545 lines
69 KiB
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545 lines
69 KiB
XML
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<mods:title id="B883D4FCF3174CB750C2804B5909BDEF">Revision of the Sundaland species of the genus Dysphaea Selys, 1853 using molecular and morphological methods, with notes on allied species (Odonata: Euphaeidae)</mods:title>
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<mods:namePart id="BE129309BCF4D3FED4CAAD461F224A8A">Dow, Rory A.</mods:namePart>
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<mods:namePart id="5C7C15195FCC281453BB04983289AD08">Stokvis, Frank R.</mods:namePart>
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<mods:title id="C0BB4758FD62ABABC5215BD457120B6D">Zootaxa</mods:title>
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<subSubSection id="C3476578FFC5FFF6FF780BA649289FE9" pageId="17" pageNumber="468" type="nomenclature">
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<paragraph id="8BE236F3FFC5FFF6FF780BA649BE9FAF" blockId="17.[151,501,151,178]" box="[151,501,151,178]" pageId="17" pageNumber="468">
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<heading id="D0AA819FFFC5FFF6FF780BA649BE9FAF" bold="true" box="[151,501,151,178]" fontSize="11" level="1" pageId="17" pageNumber="468" reason="1">
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<taxonomicName id="4C5D4D70FFC5FFF6FF780BA649BE9FAF" ID-CoL="6DSKW" authority="Selys, 1873" authorityName="Selys" authorityYear="1873" box="[151,501,151,178]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="lugens">
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<emphasis id="B929EAE1FFC5FFF6FF780BA649BE9FAF" bold="true" box="[151,501,151,178]" pageId="17" pageNumber="468">
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<emphasis id="B929EAE1FFC5FFF6FF780BA6492E9FAC" bold="true" box="[151,357,151,177]" italics="true" pageId="17" pageNumber="468">Dysphaea lugens</emphasis>
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<bibRefCitation id="EFCC4B02FFC5FFF6FE830BA949BE9FAF" author="Selys" box="[364,501,152,178]" pageId="17" pageNumber="468" refString="Selys Longchamps, E. de (1873) Troisiemes additions au synopsis de Calopterygines. Bulletins de l'Academie royale des sciences, des lettres et des beaux-arts de Belgique, Series 2, 35, 469 - 519." type="journal article" year="1873">Selys, 1873</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</paragraph>
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<paragraph id="8BE236F3FFC5FFF6FF780BEF49289FE9" blockId="17.[151,1436,222,520]" box="[151,355,222,244]" pageId="17" pageNumber="468">Selected references:</paragraph>
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<subSubSection id="C3476578FFC5FFF6FF780BCC4A1E9D15" pageId="17" pageNumber="468" type="reference_group">
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<paragraph id="8BE236F3FFC5FFF6FF780BCC4A5D9EB1" blockId="17.[151,1436,222,520]" pageId="17" pageNumber="468">
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<taxonomicName id="4C5D4D70FFC5FFF6FF780BCC49859E0E" ID-CoL="6DSKW" authority="Selys, 1873" authorityName="Selys" authorityYear="1873" box="[151,462,253,275]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="lugens">
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<emphasis id="B929EAE1FFC5FFF6FF780BCC49039E0E" box="[151,328,253,275]" italics="true" pageId="17" pageNumber="468">Dysphaea lugens</emphasis>
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<bibRefCitation id="EFCC4B02FFC5FFF6FEBC0BCC49859E0E" author="Selys" box="[339,462,253,275]" pageId="17" pageNumber="468" refString="Selys Longchamps, E. de (1873) Troisiemes additions au synopsis de Calopterygines. Bulletins de l'Academie royale des sciences, des lettres et des beaux-arts de Belgique, Series 2, 35, 469 - 519." type="journal article" year="1873">Selys, 1873</bibRefCitation>
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</taxonomicName>
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:—
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<bibRefCitation id="EFCC4B02FFC5FFF6FE030BCC4AB09E0E" author="Selys" box="[492,763,253,275]" pageId="17" pageNumber="468" refString="Selys Longchamps, E. de (1873) Troisiemes additions au synopsis de Calopterygines. Bulletins de l'Academie royale des sciences, des lettres et des beaux-arts de Belgique, Series 2, 35, 469 - 519." type="journal article" year="1873">
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Selys
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<bibRefCitation id="EFCC4B02FFC5FFF6FDC00BCC4AB09E0E" author="Longchamps" box="[559,763,253,275]" pageId="17" pageNumber="468" refString="Selys Longchamps, E. de (1873) Troisiemes additions au synopsis de Calopterygines. Bulletins de l'Academie royale des sciences, des lettres et des beaux-arts de Belgique, Series 2, 35, 469 - 519." type="journal article" year="1873" yearSuffix="p">Longchamps (1873</bibRefCitation>
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</bibRefCitation>
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: 485–486, reprint p. 21 (description of male, South Borneo, as ‘
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<taxonomicName id="4C5D4D70FFC5FFF6FF200A2D49A39E2F" ID-CoL="6DSKW" authority="De Selys" authorityName="De Selys" box="[207,488,284,306]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="lugens">
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<emphasis id="B929EAE1FFC5FFF6FF200A2D49C89E2F" box="[207,387,284,306]" italics="true" pageId="17" pageNumber="468">Dysphaea lugens,</emphasis>
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De Selys
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</taxonomicName>
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; Race de
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<taxonomicName id="4C5D4D70FFC5FFF6FDA40A2D4AE79E2F" ID-CoL="6DSLB" box="[587,684,284,306]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="dimidiata">
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<emphasis id="B929EAE1FFC5FFF6FDA40A2D4AE79E2F" box="[587,684,284,306]" italics="true" pageId="17" pageNumber="468">dimidiata</emphasis>
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</taxonomicName>
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?’);—
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<bibRefCitation id="EFCC4B02FFC5FFF6FD070A2D4D539E2F" author="Ris" box="[744,1304,284,306]" pageId="17" pageNumber="468" refString="Ris, F. (1911) Libellen von Sintang, Borneo gesammelt von Dr L. Martin. Annales de la Societe Entomologique de Belgique, 55, 231 - 255." type="journal article" year="1911">Ris (1911: 232–233; Sintang, Borneo; description of ♀)</bibRefCitation>
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;—
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<bibRefCitation id="EFCC4B02FFC5FFF6FAD70A2D4A629E4D" author="Kennedy" pageId="17" pageNumber="468" refString="Kennedy, C. H. (1920) The phylogeny of the zygopterous dragonflies as based on the evidence of the penes. The Ohio Journal of Science, 21, 19 - 29, pls. 1 - 3 excl." type="journal article" year="1920">
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Kennedy (1920: pl. 1,
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<figureCitation id="13662A76FFC5FFF6FEB40A0B49909E4D" box="[347,475,314,336]" captionStart="FIGURES 31 – 36" captionStartId="19.[151,264,1705,1728]" captionTargetBox="[157,1436,185,1683]" captionTargetId="figure@19.[157,1442,185,1684]" captionTargetPageId="19" captionText="FIGURES 31 – 36. Male anal appendages. Dorsal view: 31) D. dimidiata West Java; 32) D. lugens Bakuan, Kalimantan Barat; 33) D. ulu holotype; 34) D. vanida holotype. Dorso-lateral views: 35) D. dimidiata Terengganu; 36) D. vanida paratype Lam Khlong Gnu, Kanchanaburi." httpUri="https://zenodo.org/record/243470/files/figure.png" pageId="17" pageNumber="468">Figs. 32–33</figureCitation>
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, penis)
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</bibRefCitation>
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;—
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<bibRefCitation id="EFCC4B02FFC5FFF6FDA80A0B4C469E4D" author="Laidlaw" box="[583,1037,314,336]" pageId="17" pageNumber="468" refString="Laidlaw, F. F. (1924) A survey of the dragonfly fauna of the Malaya peninsula, with notes on that of neighbouring countries. Part II. Journal of the Malayan Branch of the Royal Asiatic Society, 2 (3), 296 - 308, pls. 3 - 5 excl." type="journal article" year="1924">Laidlaw (1924: 300, characters discussed)</bibRefCitation>
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;—Laidlaw (1931: 241, West Central Borneo);—Coomans de
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<bibRefCitation id="EFCC4B02FFC5FFF6FE2D0A684BE39E72" author="Ruiter" box="[450,936,345,367]" pageId="17" pageNumber="468" refString="Coomans de Ruiter, L. (1936). Biologische waarnemingen over beekjuffers van Borneo (Een groep van mooi gekleurde libellen). De tropische natuur, 25 (5), 69 - 78, 1 col. pl. excl." type="journal article" year="1936">
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Ruiter (1936: 74–76,
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<figureCitation id="13662A76FFC5FFF6FD4C0A684AAA9E72" box="[675,737,345,367]" captionStart="FIGURE 3" captionStartId="6.[151,250,1699,1721]" captionText="FIGURE 3. Phylogenetic reconstruction for 17 specimens of Dysphaea plus five outgroup taxa from the combined COI + 16 S + 28 S data from Bayesian Inference analysis. Posterior probability values are shown (as percentages) if less than 100 %. RMNH collection codes are shown for each specimen, with the RMNH. INS. prefix omitted." pageId="17" pageNumber="468">Fig. 3</figureCitation>
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, W. Borneo, notes)
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</bibRefCitation>
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;—
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<bibRefCitation id="EFCC4B02FFC5FFF6FC290A684D569E72" author="Lieftinck" box="[966,1309,345,367]" pageId="17" pageNumber="468" refString="Lieftinck, M. A. (1953) New dragonflies (Odonata) from Borneo, with notes on their habits and larvae. Treubia, 22 (2), 381 - 406." type="journal article" year="1953">Lieftinck (1953: 382, S. Borneo)</bibRefCitation>
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;—
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<bibRefCitation id="EFCC4B02FFC5FFF6FAD40A684AC29E93" author="Lieftinck" pageId="17" pageNumber="468" refString="Lieftinck, M. A. (1954) Handlist of Malaysian Odonata. A catalogue of the dragonflies of the Malay Peninsula, Sumatra, Java and Borneo, including the adjacent small islands. Treubia, 22 (Supplement), i - xiii + 1 - 202, 1 folded map excl." type="journal article" year="1954">Lieftinck (1954: 20, distribution, habitat, references)</bibRefCitation>
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;—
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<bibRefCitation id="EFCC4B02FFC5FFF6FD490A49496C9EB1" author="Asahina" pageId="17" pageNumber="468" refString="Asahina, S. (1985) A list of the Odonata recorded from Thailand. Part XI. Euphaeidae. Cho Cho, 8 (12), 18 - 38." type="journal article" year="1985">
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Asahina (1985: 31, 33, 34, 36,
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<figureCitation id="13662A76FFC5FFF6FC190A494C3A9E93" box="[1014,1137,376,398]" captionStart-0="FIGURES 45 – 48" captionStart-1="FIGURES 49 – 52" captionStartId-0="23.[151,264,1735,1758]" captionStartId-1="24.[151,264,1070,1093]" captionTargetBox-0="[162,1365,783,1707]" captionTargetBox-1="[151,1436,193,1049]" captionTargetId-0="figure@23.[151,1436,721,1720]" captionTargetId-1="figure@24.[151,1436,193,1049]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionText-0="FIGURES 45 – 48. Penis, lateral view: 45) D. dimidiata west Java; 46) D. lugens Selekau River, Kalimantan Barat; 47) D. ulu holotype; 48) D. vanida paratype, Khlong Nakha, Ranong." captionText-1="FIGURES 49 – 52. Habitus of female Dysphaea: 49) D. dimidiata west Java; 50) D. lugens Mount Dulit, Sarawak, photograph by Benjamin Price; 51) D. ulu paratype; 52) D. vanida paratype Kanchanaburi." httpUri-0="https://zenodo.org/record/243473/files/figure.png" httpUri-1="https://zenodo.org/record/243474/files/figure.png" pageId="17" pageNumber="468">Figs. 48–50</figureCitation>
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penis,
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<figureCitation id="13662A76FFC5FFF6FB2F0A494D479E93" box="[1216,1292,376,398]" captionStart="FIGURES 63 – 68" captionStartId="28.[151,264,1699,1722]" captionTargetBox="[151,1436,185,1683]" captionTargetId="figure@28.[151,1436,185,1684]" captionTargetPageId="28" captionText="FIGURES 63 – 68. Female synthorax, dorsal view: 63) D. dimidiata west Java; 64) D. dimidiata Sarawak; 65) D. dimidiata Johor; 66) D. lugens Gunung Sari, Kaliamantan Timur; 67) D. ulu paratype; 68) D. vanida paratype Kanchanaburi." httpUri="https://zenodo.org/record/243477/files/figure.png" pageId="17" pageNumber="468">Fig. 68</figureCitation>
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(male wings,
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<collectingCountry id="F34A7663FFC5FFF6FF280AA7496B9EB1" box="[199,288,406,428]" name="Malaysia" pageId="17" pageNumber="468">Sarawak</collectingCountry>
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)
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</bibRefCitation>
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;—
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<bibRefCitation id="EFCC4B02FFC5FFF6FEAA0AA74A5A9EB1" author="Orr" box="[325,529,406,428]" pageId="17" pageNumber="468" refString="Orr, A. G. (2003) A guide to the dragonflies of Borneo: their identification and biology. Natural History Publications (Borneo), Kota Kinabalu, x + 195 pp." type="book" year="2003">Orr (2003: 58, part)</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8BE236F3FFC5FFF6FF780A844BFD9ED6" blockId="17.[151,1436,222,520]" box="[151,950,437,459]" pageId="17" pageNumber="468">
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<taxonomicName id="4C5D4D70FFC5FFF6FF780A8449289ED6" ID-CoL="6DSLB" box="[151,355,437,459]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="dimidiata">
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<emphasis id="B929EAE1FFC5FFF6FF780A8449289ED6" box="[151,355,437,459]" italics="true" pageId="17" pageNumber="468">Dysphaea dimidiata</emphasis>
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</taxonomicName>
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[nec
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<bibRefCitation id="EFCC4B02FFC5FFF6FE740A844A599ED6" author="Selys" box="[411,530,437,459]" pageId="17" pageNumber="468" refString="Selys Longchamps, E. de (1853) Synopsis des Calopterygines. Bulletins de l'Academie royale des sciences, des lettres et des beaux-arts de Belgique, 20 (Annexe), 1 - 73." type="journal article" year="1853">Selys, 1853</bibRefCitation>
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]:—
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<bibRefCitation id="EFCC4B02FFC5FFF6FDD80A844BF99ED6" author="Orr" box="[567,946,437,459]" pageId="17" pageNumber="468" refString="Orr, A. G. (2002) Notes on the Rhinocypha cucullata Selys group from Borneo, with a description of R. viola spec. nov. (Zygoptera: Chlorocyphidae). Odonatologica, 31 (3), 287 - 295." type="journal article" year="2002">Orr (2002: 291, Central Kalimantan)</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8BE236F3FFC5FFF6FF780AE54A1E9D15" blockId="17.[151,1436,222,520]" pageId="17" pageNumber="468">
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<taxonomicName id="4C5D4D70FFC5FFF6FF780AE5491A9EF7" ID-CoL="6DSLF" box="[151,337,468,490]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="limbata">
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<emphasis id="B929EAE1FFC5FFF6FF780AE5491A9EF7" box="[151,337,468,490]" italics="true" pageId="17" pageNumber="468">Dysphaea limbata</emphasis>
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</taxonomicName>
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‘race
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<taxonomicName id="4C5D4D70FFC5FFF6FE7B0AE54A5B9EF7" ID-CoL="6DSL8" box="[404,528,468,490]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="semilimbata">
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<emphasis id="B929EAE1FFC5FFF6FE7B0AE54A5B9EF7" box="[404,528,468,490]" italics="true" pageId="17" pageNumber="468">semilimbata</emphasis>
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</taxonomicName>
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’ [nec
|
||
<bibRefCitation id="EFCC4B02FFC5FFF6FDBC0AE54A819EF7" author="Selys" box="[595,714,468,490]" pageId="17" pageNumber="468" refString="Selys Longchamps, E. de (1873) Troisiemes additions au synopsis de Calopterygines. Bulletins de l'Academie royale des sciences, des lettres et des beaux-arts de Belgique, Series 2, 35, 469 - 519." type="journal article" year="1873">Selys, 1873</bibRefCitation>
|
||
]:—
|
||
<bibRefCitation id="EFCC4B02FFC5FFF6FD010AE54A1A9D15" author="Kimmins" pageId="17" pageNumber="468" refString="Kimmins, D. E. (1936) The Odonata of the Oxford University Sarawak expedition. Journal of the Federated Malay States Museum, 18, 65 - 108." type="journal article" year="1936">Kimmins (1936: 78, specimens from River Kapah and junction of rivers Tinjar and Lejok; Mt Dulit area)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3476578FFC5FFF6FF78091E4CAA99E8" pageId="17" pageNumber="468" type="materials_examined">
|
||
<paragraph id="8BE236F3FFC5FFF6FF78091E4A9F9B28" blockId="17.[151,1437,559,2033]" pageId="17" pageNumber="468">
|
||
<emphasis id="B929EAE1FFC5FFF6FF78091E4A1E9D55" bold="true" box="[151,597,559,584]" pageId="17" pageNumber="468">
|
||
Study of the
|
||
<typeStatus id="54E68851FFC5FFF6FEDE091E492E9D55" box="[305,357,559,584]" pageId="17" pageNumber="468">type</typeStatus>
|
||
material.
|
||
<typeStatus id="54E68851FFC5FFF6FE0E091E4A1B9D55" box="[481,592,559,584]" pageId="17" pageNumber="468" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName id="4C5D4D70FFC5FFF6FDB209004A819D55" box="[605,714,560,584]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="lugens">
|
||
<emphasis id="B929EAE1FFC5FFF6FDB209004A819D55" box="[605,714,560,584]" italics="true" pageId="17" pageNumber="468">D. lugens</emphasis>
|
||
</taxonomicName>
|
||
was described from a single male specimen, which was sent to Selys by Robert McLachlan. Coll. Selys at
|
||
<collectionCode id="ED4CAE36FFC5FFF6FD7F09624A9B9D76" box="[656,720,595,619]" httpUri="http://grbio.org/cool/wzbt-fxwr" name="Institut Royal des Sciences Naturelles de Belgique" pageId="17" pageNumber="468">IRSN</collectionCode>
|
||
(Brussels) includes nine pinned male specimens of
|
||
<taxonomicName id="4C5D4D70FFC5FFF6FAC209654DD79D76" box="[1325,1436,595,619]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="lugens">
|
||
<emphasis id="B929EAE1FFC5FFF6FAC209654DD79D76" box="[1325,1436,595,619]" italics="true" pageId="17" pageNumber="468">D. lugens</emphasis>
|
||
</taxonomicName>
|
||
from Borneo (see below). Only one of them (
|
||
<figureCitation id="13662A76FFC5FFF6FD7709474A979D92" box="[664,732,630,655]" captionStart="FIGURES 5 – 8" captionStartId="13.[151,264,1156,1179]" captionTargetBox="[151,1436,193,1133]" captionTargetId="figure@13.[151,1436,193,1135]" captionTargetPageId="13" captionText="FIGURES 5 – 8. Selysian type specimens and labels: 5) D. dimidiata lectotype; 6) D. limbata lectotype; 7) D. lugens holotype; 8) D. semilimbata holotype." httpUri="https://zenodo.org/record/243465/files/figure.png" pageId="17" pageNumber="468">Fig. 7</figureCitation>
|
||
) has the same measurements given in the original description, i.e. abdomen (incl. cerci)
|
||
<quantity id="4CA59B16FFC5FFF6FE5309AB4A459DAC" box="[444,526,666,690]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="17" pageNumber="468" unit="mm" value="40.0">40 mm</quantity>
|
||
, Hw 31. It is also the only specimen with the cerci in a crossed position, which matches the single illustration of
|
||
<taxonomicName id="4C5D4D70FFC5FFF6FDE4098C4A3D9DC8" box="[523,630,701,725]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="lugens">
|
||
<emphasis id="B929EAE1FFC5FFF6FDE4098C4A3D9DC8" box="[523,630,701,725]" italics="true" pageId="17" pageNumber="468">D. lugens</emphasis>
|
||
</taxonomicName>
|
||
in the portfolio of coloured paintings of
|
||
<taxonomicName id="4C5D4D70FFC5FFF6FBAF098D4CD49DC8" box="[1088,1183,700,725]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="468" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
|
||
species in Coll. Selys, executed in the late 1800’s. In
|
||
<date id="FFE31033FFC5FFF6FE0509EE4AEA9DE5" box="[490,673,735,760]" pageId="17" pageNumber="468" value="2014-09">September 2014</date>
|
||
only a single small yellow handwritten label ‘Lansbg.’ was attached to this specimen. ‘Lansbg.’ refers to Johan Wilhelm van Lansberge (
|
||
<date id="FFE31033FFC5FFF6FC4D08324C6E9C06" box="[930,1061,771,795]" pageId="17" pageNumber="468" value="1830" valueMax="1903">1830–1903</date>
|
||
), who resided in
|
||
<collectingCountry id="F34A7663FFC5FFF6FB0508324DD79C01" box="[1258,1436,771,796]" name="Netherlands" pageId="17" pageNumber="468">the Netherlands</collectingCountry>
|
||
East Indies from
|
||
<date id="FFE31033FFC5FFF6FEB6081749BA9C22" box="[345,497,806,831]" pageId="17" pageNumber="468" value="1875" valueMax="1881">1875 to 1881</date>
|
||
. He could not have been the collector of the
|
||
<typeStatus id="54E68851FFC5FFF6FC0308174C059C22" box="[1004,1102,806,831]" pageId="17" pageNumber="468" type="holotype">holotype</typeStatus>
|
||
of species described in 1873. No doubt a mix-up in the labels had taken place at some phase after Selys’ time. This is evident also by the fact that the only identification label in Selys’ handwriting was attached to a specimen which does not match the measurements of the
|
||
<typeStatus id="54E68851FFC5FFF6FE6508BE49BB9CB5" box="[394,496,911,936]" pageId="17" pageNumber="468" type="holotype">holotype</typeStatus>
|
||
. This label reads: “
|
||
<taxonomicName id="4C5D4D70FFC5FFF6FD2208A14B769CB5" box="[717,829,912,936]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="468" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B929EAE1FFC5FFF6FD2208A14B769CB5" box="[717,829,912,936]" italics="true" pageId="17" pageNumber="468">Dysphaea</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="4C5D4D70FFC5FFF6FCA908A14C4D9CB5" authority="Selys" authorityName="Selys" box="[838,1030,911,936]" pageId="17" pageNumber="468" rank="variety" variety="lugens">
|
||
var.
|
||
<emphasis id="B929EAE1FFC5FFF6FC9708A14B8A9CB5" box="[888,961,912,936]" italics="true" pageId="17" pageNumber="468">lugens</emphasis>
|
||
Selys
|
||
</taxonomicName>
|
||
/ ♂ / Sintang”. Later someone (not Selys) has crossed out the locality name ‘Sintang’ and replaced it with ‘W.K.’, which means West-Kust (an area presently known as West Kalimantan). We have removed the wrong collector label and restored the Selysian identification label to the
|
||
<typeStatus id="54E68851FFC5FFF6FE5608C84A559B0F" box="[441,542,1017,1042]" pageId="17" pageNumber="468" type="holotype">holotype</typeStatus>
|
||
, which obviously was collected in Sintang in West Kalimantan, although in the original description reads only ‘Le sud de Bornéo’.
|
||
</paragraph>
|
||
<paragraph id="8BE236F3FFC5FFF6FF280F0E4B859975" blockId="17.[151,1437,559,2033]" pageId="17" pageNumber="468">
|
||
<emphasis id="B929EAE1FFC5FFF6FF280F0E4A629B45" bold="true" box="[199,553,1087,1112]" pageId="17" pageNumber="468">
|
||
Other material studied.
|
||
<collectionCode id="ED4CAE36FFC5FFF6FE090F0E4A629B45" box="[486,553,1087,1112]" httpUri="http://grbio.org/cool/wzbt-fxwr" name="Institut Royal des Sciences Naturelles de Belgique" pageId="17" pageNumber="468">IRSN</collectionCode>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="9D5BFD7AFFC5FFF6FDD80F714A289B4A" box="[567,611,1088,1112]" pageId="17" pageNumber="468" type="generic">2 ♂</specimenCount>
|
||
, Borneo W.[est-]K[ust]., Sintang, Coll. Clément;
|
||
<specimenCount id="9D5BFD7AFFC5FFF6FB630F714CF39B4A" box="[1164,1208,1088,1112]" pageId="17" pageNumber="468" type="generic">2 ♂</specimenCount>
|
||
, Borneo W.K., Coll. Clément;
|
||
<specimenCount id="9D5BFD7AFFC5FFF6FEE80F52497E9B66" box="[263,309,1123,1147]" pageId="17" pageNumber="468" type="generic">1 ♂</specimenCount>
|
||
, Borneo, N. Bartan, Coll. Clément;
|
||
<specimenCount id="9D5BFD7AFFC5FFF6FD390F524B4F9B66" box="[726,772,1123,1147]" pageId="17" pageNumber="468" type="generic">2 ♂</specimenCount>
|
||
, [Borneo], leg. J. W. van Lansberge;
|
||
<specimenCount id="9D5BFD7AFFC5FFF6FB5B0F524CA09B66" box="[1204,1259,1123,1147]" pageId="17" pageNumber="468" type="generic">1 ♂</specimenCount>
|
||
, N. Borneo,
|
||
<taxonomicName id="4C5D4D70FFC5FFF6FA6E0F5548A09B82" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="limbata">D. limbata</taxonomicName>
|
||
S. [nec
|
||
<bibRefCitation id="EFCC4B02FFC5FFF6FEBD0FB7499F9B82" author="Selys" box="[338,468,1158,1183]" pageId="17" pageNumber="468" refString="Selys Longchamps, E. de (1859) Additions au synopsis des Calopterygines. Bulletins de l'Academie royale des sciences, des lettres et des beaux-arts de Belgique, Series 2, 7, 437 - 451." type="journal article" year="1859">Selys 1859</bibRefCitation>
|
||
].—
|
||
<collectionCode id="ED4CAE36FFC5FFF6FE110FB74A1C9B82" LSID="urn:lsid:biocol.org:col:34992" box="[510,599,1158,1183]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="17" pageNumber="468">
|
||
<emphasis id="B929EAE1FFC5FFF6FE110FB74A1C9B82" bold="true" box="[510,599,1158,1183]" pageId="17" pageNumber="468">RMNH</emphasis>
|
||
</collectionCode>
|
||
:
|
||
<specimenCount id="9D5BFD7AFFC5FFF6FD860FB64AD29B83" box="[617,665,1159,1183]" pageId="17" pageNumber="468" type="generic">2 ♂</specimenCount>
|
||
, S. Borneo, Sampit (
|
||
<quantity id="4CA59B16FFC5FFF6FC7F0FB74BA59B82" box="[912,1006,1158,1183]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" pageId="17" pageNumber="468" unit="m" value="25.0" valueMax="50.0" valueMin="0.0">0–50 m</quantity>
|
||
), Pemantang,
|
||
<date id="FFE31033FFC5FFF6FB7B0FB64D559B82" box="[1172,1310,1158,1183]" pageId="17" pageNumber="468" value="1953-07-28">28 vii 1953</date>
|
||
, leg. M.A. Lieftinck;
|
||
<specimenCount id="9D5BFD7AFFC5FFF6FEE30F9B49739BDC" box="[268,312,1194,1218]" pageId="17" pageNumber="468" type="generic">3 ♂</specimenCount>
|
||
, S.E. Borneo, Ampah (
|
||
<quantity id="4CA59B16FFC5FFF6FDD40F9B4ADE9BDF" box="[571,661,1194,1218]" metricMagnitude="1" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" pageId="17" pageNumber="468" unit="m" value="10.0" valueMax="20.0" valueMin="0.0">0–20 m</quantity>
|
||
),
|
||
<date id="FFE31033FFC5FFF6FD460F984B579BDF" box="[681,796,1193,1218]" pageId="17" pageNumber="468" value="1948-04" valueMax="1948-05-31" valueMin="1948-04-00">iv–v 1948</date>
|
||
, leg. L.S. Liong; 1 ♀, E. Borneo, Gunung Sari, alt.
|
||
<quantity id="4CA59B16FFC5FFF6FAB30F984DDC9BDC" box="[1372,1431,1193,1218]" metricMagnitude="1" metricUnit="m" metricValue="9.5" pageId="17" pageNumber="468" unit="m" value="95.0">95 m</quantity>
|
||
,
|
||
<date id="FFE31033FFC5FFF6FF780FFC49599BF8" box="[151,274,1228,1253]" pageId="17" pageNumber="468" value="1956-08-02">2 viii 1956</date>
|
||
, leg. A.M.R. Wegner;
|
||
<specimenCount id="9D5BFD7AFFC5FFF6FDE10FFC4A729BF9" box="[526,569,1229,1252]" pageId="17" pageNumber="468" type="generic">7 ♂</specimenCount>
|
||
, E. Borneo, Tabang, Bengen River,
|
||
<date id="FFE31033FFC5FFF6FC240FFC4CD89BF8" box="[971,1171,1228,1253]" pageId="17" pageNumber="468" value="1956-08-20" valueMax="1956-10-26" valueMin="1956-08-20">20 viii–26 x 1956</date>
|
||
, leg. A.M.R. Wegner;
|
||
<specimenCount id="9D5BFD7AFFC5FFF6FA610FFC48E59A15" pageId="17" pageNumber="468" type="generic">1 ♂</specimenCount>
|
||
, E. Borneo, Kutai, Damay Kendeng Pau,
|
||
<date id="FFE31033FFC5FFF6FD4E0FC14B6F9A15" box="[673,804,1263,1288]" pageId="17" pageNumber="468" value="1938-09-17">17 ix 1938</date>
|
||
, leg. B.M. Hoeks;
|
||
<specimenCount id="9D5BFD7AFFC5FFF6FBE70FC14C729A15" box="[1032,1081,1264,1288]" pageId="17" pageNumber="468" type="generic">1 ♂</specimenCount>
|
||
, W. Borneo, M., Singkawang, Bakuan, Selakau River,
|
||
<date id="FFE31033FFC5FFF6FE4B0E224A559A36" box="[420,542,1299,1324]" pageId="17" pageNumber="468" value="1932-09-16">16 ix 1932</date>
|
||
, leg. L. Coomans de Ruiter;
|
||
<specimenCount id="9D5BFD7AFFC5FFF6FCB10E224BC29A36" box="[862,905,1299,1323]" pageId="17" pageNumber="468" type="generic">3 ♂</specimenCount>
|
||
, as above, but date
|
||
<date id="FFE31033FFC5FFF6FB8C0E224C9B9A36" box="[1123,1232,1299,1324]" pageId="17" pageNumber="468" value="1934-04-09">9 iv 1934</date>
|
||
;
|
||
<specimenCount id="9D5BFD7AFFC5FFF6FB340E224D4C9A36" box="[1243,1287,1299,1323]" pageId="17" pageNumber="468" type="generic">1 ♂</specimenCount>
|
||
, 1 ♀, Borneo Exped. Dr Nieuwenhuis, Bloe-oe,
|
||
<date id="FFE31033FFC5FFF6FDF60E064ADB9A52" box="[537,656,1335,1359]" pageId="17" pageNumber="468" value="1898-11">Nov. 1898</date>
|
||
;
|
||
<specimenCount id="9D5BFD7AFFC5FFF6FD730E064A839A53" box="[668,712,1335,1359]" pageId="17" pageNumber="468" type="generic">2 ♂</specimenCount>
|
||
, N. Borneo, Sei Bekan,
|
||
<date id="FFE31033FFC5FFF6FC390E074C2B9A52" box="[982,1120,1334,1359]" pageId="17" pageNumber="468" value="1961-08-16">16 viii 1961</date>
|
||
, Exp. P. Jauffret & R. Pujol, (ex coll. A. Heymer);
|
||
<specimenCount id="9D5BFD7AFFC5FFF6FE600E6B49F79A6C" box="[399,444,1370,1394]" pageId="17" pageNumber="468" type="generic">1 ♂</specimenCount>
|
||
, Centre Borneo, Kapuas River, Sepauk,
|
||
<date id="FFE31033FFC5FFF6FC6D0E6B4C479A6F" box="[898,1036,1369,1394]" pageId="17" pageNumber="468" value="1961-08-20">20 viii 1961</date>
|
||
, Exp. Jauffret;
|
||
<specimenCount id="9D5BFD7AFFC5FFF6FB570E6B4CAF9A6C" box="[1208,1252,1370,1394]" pageId="17" pageNumber="468" type="generic">1 ♂</specimenCount>
|
||
, Centre Borneo, Kapuas River, Environs de Silat,
|
||
<date id="FFE31033FFC5FFF6FDF50E4C4AE29A88" box="[538,681,1404,1429]" pageId="17" pageNumber="468" value="1961-08-24">24 viii 1961</date>
|
||
, Exp. Jauffret;
|
||
<specimenCount id="9D5BFD7AFFC5FFF6FCB30E4C4BC09A89" box="[860,907,1405,1429]" pageId="17" pageNumber="468" type="generic">2 ♂</specimenCount>
|
||
, N. Borneo, E. Sabah, Lahad Datu
|
||
<quantity id="4CA59B16FFC5FFF6FAC50E4D4D389A88" box="[1322,1395,1404,1429]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="17" pageNumber="468" unit="km" value="60.0">60 km</quantity>
|
||
W, Danum Valley Field Centre, Sungai Segama,
|
||
<quantity id="4CA59B16FFC5FFF6FD7F0EAE4A9C9AA5" box="[656,727,1439,1464]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="17" pageNumber="468" unit="m" value="160.0">160 m</quantity>
|
||
asl.,
|
||
<date id="FFE31033FFC5FFF6FCFE0E914B3E9AA5" box="[785,885,1440,1464]" pageId="17" pageNumber="468" value="1987-05-03">3 v 1987</date>
|
||
, leg. J. van Tol.—
|
||
<emphasis id="B929EAE1FFC5FFF6FBD20EAE4CDF9AA5" bold="true" box="[1085,1172,1439,1464]" pageId="17" pageNumber="468">
|
||
<collectionCode id="ED4CAE36FFC5FFF6FBD20EAE4CDF9AA5" box="[1085,1172,1439,1464]" pageId="17" pageNumber="468">BMNH</collectionCode>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="9D5BFD7AFFC5FFF6FB4C0E914C859AA5" box="[1187,1230,1440,1464]" pageId="17" pageNumber="468" type="generic">4 ♂</specimenCount>
|
||
, [
|
||
<collectingCountry id="F34A7663FFC5FFF6FB0F0EAE4D0D9AA5" box="[1248,1350,1439,1464]" name="Malaysia" pageId="17" pageNumber="468">Sarawak</collectingCountry>
|
||
, foot of Mt Dulit], R. Tapah, tributary of Tinjar,
|
||
<date id="FFE31033FFC5FFF6FDB80EF24AB79AC6" box="[599,764,1475,1500]" pageId="17" pageNumber="468" value="1932-09-21" valueMax="1932-09-22" valueMin="1932-09-21">21–22 ix 1932</date>
|
||
, leg. B.M. Hobby and A.W. Moore, [published by
|
||
<bibRefCitation id="EFCC4B02FFC5FFF6FADB0EF248AA9AE2" author="Kimmins" pageId="17" pageNumber="468" refString="Kimmins, D. E. (1936) The Odonata of the Oxford University Sarawak expedition. Journal of the Federated Malay States Museum, 18, 65 - 108." type="journal article" year="1936">Kimmins (1936)</bibRefCitation>
|
||
as ‘
|
||
<taxonomicName id="4C5D4D70FFC5FFF6FEFA0ED64A699AE3" box="[277,546,1510,1534]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="468" phylum="Arthropoda" rank="subSpecies" species="limbata" subSpecies="semilimbata">
|
||
<emphasis id="B929EAE1FFC5FFF6FEFA0ED64A699AE3" box="[277,546,1510,1534]" italics="true" pageId="17" pageNumber="468">D. limbata semilimbata</emphasis>
|
||
</taxonomicName>
|
||
’]; 1 ♀, [
|
||
<collectingCountry id="F34A7663FFC5FFF6FD650ED74ABB9AE2" box="[650,752,1510,1535]" name="Malaysia" pageId="17" pageNumber="468">Sarawak</collectingCountry>
|
||
, foot of Mt Dulit), Junction of rivers Tinjar and Lejok,
|
||
<date id="FFE31033FFC5FFF6FA990ED64899993F" pageId="17" pageNumber="468" value="1932-10-02">2 x 1932</date>
|
||
, leg. B.M. Hobby and A.W. Moore;
|
||
<specimenCount id="9D5BFD7AFFC5FFF6FD6C0D3B4AF9993C" box="[643,690,1546,1570]" pageId="17" pageNumber="468" type="generic">2 ♂</specimenCount>
|
||
, N. Borneo [?Sabah], Coll. McLachlan.—
|
||
<emphasis id="B929EAE1FFC5FFF6FB720D384CB9993F" bold="true" box="[1181,1266,1545,1570]" pageId="17" pageNumber="468">
|
||
<collectionCode id="ED4CAE36FFC5FFF6FB720D384CB9993F" box="[1181,1266,1545,1570]" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="17" pageNumber="468">CUMZ</collectionCode>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="9D5BFD7AFFC5FFF6FAEB0D3B4D7F993C" box="[1284,1332,1546,1570]" pageId="17" pageNumber="468" type="generic">1 ♂</specimenCount>
|
||
, 1 ♀, C. Borneo, Kalimantan,
|
||
<date id="FFE31033FFC5FFF6FE670D1D4A579958" box="[392,540,1580,1605]" pageId="17" pageNumber="468" value="1992-07" valueMax="1992-08-31" valueMin="1992-07-00">vii–viii 1992</date>
|
||
, leg. C. Jiggins, [ex coll. D.A.L. Davies].—
|
||
<emphasis id="B929EAE1FFC5FFF6FBE60D1D4C909958" bold="true" box="[1033,1243,1580,1605]" pageId="17" pageNumber="468">Coll. Hämäläinen</emphasis>
|
||
:
|
||
<specimenCount id="9D5BFD7AFFC5FFF6FB040D1C4D5C9959" box="[1259,1303,1581,1605]" pageId="17" pageNumber="468" type="generic">1 ♂</specimenCount>
|
||
, E. Borneo, Tabang, Bengen River,
|
||
<date id="FFE31033FFC5FFF6FE710D614A5A9975" box="[414,529,1615,1640]" pageId="17" pageNumber="468" value="1956-10-10">10 x 1956</date>
|
||
, leg. A.M.R. Wegner [ex coll.
|
||
<collectionCode id="ED4CAE36FFC5FFF6FC860D604B8F9975" LSID="urn:lsid:biocol.org:col:34992" box="[873,964,1617,1640]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="17" pageNumber="468">RMNH</collectionCode>
|
||
].
|
||
</paragraph>
|
||
<paragraph id="8BE236F3FFC5FFF6FF280D424CAA99E8" blockId="17.[151,1437,559,2033]" pageId="17" pageNumber="468">
|
||
<emphasis id="B929EAE1FFC5FFF6FF280D424A659996" bold="true" box="[199,558,1650,1676]" pageId="17" pageNumber="468">
|
||
Descriptive notes on
|
||
<taxonomicName id="4C5D4D70FFC5FFF6FE2F0D424A659996" box="[448,558,1650,1675]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="lugens">
|
||
<emphasis id="B929EAE1FFC5FFF6FE2F0D424A659996" bold="true" box="[448,558,1650,1675]" italics="true" pageId="17" pageNumber="468">D. lugens</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B929EAE1FFC5FFF6FDD30D424AE49991" bold="true" box="[572,687,1651,1676]" pageId="17" pageNumber="468">Diagnosis</emphasis>
|
||
. Male: A species with a proportionally longer abdomen than other Sundaland
|
||
<taxonomicName id="4C5D4D70FFC5FFF6FEFA0DA749CE99B3" box="[277,389,1686,1710]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="468" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B929EAE1FFC5FFF6FEFA0DA749CE99B3" box="[277,389,1686,1710]" italics="true" pageId="17" pageNumber="468">Dysphaea</emphasis>
|
||
</taxonomicName>
|
||
species. Opaque area on wings wider than in most
|
||
<taxonomicName id="4C5D4D70FFC5FFF6FC200DA64C1599B3" box="[975,1118,1686,1710]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="dimidiata">
|
||
<emphasis id="B929EAE1FFC5FFF6FC200DA64C1599B3" box="[975,1118,1686,1710]" italics="true" pageId="17" pageNumber="468">D. dimidiata</emphasis>
|
||
</taxonomicName>
|
||
specimens, colour of wings paler than in
|
||
<taxonomicName id="4C5D4D70FFC5FFF6FEC60D8B49FD99CC" box="[297,438,1721,1745]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="dimidiata">
|
||
<emphasis id="B929EAE1FFC5FFF6FEC60D8B49FD99CC" box="[297,438,1721,1745]" italics="true" pageId="17" pageNumber="468">D. dimidiata</emphasis>
|
||
</taxonomicName>
|
||
, brownish opaque rather than blackish opaque. Apical arms of terminal segment of penis shorter than in the other species occurring in Borneo, each with a small squarish, apical expansion
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3476578FFC5FFF2FF280DCE4C7B9FE5" lastPageId="21" lastPageNumber="472" pageId="17" pageNumber="468" type="description">
|
||
<paragraph id="8BE236F3FFC5FFF6FF280DCE494C98EC" blockId="17.[151,1437,559,2033]" pageId="17" pageNumber="468">
|
||
<emphasis id="B929EAE1FFC5FFF6FF280DCE494F9805" bold="true" box="[199,260,1791,1816]" pageId="17" pageNumber="468">Male</emphasis>
|
||
(for habitus see
|
||
<figureCitation id="13662A76FFC5FFF6FE240DCE4A579805" box="[459,540,1791,1816]" captionStart="FIGURES 5 – 8" captionStartId="13.[151,264,1156,1179]" captionTargetBox="[151,1436,193,1133]" captionTargetId="figure@13.[151,1436,193,1135]" captionTargetPageId="13" captionText="FIGURES 5 – 8. Selysian type specimens and labels: 5) D. dimidiata lectotype; 6) D. limbata lectotype; 7) D. lugens holotype; 8) D. semilimbata holotype." httpUri="https://zenodo.org/record/243465/files/figure.png" pageId="17" pageNumber="468">Figs. 7</figureCitation>
|
||
,
|
||
<figureCitation id="13662A76FFC5FFF6FDC40C314A019805" box="[555,586,1792,1816]" captionStart="FIGURES 9 – 12" captionStartId="14.[151,264,1350,1373]" captionTargetBox="[151,1436,373,1328]" captionTargetId="figure@14.[151,1436,373,1330]" captionTargetPageId="14" captionText="FIGURES 9 – 12. Habitus of male Dysphaea: 9) D. dimidiata West Java; 10) D. lugens Danum Valley, Sabah; 11) D. ulu holotype (flipped horizontally); 12) D. vanida holotype (flipped horizontally)." httpUri="https://zenodo.org/record/243466/files/figure.png" pageId="17" pageNumber="468">10</figureCitation>
|
||
). Head: Labrum, base of mandibles and clypeus shining black, frons and vertex matt black, many individuals with two small faint brownish spots anterior to median ocellus; two specimens have an additional pair of small spots, placed outside the lateral ocelli. Thorax: Matt black, with obscure brownish stripes on synthorax as in
|
||
<figureCitation id="13662A76FFC5FFF6FE530C5D4A449899" box="[444,527,1900,1925]" captionStart="FIGURES 13 – 18" captionStartId="15.[151,264,1951,1974]" captionTargetBox="[151,1436,517,1918]" captionTargetId="figure@15.[151,1436,517,1931]" captionTargetPageId="15" captionText="FIGURES 13 – 18. Synthorax male Dysphaea: 13) D. dimidiata West Java; 14) D. lugens Bakuan, Kalimantan Barat; 15) D. ulu holotype; 16) D. vanida holotype; 17) metaposternum, D. dimidiata Brunei; 18) metaposternum, D. ulu holotype." httpUri="https://zenodo.org/record/243467/files/figure.png" pageId="17" pageNumber="468">Fig. 14</figureCitation>
|
||
; in many (more mature?) specimens there are stripes only on metepisternum and metepimeron. Narrow and usually very faint bronzy antehumeral stripes extending from ca 1/3 to 1/2 length are present on many specimens. Venter of thorax without tiny tubercles on metaposternum. Legs black, partly dark brownish.
|
||
</paragraph>
|
||
<caption id="DF22667BFFC6FFF5FF780DAD4C1999F2" httpUri="https://zenodo.org/record/243469/files/figure.png" pageId="18" pageNumber="469" targetBox="[151,1436,193,1669]" targetPageId="18">
|
||
<paragraph id="8BE236F3FFC6FFF5FF780DAD4C1999F2" blockId="18.[151,1436,1692,1776]" pageId="18" pageNumber="469">
|
||
<emphasis id="B929EAE1FFC6FFF5FF780DAD491099AC" bold="true" box="[151,347,1692,1715]" pageId="18" pageNumber="469">FIGURES 25–30.</emphasis>
|
||
Wings of
|
||
<taxonomicName id="4C5D4D70FFC6FFF5FE360DAD4A7599AF" box="[473,574,1692,1714]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B929EAE1FFC6FFF5FE360DAD4A7599AF" box="[473,574,1692,1714]" italics="true" pageId="18" pageNumber="469">Dysphaea</emphasis>
|
||
</taxonomicName>
|
||
males:
|
||
<emphasis id="B929EAE1FFC6FFF5FD740DAD4AF799AE" bold="true" box="[667,700,1692,1715]" pageId="18" pageNumber="469">25)</emphasis>
|
||
<taxonomicName id="4C5D4D70FFC6FFF5FD250DAC4BF899AF" authority="Pemantang" authorityName="Pemantang" box="[714,947,1692,1714]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="lugens">
|
||
<emphasis id="B929EAE1FFC6FFF5FD250DAC4B7999AF" box="[714,818,1692,1714]" italics="true" pageId="18" pageNumber="469">D. lugens</emphasis>
|
||
Pemantang
|
||
</taxonomicName>
|
||
, Kalimantan Tengah;
|
||
<emphasis id="B929EAE1FFC6FFF5FB490DAD4C8C99AE" bold="true" box="[1190,1223,1692,1715]" pageId="18" pageNumber="469">26)</emphasis>
|
||
<taxonomicName id="4C5D4D70FFC6FFF5FB3A0DAC4DD399AF" authority="Ampah" authorityName="Ampah" box="[1237,1432,1692,1714]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="lugens">
|
||
<emphasis id="B929EAE1FFC6FFF5FB3A0DAC4D7599AF" box="[1237,1342,1692,1714]" italics="true" pageId="18" pageNumber="469">D. lugens</emphasis>
|
||
Ampah
|
||
</taxonomicName>
|
||
, Kalimantan Tengah;
|
||
<emphasis id="B929EAE1FFC6FFF5FE820D8B49C599CC" bold="true" box="[365,398,1722,1745]" pageId="18" pageNumber="469">27)</emphasis>
|
||
<taxonomicName id="4C5D4D70FFC6FFF5FE780D8D4AD999CC" authority="Selekau River" authorityName="Selekau River" box="[407,658,1723,1745]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="lugens">
|
||
<emphasis id="B929EAE1FFC6FFF5FE780D8D49B299CC" box="[407,505,1723,1745]" italics="true" pageId="18" pageNumber="469">D. lugens</emphasis>
|
||
Selekau River
|
||
</taxonomicName>
|
||
, Kalimantan Barat;
|
||
<emphasis id="B929EAE1FFC6FFF5FC8E0D8B4BC999CC" bold="true" box="[865,898,1722,1745]" pageId="18" pageNumber="469">28)</emphasis>
|
||
<taxonomicName id="4C5D4D70FFC6FFF5FC650D8D4B8099CC" box="[906,971,1723,1745]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="ulu">
|
||
<emphasis id="B929EAE1FFC6FFF5FC650D8D4B8099CC" box="[906,971,1723,1745]" italics="true" pageId="18" pageNumber="469">D. ulu</emphasis>
|
||
</taxonomicName>
|
||
paratype, Temburong, Brunei;
|
||
<emphasis id="B929EAE1FFC6FFF5FAE20D8B4D6499CC" bold="true" box="[1293,1327,1722,1745]" pageId="18" pageNumber="469">29)</emphasis>
|
||
<taxonomicName id="4C5D4D70FFC6FFF5FAD80D8D4DD799CC" box="[1335,1436,1723,1745]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="vanida" status="sp. nov.">
|
||
<emphasis id="B929EAE1FFC6FFF5FAD80D8D4DD799CC" box="[1335,1436,1723,1745]" italics="true" pageId="18" pageNumber="469">D. vanida</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B929EAE1FFC6FFF5FF780DEB48B599F2" bold="true" box="[151,254,1754,1775]" pageId="18" pageNumber="469">
|
||
<taxonomicNameLabel id="A21A579AFFC6FFF5FF780DEB48B599F2" box="[151,254,1754,1775]" pageId="18" pageNumber="469" rank="species">spec. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
paratype, Khlong Nakha, Ranong;
|
||
<emphasis id="B929EAE1FFC6FFF5FD8A0DE84ACC99ED" bold="true" box="[613,647,1753,1776]" pageId="18" pageNumber="469">30)</emphasis>
|
||
<taxonomicName id="4C5D4D70FFC6FFF5FD620DEB4ABA99F2" box="[653,753,1753,1775]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="vanida">
|
||
<emphasis id="B929EAE1FFC6FFF5FD620DEB4ABA99F2" box="[653,753,1753,1775]" italics="true" pageId="18" pageNumber="469">D. vanida</emphasis>
|
||
</taxonomicName>
|
||
paratype, Khlong Nakha, Ranong.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE236F3FFC6FFF5FF280C2D49A798D8" blockId="18.[151,1437,1820,1989]" pageId="18" pageNumber="469">
|
||
Wings: In Fw basal brownish opaque area usually extends 5–7 cells beyond nodus but sometimes as few as 2 or as many 11 cells beyond, in Hw much further apicad, usually to level of 3–7 cells proximal to pterostigma (
|
||
<figureCitation id="13662A76FFC6FFF5FA800C7148EC9861" captionStart="FIGURES 5 – 8" captionStartId="13.[151,264,1156,1179]" captionTargetBox="[151,1436,193,1133]" captionTargetId="figure@13.[151,1436,193,1135]" captionTargetPageId="13" captionText="FIGURES 5 – 8. Selysian type specimens and labels: 5) D. dimidiata lectotype; 6) D. limbata lectotype; 7) D. lugens holotype; 8) D. semilimbata holotype." httpUri="https://zenodo.org/record/243465/files/figure.png" pageId="18" pageNumber="469">Fig. 7</figureCitation>
|
||
,
|
||
<figureCitation id="13662A76FFC6FFF5FF5B0C55489A9860" box="[180,209,1892,1917]" captionStart="FIGURES 25 – 30" captionStartId="18.[151,264,1692,1715]" captionTargetBox="[151,1436,193,1669]" captionTargetId="figure@18.[151,1436,193,1671]" captionTargetPageId="18" captionText="FIGURES 25 – 30. Wings of Dysphaea males: 25) D. lugens Pemantang, Kalimantan Tengah; 26) D. lugens Ampah, Kalimantan Tengah; 27) D. lugens Selekau River, Kalimantan Barat; 28) D. ulu paratype, Temburong, Brunei; 29) D. vanida spec. nov. paratype, Khlong Nakha, Ranong; 30) D. vanida paratype, Khlong Nakha, Ranong." httpUri="https://zenodo.org/record/243469/files/figure.png" pageId="18" pageNumber="469">25</figureCitation>
|
||
) but sometimes reaches the pterostigma. The opaque patch at wing tips extends anteriorly to level of distal end of pterostigma, or a little more apicad. In some specimens there remains only a narrow hyaline area between the opaque areas (
|
||
<figureCitation id="13662A76FFC6FFF5FE8E0C9D49AB98D8" box="[353,480,1964,1989]" captionStart="FIGURES 25 – 30" captionStartId="18.[151,264,1692,1715]" captionTargetBox="[151,1436,193,1669]" captionTargetId="figure@18.[151,1436,193,1671]" captionTargetPageId="18" captionText="FIGURES 25 – 30. Wings of Dysphaea males: 25) D. lugens Pemantang, Kalimantan Tengah; 26) D. lugens Ampah, Kalimantan Tengah; 27) D. lugens Selekau River, Kalimantan Barat; 28) D. ulu paratype, Temburong, Brunei; 29) D. vanida spec. nov. paratype, Khlong Nakha, Ranong; 30) D. vanida paratype, Khlong Nakha, Ranong." httpUri="https://zenodo.org/record/243469/files/figure.png" pageId="18" pageNumber="469">Fig. 26–27</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF22667BFFC7FFF4FF780D9849FC99E1" httpUri="https://zenodo.org/record/243470/files/figure.png" pageId="19" pageNumber="470" targetBox="[157,1436,185,1683]" targetPageId="19">
|
||
<paragraph id="8BE236F3FFC7FFF4FF780D9849FC99E1" blockId="19.[151,1436,1705,1788]" pageId="19" pageNumber="470">
|
||
<emphasis id="B929EAE1FFC7FFF4FF780D98491F99A3" bold="true" box="[151,340,1705,1728]" pageId="19" pageNumber="470">FIGURES 31–36.</emphasis>
|
||
Male anal appendages. Dorsal view:
|
||
<emphasis id="B929EAE1FFC7FFF4FD3C0D984ABF99DD" bold="true" box="[723,756,1705,1728]" pageId="19" pageNumber="470">31)</emphasis>
|
||
<taxonomicName id="4C5D4D70FFC7FFF4FD140D9B4B3799A2" box="[763,892,1705,1727]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="dimidiata">
|
||
<emphasis id="B929EAE1FFC7FFF4FD140D9B4B3799A2" box="[763,892,1705,1727]" italics="true" pageId="19" pageNumber="470">D. dimidiata</emphasis>
|
||
</taxonomicName>
|
||
West Java;
|
||
<emphasis id="B929EAE1FFC7FFF4FC190D984C5C99DD" bold="true" box="[1014,1047,1705,1728]" pageId="19" pageNumber="470">32)</emphasis>
|
||
<taxonomicName id="4C5D4D70FFC7FFF4FBF10D9B4C9C99A2" authority="Bakuan" authorityName="Bakuan" box="[1054,1239,1705,1727]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="lugens">
|
||
<emphasis id="B929EAE1FFC7FFF4FBF10D9B4CCB99A2" box="[1054,1152,1705,1727]" italics="true" pageId="19" pageNumber="470">D. lugens</emphasis>
|
||
Bakuan
|
||
</taxonomicName>
|
||
, Kalimantan Barat;
|
||
<emphasis id="B929EAE1FFC7FFF4FF780DF648F399C3" bold="true" box="[151,184,1735,1758]" pageId="19" pageNumber="470">33)</emphasis>
|
||
<taxonomicName id="4C5D4D70FFC7FFF4FF2F0DF8494A99C3" box="[192,257,1736,1758]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="ulu">
|
||
<emphasis id="B929EAE1FFC7FFF4FF2F0DF8494A99C3" box="[192,257,1736,1758]" italics="true" pageId="19" pageNumber="470">D. ulu</emphasis>
|
||
</taxonomicName>
|
||
holotype;
|
||
<emphasis id="B929EAE1FFC7FFF4FE800DF649DA99C3" bold="true" box="[367,401,1735,1758]" pageId="19" pageNumber="470">34)</emphasis>
|
||
<taxonomicName id="4C5D4D70FFC7FFF4FE770DF849B699C3" box="[408,509,1736,1758]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="vanida">
|
||
<emphasis id="B929EAE1FFC7FFF4FE770DF849B699C3" box="[408,509,1736,1758]" italics="true" pageId="19" pageNumber="470">D. vanida</emphasis>
|
||
</taxonomicName>
|
||
holotype. Dorso-lateral views:
|
||
<emphasis id="B929EAE1FFC7FFF4FCAD0DF64B2899C3" bold="true" box="[834,867,1735,1758]" pageId="19" pageNumber="470">35)</emphasis>
|
||
<taxonomicName id="4C5D4D70FFC7FFF4FC840DF84BA799C3" box="[875,1004,1736,1758]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="dimidiata">
|
||
<emphasis id="B929EAE1FFC7FFF4FC840DF84BA799C3" box="[875,1004,1736,1758]" italics="true" pageId="19" pageNumber="470">D. dimidiata</emphasis>
|
||
</taxonomicName>
|
||
Terengganu;
|
||
<emphasis id="B929EAE1FFC7FFF4FB960DF64CD199C3" bold="true" box="[1145,1178,1735,1758]" pageId="19" pageNumber="470">36)</emphasis>
|
||
<taxonomicName id="4C5D4D70FFC7FFF4FB4D0DF84D4399C3" box="[1186,1288,1736,1758]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="vanida">
|
||
<emphasis id="B929EAE1FFC7FFF4FB4D0DF84D4399C3" box="[1186,1288,1736,1758]" italics="true" pageId="19" pageNumber="470">D. vanida</emphasis>
|
||
</taxonomicName>
|
||
paratype Lam Khlong Gnu, Kanchanaburi.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE236F3FFC7FFF4FF280C154C9B98B4" blockId="19.[151,1437,1828,2033]" pageId="19" pageNumber="470">
|
||
Abdomen: Matt black; S1 with obscure brownish patch; S2–5 with narrow, brown lateral stripes, stripe on S2 placed midlaterally, those on S3–5 ventrolaterally. Appendages (
|
||
<figureCitation id="13662A76FFC7FFF4FC9B0C794B8C987D" box="[884,967,1864,1889]" captionStart="FIGURES 31 – 36" captionStartId="19.[151,264,1705,1728]" captionTargetBox="[157,1436,185,1683]" captionTargetId="figure@19.[157,1442,185,1684]" captionTargetPageId="19" captionText="FIGURES 31 – 36. Male anal appendages. Dorsal view: 31) D. dimidiata West Java; 32) D. lugens Bakuan, Kalimantan Barat; 33) D. ulu holotype; 34) D. vanida holotype. Dorso-lateral views: 35) D. dimidiata Terengganu; 36) D. vanida paratype Lam Khlong Gnu, Kanchanaburi." httpUri="https://zenodo.org/record/243470/files/figure.png" pageId="19" pageNumber="470">Fig. 32</figureCitation>
|
||
) black, similar to
|
||
<taxonomicName id="4C5D4D70FFC7FFF4FB770C784D6C987D" box="[1176,1319,1864,1888]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="dimidiata">
|
||
<emphasis id="B929EAE1FFC7FFF4FB770C784D6C987D" box="[1176,1319,1864,1888]" italics="true" pageId="19" pageNumber="470">D. dimidiata</emphasis>
|
||
</taxonomicName>
|
||
; in lateral view ventral margin of cercus distinctly arched at base (
|
||
<figureCitation id="13662A76FFC7FFF4FCF30C5D4B399898" box="[796,882,1900,1925]" captionStart="FIGURES 37 – 40" captionStartId="20.[151,264,1070,1093]" captionTargetBox="[162,1392,232,1036]" captionTargetId="figure@20.[151,1436,193,1050]" captionTargetPageId="20" captionText="FIGURES 37 – 40. Male anal appendages, lateral view (scale bars 1 mm): 37) D. dimidiata west Java; 38) D. lugens Bakuan, Kalimantan Barat; 39) D. ulu holotype; 40) D. vanida holotype." httpUri="https://zenodo.org/record/243471/files/figure.png" pageId="19" pageNumber="470">Fig. 38</figureCitation>
|
||
). Penis: Terminal segment with two short apical arms, directed outwards, upwards, then downwards; transversely expanded at apex (
|
||
<figureCitation id="13662A76FFC7FFF4FBAA0CA14CD398B5" box="[1093,1176,1936,1961]" captionStart="FIGURES 41 – 44" captionStartId="21.[151,264,1790,1813]" captionTargetBox="[162,1384,617,1756]" captionTargetId="figure@21.[151,1436,555,1769]" captionTargetPageId="21" captionText="FIGURES 41 – 44. Penis, ventral view: 41) D. dimidiata west Java; 42) D. lugens Selekau River, Kalimantan Barat; 43) D. ulu holotype; 44) D. vanida paratype, Khlong Nakha, Ranong." httpUri="https://zenodo.org/record/243472/files/figure.png" pageId="19" pageNumber="470">Fig. 42</figureCitation>
|
||
,
|
||
<figureCitation id="13662A76FFC7FFF4FB4C0CA14C8898B4" box="[1187,1219,1936,1961]" captionStart="FIGURES 45 – 48" captionStartId="23.[151,264,1735,1758]" captionTargetBox="[162,1365,783,1707]" captionTargetId="figure@23.[151,1436,721,1720]" captionTargetPageId="23" captionText="FIGURES 45 – 48. Penis, lateral view: 45) D. dimidiata west Java; 46) D. lugens Selekau River, Kalimantan Barat; 47) D. ulu holotype; 48) D. vanida paratype, Khlong Nakha, Ranong." httpUri="https://zenodo.org/record/243473/files/figure.png" pageId="19" pageNumber="470">46</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE236F3FFC7FFF4FF280C844A6398EC" blockId="19.[151,1437,1828,2033]" pageId="19" pageNumber="470">
|
||
Measurements (mm): Hindwing 30–35 (usually 32–34), abdomen (apps. excl.) 37–43 (usually 39–42), cerci
|
||
<quantity id="4CA59B16FFC7FFF4FA610C84488D98ED" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="470" unit="mm" value="2.0">2 mm</quantity>
|
||
. Abdomen/Hw ratio 1.20–1.25.
|
||
</paragraph>
|
||
<caption id="DF22667BFFC0FFF3FF780F1F4B569B7E" httpUri="https://zenodo.org/record/243471/files/figure.png" pageId="20" pageNumber="471" targetBox="[162,1392,232,1036]" targetPageId="20">
|
||
<paragraph id="8BE236F3FFC0FFF3FF780F1F4B569B7E" blockId="20.[151,1436,1070,1124]" pageId="20" pageNumber="471">
|
||
<emphasis id="B929EAE1FFC0FFF3FF780F1F491E9B59" bold="true" box="[151,341,1070,1093]" pageId="20" pageNumber="471">FIGURES 37–40.</emphasis>
|
||
Male anal appendages, lateral view (scale bars 1 mm):
|
||
<emphasis id="B929EAE1FFC0FFF3FC7C0F1F4BFF9B58" bold="true" box="[915,948,1070,1093]" pageId="20" pageNumber="471">37)</emphasis>
|
||
<taxonomicName id="4C5D4D70FFC0FFF3FC530F014C769B58" box="[956,1085,1071,1093]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="dimidiata">
|
||
<emphasis id="B929EAE1FFC0FFF3FC530F014C769B58" box="[956,1085,1071,1093]" italics="true" pageId="20" pageNumber="471">D. dimidiata</emphasis>
|
||
</taxonomicName>
|
||
west Java;
|
||
<emphasis id="B929EAE1FFC0FFF3FB5A0F1F4C9C9B58" bold="true" box="[1205,1239,1070,1093]" pageId="20" pageNumber="471">38)</emphasis>
|
||
<taxonomicName id="4C5D4D70FFC0FFF3FB300F014DD39B58" authority="Bakuan" authorityName="Bakuan" box="[1247,1432,1071,1093]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="lugens">
|
||
<emphasis id="B929EAE1FFC0FFF3FB300F014D0A9B58" box="[1247,1345,1071,1093]" italics="true" pageId="20" pageNumber="471">D. lugens</emphasis>
|
||
Bakuan
|
||
</taxonomicName>
|
||
, Kalimantan Barat;
|
||
<emphasis id="B929EAE1FFC0FFF3FEB80F7C49329B79" bold="true" box="[343,377,1101,1124]" pageId="20" pageNumber="471">39)</emphasis>
|
||
<taxonomicName id="4C5D4D70FFC0FFF3FE900F7E49F59B7E" box="[383,446,1101,1124]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="ulu">
|
||
<emphasis id="B929EAE1FFC0FFF3FE900F7E49F59B7E" box="[383,446,1101,1124]" italics="true" pageId="20" pageNumber="471">D. ulu</emphasis>
|
||
</taxonomicName>
|
||
holotype;
|
||
<emphasis id="B929EAE1FFC0FFF3FDC40F7C4A069B79" bold="true" box="[555,589,1101,1124]" pageId="20" pageNumber="471">40)</emphasis>
|
||
<taxonomicName id="4C5D4D70FFC0FFF3FDBC0F7E4AFC9B7E" box="[595,695,1101,1124]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="vanida">
|
||
<emphasis id="B929EAE1FFC0FFF3FDBC0F7E4AFC9B7E" box="[595,695,1101,1124]" italics="true" pageId="20" pageNumber="471">D. vanida</emphasis>
|
||
</taxonomicName>
|
||
holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE236F3FFC0FFF3FF280FBA48BA9AD9" blockId="20.[151,1437,1163,2016]" pageId="20" pageNumber="471">
|
||
<emphasis id="B929EAE1FFC0FFF3FF280FBA49559BB9" bold="true" box="[199,286,1163,1188]" pageId="20" pageNumber="471">Female</emphasis>
|
||
(for habitus see
|
||
<figureCitation id="13662A76FFC0FFF3FE0B0FBA4A719BB9" box="[484,570,1163,1188]" captionStart="FIGURES 49 – 52" captionStartId="24.[151,264,1070,1093]" captionTargetBox="[151,1436,193,1049]" captionTargetId="figure@24.[151,1436,193,1049]" captionTargetPageId="24" captionText="FIGURES 49 – 52. Habitus of female Dysphaea: 49) D. dimidiata west Java; 50) D. lugens Mount Dulit, Sarawak, photograph by Benjamin Price; 51) D. ulu paratype; 52) D. vanida paratype Kanchanaburi." httpUri="https://zenodo.org/record/243474/files/figure.png" pageId="20" pageNumber="471">Fig. 50</figureCitation>
|
||
). Head: Labium shining black with lateral lobes largely yellow, with black hooks. Labrum shining black, with broadly yellow centre, indented by black basally. Base of mandibles largely yellow, with incomplete black stripe medially. Anteclypeus shining black with narrow yellow stripe on postclypeus along ridge. Frons matt black, yellow below antennae except for a transverse central stripe, yellow colour continuing over much of genae, and narrowly along eye margin almost to narrowest point of vertex. Antennae black and dark brown, yellow anterior mark on scape. Vertex and occiput matt black, with a transverse yellow stripe on either side of the median ocellus, and a yellow spot adjacent to each lateral ocellus (
|
||
<figureCitation id="13662A76FFC0FFF3FB050E524D099A61" box="[1258,1346,1379,1404]" captionStart="FIGURES 53 – 56" captionStartId="25.[151,264,1609,1632]" captionTargetBox="[151,1436,625,1588]" captionTargetId="figure@25.[151,1436,625,1588]" captionTargetPageId="25" captionText="FIGURES 53 – 56. Female head: 53) D. dimidiata Sarawak, dorsal view; 54) D. lugens Gunung Sari, Kalimantan Timur, dorsal view; 55) D. ulu paratype, dorsal, slightly frontal view; 56) D. vanida paratype, Kanchanaburi, dorsal, slightly frontal view." httpUri="https://zenodo.org/record/243475/files/figure.png" pageId="20" pageNumber="471">Fig. 54</figureCitation>
|
||
), in the females from Bloe-oe and Mount Dulit these marks joined on one (Dulit) or both sides; two tiny yellow spots on occiput.
|
||
</paragraph>
|
||
<paragraph id="8BE236F3FFC0FFF3FF280EFE4A289815" blockId="20.[151,1437,1163,2016]" pageId="20" pageNumber="471">
|
||
Thorax: Prothorax black with rounded large yellow spots on either side of dorsum of middle lobe, small yellow mark just anterior to central pit, pair of smaller dorsal spots on anterior lobe. Posterior part of hind lobe raised upwards to form an elongate rectangular flap; lateral parts almost perpendicular to base and central part at an oblique angle, appearing as a square tongue in dorsal view (
|
||
<figureCitation id="13662A76FFC0FFF3FCB40D0A4BF89949" box="[859,947,1595,1620]" captionStart="FIGURES 57 – 62" captionStartId="27.[151,264,1913,1936]" captionTargetBox="[155,1436,401,1899]" captionTargetId="figure@27.[155,1440,401,1900]" captionTargetPageId="27" captionText="FIGURES 57 – 62. Female prothorax, dorsal view: 57) D. dimidiata west Java; 58) D. dimidiata Sarawak; 59) D. dimidiata Tandjong Pandan, Billiton; 60) D. lugens Gunung Sari, Kaliamantan Timur; 61) D. ulu paratype; 62) D. vanida paratype, Kanchanaburi." httpUri="https://zenodo.org/record/243476/files/figure.png" pageId="20" pageNumber="471">Fig. 60</figureCitation>
|
||
); flap narrowly bordered by yellow, more broadly so at sides, with a separate paler area centrally in the female from Bloe-oe (not illustrated). Ends of flap prominent laterally. Synthorax matt black, with yellow stripes as in
|
||
<figureCitation id="13662A76FFC0FFF3FC7C0DB24BBB9981" box="[915,1008,1667,1692]" captionStart="FIGURES 63 – 68" captionStartId="28.[151,264,1699,1722]" captionTargetBox="[151,1436,185,1683]" captionTargetId="figure@28.[151,1436,185,1684]" captionTargetPageId="28" captionText="FIGURES 63 – 68. Female synthorax, dorsal view: 63) D. dimidiata west Java; 64) D. dimidiata Sarawak; 65) D. dimidiata Johor; 66) D. lugens Gunung Sari, Kaliamantan Timur; 67) D. ulu paratype; 68) D. vanida paratype Kanchanaburi." httpUri="https://zenodo.org/record/243477/files/figure.png" pageId="20" pageNumber="471">Figs. 66</figureCitation>
|
||
,
|
||
<figureCitation id="13662A76FFC0FFF3FC120DB54C579981" box="[1021,1052,1668,1692]" captionStart="FIGURES 69 – 74" captionStartId="29.[151,264,1820,1843]" captionTargetBox="[151,1436,193,1797]" captionTargetId="figure@29.[151,1436,193,1799]" captionTargetPageId="29" captionText="FIGURES 69 – 74. Female synthorax, lateral view: 69) D. dimidiata west Java; 70) D. dimidiata Sarawak; 71) D. dimidiata Johor; 72) D. lugens Mount Dulit, Sarawak, photograph by Benjamin Price; 73) D. ulu paratype; 74) D. vanida paratype Kanchanaburi." httpUri="https://zenodo.org/record/243478/files/figure.png" pageId="20" pageNumber="471">72</figureCitation>
|
||
. Yellow stripes on mesepisternum and mesepimeron forming loops. Metepisternum and metepimeron mostly yellow. Legs black, yellow markings on both apical and posterior sides of coxae, small yellow mark upper anterior femur, large yellow mark upper ca 1/3 middle femur and ca 2/3 posterior femur.
|
||
</paragraph>
|
||
<paragraph id="8BE236F3FFC0FFF3FF280C224D6A986E" blockId="20.[151,1437,1163,2016]" pageId="20" pageNumber="471">
|
||
Wings: Hyaline with slight brownish tint around costal and subcostal space from base to nodus in both wings and faintly darkened area at extreme wing tips (
|
||
<figureCitation id="13662A76FFC0FFF3FD580C064B46984D" box="[695,781,1847,1872]" captionStart="FIGURES 49 – 52" captionStartId="24.[151,264,1070,1093]" captionTargetBox="[151,1436,193,1049]" captionTargetId="figure@24.[151,1436,193,1049]" captionTargetPageId="24" captionText="FIGURES 49 – 52. Habitus of female Dysphaea: 49) D. dimidiata west Java; 50) D. lugens Mount Dulit, Sarawak, photograph by Benjamin Price; 51) D. ulu paratype; 52) D. vanida paratype Kanchanaburi." httpUri="https://zenodo.org/record/243474/files/figure.png" pageId="20" pageNumber="471">Fig. 50</figureCitation>
|
||
). Fw with 32–36 antenodals in first row; Hw with 28–31. Quadrangle with 3–5 crossveins. Pterostigma long, covering 10–12 underlying cells in Fw,
|
||
<quantity id="4CA59B16FFC0FFF3FB780C6D4CA79869" box="[1175,1260,1883,1908]" metricMagnitude="-1" metricUnit="m" metricValue="2.413" metricValueMax="2.54" metricValueMin="2.286" pageId="20" pageNumber="471" unit="in" value="9.5" valueMax="10.0" valueMin="9.0">9–10 in</quantity>
|
||
Hw.
|
||
</paragraph>
|
||
<paragraph id="8BE236F3FFC0FFF2FF280C4E4C109FC8" blockId="20.[151,1437,1163,2016]" lastBlockId="21.[151,1437,151,501]" lastPageId="21" lastPageNumber="472" pageId="20" pageNumber="471">Abdomen: Matt black, with pale markings as follows: S1 mostly yellow laterally, extending upwards and downwards at apex of segment. S2–7 with yellow lateral stripe, becoming orange on S6–7, broadest on S2–3 and gradually narrowing towards apical segments, narrowly interrupted at near base from S4 or S6. Separate small lateral spots at apex of S6–7. Faint, interrupted, narrow orange lateral stripe on S8, absent on one female, only apical part present on another, irregular orange lateral mark on S9. Appendages black.</paragraph>
|
||
<paragraph id="8BE236F3FFC1FFF2FF280BD14C7B9FE5" blockId="21.[151,1437,151,501]" box="[199,1072,223,248]" pageId="21" pageNumber="472">Measurements (mm): Hw 30.5–34, abdomen (apps. excl.) 33.5–35, cerci ca 1.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3476578FFC1FFF2FF280A354D689EE8" pageId="21" pageNumber="472" type="discussion">
|
||
<paragraph id="8BE236F3FFC1FFF2FF280A354D689EE8" blockId="21.[151,1437,151,501]" pageId="21" pageNumber="472">
|
||
<emphasis id="B929EAE1FFC1FFF2FF280A3549709E00" bold="true" box="[199,315,260,285]" pageId="21" pageNumber="472">Remarks.</emphasis>
|
||
Unfortunately no fresh specimens of
|
||
<taxonomicName id="4C5D4D70FFC1FFF2FD1F0A344B159E01" box="[752,862,260,284]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="lugens">
|
||
<emphasis id="B929EAE1FFC1FFF2FD1F0A344B159E01" box="[752,862,260,284]" italics="true" pageId="21" pageNumber="472">D. lugens</emphasis>
|
||
</taxonomicName>
|
||
were available for the molecular study. However, due to the considerable structural differences, the specific status of
|
||
<taxonomicName id="4C5D4D70FFC1FFF2FC730A184C419E5D" box="[924,1034,296,320]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="lugens">
|
||
<emphasis id="B929EAE1FFC1FFF2FC730A184C419E5D" box="[924,1034,296,320]" italics="true" pageId="21" pageNumber="472">D. lugens</emphasis>
|
||
</taxonomicName>
|
||
is not in doubt. The association of female specimens with
|
||
<taxonomicName id="4C5D4D70FFC1FFF2FE4B0A7C4A5A9E79" box="[420,529,332,356]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="lugens">
|
||
<emphasis id="B929EAE1FFC1FFF2FE4B0A7C4A5A9E79" box="[420,529,332,356]" italics="true" pageId="21" pageNumber="472">D. lugens</emphasis>
|
||
</taxonomicName>
|
||
is a supposition based on occurrence at the same sites and differences from
|
||
<taxonomicName id="4C5D4D70FFC1FFF2FA6E0A7C49489E95" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="dimidiata">
|
||
<emphasis id="B929EAE1FFC1FFF2FA6E0A7C49489E95" italics="true" pageId="21" pageNumber="472">D. dimidiata</emphasis>
|
||
</taxonomicName>
|
||
females. The fact that some males bear faint markings on the vertex in equivalent positions to the females (never seen in the other species occurring in Borneo) lends support to the association of the sexes. The description by
|
||
<bibRefCitation id="EFCC4B02FFC1FFF2FEAE0A8649B89ECD" author="Ris" box="[321,499,439,464]" pageId="21" pageNumber="472" refString="Ris, F. (1911) Libellen von Sintang, Borneo gesammelt von Dr L. Martin. Annales de la Societe Entomologique de Belgique, 55, 231 - 255." type="journal article" year="1911">Ris (1911: 232)</bibRefCitation>
|
||
of female
|
||
<taxonomicName id="4C5D4D70FFC1FFF2FD9F0A884A969ECD" box="[624,733,440,464]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="lugens">
|
||
<emphasis id="B929EAE1FFC1FFF2FD9F0A884A969ECD" box="[624,733,440,464]" italics="true" pageId="21" pageNumber="472">D. lugens</emphasis>
|
||
</taxonomicName>
|
||
mentions “between antenna and ocelli weakly reddish-brown spots”. It seems likely that the markings on the vertex of males are age dependent, fading with maturity.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |