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<document id="846DBBA6D7FA2069EBF6E3F2A075CF2B" ID-DOI="10.26028/cybium/2021-451-006" ID-ISSN="2101-0315" ID-Zenodo-Dep="10493556" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.tables_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1705002004153" checkinUser="felipe" docAuthor="Keith, Philippe, Delrieu-Trottin, Erwan, Utama, Ilham V., Sauri, Sopian, Busson, Frédéric, Nurjirana, Wowor, Daisy, Dahruddin, Hadi &amp; Hubert, Nicolas" docDate="2021" docId="03F487F9FFF3FFB3FF07BAC118627B2D" docLanguage="en" docName="Cybium.45.1.53-58.pdf" docOrigin="Cybium 45 (1)" docSource="http://dx.doi.org/10.26028/cybium/2021-451-006" docStyle="DocumentStyle:EAB95795642928CB986F0923DE58F0B7.1:Cybium.2019-.journal_article" docStyleId="EAB95795642928CB986F0923DE58F0B7" docStyleName="Cybium.2019-.journal_article" docStyleVersion="1" docTitle="Schismatogobius limmoni Keith &amp; Delrieu-Trottin &amp; Utama &amp; Sauri &amp; Busson &amp; Nurjirana &amp; Wowor &amp; Dahruddin &amp; Hubert 2021, n. sp." docType="treatment" docVersion="2" lastPageNumber="58" masterDocId="FFCDFF81FFF1FFB6FFB1B8001A7F785C" masterDocTitle="A new species of Schismatogobius (Teleostei: Gobiidae) from Sulawesi (Indonesia)" masterLastPageNumber="58" masterPageNumber="53" pageNumber="55" updateTime="1711984439773" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="B9F40B3A523603175D33E30A50BF1A14" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="5D8558BC498AEBE741906C1E8BD2A01F">A new species of Schismatogobius (Teleostei: Gobiidae) from Sulawesi (Indonesia)</mods:title>
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<mods:name id="18F31086F69E8437B5E2BB7AAC66FF4D" type="personal">
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<mods:namePart id="47C9C96A48BA4CEBBD7DAD8CC6AB6BDF">Keith, Philippe</mods:namePart>
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<mods:namePart id="6F8E7B7FA2E5351A90FB331483F0D9A8">Delrieu-Trottin, Erwan</mods:namePart>
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<mods:namePart id="F757040AC1C387E2FE2A7F702C6F3123">Utama, Ilham V.</mods:namePart>
</mods:name>
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<mods:namePart id="A6835D53B52C7FF63AB70867171C72F4">Sauri, Sopian</mods:namePart>
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<mods:namePart id="CB92BC62E777559EECC6A272E90ED236">Busson, Frédéric</mods:namePart>
</mods:name>
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<mods:namePart id="0B784F7E06CD760B64DF65587A74CEE3">Nurjirana</mods:namePart>
</mods:name>
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<mods:namePart id="9DDC65ADA38ACBA714D389D7C0947151">Wowor, Daisy</mods:namePart>
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<mods:namePart id="766A7F3B03909769C3ABC0423CC22699">Dahruddin, Hadi</mods:namePart>
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<mods:namePart id="DB1BC739AAA65B24FE8E8022705ABE13">Hubert, Nicolas</mods:namePart>
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<mods:typeOfResource id="5364D675D3CDE78C1DA29265860BE822">text</mods:typeOfResource>
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<mods:title id="E77D75FF17D354DE46ADD0DCD5BC07BF">Cybium</mods:title>
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<mods:date id="036A50474F747B700A4DDDAAB4052578">2021</mods:date>
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<mods:classification id="43407643070BF2B8372A910F880CD77A">journal article</mods:classification>
<mods:identifier id="91CC4A139ACFE17AEE0A549B09984437" type="DOI">10.26028/cybium/2021-451-006</mods:identifier>
<mods:identifier id="022E08BBF7CF767163E71A018D68437D" type="ISSN">2101-0315</mods:identifier>
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<treatment id="03F487F9FFF3FFB3FF07BAC118627B2D" ID-DOI="http://doi.org/10.5281/zenodo.10493566" ID-Zenodo-Dep="10493566" LSID="urn:lsid:plazi:treatment:03F487F9FFF3FFB3FF07BAC118627B2D" httpUri="http://treatment.plazi.org/id/03F487F9FFF3FFB3FF07BAC118627B2D" lastPageId="5" lastPageNumber="58" pageId="2" pageNumber="55">
<subSubSection id="C3476564FFF3FFB4FF07BAC118AA7A86" box="[182,725,705,730]" pageId="2" pageNumber="55" type="nomenclature">
<paragraph id="8BE236EFFFF3FFB4FF07BAC118AA7A86" blockId="2.[182,725,705,730]" box="[182,725,705,730]" pageId="2" pageNumber="55">
<heading id="D0AA8183FFF3FFB4FF07BAC118AA7A86" bold="true" box="[182,725,705,730]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="55" reason="2">
<emphasis id="B929EAFDFFF3FFB4FF07BAC118AA7A86" bold="true" box="[182,725,705,730]" pageId="2" pageNumber="55">
<taxonomicName id="4C5D4D6CFFF3FFB4FF07BAC11BAC7A85" authority="Keith &amp; Delrieu-Trottin &amp; Utama &amp; Sauri &amp; Busson &amp; Nurjirana &amp; Wowor &amp; Dahruddin &amp; Hubert, 2021" authorityName="Keith &amp; Delrieu-Trottin &amp; Utama &amp; Sauri &amp; Busson &amp; Nurjirana &amp; Wowor &amp; Dahruddin &amp; Hubert" authorityYear="2021" box="[182,467,705,729]" family="Gobiidae" genus="Schismatogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="limmoni" status="sp. nov.">
<emphasis id="B929EAFDFFF3FFB4FF07BAC11BAC7A85" bold="true" box="[182,467,705,729]" italics="true" pageId="2" pageNumber="55">Schismatogobius limmoni</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21A5786FFF3FFB4FE6EBAC218657A86" box="[479,538,706,730]" pageId="2" pageNumber="55" rank="species">n. sp.</taxonomicNameLabel>
Keith &amp; Hubert
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3476564FFF3FFB4FEDDBAE318607AA7" box="[364,543,739,763]" pageId="2" pageNumber="55" type="description">
<paragraph id="8BE236EFFFF3FFB4FEDDBAE318607AA7" blockId="2.[364,543,739,763]" box="[364,543,739,763]" pageId="2" pageNumber="55">
(
<figureCitation id="13662A6AFFF3FFB4FEC2BAE31BB07AA7" box="[371,463,739,763]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStartId-0="3.[132,195,1837,1859]" captionStartId-1="4.[132,198,1861,1883]" captionStartId-2="4.[813,881,420,442]" captionStartId-3="4.[813,880,710,732]" captionTargetBox-0="[132,1455,217,1816]" captionTargetBox-1="[140,767,1435,1838]" captionTargetBox-2="[813,1456,215,403]" captionTargetBox-3="[813,1456,498,694]" captionTargetId-0="figure-93@3.[132,1455,215,1820]" captionTargetId-1="figure-654@4.[132,775,1426,1844]" captionTargetId-2="figure-701@4.[813,1456,215,403]" captionTargetId-3="figure-671@4.[813,1456,498,694]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="4" captionTargetPageId-3="4" captionText-0="Figure 1. Most likely ML tree inferred using the TIM+I model (lnL = 3711.2906, I = 0.6520, f(A) = 0.2382, f(C) = 0.3018, f(G) = 0.1692, f(T) = 0.2909, AC = GT = 1.0, AT = CG = 0.6842, AG = 12.0806, CT = 6.0413). BP are given above each branch. Schismatogobius limmoni branch is highlighted in bold." captionText-1="Figure 2. Schematic drawing of the ventral surface of head of Schismatogobius limmoni." captionText-2="Figure 3. Schismatogobius limmoni n. sp., male, holotype MZB.25491 (SL 22.2 mm; BIF 10059) (Photo N. Hubert)." captionText-3="Figure 4. Schismatogobius limmoni n. sp., female, paratype in MNHN-IC 2020-0176 (SL 24.7 mm; BIF 10054) (Photo N. Hubert)." figureDoi-0="http://doi.org/10.5281/zenodo.10493558" figureDoi-1="http://doi.org/10.5281/zenodo.10493564" figureDoi-2="http://doi.org/10.5281/zenodo.10493560" figureDoi-3="http://doi.org/10.5281/zenodo.10493562" httpUri-0="https://zenodo.org/record/10493558/files/figure.png" httpUri-1="https://zenodo.org/record/10493564/files/figure.png" httpUri-2="https://zenodo.org/record/10493560/files/figure.png" httpUri-3="https://zenodo.org/record/10493562/files/figure.png" pageId="2" pageNumber="55">Figs 1-4</figureCitation>
;
<tableCitation id="C6DF0354FFF3FFB4FE68BAE318697AA6" box="[473,534,739,763]" captionStart="Table I" captionStartId="2.[813,868,217,239]" captionTargetId="graphics-749@2.[906,1361,436,1101]" captionTargetPageId="2" captionText="Table I. Morphomeristics of the new Schismatogobius species. Morphometrics are given as percentages of standard length, except JL/HL. P, Pectoral rays; D, Dorsal rays; A, Anal rays; C, Caudal branched rays; PDL, Predorsal length (% SL); PAL, Preanal length (% SL); HL, Head length (% SL); JL, jaw length (% SL); CPD, Caudal peduncle depth (% SL); Pect-L, Pectoral fin length (% SL); BDa, Body depth at anus (% SL); SDFL, Second dorsal fin length (% SL); AFL, Anal fin length (% SL); CFL, Caudal fin length (% SL)." httpUri="http://table.plazi.org/id/DF226667FFF3FFB4FC9CB8D9190679F3" pageId="2" pageNumber="55" tableUuid="DF226667FFF3FFB4FC9CB8D9190679F3">Tab. I</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3476564FFF3FFB4FF35BB231B587CF8" pageId="2" pageNumber="55" type="materials_examined">
<paragraph id="8BE236EFFFF3FFB4FF35BB231B247B67" blockId="2.[132,775,803,1188]" box="[132,347,803,827]" pageId="2" pageNumber="55">
<emphasis id="B929EAFDFFF3FFB4FF35BB231B247B67" bold="true" box="[132,347,803,827]" pageId="2" pageNumber="55">Material examined</emphasis>
</paragraph>
<paragraph id="8BE236EFFFF3FFB4FF1BBB441B397B20" blockId="2.[132,775,803,1188]" pageId="2" pageNumber="55">
<materialsCitation id="3B353CB2FFF3FFB4FF1BBB441B407B21" location="Sulawesi" pageId="2" pageNumber="55" specimenCount="7" stateProvince="Sulawesi Utara">
<specimenCount id="9D5BFD66FFF3FFB4FF1BBB441B107B00" box="[170,367,836,860]" count="7" pageId="2" pageNumber="55" type="generic">Seven specimens</specimenCount>
from
<collectingRegion id="4999F80DFFF3FFB4FE04BB4418637B00" box="[437,540,836,860]" country="Indonesia" name="Sulawesi Utara" pageId="2" pageNumber="55">Sulawesi</collectingRegion>
with a size range of
<quantity id="4CA59B0AFFF3FFB4FF35BB651B657B20" box="[132,282,869,893]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" metricValueMax="2.63" metricValueMin="2.17" pageId="2" pageNumber="55" unit="mm" value="24.0" valueMax="26.3" valueMin="21.7">21.7-26.3 mm</quantity>
SL
</materialsCitation>
.
</paragraph>
<paragraph id="8BE236EFFFF3FFB4FF1BBB861AA87C5C" blockId="2.[132,775,803,1188]" pageId="2" pageNumber="55">
<materialsCitation id="3B353CB2FFF3FFB4FF1BBB861AA87C5C" collectingDate="2019-07-12" collectionCode="MZB" country="Indonesia" county="Hubert" location="Palu" municipality="Wera Waterfall Nature Park" pageId="2" pageNumber="55" specimenCount="1" stateProvince="Sulawesi Utara" typeStatus="holotype">
<emphasis id="B929EAFDFFF3FFB4FF1BBB861B727BC2" box="[170,269,902,926]" italics="true" pageId="2" pageNumber="55">
<typeStatus id="54E6884DFFF3FFB4FF1BBB861B727BC2" box="[170,269,902,926]" pageId="2" pageNumber="55" type="holotype">Holotype</typeStatus>
</emphasis>
.
<collectionCode id="ED4CAE2AFFF3FFB4FE9FBB851B147BC1" box="[302,363,901,925]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="55" type="Museum">MZB</collectionCode>
.25491, male (
<quantity id="4CA59B0AFFF3FFB4FDB0BB86181C7BC1" box="[513,611,902,926]" metricMagnitude="-2" metricUnit="m" metricValue="2.2199999999999998" pageId="2" pageNumber="55" unit="mm" value="22.2">22.2 mm</quantity>
SL);
<collectingRegion id="4999F80DFFF3FFB4FD2EBB86197C7BC2" box="[671,771,902,926]" country="Indonesia" name="Sulawesi Utara" pageId="2" pageNumber="55">Sulawesi</collectingRegion>
,
<collectingRegion id="4999F80DFFF3FFB4FF35BBA61B3E7BE2" box="[132,321,934,958]" country="Indonesia" name="Sulawesi Tengah" pageId="2" pageNumber="55">Sulawesi Tengah</collectingRegion>
,
<location id="8E826034FFF3FFB4FEFCBBA61BFE7BE2" LSID="urn:lsid:plazi:treatment:03F487F9FFF3FFB3FF07BAC118627B2D:8E826034FFF3FFB4FEFCBBA61BFE7BE2" box="[333,385,934,958]" country="Indonesia" county="Hubert" municipality="Wera Waterfall Nature Park" name="Palu" pageId="2" pageNumber="55" stateProvince="Sulawesi Utara">Palu</location>
,
<collectingMunicipality id="6B86AC95FFF3FFB4FE3DBBA718BF7BE2" box="[396,704,934,958]" pageId="2" pageNumber="55">Wera Waterfall Nature Park</collectingMunicipality>
,
<collectingCountry id="F34A767FFFF3FFB4FD7ABBA61ABE7B83" name="Indonesia" pageId="2" pageNumber="55">Indonesia</collectingCountry>
,
<date id="FFE3102FFFF3FFB4FF7FBBC71B237B83" box="[206,348,967,991]" pageId="2" pageNumber="55" value="2019-07-12">
<collectingDate id="EFA7E9C7FFF3FFB4FF7FBBC71B237B83" box="[206,348,967,991]" pageId="2" pageNumber="55" value="2019-07-12">12 Jul. 2019</collectingDate>
</date>
,
<collectingCounty id="62834E63FFF3FFB4FED9BBC71BC97B83" box="[360,438,967,991]" pageId="2" pageNumber="55">Hubert</collectingCounty>
<emphasis id="B929EAFDFFF3FFB4FE0FBBC81B877B83" box="[446,504,967,991]" italics="true" pageId="2" pageNumber="55">et al.</emphasis>
coll. (field number BIF 10059).
</materialsCitation>
</paragraph>
<paragraph id="8BE236EFFFF3FFB4FF1BBC091B587CF8" blockId="2.[132,775,803,1188]" pageId="2" pageNumber="55">
<materialsCitation id="3B353CB2FFF3FFB4FF1BBC09197D7C7D" box="[170,770,1033,1057]" collectionCode="MZB" pageId="2" pageNumber="55" specimenCount="4" specimenCount-female="4" typeStatus="paratype">
<emphasis id="B929EAFDFFF3FFB4FF1BBC091B697C7C" box="[170,278,1033,1056]" italics="true" pageId="2" pageNumber="55">
<typeStatus id="54E6884DFFF3FFB4FF1BBC091B697C7C" box="[170,278,1033,1056]" pageId="2" pageNumber="55" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<collectionCode id="ED4CAE2AFFF3FFB4FE87BC091B0D7C7D" box="[310,370,1033,1057]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="55" type="Museum">MZB</collectionCode>
.25492,
<specimenCount id="9D5BFD66FFF3FFB4FE77BC0918527C7D" box="[454,557,1033,1057]" count="4" pageId="2" pageNumber="55" type="female">4 females</specimenCount>
(
<quantity id="4CA59B0AFFF3FFB4FD8BBC0918AE7C7C" box="[570,721,1033,1057]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" metricValueMax="2.63" metricValueMin="2.17" pageId="2" pageNumber="55" unit="mm" value="24.0" valueMax="26.3" valueMin="21.7">21.7-26.3 mm</quantity>
SL)
</materialsCitation>
;
<materialsCitation id="3B353CB2FFF3FFB4FF35BC2A1B4C7C3E" collectingDate="2019-07-12" collectionCode="MNHN-IC" country="Indonesia" county="Hubert" location="Palu" municipality="Wera Waterfall Nature Park" pageId="2" pageNumber="55" specimenCode="MNHN-IC 2020-0176" specimenCount="2" specimenCount-female="2" stateProvince="Sulawesi Utara" typeStatus="paratype">same data as for holotype (field numbers BIF 10050, 10058, 10060, 10061).</materialsCitation>
<materialsCitation id="3B353CB2FFF3FFB4FE8DBC4A1B5C7CF8" collectingDate="2019-07-12" collectionCode="MNHN-IC" country="Indonesia" county="Hubert" location="Palu" municipality="Wera Waterfall Nature Park" pageId="2" pageNumber="55" specimenCode="MNHN-IC 2020-0176" specimenCount="2" specimenCount-female="2" stateProvince="Sulawesi Utara" typeStatus="paratype">
<specimenCode id="DBFB9E94FFF3FFB4FE8DBC4A18427C3E" box="[316,573,1098,1122]" collectionCode="MNHN-IC" pageId="2" pageNumber="55">MNHN-IC 2020-0176</specimenCode>
,
<specimenCount id="9D5BFD66FFF3FFB4FDFBBC4A18C67C3E" box="[586,697,1098,1122]" count="2" pageId="2" pageNumber="55" type="female">2 females</specimenCount>
(
<quantity id="4CA59B0AFFF3FFB4FD7BBC4A1A9B7CDF" metricMagnitude="-2" metricUnit="m" metricValue="2.475" metricValueMax="2.48" metricValueMin="2.4699999999999998" pageId="2" pageNumber="55" unit="mm" value="24.75" valueMax="24.8" valueMin="24.7">24.7- 24.8 mm</quantity>
SL); same data as for holotype (field numbers BIF 10054, 10055)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3476564FFF3FFB4FF35BCCC187C7E50" pageId="2" pageNumber="55" type="diagnosis">
<paragraph id="8BE236EFFFF3FFB4FF35BCCC1A8D7CB8" blockId="2.[132,775,1228,1548]" box="[132,242,1228,1252]" pageId="2" pageNumber="55">
<emphasis id="B929EAFDFFF3FFB4FF35BCCC1A8D7CB8" bold="true" box="[132,242,1228,1252]" pageId="2" pageNumber="55">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BE236EFFFF3FFB4FF1BBCEE187C7E50" blockId="2.[132,775,1228,1548]" pageId="2" pageNumber="55">
A small
<taxonomicName id="4C5D4D6CFFF3FFB4FEB6BCED1BBC7D59" authorityName="Keith &amp; Delrieu-Trottin &amp; Utama &amp; Sauri &amp; Busson &amp; Nurjirana &amp; Wowor &amp; Dahruddin &amp; Hubert" authorityYear="2021" box="[263,451,1261,1285]" family="Gobiidae" genus="Schismatogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
<emphasis id="B929EAFDFFF3FFB4FEB6BCED1BBC7D59" box="[263,451,1261,1285]" italics="true" pageId="2" pageNumber="55">Schismatogobius</emphasis>
</taxonomicName>
. 16-17 pectoral rays (mostly 17); pectoral fins with a medium black triangular wedge dorsally usually with 2 vertical rows of dark spots below it. First dorsal fin membrane posterior to spine 6 not connected to base of spine of second dorsal fin. Ventral surface of head in male blackish or greyish in the medium part. Frenum whitish. Ventral surface of head in female mainly whitish. Frenum whitish. A single mitochondrial lineage was observed for this species (BOLD: AEA7944).
</paragraph>
</subSubSection>
<subSubSection id="C3476564FFF3FFB2FF35BE341F037D6E" lastPageId="4" lastPageNumber="57" pageId="2" pageNumber="55" type="description">
<paragraph id="8BE236EFFFF3FFB4FF35BE341B777E10" blockId="2.[132,776,1588,1908]" box="[132,264,1588,1612]" pageId="2" pageNumber="55">
<emphasis id="B929EAFDFFF3FFB4FF35BE341B777E10" bold="true" box="[132,264,1588,1612]" pageId="2" pageNumber="55">Description</emphasis>
</paragraph>
<paragraph id="8BE236EFFFF3FFB4FF1BBE551F5D7CF1" blockId="2.[132,776,1588,1908]" lastBlockId="2.[813,1456,1141,1908]" pageId="2" pageNumber="55">
A small-sized
<taxonomicName id="4C5D4D6CFFF3FFB4FEE7BE5518637E31" authorityName="Keith &amp; Delrieu-Trottin &amp; Utama &amp; Sauri &amp; Busson &amp; Nurjirana &amp; Wowor &amp; Dahruddin &amp; Hubert" authorityYear="2021" box="[342,540,1621,1645]" family="Gobiidae" genus="Schismatogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
<emphasis id="B929EAFDFFF3FFB4FEE7BE5518637E31" box="[342,540,1621,1645]" italics="true" pageId="2" pageNumber="55">Schismatogobius</emphasis>
</taxonomicName>
(average adult size &lt;
<quantity id="4CA59B0AFFF3FFB4FF2CBE761A917ED1" box="[157,238,1654,1678]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="55" unit="mm" value="27.0">27 mm</quantity>
SL). Body naked, slender, almost circular in cross-section. Head (27.8-29.8 %SL) rounded, snout rather pointed. Mouth oblique, lower lip more prominent. Upper jaw length in male (13.5 %SL) much greater than in female (10.5-11.7 %SL); jaw length 45.5 % of HL in male (
<typeStatus id="54E6884DFFF3FFB4FD7ABEF919787F4C" box="[715,775,1785,1809]" pageId="2" pageNumber="55" type="holotype">
<typeStatus id="54E6884DFFF3FFB4FF35BF1A1ACA7F6E" box="[132,181,1818,1842]" pageId="2" pageNumber="55">holotype</typeStatus>
</typeStatus>
) and 36.5-39% of HL in females. Lower jaw reaching vertical of 1/2 to 2/3 of the eye in female and&gt; 2/
<quantity id="4CA59B0AFFF3FFB4FD2FBF3B18BA7F0E" box="[670,709,1850,1875]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="2" pageNumber="55" unit="in" value="3.0">3 in</quantity>
male. Eyes high on head, close together with interorbital width about equal to 2/3 eye diameter. Anterior nostril short and tube-like. Morphomeristic data given in table I.
</paragraph>
<paragraph id="8BE236EFFFF3FFB4FCE2BCB619E17E0D" blockId="2.[813,1456,1141,1908]" pageId="2" pageNumber="55">Dorsal fins VI-I,8-9, membrane in first dorsal fin posterior to spine 6 not connected to base of spine of second dorsal fin. D1 with all spines about equal in length. Anal fin I,8-9, origin directly opposite to second dorsal fin origin. Caudal fin with 10-12 branched rays, posterior margin straight. Pectoral fins oblong with posterior margin rounded and 16(2)-17(5) rays, ventralmost ray unbranched. Pelvic fins always I,5, with both fins joined together on their entire length between fifth rays to form a strong cup-like disc and a well developed and lobed fraenum between spines; fins not extending beyond anus. Body depth at anus 14.1-15.2%SL, predorsal length 41.1-45.6 %SL and preanal length 62.9- 68.4 %SL.</paragraph>
<paragraph id="8BE236EFFFF3FFB4FCE2BE5A1EB47EAE" blockId="2.[813,1456,1141,1908]" pageId="2" pageNumber="55">Tongue bilobed. Teeth in upper jaw in two rows (first row 12-17), conical and slightly recurved, mostly caniniform in frontal position in male. Teeth in lower jaw in 2-3 rows (first row 12-17), all teeth conical; only outer row with slightly enlarged and somewhat recurved teeth.</paragraph>
<paragraph id="8BE236EFFFF3FFB2FCE2BEFB1B0479ED" blockId="2.[813,1456,1141,1908]" lastBlockId="4.[132,775,217,561]" lastPageId="4" lastPageNumber="57" pageId="2" pageNumber="55">Cephalic sensory pore system always with pores B, D, F, K, L, N and O, pore D singular with all other pores paired; oculoscapular canal absent between pores F and K. Anterior interorbital extension of anterior oculoscapular canal with double terminal pores B slightly posterior to posterior nostril. D pore at rear of intertobital. Posterior extension of anterior oculoscapular canal terminating laterally on each side of head at pore F, just behind posterior edge of eye. Posterior oculoscapular canal with 2 terminal pores, K and L; preopercular canal with 2 pores, N and O. Cutaneous sensory papillae not well developed.</paragraph>
<caption id="DF226667FFF2FFB5FF35BF2D1BAD7F2F" ID-DOI="http://doi.org/10.5281/zenodo.10493558" ID-Zenodo-Dep="10493558" httpUri="https://zenodo.org/record/10493558/files/figure.png" pageId="3" pageNumber="56" startId="3.[132,195,1837,1859]" targetBox="[132,1455,217,1816]" targetPageId="3" targetType="figure">
<paragraph id="8BE236EFFFF2FFB5FF35BF2D1BAD7F2F" blockId="3.[132,1456,1837,1907]" pageId="3" pageNumber="56">
Figure 1. Most likely ML tree inferred using the TIM+I model (lnL = 3711.2906, I = 0.6520, f(A) = 0.2382, f(C) = 0.3018, f(G) = 0.1692, f(T) = 0.2909, AC = GT = 1.0, AT = CG = 0.6842, AG = 12.0806, CT = 6.0413). BP are given above each branch.
<taxonomicName id="4C5D4D6CFFF2FFB5FB6CBF451ACB7F2E" authorityName="Keith &amp; Delrieu-Trottin &amp; Utama &amp; Sauri &amp; Busson &amp; Nurjirana &amp; Wowor &amp; Dahruddin &amp; Hubert" authorityYear="2021" family="Gobiidae" genus="Schismatogobius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="limmoni">
<emphasis id="B929EAFDFFF2FFB5FB6CBF451ACB7F2E" italics="true" pageId="3" pageNumber="56">Schismatogobius limmoni</emphasis>
</taxonomicName>
branch is highlighted in bold.
</paragraph>
</caption>
<caption id="DF226667FFF5FFB2FC9CB9A41F26798E" ID-DOI="http://doi.org/10.5281/zenodo.10493560" ID-Zenodo-Dep="10493560" httpUri="https://zenodo.org/record/10493560/files/figure.png" pageId="4" pageNumber="57" startId="4.[813,881,420,442]" targetBox="[813,1456,215,403]" targetPageId="4" targetType="figure">
<paragraph id="8BE236EFFFF5FFB2FC9CB9A41F26798E" blockId="4.[813,1455,420,466]" pageId="4" pageNumber="57">
Figure 3.
<taxonomicName id="4C5D4D6CFFF5FFB2FC1DB9A41EC179E5" authority="Keith &amp; Delrieu-Trottin &amp; Utama &amp; Sauri &amp; Busson &amp; Nurjirana &amp; Wowor &amp; Dahruddin &amp; Hubert, 2021" authorityName="Keith &amp; Delrieu-Trottin &amp; Utama &amp; Sauri &amp; Busson &amp; Nurjirana &amp; Wowor &amp; Dahruddin &amp; Hubert" authorityYear="2021" box="[940,1214,420,441]" family="Gobiidae" genus="Schismatogobius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="57" phylum="Chordata" rank="species" species="limmoni" status="sp. nov.">
<emphasis id="B929EAFDFFF5FFB2FC1DB9A41EC179E5" box="[940,1214,420,441]" italics="true" pageId="4" pageNumber="57">Schismatogobius limmoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A5786FFF5FFB2FB77B9A41F7B79E5" box="[1222,1284,420,441]" pageId="4" pageNumber="57" rank="species">n. sp.</taxonomicNameLabel>
, male, holotype MZB.25491 (SL 22.2 mm; BIF 10059) (Photo N. Hubert).
</paragraph>
</caption>
<paragraph id="8BE236EFFFF5FFB2FF1BB9B91BCC7A6D" blockId="4.[132,775,217,561]" pageId="4" pageNumber="57">Sexual dimorphism fairly well developed with male having jaws longer than female and a different colour pattern on ventral surface of head. Urogenital papilla oval in female and slightly pointed in male.</paragraph>
<paragraph id="8BE236EFFFF5FFB2FF35BA591B087A2D" blockId="4.[132,775,601,1169]" box="[132,375,601,625]" pageId="4" pageNumber="57">
<emphasis id="B929EAFDFFF5FFB2FF35BA591B087A2D" box="[132,375,601,625]" italics="true" pageId="4" pageNumber="57">Colour in preservation</emphasis>
</paragraph>
<paragraph id="8BE236EFFFF5FFB2FF1BBA791BD57CCD" blockId="4.[132,775,601,1169]" pageId="4" pageNumber="57">
Five vertical black bands in dorsal view. The first one on the nape, the second one below the first dorsal fin, the third and fourth below the second dorsal fin and the fifth band at hypural crease. These bands are not uniform but with dorsal or lateral brown to greyish spots. Colour of body between each band whitish to greyish. Head and cheeks usually mottled with closely spaced black or grey spots and markings. Inferior half of lateral part of head brownish in male. Ventral surface of head in male blackish or greyish in the medium part. Fraenum whitish (
<figureCitation id="13662A6AFFF5FFB2FE3ABB991BAE7BED" box="[395,465,921,945]" captionStart="Figure 2" captionStartId="4.[132,198,1861,1883]" captionTargetBox="[140,767,1435,1838]" captionTargetId="figure-654@4.[132,775,1426,1844]" captionTargetPageId="4" captionText="Figure 2. Schematic drawing of the ventral surface of head of Schismatogobius limmoni." figureDoi="http://doi.org/10.5281/zenodo.10493564" httpUri="https://zenodo.org/record/10493564/files/figure.png" pageId="4" pageNumber="57">Fig. 2</figureCitation>
). Ventral surface of head in female whitish, lightly spotted of brown. Fraenum whitish (
<figureCitation id="13662A6AFFF5FFB2FF05BBD91A877BAD" box="[180,248,985,1009]" captionStart="Figure 2" captionStartId="4.[132,198,1861,1883]" captionTargetBox="[140,767,1435,1838]" captionTargetId="figure-654@4.[132,775,1426,1844]" captionTargetPageId="4" captionText="Figure 2. Schematic drawing of the ventral surface of head of Schismatogobius limmoni." figureDoi="http://doi.org/10.5281/zenodo.10493564" httpUri="https://zenodo.org/record/10493564/files/figure.png" pageId="4" pageNumber="57">Fig. 2</figureCitation>
). Belly whitish to greyish. First dorsal and second dorsal fins with rows of black spots on rays. Caudal fin mainly translucent with two black spots posteriorly. A black band at hypural base. Pectoral fins translucent with a medium black triangular wedge dorsally usually with 2 vertical rows of dark spots below it.
</paragraph>
<caption id="DF226667FFF5FFB2FC9CBAC619017B50" ID-DOI="http://doi.org/10.5281/zenodo.10493562" ID-Zenodo-Dep="10493562" httpUri="https://zenodo.org/record/10493562/files/figure.png" pageId="4" pageNumber="57" startId="4.[813,880,710,732]" targetBox="[813,1456,498,694]" targetPageId="4" targetType="figure">
<paragraph id="8BE236EFFFF5FFB2FC9CBAC619017B50" blockId="4.[813,1455,710,780]" pageId="4" pageNumber="57">
Figure 4.
<taxonomicName id="4C5D4D6CFFF5FFB2FC16BAC61ECD7A87" authority="Keith &amp; Delrieu-Trottin &amp; Utama &amp; Sauri &amp; Busson &amp; Nurjirana &amp; Wowor &amp; Dahruddin &amp; Hubert, 2021" authorityName="Keith &amp; Delrieu-Trottin &amp; Utama &amp; Sauri &amp; Busson &amp; Nurjirana &amp; Wowor &amp; Dahruddin &amp; Hubert" authorityYear="2021" box="[935,1202,710,731]" family="Gobiidae" genus="Schismatogobius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="57" phylum="Chordata" rank="species" species="limmoni" status="sp. nov.">
<emphasis id="B929EAFDFFF5FFB2FC16BAC61ECD7A87" box="[935,1202,710,731]" italics="true" pageId="4" pageNumber="57">Schismatogobius limmoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A5786FFF5FFB2FB0BBAC71E8A7A80" box="[1210,1269,711,732]" pageId="4" pageNumber="57" rank="species">n. sp.</taxonomicNameLabel>
, female, paratype
<emphasis id="B929EAFDFFF5FFB2FC9CBADF193F7AA8" box="[813,832,735,756]" italics="true" pageId="4" pageNumber="57">in</emphasis>
MNHN-IC 2020-0176 (SL 24.7 mm; BIF 10054) (Photo N. Hubert).
</paragraph>
</caption>
<paragraph id="8BE236EFFFF5FFB2FF35BCB91BFD7C8D" blockId="4.[132,775,1209,1393]" box="[132,386,1209,1233]" pageId="4" pageNumber="57">
<emphasis id="B929EAFDFFF5FFB2FF35BCB91B5D7C8D" box="[132,290,1209,1233]" italics="true" pageId="4" pageNumber="57">Colour in life (</emphasis>
<figureCitation id="13662A6AFFF5FFB2FE93BCB91B067C8D" box="[290,377,1209,1233]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStartId-0="4.[813,881,420,442]" captionStartId-1="4.[813,880,710,732]" captionTargetBox-0="[813,1456,215,403]" captionTargetBox-1="[813,1456,498,694]" captionTargetId-0="figure-701@4.[813,1456,215,403]" captionTargetId-1="figure-671@4.[813,1456,498,694]" captionTargetPageId-0="4" captionTargetPageId-1="4" captionText-0="Figure 3. Schismatogobius limmoni n. sp., male, holotype MZB.25491 (SL 22.2 mm; BIF 10059) (Photo N. Hubert)." captionText-1="Figure 4. Schismatogobius limmoni n. sp., female, paratype in MNHN-IC 2020-0176 (SL 24.7 mm; BIF 10054) (Photo N. Hubert)." figureDoi-0="http://doi.org/10.5281/zenodo.10493560" figureDoi-1="http://doi.org/10.5281/zenodo.10493562" httpUri-0="https://zenodo.org/record/10493560/files/figure.png" httpUri-1="https://zenodo.org/record/10493562/files/figure.png" pageId="4" pageNumber="57">Figs 3-4</figureCitation>
)
</paragraph>
<paragraph id="8BE236EFFFF5FFB2FF1BBCD91F037D6E" blockId="4.[132,775,1209,1393]" lastBlockId="4.[813,1456,826,1330]" pageId="4" pageNumber="57">Five vertical blackish bands in dorsal view. The first one on the nape, the second one below the first dorsal fin, the third and fourth below the second dorsal fin and the fifth band at hypural crease. These bands are not uniformly black but with dorsal or lateral greyish spots. Colour of body, on the upper part and between each band greyish with some pinkish to orange areas, particularly between the second and the third bands. On the inferior longitudinal part, the body is mottled black and grey with three aligned horizontal ovoid whitish to silvery spots on the posterior part. Head and cheeks mottled with closely spaced black or pink-orange spots and markings, and an overall shade of greyish-yellow. Sub-inferior part of head blackish. Ventral surface of head in male blackish or greyish in the medium part. Fraenum whitish. Ventral surface of head in female whitish. Fraenum whitish. Belly whitish to greyish. First and second dorsal fins with rows of black spots on rays. Caudal fin mainly translucent with two median white spots dorsally and ventrally, two black spots posteriorly and a large white spot at hypural base. Pectoral fins translucent with a medium black triangular wedge dorsally usually with 2 vertical rows of dark spots below it.</paragraph>
</subSubSection>
<subSubSection id="C3476564FFF5FFB2FC9CBD5A1E3C7D8E" pageId="4" pageNumber="57" type="biology_ecology">
<paragraph id="8BE236EFFFF5FFB2FC9CBD5A19FB7D2E" blockId="4.[813,900,1370,1394]" box="[813,900,1370,1394]" pageId="4" pageNumber="57">
<heading id="D0AA8183FFF5FFB2FC9CBD5A19FB7D2E" bold="true" box="[813,900,1370,1394]" fontSize="9" level="2" pageId="4" pageNumber="57" reason="3">
<emphasis id="B929EAFDFFF5FFB2FC9CBD5A19FB7D2E" bold="true" box="[813,900,1370,1394]" pageId="4" pageNumber="57">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8BE236EFFFF5FFB2FCE2BD7A1E3C7D8E" blockId="4.[813,1456,1402,1490]" pageId="4" pageNumber="57">
<taxonomicName id="4C5D4D6CFFF5FFB2FCE2BD7A1E147DCE" authorityName="Keith &amp; Delrieu-Trottin &amp; Utama &amp; Sauri &amp; Busson &amp; Nurjirana &amp; Wowor &amp; Dahruddin &amp; Hubert" authorityYear="2021" box="[851,1131,1402,1426]" family="Gobiidae" genus="Schismatogobius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="57" phylum="Chordata" rank="species" species="limmoni">
<emphasis id="B929EAFDFFF5FFB2FCE2BD7A1E147DCE" box="[851,1131,1402,1426]" italics="true" pageId="4" pageNumber="57">Schismatogobius limmoni</emphasis>
</taxonomicName>
has been collected in a freshwater stream with moderate to fast flow in shallow areas of boulders, gravel and sand.
</paragraph>
</subSubSection>
<subSubSection id="C3476564FFF5FFB2FC9CBDFA1FF57ECE" pageId="4" pageNumber="57" type="etymology">
<paragraph id="8BE236EFFFF5FFB2FC9CBDFA19D77E4E" blockId="4.[813,1455,1530,1682]" box="[813,936,1530,1554]" pageId="4" pageNumber="57">
<emphasis id="B929EAFDFFF5FFB2FC9CBDFA19D77E4E" bold="true" box="[813,936,1530,1554]" pageId="4" pageNumber="57">Etymology</emphasis>
</paragraph>
<paragraph id="8BE236EFFFF5FFB2FCE2BE1A1FF57ECE" blockId="4.[813,1455,1530,1682]" pageId="4" pageNumber="57">
The new species is named
<taxonomicName id="4C5D4D6CFFF5FFB2FBDEBE1A1EB87E6E" authorityName="Keith &amp; Delrieu-Trottin &amp; Utama &amp; Sauri &amp; Busson &amp; Nurjirana &amp; Wowor &amp; Dahruddin &amp; Hubert" authorityYear="2021" box="[1135,1223,1562,1586]" family="Gobiidae" genus="Schismatogobius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="57" phylum="Chordata" rank="species" species="limmoni">
<emphasis id="B929EAFDFFF5FFB2FBDEBE1A1EB87E6E" box="[1135,1223,1562,1586]" italics="true" pageId="4" pageNumber="57">limmoni</emphasis>
</taxonomicName>
in dedication to Gino Limmon, from Universitas Pattimura, for all his work to improve our knowledge on Indonesian fishes and its application for the conservation of the Indonesian aquatic fauna.
</paragraph>
</subSubSection>
<subSubSection id="C3476564FFF5FFB3FC9CBEBA18B17AED" lastPageId="5" lastPageNumber="58" pageId="4" pageNumber="57" type="diagnosis">
<paragraph id="8BE236EFFFF5FFB2FC9CBEBA19EA7E8E" blockId="4.[813,917,1722,1746]" box="[813,917,1722,1746]" pageId="4" pageNumber="57">
<heading id="D0AA8183FFF5FFB2FC9CBEBA19EA7E8E" bold="true" box="[813,917,1722,1746]" fontSize="9" level="2" pageId="4" pageNumber="57" reason="3">
<emphasis id="B929EAFDFFF5FFB2FC9CBEBA19EA7E8E" bold="true" box="[813,917,1722,1746]" pageId="4" pageNumber="57">Affinities</emphasis>
</heading>
</paragraph>
<paragraph id="8BE236EFFFF5FFB3FCE2BEDA18B17AED" blockId="4.[813,1456,1754,1906]" lastBlockId="5.[132,775,217,690]" lastPageId="5" lastPageNumber="58" pageId="4" pageNumber="57">
<taxonomicName id="4C5D4D6CFFF5FFB2FCE2BEDA1E077EAE" authorityName="Keith &amp; Delrieu-Trottin &amp; Utama &amp; Sauri &amp; Busson &amp; Nurjirana &amp; Wowor &amp; Dahruddin &amp; Hubert" authorityYear="2021" box="[851,1144,1754,1778]" family="Gobiidae" genus="Schismatogobius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="57" phylum="Chordata" rank="species" species="limmoni">
<emphasis id="B929EAFDFFF5FFB2FCE2BEDA1E077EAE" box="[851,1144,1754,1778]" italics="true" pageId="4" pageNumber="57">Schismatogobius limmoni</emphasis>
</taxonomicName>
differs from the other species sequenced and present in the area studied by displaying distinct mitochondrial lineages from its closest relatives and high TIM+I genetic distances to its relatives at
<emphasis id="B929EAFDFFF5FFB2FAF8BF3A1F087F0D" box="[1353,1399,1850,1873]" italics="true" pageId="4" pageNumber="57">COI</emphasis>
gene (0.6749 to its closest relative
<taxonomicName id="4C5D4D6CFFF5FFB2FBC6BF5A1F707F2E" authorityName="Keith, Darhuddin, Limmon &amp; Hubert" authorityYear="2018" box="[1143,1295,1882,1906]" family="Gobiidae" genus="Schismatogobius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="57" phylum="Chordata" rank="species" species="sapoliensis">
<emphasis id="B929EAFDFFF5FFB2FBC6BF5A1F707F2E" box="[1143,1295,1882,1906]" italics="true" pageId="4" pageNumber="57">S. sapoliensis</emphasis>
</taxonomicName>
). It is also the second only known species of the area studied (with
<taxonomicName id="4C5D4D6CFFF4FFB3FD03B8DA1AB5794D" authorityName="Keith, Darhuddin, Limmon &amp; Hubert" authorityYear="2018" family="Gobiidae" genus="Schismatogobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="58" phylum="Chordata" rank="species" species="sapoliensis">
<emphasis id="B929EAFDFFF4FFB3FD03B8DA1AB5794D" italics="true" pageId="5" pageNumber="58">S. sapoliensis</emphasis>
</taxonomicName>
) with 16-17 pectorals rays and having a broad black triangular wedge dorsally on pectoral fins. Indeed,
<emphasis id="B929EAFDFFF4FFB3FD0AB91A1B5D790D" italics="true" pageId="5" pageNumber="58">
<taxonomicName id="4C5D4D6CFFF4FFB3FD0AB91A1ADE790D" authorityName="Keith, Lord, Hadiaty &amp; Hubert" authorityYear="2017" family="Gobiidae" genus="Schismatogobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="58" phylum="Chordata" rank="species" species="saurii">S. saurii</taxonomicName>
,
<taxonomicName id="4C5D4D6CFFF4FFB3FF1EB93A1B5D790D" authorityName="Keith, Hubert, Limmon &amp; Darhuddin" authorityYear="2017" box="[175,290,313,337]" family="Gobiidae" genus="Schismatogobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="58" phylum="Chordata" rank="species" species="bussoni">S. bussoni</taxonomicName>
</emphasis>
and
<taxonomicName id="4C5D4D6CFFF4FFB3FEEBB93A1BAC790D" authorityName="de Beaufort" authorityYear="1912" box="[346,467,313,337]" family="Gobiidae" genus="Schismatogobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="58" phylum="Chordata" rank="species" species="bruynisi">
<emphasis id="B929EAFDFFF4FFB3FEEBB93A1BAC790D" box="[346,467,313,337]" italics="true" pageId="5" pageNumber="58">S. bruynisi</emphasis>
</taxonomicName>
have 14-15 pectorals rays,
<taxonomicName id="4C5D4D6CFFF4FFB3FF35B95A1B57792D" authorityName="Keith, Darhuddin, Sukmono &amp; Hubert" authorityYear="2017" box="[132,296,345,369]" family="Gobiidae" genus="Schismatogobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="58" phylum="Chordata" rank="species" species="risdawatiae">
<emphasis id="B929EAFDFFF4FFB3FF35B95A1B57792D" box="[132,296,345,369]" italics="true" pageId="5" pageNumber="58">S. risdawatiae</emphasis>
</taxonomicName>
has 15-16 pectorals rays, and
<taxonomicName id="4C5D4D6CFFF4FFB3FD3BB95A1978792D" baseAuthorityName="Herre" baseAuthorityYear="1927" box="[650,775,346,369]" family="Gobiidae" genus="Schismatogobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="58" phylum="Chordata" rank="species" species="insignus">
<emphasis id="B929EAFDFFF4FFB3FD3BB95A1978792D" box="[650,775,346,369]" italics="true" pageId="5" pageNumber="58">S. insignus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5D4D6CFFF4FFB3FF03B97A1B4879CD" authorityName="Keith, Lord &amp; Hubert" authorityYear="2017" box="[178,311,377,401]" family="Gobiidae" genus="Schismatogobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="58" phylum="Chordata" rank="species" species="arscuttoli">
<emphasis id="B929EAFDFFF4FFB3FF03B97A1B4879CD" box="[178,311,377,401]" italics="true" pageId="5" pageNumber="58">S. arscuttoli</emphasis>
</taxonomicName>
have 16-17 pectorals rays but the pectoral fins are striped with rows of dark spots (
<bibRefCitation id="EFCC4B1EFFF4FFB3FD88B999197C79ED" author="KEITH P. &amp; LORD C. &amp; DARHUDDIN H. &amp; LIMMON G. &amp; SUKMONO T. &amp; HADIATY R. &amp; HUBERT N." box="[569,771,409,433]" pageId="5" pageNumber="58" pagination="195 - 211" refId="ref4664" refString="KEITH P., LORD C., DARHUDDIN H., LIMMON G., SUKMONO T., HADIATY R. &amp; HUBERT N., 2017 b. - Schismatogobius (Gobiidae) from Indonesia, with description of four new species. Cybium, 41 (2): 195 - 211. DOI: 0.26028 / cybium / 2017 - 412 - 010" type="journal article" year="2017">
Keith
<emphasis id="B929EAFDFFF4FFB3FDCEB99A18CA79ED" box="[639,693,409,433]" italics="true" pageId="5" pageNumber="58">et al.</emphasis>
2017b
</bibRefCitation>
,
<bibRefCitation id="EFCC4B1EFFF4FFB3FF35B9B91AC0798D" author="KEITH P. &amp; DAHRUDDIN H. &amp; LIMMON G. &amp; HUBERT N." box="[132,191,441,465]" pageId="5" pageNumber="58" pagination="195 - 200" refId="ref4732" refString="KEITH P., DAHRUDDIN H., LIMMON G. &amp; HUBERT N., 2018. - A new species of Schismatogobius (Teleostei: Gobiidae) from Halmahera (Indonesia). Cybium, 42 (2): 195 - 200. DOI: 10.26028 / cybium / 2018 - 422 - 007" type="journal article" year="2018">2018</bibRefCitation>
). It differs from
<taxonomicName id="4C5D4D6CFFF4FFB3FEC3B9BA1879798D" authorityName="Keith, Darhuddin, Limmon &amp; Hubert" authorityYear="2018" box="[370,518,441,465]" family="Gobiidae" genus="Schismatogobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="58" phylum="Chordata" rank="species" species="sapoliensis">
<emphasis id="B929EAFDFFF4FFB3FEC3B9BA1879798D" box="[370,518,441,465]" italics="true" pageId="5" pageNumber="58">S. sapoliensis</emphasis>
</taxonomicName>
in having a greater jaw length/head length ratio in female (36.5-39%
<emphasis id="B929EAFDFFF4FFB3FDC1B9DA18F979AD" box="[624,646,474,497]" italics="true" pageId="5" pageNumber="58">vs</emphasis>
32-36.7%), a greater jaw length in female (10.5-11.7%
<emphasis id="B929EAFDFFF4FFB3FDE8B9FA18107A4D" box="[601,623,506,529]" italics="true" pageId="5" pageNumber="58">vs</emphasis>
9.1-11% SL), pectoral fins (usually 17 rays
<emphasis id="B929EAFDFFF4FFB3FE77BA1A1BA37A6D" box="[454,476,538,561]" italics="true" pageId="5" pageNumber="58">vs</emphasis>
usually 16) with a medium black triangular wedge dorsally usually with 2 vertical rows of dark spots below it
<emphasis id="B929EAFDFFF4FFB3FEC0BA5A1BCA7A2D" box="[369,437,602,625]" italics="true" pageId="5" pageNumber="58">versus</emphasis>
pectoral fins with a large transverse medium black band crossing usually all the fin, and a greater preanal length (62.9-68.4
<emphasis id="B929EAFDFFF4FFB3FE57BA9A1B837AED" box="[486,508,666,689]" italics="true" pageId="5" pageNumber="58">vs</emphasis>
61.4-62.8) (
<tableCitation id="C6DF0354FFF4FFB3FD30BA9918C17AED" box="[641,702,665,689]" captionStart="Table I" captionStartId="2.[813,868,217,239]" captionTargetId="graphics-749@2.[906,1361,436,1101]" captionTargetPageId="2" captionText="Table I. Morphomeristics of the new Schismatogobius species. Morphometrics are given as percentages of standard length, except JL/HL. P, Pectoral rays; D, Dorsal rays; A, Anal rays; C, Caudal branched rays; PDL, Predorsal length (% SL); PAL, Preanal length (% SL); HL, Head length (% SL); JL, jaw length (% SL); CPD, Caudal peduncle depth (% SL); Pect-L, Pectoral fin length (% SL); BDa, Body depth at anus (% SL); SDFL, Second dorsal fin length (% SL); AFL, Anal fin length (% SL); CFL, Caudal fin length (% SL)." httpUri="http://table.plazi.org/id/DF226667FFF3FFB4FC9CB8D9190679F3" pageId="5" pageNumber="58" tableUuid="DF226667FFF3FFB4FC9CB8D9190679F3">Tab. I</tableCitation>
).
</paragraph>
</subSubSection>
<caption id="DF226667FFF5FFB2FF35BF451BFE7F2E" ID-DOI="http://doi.org/10.5281/zenodo.10493564" ID-Zenodo-Dep="10493564" httpUri="https://zenodo.org/record/10493564/files/figure.png" pageId="4" pageNumber="57" startId="4.[132,198,1861,1883]" targetBox="[140,767,1435,1838]" targetPageId="4" targetType="figure">
<paragraph id="8BE236EFFFF5FFB2FF35BF451BFE7F2E" blockId="4.[132,775,1861,1906]" pageId="4" pageNumber="57">
Figure 2. Schematic drawing of the ventral surface of head of
<taxonomicName id="4C5D4D6CFFF5FFB2FF35BF5D1B047F2E" authorityName="Keith &amp; Delrieu-Trottin &amp; Utama &amp; Sauri &amp; Busson &amp; Nurjirana &amp; Wowor &amp; Dahruddin &amp; Hubert" authorityYear="2021" box="[132,379,1885,1906]" family="Gobiidae" genus="Schismatogobius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="57" phylum="Chordata" rank="species" species="limmoni">
<emphasis id="B929EAFDFFF5FFB2FF35BF5D1B047F2E" box="[132,379,1885,1906]" italics="true" pageId="4" pageNumber="57">Schismatogobius limmoni</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C3476564FFF4FFB3FF35BAD918A27B4D" box="[132,733,729,785]" pageId="5" pageNumber="58" type="distribution">
<paragraph id="8BE236EFFFF4FFB3FF35BAD918A27B4D" blockId="5.[132,733,729,785]" box="[132,733,729,785]" pageId="5" pageNumber="58">
<emphasis id="B929EAFDFFF4FFB3FF35BAD91B707AAD" bold="true" box="[132,271,729,753]" pageId="5" pageNumber="58">Distribution</emphasis>
This species is known only from
<collectingRegion id="4999F80DFFF4FFB3FDB9BAF918147B4D" box="[520,619,761,785]" country="Indonesia" name="Sulawesi Utara" pageId="5" pageNumber="58">Sulawesi</collectingRegion>
,
<collectingCountry id="F34A767FFFF4FFB3FDC4BAF918A27B4D" box="[629,733,761,785]" name="Indonesia" pageId="5" pageNumber="58">Indonesia</collectingCountry>
</paragraph>
</subSubSection>
<subSubSection id="C3476564FFF4FFB3FF35BB3918627B2D" pageId="5" pageNumber="58" type="materials_examined">
<paragraph id="8BE236EFFFF4FFB3FF35BB3918627B2D" blockId="5.[132,541,825,881]" pageId="5" pageNumber="58">
<emphasis id="B929EAFDFFF4FFB3FF35BB391BFF7B0D" bold="true" box="[132,384,825,849]" pageId="5" pageNumber="58">Comparative material</emphasis>
See
<bibRefCitation id="EFCC4B1EFFF4FFB3FF66BB591BE87B2D" author="KEITH P. &amp; LORD C. &amp; LARSON H. K." box="[215,407,857,881]" pageId="5" pageNumber="58" pagination="45 - 66" refId="ref4602" refString="KEITH P., LORD C. &amp; LARSON H. K., 2017 a. - Review of Schismatogobius (Teleostei: Gobiidae) from Papua New Guinea to Samoa, with description of seven new species. Cybium, 41 (1): 45 - 66. DOI: 10.26028 / cybium / 2017 - 411 - 005" type="journal article" year="2017">
Keith
<emphasis id="B929EAFDFFF4FFB3FEA9BB5A1B327B2D" box="[280,333,857,881]" italics="true" pageId="5" pageNumber="58">et al.</emphasis>
2017a
</bibRefCitation>
, b and 2018.
</paragraph>
</subSubSection>
</treatment>
</document>