261,1287]" pageId="14" pageNumber="69">Male</emphasis>
(
<figureCitation id="13682A74FFB9FFD5FEF33AF9FE468D11" box="[281,426,1260,1287]" captionStart="FIGURE 20" captionStartId="13.[151,250,1878,1902]" captionTargetBox="[151,1436,987,1854]" captionTargetId="figure-223@13.[151,1436,987,1854]" captionTargetPageId="13" captionText="FIGURE 20. Utivarachna ichneumon Deeleman-Reinhold, 2001, male, holotype (RNMH.ARA.15223). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view; E label. Scale bars: AC, 1 mm; D, 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.8324304" httpUri="https://zenodo.org/record/8324304/files/figure.png" pageId="14" pageNumber="69">Fig. 20AD</figureCitation>
). Carapace entirely granulated, dorsally oval; posterior stalk slightly shorter than 1/2 carapace length; PER recurved in dorsal view; ocular tubercle accompanied with PLE; thoracic lateral margins undulating. Chelicerae granulated, with five prolateral and five retrolateral teeth. Ventral clypeus margin with small wedge-shaped projection between chelicerae. Pedicel short. Abdomen elongate oval; anterior two-thirds covered with dorsal scutum, weakly developed. Leg I and II elongated, covered with spherical granulation; ventral sides of tibia, metatarsus and tarsus bearing cusps.
</paragraph>
<paragraph id="8BEC36F1FFB9FFD5FF2D3BD1FDDD8E84" blockId="14.[151,1437,828,2043]" pageId="14" pageNumber="69">
Palp (
<figureCitation id="13682A74FFB9FFD5FEEC3BD1FE7C8DC9" box="[262,400,1476,1503]" captionStart="FIGURE 21" captionStartId="14.[151,250,686,710]" captionTargetBox="[151,1435,181,662]" captionTargetId="figure-17@14.[151,1435,181,662]" captionTargetPageId="14" captionText="FIGURE 21. Utivarachna ichneumon, male, holotype (RNMH.ARA.15223). A left palp, prolateral view; B same, ventral view; C same, retrolateral view; D same, dorsal view. Scale bars: 0.50 mm. E—embolus, Sd—sperm duct, St—subtegulum." figureDoi="http://doi.org/10.5281/zenodo.8324308" httpUri="https://zenodo.org/record/8324308/files/figure.png" pageId="14" pageNumber="69">Fig. 21AD</figureCitation>
). Cymbium slender oval; length almost twice as wide. Bulb oval; posterior half of bulb weakly sclerotized with subtegulum, and anterior half membranous. Embolus coiled; basal part of embolus accompanied by slender sclerotized plate covered by membranous surface of bulb. Sperm duct running from retrolateral side of bulb, U-curved along posterior edge of bulb, then curving to prolateral side. RTA strongly extended, apically developed into semi-transparent plate with apical claw; prolateral corner of semi-transparent plate developed, and anterior margin transforming into short claw.
</paragraph>
<paragraph id="8BEC36F1FFB9FFD5FF2D3889FF0F8EE8" blockId="14.[151,1437,828,2043]" pageId="14" pageNumber="69">
Coloration and setation (
<figureCitation id="13682A74FFB9FFD5FE033889FD958EA1" box="[489,633,1692,1719]" captionStart="FIGURE 20" captionStartId="13.[151,250,1878,1902]" captionTargetBox="[151,1436,987,1854]" captionTargetId="figure-223@13.[151,1436,987,1854]" captionTargetPageId="13" captionText="FIGURE 20. Utivarachna ichneumon Deeleman-Reinhold, 2001, male, holotype (RNMH.ARA.15223). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view; E label. Scale bars: AC, 1 mm; D, 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.8324304" httpUri="https://zenodo.org/record/8324304/files/figure.png" pageId="14" pageNumber="69">Fig. 20AD</figureCitation>
). Carapace and chelicerae brown, covered with fine setae. Abdomen cream, with pale brown dorsal scutum, entirely covered with small fine setae. All legs pale brown; leg I darker than others.
</paragraph>
<paragraph id="8BEC36F1FFB9FFD5FF2D391CFC9E8F7D" blockId="14.[151,1437,828,2043]" pageId="14" pageNumber="69">
<emphasis id="B927EAE3FFB9FFD5FF2D391CFEF28F35" bold="true" box="[199,286,1801,1827]" pageId="14" pageNumber="69">Female</emphasis>
(
<figureCitation id="13682A74FFB9FFD5FEDB391DFE2C8F35" box="[305,448,1800,1827]" captionStart="FIGURE 22" captionStartId="15.[151,250,836,860]" captionTargetBox="[153,1419,181,806]" captionTargetId="figure-17@15.[151,1436,181,812]" captionTargetPageId="15" captionText="FIGURE 22. Utivarachna ichneumon Deeleman-Reinhold, 2001, female (DV53). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view. Scale bars AD: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8324310" httpUri="https://zenodo.org/record/8324310/files/figure.png" pageId="14" pageNumber="69">Fig. 22AD</figureCitation>
). Somatic characters almost same as in males, except for abdomen shape and shorter posterior stalk; posterior stalk almost as long as 1/4 carapace length. Chelicerae granulated, with five or six prolateral and five retrolateral teeth. Abdomen oval, without dorsal scutum.
</paragraph>
<paragraph id="8BEC36F1FFB9FFD5FF2D3961FC0B8FC0" blockId="14.[151,1437,828,2043]" pageId="14" pageNumber="69">
Copulatory organs (
<figureCitation id="13682A74FFB9FFD5FE453961FDD18F99" box="[431,573,1908,1935]" captionStart="FIGURE 23" captionStartId="15.[151,250,1229,1253]" captionTargetBox="[151,1435,921,1205]" captionTargetId="figure-54@15.[151,1435,921,1205]" captionTargetPageId="15" captionText="FIGURE 23. Utivarachna ichneumon, female (DV53). A epigyne, ventral view; B internal organs, ventral view; C same, dorsal view. Scale bars: 0.50 mm. Br—bursa, Fd—fertilization duct, S—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.8324314" httpUri="https://zenodo.org/record/8324314/files/figure.png" pageId="14" pageNumber="69">Fig. 23AC</figureCitation>
). Copulatory atrium longer than wide. Copulatory ducts curved into loop anteriorly, then intricately coiled. Bursae long, tube-like shaped, extending posteriorly. Spermathecae globular. Fertilization ducts straight, extending from spermathecae towards posterior.
</paragraph>
<paragraph id="8BEC36F1FFB9FFD5FF2D39F5FC678FEC" blockId="14.[151,1437,828,2043]" box="[199,907,2016,2043]" pageId="14" pageNumber="69">
Coloration and setation (
<figureCitation id="13682A74FFB9FFD5FE3639F5FD848FED" box="[476,616,2016,2043]" captionStart="FIGURE 22" captionStartId="15.[151,250,836,860]" captionTargetBox="[153,1419,181,806]" captionTargetId="figure-17@15.[151,1436,181,812]" captionTargetPageId="15" captionText="FIGURE 22. Utivarachna ichneumon Deeleman-Reinhold, 2001, female (DV53). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view. Scale bars AD: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8324310" httpUri="https://zenodo.org/record/8324310/files/figure.png" pageId="14" pageNumber="69">Fig. 22AD</figureCitation>
). Almost same as in male.
</paragraph>
</subSubSection>
<caption id="DF2C6679FFB8FFD4FF7D3D51FB9D8B96" ID-DOI="http://doi.org/10.5281/zenodo.8324310" ID-Zenodo-Dep="8324310" httpUri="https://zenodo.org/record/8324310/files/figure.png" pageId="15" pageNumber="70" startId="15.[151,250,836,860]" targetBox="[153,1419,181,806]" targetPageId="15" targetType="figure">
<paragraph id="8BEC36F1FFB8FFD4FF7D3D51FB9D8B96" blockId="15.[151,1436,836,897]" pageId="15" pageNumber="70">
<emphasis id="B927EAE3FFB8FFD4FF7D3D51FEF38B4A" bold="true" box="[151,287,836,860]" pageId="15" pageNumber="70">FIGURE 22.</emphasis>
<taxonomicName id="4C534D72FFB8FFD4FECE3D50FCC58B4B" authority="Deeleman-Reinhold, 2001" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[292,809,837,861]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="70" phylum="Arthropoda" rank="species" species="ichneumon">
<emphasis id="B927EAE3FFB8FFD4FECE3D50FDF48B4B" box="[292,536,837,861]" italics="true" pageId="15" pageNumber="70">Utivarachna ichneumon</emphasis>
<bibRefCitation id="EFC24B00FFB8FFD4FDF73D50FCC58B4B" author="Deeleman-Reinhold, C. L." box="[541,809,837,861]" pageId="15" pageNumber="70" refId="ref8882" refString="Deeleman-Reinhold, C. L. (2001) Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]). Brill, Leiden, 591 pp. https: // doi. org / 10.1163 / 9789004475588" type="book" year="2001">Deeleman-Reinhold, 2001</bibRefCitation>
</taxonomicName>
, female (DV53).
<emphasis id="B927EAE3FFB8FFD4FC3D3D51FC058B4A" bold="true" box="[983,1001,836,860]" pageId="15" pageNumber="70">A</emphasis>
habitus, dorsal view;
<emphasis id="B927EAE3FFB8FFD4FB293D50FB388B4B" bold="true" box="[1219,1236,837,861]" pageId="15" pageNumber="70">B</emphasis>
same, ventral view;
<emphasis id="B927EAE3FFB8FFD4FF7D3D7DFF458B96" bold="true" box="[151,169,872,896]" pageId="15" pageNumber="70">C</emphasis>
cephalic part and chelicerae, frontal view;
<emphasis id="B927EAE3FFB8FFD4FDB53D7CFD9D8B97" bold="true" box="[607,625,873,897]" pageId="15" pageNumber="70">D</emphasis>
leg I, left, prolateral view. Scale bars AD: 1 mm.
</paragraph>
</caption>
<caption id="DF2C6679FFB8FFD4FF7D3AD8FC508D1F" ID-DOI="http://doi.org/10.5281/zenodo.8324314" ID-Zenodo-Dep="8324314" httpUri="https://zenodo.org/record/8324314/files/figure.png" pageId="15" pageNumber="70" startId="15.[151,250,1229,1253]" targetBox="[151,1435,921,1205]" targetPageId="15" targetType="figure">
<paragraph id="8BEC36F1FFB8FFD4FF7D3AD8FC508D1F" blockId="15.[151,1436,1229,1289]" pageId="15" pageNumber="70">
<emphasis id="B927EAE3FFB8FFD4FF7D3AD8FEF38CF3" bold="true" box="[151,287,1229,1253]" pageId="15" pageNumber="70">FIGURE 23.</emphasis>
<taxonomicName id="4C534D72FFB8FFD4FECE3AD8FDF48CF3" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[292,536,1229,1253]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="70" phylum="Arthropoda" rank="species" species="ichneumon">
<emphasis id="B927EAE3FFB8FFD4FECE3AD8FDF48CF3" box="[292,536,1229,1253]" italics="true" pageId="15" pageNumber="70">Utivarachna ichneumon</emphasis>
</taxonomicName>
, female (DV53).
<emphasis id="B927EAE3FFB8FFD4FD233AD8FD378CF3" bold="true" box="[713,731,1229,1253]" pageId="15" pageNumber="70">A</emphasis>
epigyne, ventral view;
<emphasis id="B927EAE3FFB8FFD4FC2C3AD8FC3B8CF3" bold="true" box="[966,983,1229,1253]" pageId="15" pageNumber="70">B</emphasis>
internal organs, ventral view;
<emphasis id="B927EAE3FFB8FFD4FAED3AD8FAF58CF3" bold="true" box="[1287,1305,1229,1253]" pageId="15" pageNumber="70">C</emphasis>
same, dorsal view. Scale bars: 0.50 mm. Br—bursa, Fd—fertilization duct, S—spermatheca.
</paragraph>
</caption>
<subSubSection id="C349657AFFB8FFD4FF2D3B4EFCA78D63" box="[199,843,1371,1398]" pageId="15" pageNumber="70" type="distribution">
<paragraph id="8BEC36F1FFB8FFD4FF2D3B4EFCA78D63" blockId="15.[151,1436,1371,1541]" box="[199,843,1371,1398]" pageId="15" pageNumber="70">
<emphasis id="B927EAE3FFB8FFD4FF2D3B4EFE8D8D63" bold="true" box="[199,353,1371,1397]" pageId="15" pageNumber="70">Distribution.</emphasis>
<collectingRegion id="4997F813FFB8FFD4FE8D3B4EFE478D63" box="[359,427,1371,1397]" country="Malaysia" name="Sabah" pageId="15" pageNumber="70">Sabah</collectingRegion>
(known only from the
<typeStatus id="54E88853FFB8FFD4FD453B49FD338D60" box="[687,735,1372,1398]" pageId="15" pageNumber="70">type</typeStatus>
locality).
</paragraph>
</subSubSection>
<subSubSection id="C349657AFFB8FFD4FF2D3B6AFEBA8E13" pageId="15" pageNumber="70" type="discussion">
<paragraph id="8BEC36F1FFB8FFD4FF2D3B6AFEBA8E13" blockId="15.[151,1436,1371,1541]" pageId="15" pageNumber="70">
<emphasis id="B927EAE3FFB8FFD4FF2D3B6AFED78D8F" bold="true" box="[199,315,1407,1433]" pageId="15" pageNumber="70">Remarks.</emphasis>
The non-type material examined in the present study was collected from the
<typeStatus id="54E88853FFB8FFD4FB453B95FB338D8C" box="[1199,1247,1408,1434]" pageId="15" pageNumber="70">type</typeStatus>
locality, Danum Valley,
<collectingRegion id="4997F813FFB8FFD4FF003BB6FE878DA8" box="[234,363,1443,1470]" country="Malaysia" name="Sabah" pageId="15" pageNumber="70">Sabah state</collectingRegion>
. The conspecificity of male and female specimens was based on the long stalk of carapace. Other
<taxonomicName id="4C534D72FFB8FFD4FF7D3BD2FE918DF7" box="[151,381,1479,1505]" pageId="15" pageNumber="70">
<emphasis id="B927EAE3FFB8FFD4FF7D3BD2FEC98DF7" box="[151,293,1479,1505]" italics="true" pageId="15" pageNumber="70">Utivarachna</emphasis>
species
</taxonomicName>
with long posterior stalks were not collected from the study site. The female is described here for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>