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<document id="DE55E31FAF3F7720BD201B8803775930" ID-DOI="10.11646/zootaxa.4048.1.3" ID-GBIF-Dataset="0530d3f6-2d76-46b5-a389-084d7ba41850" ID-ISSN="1175-5326" ID-Zenodo-Dep="290090" ID-ZooBank="A5C40F83-2369-41E7-BB51-A848E77F5A74" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461342079078" checkinUser="plazi" docAuthor="Rovito, Sean M. &amp; Parra-Olea, Gabriela" docDate="2015" docId="03FA87CFFFCE0057E68EFBABBBC8F849" docLanguage="en" docName="zt04048p074.pdf" docOrigin="Zootaxa 4048 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chiropterotriton infernalis Rovito &amp; Parra-Olea, 2015, sp. nov." docType="treatment" docVersion="5" lastPageNumber="68" masterDocId="FFC3FFB7FFC6005CE619FFC8BA2CFF86" masterDocTitle="Two new species of Chiropterotriton (Caudata: Plethodontidae) from northern Mexico" masterLastPageNumber="74" masterPageNumber="57" pageNumber="65" updateTime="1698620376902" updateUser="plazi">
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<mods:title id="6C773A7820267966C5F12351203A7314">Two new species of Chiropterotriton (Caudata: Plethodontidae) from northern Mexico</mods:title>
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<mods:namePart id="D20E07D9E83EC204EFEAD87775B94C2C">Rovito, Sean M.</mods:namePart>
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<mods:namePart id="BA25076307124B7D5ECCBB1AE6FB52A2">Parra-Olea, Gabriela</mods:namePart>
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<mods:title id="E33C255E419D5B5775A53B7743EEA0CC">Zootaxa</mods:title>
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<mods:date id="AF638DB17FC035B47449930E9A1E9B9E">2015</mods:date>
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<treatment id="03FA87CFFFCE0057E68EFBABBBC8F849" ID-DOI="http://doi.org/10.5281/zenodo.5614620" ID-GBIF-Taxon="127694327" ID-Zenodo-Dep="5614620" LSID="urn:lsid:plazi:treatment:03FA87CFFFCE0057E68EFBABBBC8F849" httpUri="http://treatment.plazi.org/id/03FA87CFFFCE0057E68EFBABBBC8F849" lastPageId="11" lastPageNumber="68" pageId="8" pageNumber="65">
<subSubSection id="C3496552FFCE0054E68EFBABBB4BFB45" pageId="8" pageNumber="65" type="nomenclature">
<paragraph id="8BEC36D9FFCE0054E68EFBABB813FBF8" blockId="8.[151,949,1123,1219]" box="[151,575,1123,1150]" pageId="8" pageNumber="65">
<heading id="D0A481B5FFCE0054E68EFBABB813FBF8" bold="true" box="[151,575,1123,1150]" fontSize="11" level="1" pageId="8" pageNumber="65" reason="1">
<emphasis id="B927EACBFFCE0054E68EFBABB813FBF8" bold="true" box="[151,575,1123,1150]" pageId="8" pageNumber="65">
<taxonomicName id="4C534D5AFFCE0054E68EFBABBBF6FBFB" ID-CoL="5Y6LR" box="[151,474,1123,1149]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="infernalis" status="sp. nov.">
<emphasis id="B927EACBFFCE0054E68EFBABBBF6FBFB" bold="true" box="[151,474,1123,1149]" italics="true" pageId="8" pageNumber="65">Chiropterotriton infernalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21457B0FFCE0054E7F8FBACB813FBF8" box="[481,575,1124,1150]" pageId="8" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEC36D9FFCE0054E68EFB4EBB4BFB45" blockId="8.[151,949,1123,1219]" pageId="8" pageNumber="65">
Sistema Purificación Salamander, Salamandra del Sistema Purificación (
<figureCitation id="13682A5CFFCE0054E686FB62BAC7FB45" box="[159,235,1194,1219]" captionStart="FIGURE 3" captionStartId="10.[151,250,1928,1950]" captionTargetBox="[207,1375,193,1896]" captionTargetId="figure@10.[207,1376,193,1897]" captionTargetPageId="10" captionText="FIGURE 3. Photos in life of Chiropterotriton from Tamaulipas, Nuevo León, and San Luis Potosí. A) holotype of Chiropterotriton cieloensis (IBH 29561), B) C. cracens (IBH 28178), C) C. multidentatus (IBH 28177), D) holotype of C. infernalis (MVZ 269665)., E) C. miquihuanus (MVZ 269632)." httpUri="https://zenodo.org/record/290093/files/figure.png" pageId="8" pageNumber="65">Figs. 3</figureCitation>
D, 4B, 5B)
</paragraph>
</subSubSection>
<subSubSection id="C3496552FFCE0054E68EFB3BBE3EFAA1" pageId="8" pageNumber="65" type="reference_group">
<paragraph id="8BEC36D9FFCE0054E68EFB3BB90BFA8E" blockId="8.[151,1042,1266,1319]" box="[151,807,1266,1288]" pageId="8" pageNumber="65">
<taxonomicName id="4C534D5AFFCE0054E68EFB3BBBE2FA8E" ID-CoL="5Y6KY" box="[151,462,1267,1288]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="multidentatus">
<emphasis id="B927EACBFFCE0054E68EFB3BBBE2FA8E" box="[151,462,1267,1288]" italics="true" pageId="8" pageNumber="65">Chiropterotriton multidentatus</emphasis>
</taxonomicName>
(part):
<bibRefCitation id="EFC24B28FFCE0054E403FB3AB88DFA8E" author="Martin" box="[538,673,1266,1288]" pageId="8" pageNumber="65" refString="Martin, P. S. (1958) A biogeography of reptiles and amphibians in the Gomez Farias Region, Tamaulipas, Mexico. Miscellaneous Publications of the Museum of Zoology, University of Michigan, 101, 1 - 102." type="journal article" year="1958">Martin, 1958</bibRefCitation>
;
<bibRefCitation id="EFC24B28FFCE0054E4B2FB3AB90FFA8E" author="Rabb" box="[683,803,1266,1288]" pageId="8" pageNumber="65" refString="Rabb, G. B. (1958) On certain Mexican salamanders of the plethodontid genus Chiropterotriton. Occasional Papers of the Museum of Zoology, University of Michigan, 587, 1 - 37." type="journal article" year="1958">Rabb, 1958</bibRefCitation>
.
</paragraph>
<paragraph id="8BEC36D9FFCE0054E68EFAD9BE3EFAA1" blockId="8.[151,1042,1266,1319]" box="[151,1042,1297,1319]" pageId="8" pageNumber="65">
<emphasis id="B927EACBFFCE0054E68EFAD9BBCAFAA0" box="[151,486,1297,1318]" italics="true" pageId="8" pageNumber="65">
<taxonomicName id="4C534D5AFFCE0054E68EFAD9BB12FAA0" ID-CoL="3NCW" box="[151,318,1297,1318]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="65" phylum="Chordata" rank="genus">Chiropterotriton</taxonomicName>
cf
<taxonomicName id="4C534D5AFFCE0054E745FAD9BBCAFAA0" ID-CoL="5Y6KY" box="[348,486,1297,1318]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="multidentatus">multidentatus</taxonomicName>
</emphasis>
(part):
<bibRefCitation id="EFC24B28FFCE0054E42BFAD9BE3EFAA1" author="Campbell" box="[562,1042,1297,1319]" pageId="8" pageNumber="65" refString="Campbell, J. A., Streicher, J. W., Cox, C. L. &amp; Brodie, E. D. Jr. (2014) A new salamander of the genus Chiropterotriton (Caudata: Plethodontidae) from the Sierra Madre Oriental of Tamaulipas, Mexico. South American Journal of Herpetology, 9, 228 - 234. http: // dx. doi. org / 10.2994 / SAJH-D- 14 - 00042.1" type="journal article" year="2014">Campbell, Streicher, Cox, and Brodie, Jr., 2014</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3496552FFCE0054E68EFA9CBE9AF936" pageId="8" pageNumber="65" type="materials_examined">
<paragraph id="8BEC36D9FFCE0054E68EFA9CB871FA5E" blockId="8.[151,1437,1364,2037]" pageId="8" pageNumber="65">
<emphasis id="B927EACBFFCE0054E68EFA9CBB27FAEB" bold="true" box="[151,267,1364,1389]" pageId="8" pageNumber="65">
<typeStatus id="54E8887BFFCE0054E68EFA9CBB2AFAEB" box="[151,262,1364,1389]" pageId="8" pageNumber="65" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED42AE1CFFCE0054E70DFA9DBB7EFAEA" LSID="urn:lsid:biocol.org:col:34777" box="[276,338,1365,1388]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34777" name="Museum of Vertebrate Zoology, University of California Berkeley" pageId="8" pageNumber="65">MVZ</collectionCode>
269665 (original field number SMR2033), an adult male from the Cueva del Brinco, Conrado Castillo, ca.
<quantity id="4CAB9B3CFFCE0054E735FABFBBA1FA16" box="[300,397,1399,1424]" metricMagnitude="4" metricUnit="m" metricValue="4.35" pageId="8" pageNumber="65" unit="km" value="43.5">43.5 km</quantity>
SW (by rd) of Ejido Guayabas, Municipio Hidalgo, Tamaulipas,
<collectingCountry id="F3447649FFCE0054E280FABFBEDDFA16" box="[1177,1265,1399,1424]" name="Mexico" pageId="8" pageNumber="65">Mexico</collectingCountry>
,
<geoCoordinate id="EE67501EFFCE0054E318FABFBFB4FA09" box="[1281,1432,1399,1424]" direction="north" orientation="latitude" pageId="8" pageNumber="65" precision="1" value="23.95949">23.95949° N</geoCoordinate>
,
<geoCoordinate id="EE67501EFFCE0054E68EFA54BB01FA32" box="[151,301,1436,1460]" direction="west" orientation="longitude" pageId="8" pageNumber="65" precision="1" value="-99.47447">99.47447° W</geoCoordinate>
(WGS84 datum),
<quantity id="4CAB9B3CFFCE0054E7E3FA54B862FA32" box="[506,590,1436,1460]" metricMagnitude="3" metricUnit="m" metricValue="1.92" pageId="8" pageNumber="65" unit="m" value="1920.0">1920 m</quantity>
elevation, collected
<date id="FFED1019FFCE0054E52CFA54B9DCFA32" box="[821,1008,1436,1461]" pageId="8" pageNumber="65" value="2012-10-11">11 October 2012</date>
by Sean M. Rovito, Mariana Meneses Marín and Rafael Barzalobre Gerónimo.
</paragraph>
<paragraph id="8BEC36D9FFCE0054E6DEFA2CBB71F90B" blockId="8.[151,1437,1364,2037]" pageId="8" pageNumber="65">
<emphasis id="B927EACBFFCE0054E6DEFA2CBB6AFA7B" bold="true" box="[199,326,1508,1533]" pageId="8" pageNumber="65">
<typeStatus id="54E8887BFFCE0054E6DEFA2CBB6DFA7B" box="[199,321,1508,1533]" pageId="8" pageNumber="65" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9D55FD50FFCE0054E756FA2CBBF1FA7B" box="[335,477,1508,1533]" pageId="8" pageNumber="65" type="generic">8 specimens</specimenCount>
, all from the Sistema Purificación, Tamaulipas,
<collectingCountry id="F3447649FFCE0054E210FA2CBE4EFA7B" box="[1033,1122,1508,1533]" name="Mexico" pageId="8" pageNumber="65">Mexico</collectingCountry>
. Seven males: KU 182340, Sumidero de Oyamel; KU 182341, Viento Bajo; KU 182342, Pozo de Juan Fuentes Pérez, Conrado Castillo; KU 187773, Cueva X, Conrado Castillo; KU 187776, Cueva del Moro, Yerbabuena;
<collectionCode id="ED42AE1CFFCE0054E242F9E5BE9EF9C2" box="[1115,1202,1581,1604]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="8" pageNumber="65">UMMZ</collectionCode>
<date id="FFED1019FFCE0054E2A7F9E4BF3DF9C3" box="[1214,1297,1580,1605]" pageId="8" pageNumber="65" value="25-11-13">111325</date>
<date id="FFED1019FFCE0054E306F9E4BF5FF9C3" box="[1311,1395,1580,1605]" pageId="8" pageNumber="65" value="26-11-13">111326</date>
, El Chihue,
<quantity id="4CAB9B3CFFCE0054E6E1F998BB6DF9EE" box="[248,321,1615,1640]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="65" unit="km" value="17.0">17 km</quantity>
by road SE of Revillagigedo, ca.
<quantity id="4CAB9B3CFFCE0054E4D3F998B90DF9E1" box="[714,801,1616,1640]" metricMagnitude="3" metricUnit="m" metricValue="1.89" pageId="8" pageNumber="65" unit="m" value="1890.0">1890 m</quantity>
elevation (
<quantity id="4CAB9B3CFFCE0054E5BBF987B9D5F9EE" box="[930,1017,1615,1640]" metricMagnitude="3" metricUnit="m" metricValue="1.8897599999999997" pageId="8" pageNumber="65" unit="ft" value="6200.0">6200 ft</quantity>
). One female: KU187774, Cueva X, Conrado Castillo.
</paragraph>
<paragraph id="8BEC36D9FFCE0054E6DEF95FBE9AF936" blockId="8.[151,1437,1364,2037]" box="[199,1206,1687,1712]" pageId="8" pageNumber="65">
<emphasis id="B927EACBFFCE0054E6DEF95FBB95F936" bold="true" box="[199,441,1687,1712]" pageId="8" pageNumber="65">Referred specimens.</emphasis>
<collectionCode id="ED42AE1CFFCE0054E7D9F950BBDCF929" box="[448,496,1688,1711]" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="8" pageNumber="65">IBH</collectionCode>
29575, KU 182338, 182339,
<collectionCode id="ED42AE1CFFCE0054E527F950B9B9F929" box="[830,917,1688,1711]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="8" pageNumber="65">UMMZ</collectionCode>
<date id="FFED1019FFCE0054E585F950B9DDF936" box="[924,1009,1688,1712]" pageId="8" pageNumber="65" value="23-11-13">111323</date>
,
<date id="FFED1019FFCE0054E5E5F950BE7DF936" box="[1020,1105,1688,1712]" pageId="8" pageNumber="65" value="24-11-13">111324</date>
,
<date id="FFED1019FFCE0054E245F950BE9EF936" box="[1116,1202,1688,1712]" pageId="8" pageNumber="65" value="27-11-13">111327</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C3496552FFCE0054E6DEF974BFB0F872" pageId="8" pageNumber="65" type="diagnosis">
<paragraph id="8BEC36D9FFCE0054E6DEF974BFB0F872" blockId="8.[151,1437,1364,2037]" pageId="8" pageNumber="65">
<emphasis id="B927EACBFFCE0054E6DEF974BB6DF953" bold="true" box="[199,321,1724,1749]" pageId="8" pageNumber="65">Diagnosis.</emphasis>
Assigned to
<taxonomicName id="4C534D5AFFCE0054E7C1F974B8BDF952" box="[472,657,1724,1748]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="65" phylum="Chordata" rank="genus">
<emphasis id="B927EACBFFCE0054E7C1F974B8BDF952" box="[472,657,1724,1748]" italics="true" pageId="8" pageNumber="65">Chiropterotriton</emphasis>
</taxonomicName>
based on presence of sublingual fold, extensively webbed feet, long tail, and mtDNA sequence data analysis. Distinguished from all other species of
<taxonomicName id="4C534D5AFFCE0054E27FF928BF33F97E" box="[1126,1311,1760,1784]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="65" phylum="Chordata" rank="genus">
<emphasis id="B927EACBFFCE0054E27FF928BF33F97E" box="[1126,1311,1760,1784]" italics="true" pageId="8" pageNumber="65">Chiropterotriton</emphasis>
</taxonomicName>
except
<taxonomicName id="4C534D5AFFCE0054E39BF928BB2FF89A" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="magnipes">
<emphasis id="B927EACBFFCE0054E39BF928BB2FF89A" italics="true" pageId="8" pageNumber="65">C. magnipes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C534D5AFFCE0054E727F8CCBBE6F89A" box="[318,458,1796,1820]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="mosaueri">
<emphasis id="B927EACBFFCE0054E727F8CCBBE6F89A" box="[318,458,1796,1820]" italics="true" pageId="8" pageNumber="65">C. mosaueri</emphasis>
</taxonomicName>
by relative size of feet for males (FW/SVL
<taxonomicName id="4C534D5AFFCE0054E5C9F8CCBE73F89A" box="[976,1119,1796,1820]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="infernalis">
<emphasis id="B927EACBFFCE0054E5C9F8CCBE73F89A" box="[976,1119,1796,1820]" italics="true" pageId="8" pageNumber="65">C. infernalis</emphasis>
</taxonomicName>
males 0.115±0.011, female 0.092 vs.
<taxonomicName id="4C534D5AFFCE0054E708F8E0BB8AF8C6" box="[273,422,1832,1856]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="cieloensis">
<emphasis id="B927EACBFFCE0054E708F8E0BB8AF8C6" box="[273,422,1832,1856]" italics="true" pageId="8" pageNumber="65">C. cieloensis</emphasis>
</taxonomicName>
males 0.098±0.0093, females 0.098±0.118;
<taxonomicName id="4C534D5AFFCE0054E5A3F8E0BE14F8C6" box="[954,1080,1832,1856]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="cracens">
<emphasis id="B927EACBFFCE0054E5A3F8E0BE14F8C6" box="[954,1080,1832,1856]" italics="true" pageId="8" pageNumber="65">C. cracens</emphasis>
</taxonomicName>
males 0.087±0.0064, females 0.090±0.014;
<taxonomicName id="4C534D5AFFCE0054E72DF884BBCBF8E2" box="[308,487,1868,1892]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="miquihuanus">
<emphasis id="B927EACBFFCE0054E72DF884BBCBF8E2" box="[308,487,1868,1892]" italics="true" pageId="8" pageNumber="65">C. miquihuanus</emphasis>
</taxonomicName>
males 0.072±0.0069, females 0.073±0.0075;
<taxonomicName id="4C534D5AFFCE0054E5F4F884BE84F8E2" box="[1005,1192,1868,1892]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="multidentatus">
<emphasis id="B927EACBFFCE0054E5F4F884BE84F8E2" box="[1005,1192,1868,1892]" italics="true" pageId="8" pageNumber="65">C. multidentatus</emphasis>
</taxonomicName>
males 0.107±0.0076, females 0.101±0.013;
<taxonomicName id="4C534D5AFFCE0054E788F8B8B82FF801" box="[401,515,1904,1927]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="priscus">
<emphasis id="B927EACBFFCE0054E788F8B8B82FF801" box="[401,515,1904,1927]" italics="true" pageId="8" pageNumber="65">C. priscus</emphasis>
</taxonomicName>
males 0.084±0.0068, females 0.084±0.0064). Distinguished from
<taxonomicName id="4C534D5AFFCE0054E2F4F8B8BF54F801" box="[1261,1400,1904,1927]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="magnipes">
<emphasis id="B927EACBFFCE0054E2F4F8B8BF54F801" box="[1261,1400,1904,1927]" italics="true" pageId="8" pageNumber="65">C. magnipes</emphasis>
</taxonomicName>
by less extensive foot webbing (webbing reaches onto penultimate phalange in
<taxonomicName id="4C534D5AFFCE0054E5EAF85CBEACF82A" box="[1011,1152,1940,1964]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="infernalis">
<emphasis id="B927EACBFFCE0054E5EAF85CBEACF82A" box="[1011,1152,1940,1964]" italics="true" pageId="8" pageNumber="65">C. infernalis</emphasis>
</taxonomicName>
vs. all digits fully within web for
<taxonomicName id="4C534D5AFFCE0054E6ECF870BBADF856" box="[245,385,1976,2000]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="magnipes">
<emphasis id="B927EACBFFCE0054E6ECF870BBADF856" box="[245,385,1976,2000]" italics="true" pageId="8" pageNumber="65">C. magnipes</emphasis>
</taxonomicName>
), fewer premaxillary/maxillary and vomerine teeth (MT males:
<taxonomicName id="4C534D5AFFCE0054E24FF870BECEF856" box="[1110,1250,1976,2000]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="infernalis">
<emphasis id="B927EACBFFCE0054E24FF870BECEF856" box="[1110,1250,1976,2000]" italics="true" pageId="8" pageNumber="65">C. infernalis</emphasis>
</taxonomicName>
49.3±7.11 vs
<taxonomicName id="4C534D5AFFCE0054E39BF870BB2FF872" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="magnipes">
<emphasis id="B927EACBFFCE0054E39BF870BB2FF872" italics="true" pageId="8" pageNumber="65">C. magnipes</emphasis>
</taxonomicName>
79.1±5.59; females:
<taxonomicName id="4C534D5AFFCE0054E7E9F814B851F872" box="[496,637,2012,2036]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="infernalis">
<emphasis id="B927EACBFFCE0054E7E9F814B851F872" box="[496,637,2012,2036]" italics="true" pageId="8" pageNumber="65">C. infernalis</emphasis>
</taxonomicName>
53 vs.
<taxonomicName id="4C534D5AFFCE0054E4D6F814B970F872" box="[719,860,2012,2036]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="magnipes">
<emphasis id="B927EACBFFCE0054E4D6F814B970F872" box="[719,860,2012,2036]" italics="true" pageId="8" pageNumber="65">C. magnipes</emphasis>
</taxonomicName>
93.6±22.14; VT males:
<taxonomicName id="4C534D5AFFCE0054E276F814BED7F872" box="[1135,1275,2012,2036]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="infernalis">
<emphasis id="B927EACBFFCE0054E276F814BED7F872" box="[1135,1275,2012,2036]" italics="true" pageId="8" pageNumber="65">C. infernalis</emphasis>
</taxonomicName>
17.6±3.16 vs.
</paragraph>
</subSubSection>
<subSubSection id="C3496552FFCF0055E68EFF50BB02FCB1" pageId="9" pageNumber="66" type="reference_group">
<paragraph id="8BEC36D9FFCF0055E68EFF50BB02FCB1" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="66">
<taxonomicName id="4C534D5AFFCF0055E68EFF50BB0FFF29" ID-CoL="5Y6LS" box="[151,291,152,175]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="magnipes">
<emphasis id="B927EACBFFCF0055E68EFF50BB0FFF29" box="[151,291,152,175]" italics="true" pageId="9" pageNumber="66">C. magnipes</emphasis>
</taxonomicName>
29.5±6.23; females:
<taxonomicName id="4C534D5AFFCF0055E417FF50B8B6FF36" ID-CoL="5Y6LR" box="[526,666,152,176]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="infernalis">
<emphasis id="B927EACBFFCF0055E417FF50B8B6FF36" box="[526,666,152,176]" italics="true" pageId="9" pageNumber="66">C. infernalis</emphasis>
</taxonomicName>
15 vs.
<taxonomicName id="4C534D5AFFCF0055E4F3FF50B959FF29" ID-CoL="5Y6LS" box="[746,885,152,175]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="magnipes">
<emphasis id="B927EACBFFCF0055E4F3FF50B959FF29" box="[746,885,152,175]" italics="true" pageId="9" pageNumber="66">C. magnipes</emphasis>
</taxonomicName>
females 38.3±9.74), and darker coloration (
<bibRefCitation id="EFC24B28FFCF0055E346FF5FBAFFFF53" author="Rabb" pageId="9" pageNumber="66" refString="Rabb, G. B. (1965) A new salamander of the genus Chiropterotriton (Caudata: Plethodontidae) from Mexico. Breviora, 235, 1 - 8." type="journal article" year="1965">Rabb 1965</bibRefCitation>
). Differs from
<taxonomicName id="4C534D5AFFCF0055E765FF74B829FF52" ID-CoL="5Y6M5" box="[380,517,188,212]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="mosaueri">
<emphasis id="B927EACBFFCF0055E765FF74B829FF52" box="[380,517,188,212]" italics="true" pageId="9" pageNumber="66">C. mosaueri</emphasis>
</taxonomicName>
by having a relatively shorter tail in males (TL/SVL males:
<taxonomicName id="4C534D5AFFCF0055E2ABFF74BF12FF52" ID-CoL="5Y6LR" box="[1202,1342,188,212]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="infernalis">
<emphasis id="B927EACBFFCF0055E2ABFF74BF12FF52" box="[1202,1342,188,212]" italics="true" pageId="9" pageNumber="66">C. infernalis</emphasis>
</taxonomicName>
1.2±0.1 vs.
<taxonomicName id="4C534D5AFFCF0055E6DFFF28BB74FF71" ID-CoL="5Y6M5" box="[198,344,224,247]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="mosaueri">
<emphasis id="B927EACBFFCF0055E6DFFF28BB74FF71" box="[198,344,224,247]" italics="true" pageId="9" pageNumber="66">C. mosaueri</emphasis>
</taxonomicName>
1.3±0.16; females:
<taxonomicName id="4C534D5AFFCF0055E448FF28B8CAFF7E" ID-CoL="5Y6LR" box="[593,742,224,248]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="infernalis">
<emphasis id="B927EACBFFCF0055E448FF28B8CAFF7E" box="[593,742,224,248]" italics="true" pageId="9" pageNumber="66">C. infernalis</emphasis>
</taxonomicName>
1.2 vs.
<taxonomicName id="4C534D5AFFCF0055E54EFF28B9C5FF71" ID-CoL="5Y6M5" box="[855,1001,224,247]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="mosaueri">
<emphasis id="B927EACBFFCF0055E54EFF28B9C5FF71" box="[855,1001,224,247]" italics="true" pageId="9" pageNumber="66">C. mosaueri</emphasis>
</taxonomicName>
1.4) (
<bibRefCitation id="EFC24B28FFCF0055E225FF17BEC0FF7E" author="Woodall" box="[1084,1260,223,248]" pageId="9" pageNumber="66" refString="Woodall, H. T. (1941) A new Mexican salamander of the genus Oedipus. Occasional Papers of the Museum of Zoology, University of Michigan, 444, 1 - 4." type="journal article" year="1941">Woodall, 1941</bibRefCitation>
) and from
<taxonomicName id="4C534D5AFFCF0055E39BFF28BB1CFE9A" ID-CoL="5Y6KY" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="multidentatus">
<emphasis id="B927EACBFFCF0055E39BFF28BB1CFE9A" italics="true" pageId="9" pageNumber="66">C. multidentatus</emphasis>
</taxonomicName>
by having a relatively longer tail (TL/SVL
<taxonomicName id="4C534D5AFFCF0055E537FECCB9C5FE9A" ID-CoL="5Y6KY" box="[814,1001,260,284]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="multidentatus">
<emphasis id="B927EACBFFCF0055E537FECCB9C5FE9A" box="[814,1001,260,284]" italics="true" pageId="9" pageNumber="66">C. multidentatus</emphasis>
</taxonomicName>
: males 1.1±0.10, females 1.0±0.081). Additionally, differs from
<emphasis id="B927EACBFFCF0055E7DCFEE0B85EFEE2" italics="true" pageId="9" pageNumber="66">
<taxonomicName id="4C534D5AFFCF0055E7DCFEE0B87CFEC6" ID-CoL="5Y6LQ" box="[453,592,296,320]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="arboreus">C. arboreus</taxonomicName>
,
<taxonomicName id="4C534D5AFFCF0055E444FEE0B928FEC6" ID-CoL="5Y6LM" box="[605,772,296,320]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="chiropterus">C. chiropterus</taxonomicName>
,
<taxonomicName id="4C534D5AFFCF0055E508FEE0B9E3FEC6" ID-CoL="5Y6M7" box="[785,975,296,320]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="chondrostega">C. chondrostega</taxonomicName>
,
<taxonomicName id="4C534D5AFFCF0055E5C4FEE0BE76FEC6" ID-CoL="5Y6M8" box="[989,1114,296,320]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="cracens">C. cracens</taxonomicName>
,
<taxonomicName id="4C534D5AFFCF0055E271FEE0BF28FEC6" ID-CoL="5Y6M9" box="[1128,1284,296,320]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="dimidiatus">C. dimidiatus</taxonomicName>
,
<taxonomicName id="4C534D5AFFCF0055E30BFEE0BF59FEC6" ID-CoL="5Y6M3" box="[1298,1397,296,320]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="lavae">C. lavae</taxonomicName>
,
<taxonomicName id="4C534D5AFFCF0055E39AFEE0BB07FEE2" ID-CoL="5Y6LT" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="miquihuanus">C. miquihuanus</taxonomicName>
,
<taxonomicName id="4C534D5AFFCF0055E72FFE84BBDFFEE2" ID-CoL="5Y6KY" box="[310,499,332,356]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="multidentatus">C. multidentatus</taxonomicName>
,
<taxonomicName id="4C534D5AFFCF0055E7E7FE84B85EFEE2" ID-CoL="5Y6LG" box="[510,626,332,356]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="orculus">C. orculus</taxonomicName>
</emphasis>
,
<emphasis id="B927EACBFFCF0055E498FE84B8D5FEE2" box="[641,761,332,356]" italics="true" pageId="9" pageNumber="66">
<taxonomicName id="4C534D5AFFCF0055E498FE84B8DAFEE2" ID-CoL="5Y6LB" box="[641,758,332,356]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="priscus">C. priscus</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C534D5AFFCF0055E528FE84B995FEE2" ID-CoL="5Y6BF" box="[817,953,332,356]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="terrestris">
<emphasis id="B927EACBFFCF0055E528FE84B995FEE2" box="[817,953,332,356]" italics="true" pageId="9" pageNumber="66">C. terrestris</emphasis>
</taxonomicName>
by more extensive foot webbing (webbing extends onto penultimate phalange of third toe on foot of
<taxonomicName id="4C534D5AFFCF0055E536FEB8B991FE0E" ID-CoL="5Y6LR" box="[815,957,368,392]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="infernalis">
<emphasis id="B927EACBFFCF0055E536FEB8B991FE0E" box="[815,957,368,392]" italics="true" pageId="9" pageNumber="66">C. infernalis</emphasis>
</taxonomicName>
vs. to base of penultimate phalange in
<taxonomicName id="4C534D5AFFCF0055E39BFEB8BB9DFE2A" ID-CoL="5Y6LQ" authority="Taylor, 1941" authorityName="Taylor" authorityYear="1941" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="arboreus">
<emphasis id="B927EACBFFCF0055E39BFEB8BAD7FE2A" italics="true" pageId="9" pageNumber="66">C. arboreus</emphasis>
(
<bibRefCitation id="EFC24B28FFCF0055E708FE5CBB85FE2A" author="Taylor" box="[273,425,404,429]" pageId="9" pageNumber="66" refString="Taylor, E. H. (1941) New plethodont salamanders from Mexico. Herpetologica, 2, 57 - 65." type="journal article" year="1941">Taylor, 1941</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C534D5AFFCF0055E7EEFE5CB931FE2B" ID-CoL="5Y6LB" authority="Rabb, 1956" authorityName="Rabb" authorityYear="1956" box="[503,797,404,429]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="priscus">
<emphasis id="B927EACBFFCF0055E7EEFE5CB85DFE2A" box="[503,625,404,428]" italics="true" pageId="9" pageNumber="66">C. priscus</emphasis>
(Rabb, 1956)
</taxonomicName>
; only modest webbing present in
<emphasis id="B927EACBFFCF0055E2DDFE5CBEFAFE56" italics="true" pageId="9" pageNumber="66">
<taxonomicName id="4C534D5AFFCF0055E2DDFE5CBF43FE2A" ID-CoL="5Y6LM" box="[1220,1391,404,428]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="chiropterus">C. chiropterus</taxonomicName>
,
<taxonomicName id="4C534D5AFFCF0055E39BFE5CBB1DFE56" ID-CoL="5Y6M7" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="chondrostega">C. chondrostega</taxonomicName>
,
<taxonomicName id="4C534D5AFFCF0055E724FE70BB94FE56" ID-CoL="5Y6M8" box="[317,440,440,464]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="cracens">C. cracens</taxonomicName>
,
<taxonomicName id="4C534D5AFFCF0055E7DDFE70B873FE56" ID-CoL="5Y6M9" box="[452,607,440,464]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="dimidiatus">C. dimidiatus</taxonomicName>
,
<taxonomicName id="4C534D5AFFCF0055E472FE70B8E0FE56" ID-CoL="5Y6M3" box="[619,716,440,464]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="lavae">C. lavae</taxonomicName>
,
<taxonomicName id="4C534D5AFFCF0055E4C1FE70B9A1FE56" ID-CoL="5Y6LT" box="[728,909,440,464]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="miquihuanus">C. miquihuanus</taxonomicName>
,
<taxonomicName id="4C534D5AFFCF0055E580FE70BE7AFE56" ID-CoL="5Y6KY" box="[921,1110,440,464]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="multidentatus">C. multidentatus</taxonomicName>
,
<taxonomicName id="4C534D5AFFCF0055E27BFE70BEFAFE56" ID-CoL="5Y6LG" box="[1122,1238,440,464]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="orculus">C. orculus</taxonomicName>
</emphasis>
, and
<taxonomicName id="4C534D5AFFCF0055E30CFE70BB83FE72" ID-CoL="5Y6BF" authority="Campbell et al., 2014" authorityName="Campbell et al." authorityYear="2014" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="terrestris">
<emphasis id="B927EACBFFCF0055E30CFE70BFB7FE49" box="[1301,1435,440,463]" italics="true" pageId="9" pageNumber="66">C. terrestris</emphasis>
(
<bibRefCitation id="EFC24B28FFCF0055E687FE14BBB1FE72" author="Campbell" box="[158,413,476,501]" pageId="9" pageNumber="66" refString="Campbell, J. A., Streicher, J. W., Cox, C. L. &amp; Brodie, E. D. Jr. (2014) A new salamander of the genus Chiropterotriton (Caudata: Plethodontidae) from the Sierra Madre Oriental of Tamaulipas, Mexico. South American Journal of Herpetology, 9, 228 - 234. http: // dx. doi. org / 10.2994 / SAJH-D- 14 - 00042.1" type="journal article" year="2014">
Campbell
<emphasis id="B927EACBFFCF0055E70FFE15BB7DFE72" box="[278,337,476,500]" italics="true" pageId="9" pageNumber="66">et al.</emphasis>
, 2014
</bibRefCitation>
))
</taxonomicName>
. Differs from
<taxonomicName id="4C534D5AFFCF0055E44DFE14B925FE72" ID-CoL="5Y6LT" box="[596,777,476,500]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="miquihuanus">
<emphasis id="B927EACBFFCF0055E44DFE14B925FE72" box="[596,777,476,500]" italics="true" pageId="9" pageNumber="66">C. miquihuanus</emphasis>
</taxonomicName>
by much smaller external nares (NL males:
<taxonomicName id="4C534D5AFFCF0055E314FE14BFB0FE72" ID-CoL="5Y6LR" box="[1293,1436,476,500]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="infernalis">
<emphasis id="B927EACBFFCF0055E314FE14BFB0FE72" box="[1293,1436,476,500]" italics="true" pageId="9" pageNumber="66">C. infernalis</emphasis>
</taxonomicName>
0.24±0.03 vs.
<taxonomicName id="4C534D5AFFCF0055E759FDC8BBDAFD9E" ID-CoL="5Y6LT" box="[320,502,512,536]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="miquihuanus">
<emphasis id="B927EACBFFCF0055E759FDC8BBDAFD9E" box="[320,502,512,536]" italics="true" pageId="9" pageNumber="66">C. miquihuanus</emphasis>
</taxonomicName>
0.82±0.07; females:
<taxonomicName id="4C534D5AFFCF0055E4E9FDC8B9ACFD9E" ID-CoL="5Y6LR" box="[752,896,512,536]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="infernalis">
<emphasis id="B927EACBFFCF0055E4E9FDC8B9ACFD9E" box="[752,896,512,536]" italics="true" pageId="9" pageNumber="66">C. infernalis</emphasis>
</taxonomicName>
0.2 vs.
<taxonomicName id="4C534D5AFFCF0055E5FEFDC8BEB1FD9E" ID-CoL="5Y6LT" box="[999,1181,512,536]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="miquihuanus">
<emphasis id="B927EACBFFCF0055E5FEFDC8BEB1FD9E" box="[999,1181,512,536]" italics="true" pageId="9" pageNumber="66">C. miquihuanus</emphasis>
</taxonomicName>
0.85±0.04) and more premaxillary-maxillary teeth (MT males:
<taxonomicName id="4C534D5AFFCF0055E49AFDECB939FDBA" ID-CoL="5Y6LR" box="[643,789,548,572]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="infernalis">
<emphasis id="B927EACBFFCF0055E49AFDECB939FDBA" box="[643,789,548,572]" italics="true" pageId="9" pageNumber="66">C. infernalis</emphasis>
</taxonomicName>
49.2±7.11 vs.
<taxonomicName id="4C534D5AFFCF0055E5D5FDECBEA8FDBA" ID-CoL="5Y6LT" box="[972,1156,548,572]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="miquihuanus">
<emphasis id="B927EACBFFCF0055E5D5FDECBEA8FDBA" box="[972,1156,548,572]" italics="true" pageId="9" pageNumber="66">C. miquihuanus</emphasis>
</taxonomicName>
27.5±6.07; females:
<taxonomicName id="4C534D5AFFCF0055E39BFDECBB2FFDE6" ID-CoL="5Y6LR" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="infernalis">
<emphasis id="B927EACBFFCF0055E39BFDECBB2FFDE6" italics="true" pageId="9" pageNumber="66">C. infernalis</emphasis>
</taxonomicName>
53, vs.
<taxonomicName id="4C534D5AFFCF0055E77DFD80B836FDE6" ID-CoL="5Y6LT" box="[356,538,584,608]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="miquihuanus">
<emphasis id="B927EACBFFCF0055E77DFD80B836FDE6" box="[356,538,584,608]" italics="true" pageId="9" pageNumber="66">C. miquihuanus</emphasis>
</taxonomicName>
43.3±7.7), and from
<taxonomicName id="4C534D5AFFCF0055E50DFD80B959FDE6" ID-CoL="5Y6M3" box="[788,885,584,608]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="lavae">
<emphasis id="B927EACBFFCF0055E50DFD80B959FDE6" box="[788,885,584,608]" italics="true" pageId="9" pageNumber="66">C. lavae</emphasis>
</taxonomicName>
by having more vomerine teeth (VT males:
<taxonomicName id="4C534D5AFFCF0055E39BFD80BB2FFD02" ID-CoL="5Y6LR" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="infernalis">
<emphasis id="B927EACBFFCF0055E39BFD80BB2FFD02" italics="true" pageId="9" pageNumber="66">C. infernalis</emphasis>
</taxonomicName>
males 17.6±3.16, female 15 vs. approximately
<quantity id="4CAB9B3CFFCF0055E534FDA4B945FD03" box="[813,873,620,645]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="9" pageNumber="66" unit="in" value="12.0">12 in</quantity>
<taxonomicName id="4C534D5AFFCF0055E56AFDA4BEA2FD02" ID-CoL="5Y6M3" authority="Taylor, 1942" authorityName="Taylor" authorityYear="1942" box="[883,1166,620,645]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="lavae">
<emphasis id="B927EACBFFCF0055E56AFDA4B9F8FD02" box="[883,980,620,644]" italics="true" pageId="9" pageNumber="66">C. lavae</emphasis>
<bibRefCitation id="EFC24B28FFCF0055E5C4FDA4BEA2FD02" author="Taylor" box="[989,1166,620,645]" pageId="9" pageNumber="66" refString="Taylor, E. H. (1942) New Caudata and Salientia from Mexico. University of Kansas Science Bulletin, 28, 295 - 323." type="journal article" year="1942">(Taylor, 1942))</bibRefCitation>
</taxonomicName>
. Distinguished from
<emphasis id="B927EACBFFCF0055E39BFDA4BBE9FD2E" italics="true" pageId="9" pageNumber="66">
<taxonomicName id="4C534D5AFFCF0055E39BFDA4BB1DFD2E" ID-CoL="5Y6M7" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="chondrostega">C. chondrostega</taxonomicName>
,
<taxonomicName id="4C534D5AFFCF0055E758FD58BBEDFD2E" ID-CoL="5Y6M8" box="[321,449,656,680]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="cracens">C. cracens</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C534D5AFFCF0055E41FFD58B88EFD2E" ID-CoL="5Y6M9" box="[518,674,656,680]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="dimidiatus">
<emphasis id="B927EACBFFCF0055E41FFD58B88EFD2E" box="[518,674,656,680]" italics="true" pageId="9" pageNumber="66">C. dimidiatus</emphasis>
</taxonomicName>
by larger size (SVL males:
<taxonomicName id="4C534D5AFFCF0055E5E5FD58BEA1FD2E" ID-CoL="5Y6LR" box="[1020,1165,656,680]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="infernalis">
<emphasis id="B927EACBFFCF0055E5E5FD58BEA1FD2E" box="[1020,1165,656,680]" italics="true" pageId="9" pageNumber="66">C. infernalis</emphasis>
</taxonomicName>
36.4±3.84;
<taxonomicName id="4C534D5AFFCF0055E306FD58BFB0FD2E" ID-CoL="5Y6M8" box="[1311,1436,656,680]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="cracens">
<emphasis id="B927EACBFFCF0055E306FD58BFB0FD2E" box="[1311,1436,656,680]" italics="true" pageId="9" pageNumber="66">C. cracens</emphasis>
</taxonomicName>
25.7±1.98; females:
<taxonomicName id="4C534D5AFFCF0055E767FD7CB827FD4A" ID-CoL="5Y6LR" box="[382,523,692,716]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="infernalis">
<emphasis id="B927EACBFFCF0055E767FD7CB827FD4A" box="[382,523,692,716]" italics="true" pageId="9" pageNumber="66">C. infernalis</emphasis>
</taxonomicName>
29.7,
<taxonomicName id="4C534D5AFFCF0055E44BFD7CB8E6FD4A" ID-CoL="5Y6M8" box="[594,714,692,716]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="cracens">
<emphasis id="B927EACBFFCF0055E44BFD7CB8E6FD4A" box="[594,714,692,716]" italics="true" pageId="9" pageNumber="66">C. cracens</emphasis>
</taxonomicName>
27.4±3.30;
<taxonomicName id="4C534D5AFFCF0055E54BFD7CBE27FD4A" ID-CoL="5Y6M7" box="[850,1035,692,716]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="chondrostega">
<emphasis id="B927EACBFFCF0055E54BFD7CBE27FD4A" box="[850,1035,692,716]" italics="true" pageId="9" pageNumber="66">C. chondrostega</emphasis>
</taxonomicName>
maximum SVL 31.0 (
<bibRefCitation id="EFC24B28FFCF0055E311FD7CBFA3FD4B" author="Rabb" box="[1288,1423,692,717]" pageId="9" pageNumber="66" refString="Rabb, G. B. (1958) On certain Mexican salamanders of the plethodontid genus Chiropterotriton. Occasional Papers of the Museum of Zoology, University of Michigan, 587, 1 - 37." type="journal article" year="1958">Rabb, 1958</bibRefCitation>
),
<taxonomicName id="4C534D5AFFCF0055E68EFD10BB1AFD76" ID-CoL="5Y6M9" box="[151,310,728,752]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="dimidiatus">
<emphasis id="B927EACBFFCF0055E68EFD10BB1AFD76" box="[151,310,728,752]" italics="true" pageId="9" pageNumber="66">C. dimidiatus</emphasis>
</taxonomicName>
maximum SVL 29
<bibRefCitation id="EFC24B28FFCF0055E42CFD1FB8F2FD76" author="Rabb" box="[565,734,727,752]" pageId="9" pageNumber="66" refString="Rabb, G. B. (1958) On certain Mexican salamanders of the plethodontid genus Chiropterotriton. Occasional Papers of the Museum of Zoology, University of Michigan, 587, 1 - 37." type="journal article" year="1958">(Rabb, 1958))</bibRefCitation>
. Has more premaxillary-maxillary teeth in males than
<taxonomicName id="4C534D5AFFCF0055E39BFD10BB3BFC92" ID-CoL="5Y6LM" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="chiropterus">
<emphasis id="B927EACBFFCF0055E39BFD10BB3BFC92" italics="true" pageId="9" pageNumber="66">C. chiropterus</emphasis>
</taxonomicName>
(fewer than 12 MT in
<taxonomicName id="4C534D5AFFCF0055E430FD34B9CAFC92" ID-CoL="5Y6LM" authority="Campbell et al., 2014" authorityName="Campbell et al." authorityYear="2014" box="[553,998,764,789]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="chiropterus">
<emphasis id="B927EACBFFCF0055E430FD34B8E1FC92" box="[553,717,764,788]" italics="true" pageId="9" pageNumber="66">C. chiropterus</emphasis>
(
<bibRefCitation id="EFC24B28FFCF0055E4C6FD34B9F2FC92" author="Campbell" box="[735,990,764,789]" pageId="9" pageNumber="66" refString="Campbell, J. A., Streicher, J. W., Cox, C. L. &amp; Brodie, E. D. Jr. (2014) A new salamander of the genus Chiropterotriton (Caudata: Plethodontidae) from the Sierra Madre Oriental of Tamaulipas, Mexico. South American Journal of Herpetology, 9, 228 - 234. http: // dx. doi. org / 10.2994 / SAJH-D- 14 - 00042.1" type="journal article" year="2014">
Campbell
<emphasis id="B927EACBFFCF0055E541FD35B9BFFC92" box="[856,915,764,788]" italics="true" pageId="9" pageNumber="66">et al.</emphasis>
, 2014
</bibRefCitation>
)
</taxonomicName>
and more MT for both sexes than
<taxonomicName id="4C534D5AFFCF0055E39BFD34BB04FCBE" ID-CoL="5Y6LT" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="miquihuanus">
<emphasis id="B927EACBFFCF0055E39BFD34BB04FCBE" italics="true" pageId="9" pageNumber="66">C. miquihuanus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3496552FFCF0057E6DEFC8CBEE7F8B8" lastPageId="11" lastPageNumber="68" pageId="9" pageNumber="66" type="description">
<paragraph id="8BEC36D9FFCF0055E6DEFC8CBBCBFBFA" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="66">
<emphasis id="B927EACBFFCF0055E6DEFC8CB823FCDB" bold="true" box="[199,527,836,861]" pageId="9" pageNumber="66">
Description of the
<typeStatus id="54E8887BFFCF0055E7B8FC8CB826FCDB" box="[417,522,836,861]" pageId="9" pageNumber="66" type="holotype">holotype</typeStatus>
.
</emphasis>
Moderately sized adult male (SVL 39.3) with short trunk and very long (TL/SVL 1.32), thin tapering rounded tail. Large, wide head (HW/HL 0.61) with protruding eyes that extend just beyond margin of jaw when viewed from above. Nostril small (NL 0.23), oval shaped (NL/NW 1.44). Head flattened (head depth 2.7), snout blunt, nasolabial protuberances well developed. Oval-shaped mental gland present. Premaxillary teeth (3) pierce lip, vomerine teeth (21) arranged in a row extending nearly to outer margin of choanae. Very long limbs (LI -2), hands and feet very large and wide (FW 5.0) and webbed, with terminal and part of penultimate phalanges of third toe extending out of webbing on the hand and foot. Toe tips rounded with very well developed subterminal pads. Digits in order of increasing length: manus I-II~IV-III, pes I-V~II-IV-III. Phalangeal formulae: manus 1-2-3-2, pes 1-2-3-3-2.
</paragraph>
<paragraph id="8BEC36D9FFCF0055E6DEFB4FB86EFAD2" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="66">
<emphasis id="B927EACBFFCF0055E6DEFB4FBE36FB26" bold="true" box="[199,1050,1159,1184]" pageId="9" pageNumber="66">
Measurements (in mm), limb interval and tooth counts of the
<typeStatus id="54E8887BFFCF0055E5B5FB4FBE39FB26" box="[940,1045,1159,1184]" pageId="9" pageNumber="66" type="holotype">holotype</typeStatus>
.
</emphasis>
SVL 39.3, TL 51.9, AX 20.4, HL 10.1, HW 6.2, head depth at posterior angle of jaw 2.7, snout projection beyond mandible 0.4, IO 2.2, EN 1.7, IN 2.0, eyelid length 2.8, eyelid width 1.4, horizontal orbit diameter 2.1, FLL 11.6, HLL 12.9, snout to forelimb 13.1, width of right hand 3.8, FW 5.0, T3 1.5, T5 1.4, snout to anterior angle of vent 37.6, tail width at base 2.1, tail depth at base 2.5, shoulder width 4.6. NL 0.23, NW 0.16. Adpressed limbs overlap by 2 costal folds. Premaxillary teeth 3, maxillary teeth 53, vomerine teeth 21.
</paragraph>
<paragraph id="8BEC36D9FFCF0055E6DEFA97BE6AF9D6" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="66">
<emphasis id="B927EACBFFCF0055E6DEFA97BB12FAFE" bold="true" box="[199,318,1375,1400]" pageId="9" pageNumber="66">Variation:</emphasis>
The three largest individuals in the
<typeStatus id="54E8887BFFCF0055E4C9FAA9B92CFAFF" box="[720,768,1377,1401]" pageId="9" pageNumber="66">type</typeStatus>
series are approximately the same size (38.0
<quantity id="4CAB9B3CFFCF0055E319FA97BF4AFAFE" box="[1280,1382,1375,1400]" metricMagnitude="-2" metricUnit="m" metricValue="4.16" pageId="9" pageNumber="66" unit="mm" value="41.6">41.6 mm</quantity>
) and are sexually mature, suggesting that this may be near the maximum size for males of the species. The only female
<typeStatus id="54E8887BFFCF0055E68EFA61BADBFA47" box="[151,247,1449,1473]" pageId="9" pageNumber="66" type="paratype">paratype</typeStatus>
is substantially smaller than the
<typeStatus id="54E8887BFFCF0055E47BFA6FB8E8FA46" box="[610,708,1447,1472]" pageId="9" pageNumber="66" type="holotype">holotype</typeStatus>
(SVL
<quantity id="4CAB9B3CFFCF0055E516FA60B95BFA46" box="[783,887,1448,1472]" metricMagnitude="-2" metricUnit="m" metricValue="2.9699999999999998" pageId="9" pageNumber="66" unit="mm" value="29.7">29.7 mm</quantity>
), but the maximum size of females is likely equal to or larger than that of the
<typeStatus id="54E8887BFFCF0055E7DCFA04B807FA63" box="[453,555,1484,1509]" pageId="9" pageNumber="66" type="holotype">holotype</typeStatus>
. Compared to the
<typeStatus id="54E8887BFFCF0055E4ECFA04B976FA63" box="[757,858,1484,1509]" pageId="9" pageNumber="66" type="holotype">holotype</typeStatus>
, the limbs of the rest of the
<typeStatus id="54E8887BFFCF0055E294FA05BE91FA63" box="[1165,1213,1485,1509]" pageId="9" pageNumber="66">type</typeStatus>
series are relatively shorter, with the largest limb interval (relatively shortest limbs) equal to 1 costal fold of separation between adpressed limbs. Counts of maxillary and vomerine teeth vary substantially within this species; the largest male
<typeStatus id="54E8887BFFCF0055E68EF9F1BADBF9D7" box="[151,247,1593,1617]" pageId="9" pageNumber="66" type="paratype">paratype</typeStatus>
(UMMZ
<date id="FFED1019FFCF0055E77CF9FFBB96F9D6" box="[357,442,1591,1616]" pageId="9" pageNumber="66" value="25-11-13">111325</date>
, SVL
<quantity id="4CAB9B3CFFCF0055E418F9FFB84BF9D6" box="[513,615,1591,1616]" metricMagnitude="-2" metricUnit="m" metricValue="4.16" pageId="9" pageNumber="66" unit="mm" value="41.6">41.6 mm</quantity>
) has only 36 premaxillary-maxillary teeth.
</paragraph>
<paragraph id="8BEC36D9FFCF0055E6DEF994B879F8CA" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="66">
<emphasis id="B927EACBFFCF0055E6DEF994B8D6F9F3" bold="true" box="[199,762,1628,1653]" pageId="9" pageNumber="66">
Coloration of the
<typeStatus id="54E8887BFFCF0055E780F994B82CF9F3" box="[409,512,1628,1653]" pageId="9" pageNumber="66" type="holotype">holotype</typeStatus>
in life (from photos).
</emphasis>
Upper side of head and dorsum from rostrum to insertion of forelimbs Pratts Rufous (72) with scattered Clay Color (20) speckles, slightly lighter from snout to interocular region. Dorsum Sepia (286) with Smoke Gray (266) speckles. Upper side of tail Fawn Color (258) with Tawny Olive (17) speckles. Upper side of limbs Mahogany Red (34) with Medium Chrome Orange (75) mottling. Upper side of hands and feet Medium Fawn Color (256) with Smoky White (261) flecks. Toe tips Dark Rose (238). Gular region and underside of limbs Flesh Ocher (57), venter Dusky Brown (285) with scattered pale flecks. Underside of tail Sepia (279). Iris Yellow Ocher (14).
</paragraph>
<paragraph id="8BEC36D9FFCF0055E6DEF89FB99FF85B" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="66">
<emphasis id="B927EACBFFCF0055E6DEF89FB8ACF8F6" bold="true" box="[199,640,1879,1904]" pageId="9" pageNumber="66">
Coloration of the
<typeStatus id="54E8887BFFCF0055E781F89FBBD3F8F6" box="[408,511,1879,1904]" pageId="9" pageNumber="66" type="holotype">holotype</typeStatus>
in alcohol.
</emphasis>
Dorsum medium brown with pale flecks, becoming more numerous on head. Lighter brown on sides of head and from snout to interorbital region. Dorsal surface of limbs grey-brown with pale yellowish mottling, upper surface of feet greyish. Ventral surface greyish very pale yellow, with some very pale yellow mottling on gular region. Underside of feet light grey.
</paragraph>
<caption id="DF2C6651FFCC0056E68EF840B938F85A" httpUri="https://zenodo.org/record/290093/files/figure.png" pageId="10" pageNumber="67" targetBox="[207,1375,193,1896]" targetPageId="10">
<paragraph id="8BEC36D9FFCC0056E68EF840B938F85A" blockId="10.[151,1436,1928,2012]" pageId="10" pageNumber="67">
<emphasis id="B927EACBFFCC0056E68EF840BB36F818" bold="true" box="[151,282,1928,1950]" pageId="10" pageNumber="67">FIGURE 3.</emphasis>
Photos in life of
<taxonomicName id="4C534D5AFFCC0056E7F7F840B8B9F81B" box="[494,661,1928,1949]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="67" phylum="Chordata" rank="genus">
<emphasis id="B927EACBFFCC0056E7F7F840B8B9F81B" box="[494,661,1928,1949]" italics="true" pageId="10" pageNumber="67">Chiropterotriton</emphasis>
</taxonomicName>
from Tamaulipas, Nuevo León, and San Luis Potosí. A) holotype of
<taxonomicName id="4C534D5AFFCC0056E68EF86FBB87F83A" box="[151,427,1959,1980]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="67" phylum="Chordata" rank="species" species="cieloensis">
<emphasis id="B927EACBFFCC0056E68EF86FBB87F83A" box="[151,427,1959,1980]" italics="true" pageId="10" pageNumber="67">Chiropterotriton cieloensis</emphasis>
</taxonomicName>
(IBH 29561), B)
<taxonomicName id="4C534D5AFFCC0056E472F860B8F6F83B" box="[619,730,1960,1981]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="67" phylum="Chordata" rank="species" species="cracens">
<emphasis id="B927EACBFFCC0056E472F860B8F6F83B" box="[619,730,1960,1981]" italics="true" pageId="10" pageNumber="67">C. cracens</emphasis>
</taxonomicName>
(IBH 28178), C)
<taxonomicName id="4C534D5AFFCC0056E580F860BE68F83A" box="[921,1092,1959,1981]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="67" phylum="Chordata" rank="species" species="multidentatus">
<emphasis id="B927EACBFFCC0056E580F860BE68F83A" box="[921,1092,1959,1981]" italics="true" pageId="10" pageNumber="67">C. multidentatus</emphasis>
</taxonomicName>
(IBH 28177), D) holotype of
<taxonomicName id="4C534D5AFFCC0056E39DF860BAD4F85D" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="67" phylum="Chordata" rank="species" species="infernalis">
<emphasis id="B927EACBFFCC0056E39DF860BAD4F85D" italics="true" pageId="10" pageNumber="67">C. infernalis</emphasis>
</taxonomicName>
(MVZ 269665)., E)
<taxonomicName id="4C534D5AFFCC0056E7D2F80EB847F85D" box="[459,619,1990,2011]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="67" phylum="Chordata" rank="species" species="miquihuanus">
<emphasis id="B927EACBFFCC0056E7D2F80EB847F85D" box="[459,619,1990,2011]" italics="true" pageId="10" pageNumber="67">C. miquihuanus</emphasis>
</taxonomicName>
(MVZ 269632).
</paragraph>
</caption>
<caption id="DF2C6651FFCD0057E68EFAA0B854FA3D" httpUri="https://zenodo.org/record/290094/files/figure.png" pageId="11" pageNumber="68" targetBox="[161,1425,205,1362]" targetPageId="11">
<paragraph id="8BEC36D9FFCD0057E68EFAA0B854FA3D" blockId="11.[151,1436,1384,1467]" pageId="11" pageNumber="68">
<emphasis id="B927EACBFFCD0057E68EFAA0BB38FAF8" bold="true" box="[151,276,1384,1406]" pageId="11" pageNumber="68">FIGURE 4.</emphasis>
Head morphology of
<taxonomicName id="4C534D5AFFCD0057E7E1FAA0B8B3FAFB" box="[504,671,1384,1405]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="68" phylum="Chordata" rank="genus">
<emphasis id="B927EACBFFCD0057E7E1FAA0B8B3FAFB" box="[504,671,1384,1405]" italics="true" pageId="11" pageNumber="68">Chiropterotriton</emphasis>
</taxonomicName>
. A)
<taxonomicName id="4C534D5AFFCD0057E4D6FAA0BE2DFAFB" box="[719,1025,1384,1405]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="68" phylum="Chordata" rank="species" species="miquihuanus">
<emphasis id="B927EACBFFCD0057E4D6FAA0BE2DFAFB" box="[719,1025,1384,1405]" italics="true" pageId="11" pageNumber="68">Chiropterotriton miquihuanus</emphasis>
</taxonomicName>
(MVZ 269643), B)
<taxonomicName id="4C534D5AFFCD0057E2CDFAA0BF78FAFB" box="[1236,1364,1384,1405]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="68" phylum="Chordata" rank="species" species="infernalis">
<emphasis id="B927EACBFFCD0057E2CDFAA0BF78FAFB" box="[1236,1364,1384,1405]" italics="true" pageId="11" pageNumber="68">C. infernalis</emphasis>
</taxonomicName>
(MVZ 269665), C)
<taxonomicName id="4C534D5AFFCD0057E702FA4FBBA8FA1A" box="[283,388,1415,1436]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="68" phylum="Chordata" rank="species" species="priscus">
<emphasis id="B927EACBFFCD0057E702FA4FBBA8FA1A" box="[283,388,1415,1436]" italics="true" pageId="11" pageNumber="68">C. priscus</emphasis>
</taxonomicName>
(MVZ 269656), D)
<taxonomicName id="4C534D5AFFCD0057E442FA4FB929FA1A" box="[603,773,1415,1436]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="68" phylum="Chordata" rank="species" species="multidentatus">
<emphasis id="B927EACBFFCD0057E442FA4FB929FA1A" box="[603,773,1415,1436]" italics="true" pageId="11" pageNumber="68">C. multidentatus</emphasis>
</taxonomicName>
(MVZ 163911), E)
<taxonomicName id="4C534D5AFFCD0057E5CEFA4FBE69FA1A" box="[983,1093,1415,1436]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="68" phylum="Chordata" rank="species" species="cracens">
<emphasis id="B927EACBFFCD0057E5CEFA4FBE69FA1A" box="[983,1093,1415,1436]" italics="true" pageId="11" pageNumber="68">C. cracens</emphasis>
</taxonomicName>
(MVZ 173388), F)
<taxonomicName id="4C534D5AFFCD0057E300FA4FBFB0FA1A" box="[1305,1436,1415,1436]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="68" phylum="Chordata" rank="species" species="cieloensis">
<emphasis id="B927EACBFFCD0057E300FA4FBFB0FA1A" box="[1305,1436,1415,1436]" italics="true" pageId="11" pageNumber="68">C. cieloensis</emphasis>
</taxonomicName>
(MVZ158571). Scale bars correspond to 1 mm.
</paragraph>
</caption>
<paragraph id="8BEC36D9FFCD0057E6DEFA2ABEE7F8B8" blockId="11.[151,1436,1506,1999]" pageId="11" pageNumber="68">
<emphasis id="B927EACBFFCD0057E6DEFA2ABBDEFA7D" bold="true" box="[199,498,1506,1531]" pageId="11" pageNumber="68">Habitat and distribution.</emphasis>
All specimens of this species have been found in caves of the Sistema Purificación, the longest known cave system in
<collectingCountry id="F3447649FFCD0057E407F9CEB85BF999" box="[542,631,1542,1567]" name="Mexico" pageId="11" pageNumber="68">Mexico</collectingCountry>
. The two specimens for which we have detailed collecting information (the
<typeStatus id="54E8887BFFCD0057E6D0F9E2BB07F9C5" box="[201,299,1578,1603]" pageId="11" pageNumber="68" type="holotype">holotype</typeStatus>
and IBH 29575) were collected inside caves fairly near to the entrances. The habitat around the caves consists of mixed pine, oak, and
<taxonomicName id="4C534D5AFFCD0057E43DF986B89AF9E0" box="[548,694,1614,1638]" class="Magnoliopsida" family="Altingiaceae" genus="Liquidambar" kingdom="Plantae" order="Saxifragales" pageId="11" pageNumber="68" phylum="Tracheophyta" rank="genus">
<emphasis id="B927EACBFFCD0057E43DF986B89AF9E0" box="[548,694,1614,1638]" italics="true" pageId="11" pageNumber="68">Liquidambar</emphasis>
</taxonomicName>
forest with limestone rock outcrops containing small caves or crevices. The Sistema Purificación remains relatively poorly explored for salamanders, and the species doubtless occurs in many caves in the region of the
<typeStatus id="54E8887BFFCD0057E47CF95FB8B9F929" box="[613,661,1687,1711]" pageId="11" pageNumber="68">type</typeStatus>
locality. Although the
<typeStatus id="54E8887BFFCD0057E58CF95FB9E9F929" box="[917,965,1687,1711]" pageId="11" pageNumber="68">type</typeStatus>
series and all referred specimens are from Tamaulipas, the
<typeStatus id="54E8887BFFCD0057E749F973BBACF955" box="[336,384,1723,1747]" pageId="11" pageNumber="68">type</typeStatus>
locality is less than
<quantity id="4CAB9B3CFFCD0057E47EF973B88DF955" box="[615,673,1722,1747]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="68" unit="km" value="2.0">2 km</quantity>
from the border with Nuevo León, and the species almost certainly occurs in that state as well. T. Burkhardt (pers. comm.) found salamanders that we tentatively assign to this species to be abundant in a cave in Nuevo León east of Zaragoza, suggesting that it occurs somewhat more widely in the region. Both
<taxonomicName id="4C534D5AFFCD0057E733F8EEB817F8B8" box="[298,571,1830,1854]" class="Amphibia" family="Plethodontidae" genus="Aquiloeurycea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="68" phylum="Chordata" rank="species" species="galeanae">
<emphasis id="B927EACBFFCD0057E733F8EEB817F8B8" box="[298,571,1830,1854]" italics="true" pageId="11" pageNumber="68">Aquiloeurycea galeanae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C534D5AFFCD0057E468F8EFB8DAF8B8" box="[625,758,1830,1854]" class="Amphibia" family="Plethodontidae" genus="Aquiloeurycea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="68" phylum="Chordata" rank="species" species="scandens">
<emphasis id="B927EACBFFCD0057E468F8EFB8DAF8B8" box="[625,758,1830,1854]" italics="true" pageId="11" pageNumber="68">A. scandens</emphasis>
</taxonomicName>
occur at the
<typeStatus id="54E8887BFFCD0057E59EF8EFB99BF8B9" box="[903,951,1831,1855]" pageId="11" pageNumber="68">type</typeStatus>
locality of
<taxonomicName id="4C534D5AFFCD0057E221F8EFBEE8F8B8" box="[1080,1220,1830,1854]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="68" phylum="Chordata" rank="species" species="infernalis">
<emphasis id="B927EACBFFCD0057E221F8EFBEE8F8B8" box="[1080,1220,1830,1854]" italics="true" pageId="11" pageNumber="68">C. infernalis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3496552FFCD0057E6DEF882BBC8F849" pageId="11" pageNumber="68" type="etymology">
<paragraph id="8BEC36D9FFCD0057E6DEF882BBC8F849" blockId="11.[151,1436,1506,1999]" pageId="11" pageNumber="68">
<emphasis id="B927EACBFFCD0057E6DEF882BB62F8E5" bold="true" box="[199,334,1866,1891]" pageId="11" pageNumber="68">Etymology.</emphasis>
The specific epithet, Latin for infernal, hellish, or lower, makes reference to the cave-dwelling subterranean habits of this species. The incredible extent of the Sistema Purificación cave system, parts of which remain unexplored, gives the sensation of entering into the underworld when searching for salamanders at localities such as the Cueva del Brinco.
</paragraph>
</subSubSection>
</treatment>
</document>