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<document id="06D516630B56BA8FA48B5672C32C71B5" ID-CLB-Dataset="29182" ID-DOI="10.11646/zootaxa.4471.1.6" ID-GBIF-Dataset="3e683ef4-68f0-4189-8578-e3ae30d0a56d" ID-ISSN="1175-5326" ID-Zenodo-Dep="1454471" ID-ZooBank="C7BF13B6-F212-4D89-8D75-6908101C83E2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539175574369" checkinUser="plazi" docAuthor="Hofmann, Sylvia, Mebert, Konrad, Schulz, Klaus-Dieter, Helfenberger, Notker, Göçmen, Bayram &amp; Böhme, Wolfgang" docDate="2018" docId="03F487D0FFC5A54F2CC4FF4F35A4B8F7" docLanguage="en" docName="zootaxa.4471.1.6.pdf" docOrigin="Zootaxa 4471 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zamenis hohenackeri" docType="key" docVersion="7" lastPageNumber="147" masterDocId="FFCDFFA8FFCFA5452C53FFD7303FBB36" masterDocTitle="A new subspecies of Zamenis hohenackeri (Strauch, 1873) (Serpentes: Colubridae) based on morphological and molecular data" masterLastPageNumber="153" masterPageNumber="137" pageNumber="147" updateTime="1698571403142" updateUser="ExternalLinkService">
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<mods:title id="C4537415633D0E98063446FA0C3BE006">A new subspecies of Zamenis hohenackeri (Strauch, 1873) (Serpentes: Colubridae) based on morphological and molecular data</mods:title>
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<mods:namePart id="D214902F3BA5FB8CD50AEE7EF532C98C">Mebert, Konrad</mods:namePart>
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<mods:namePart id="4226F2CF7F6C1113A27CB13F05A87BD0">Helfenberger, Notker</mods:namePart>
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<mods:namePart id="98D3AB7DC380F11320AB949AB9EDC3A6">Göçmen, Bayram</mods:namePart>
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<mods:namePart id="5CB9E2E91E1506ABFA9084C5F4228E91">Böhme, Wolfgang</mods:namePart>
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<treatment id="03F487D0FFC5A54F2CC4FF4F35A4B8F7" ID-DOI="http://doi.org/10.5281/zenodo.6490605" ID-GBIF-Taxon="148680599" ID-Zenodo-Dep="6490605" LSID="urn:lsid:plazi:treatment:03F487D0FFC5A54F2CC4FF4F35A4B8F7" httpUri="http://treatment.plazi.org/id/03F487D0FFC5A54F2CC4FF4F35A4B8F7" lastPageNumber="147" pageId="10" pageNumber="147">
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<paragraph id="8BE236C6FFC5A54F2CC4FF4F337BBB87" blockId="10.[151,836,151,178]" box="[151,836,151,178]" pageId="10" pageNumber="147">
<heading id="D0AA81AAFFC5A54F2CC4FF4F337BBB87" bold="true" box="[151,836,151,178]" fontSize="11" level="1" pageId="10" pageNumber="147" reason="1">
<emphasis id="B929EAD4FFC5A54F2CC4FF4F337BBB87" bold="true" box="[151,836,151,178]" pageId="10" pageNumber="147">
Key to snake taxa of
<taxonomicName id="4C5D4D45FFC5A54F2DCFFF40329DBB87" box="[412,674,151,177]" class="Reptilia" family="Colubridae" genus="Zamenis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="hohenackeri">
<emphasis id="B929EAD4FFC5A54F2DCFFF40329DBB87" bold="true" box="[412,674,151,177]" italics="true" pageId="10" pageNumber="147">Zamenis hohenackeri</emphasis>
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and
<taxonomicName id="4C5D4D45FFC5A54F2EB3FF40337BBB87" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[736,836,151,177]" class="Reptilia" family="Colubridae" genus="Zamenis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="situla">
<emphasis id="B929EAD4FFC5A54F2EB3FF40337BBB87" bold="true" box="[736,836,151,177]" italics="true" pageId="10" pageNumber="147">Z. situla</emphasis>
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<paragraph id="8BE236C6FFC5A54F2CC4FF08337EBBC1" blockId="10.[151,833,222,247]" box="[151,833,222,247]" pageId="10" pageNumber="147">
For exampleS of principal colour morphS and taxa See
<figureCitation id="13662A43FFC5A54F2EABFF09337EBBC1" box="[760,833,222,247]" captionStart="FIGURE 5" captionStartId="10.[151,250,1867,1889]" captionTargetBox="[150,1436,1045,1845]" captionTargetId="figure@10.[150,1436,1045,1845]" captionTargetPageId="10" captionText="FIGURE 5. PRINcIpal bodY coloUR pattERN of Zamenis situla aNd Z. hohenackeri taxa: A) Z. situla blotchEd, AgoRa, IzmIR (local- ItY 139, FIg. 1); B) Z. situla StRIpEd, IzmIR (localItY 139, FIg. 1); C) Z. h. lyciensis ssp. nov. KIRkpINaR KaRaNfIll, ARIbElENI YaYlaSI, MUğla (localItY 52, FIg. 1); D) Z. h. tauricus Aladaǧ, MERSIN (localItY 85, FIg. 1); E) Z. h. hohenackeri GüNINdI, KaRS (localItY 18, FIg. 1). SEE moRE ExamplES IN SUpplEmENtaRY FIgS. 13." httpUri="https://zenodo.org/record/1454481/files/figure.png" pageId="10" pageNumber="147">Fig. 5.</figureCitation>
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<paragraph id="8BE236C6FFC5A54F2CC4FEF235A4BAF8" blockId="10.[151,1436,293,961]" pageId="10" pageNumber="147">
<keyLead id="30AC95F3FFC5A54F2CC4FEF235A4BAF8" pageId="10" pageNumber="147">
1a Upper side of head with three groups of distinct markings (
<figureCitation id="13662A43FFC5A54F2F4DFEF23365BA0D" box="[798,858,293,315]" captionStart="FIGURE 6" captionStartId="12.[151,250,1840,1862]" captionTargetBox="[150,1436,1309,1819]" captionTargetId="figure@12.[150,1436,1309,1819]" captionTargetPageId="12" captionText="FIGURE 6. CompaRISoN of hEad coloUR pattERN dIffERENcES bEtwEEN (a) Zamenis hohenackeri lyciensis ssp. nov. KIRkpINaR KaRaNfIll, MUğla, aNd (b) Z. situla, LakE YlIkI, GREEcE." httpUri="https://zenodo.org/record/1454483/files/figure.png" pageId="10" pageNumber="147">Fig. 6</figureCitation>
): 1) the anterior end of the dorsal band or equivalent horseshoe-shaped nuchal blotch reaching as a thick line along the parietal suture up/into to the frontal plate, 2) upper ends of the bilateral postocular line beginning above each eye on the supraoculars/parietals/frontal, 3) a thick line reaching across the suture of the frontal/prefrontals, connecting the eyes, but in some specimens being dissolved into broad blotches; 2527 dorsals at mid-body; dorsal blotches usually rusty, reddish or orangish in juveniles and adults, 35 dorsal in width along the body axis; dorsal blotches can form two paravertebral lines; 220255 ventrals...............................
<taxonomicName id="4C5D4D45FFC5A54F295DFE6E35A4BAF8" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1294,1435,441,462]" class="Reptilia" family="Colubridae" genus="Zamenis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="situla">
<emphasis id="B929EAD4FFC5A54F295DFE6E35A4BAF8" box="[1294,1435,441,462]" italics="true" pageId="10" pageNumber="147">Zamenis situla</emphasis>
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<paragraph id="8BE236C6FFC5A54F2CC4FE0235A4B975" blockId="10.[151,1436,293,961]" pageId="10" pageNumber="147">
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1b Upper side of head usually without distinct black markings (
<figureCitation id="13662A43FFC5A54F2F7AFE02335BBADD" box="[809,868,469,491]" captionStart="FIGURE 6" captionStartId="12.[151,250,1840,1862]" captionTargetBox="[150,1436,1309,1819]" captionTargetId="figure@12.[150,1436,1309,1819]" captionTargetPageId="12" captionText="FIGURE 6. CompaRISoN of hEad coloUR pattERN dIffERENcES bEtwEEN (a) Zamenis hohenackeri lyciensis ssp. nov. KIRkpINaR KaRaNfIll, MUğla, aNd (b) Z. situla, LakE YlIkI, GREEcE." httpUri="https://zenodo.org/record/1454483/files/figure.png" pageId="10" pageNumber="147">Fig. 6</figureCitation>
); 1) the nuchal blotch barely reaching the parietals or continuing along the suture only as a thin line, sometimes with a blotch on each parietal, 2) upper end of the thick postocular line beginning at mid-level (postocular) of the eye, not above, 3) no line across the snout anterior the eyes; 23, rarely 25 dorsals at mid-body............................................................................................ 2
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<keyStep id="30A92E63FFC5A54F2CC4FD9C35A3B9E0" pageId="10" pageNumber="147">
<paragraph id="8BE236C6FFC5A54F2CC4FD9C35A4B948" blockId="10.[151,1436,293,961]" pageId="10" pageNumber="147">
<keyLead id="30AC95F3FFC5A54F2CC4FD9C35A4B948" pageId="10" pageNumber="147">
2a Dorsal blotches 13 dorsals in width (along body axis,
<figureCitation id="13662A43FFC5A54F2EA9FD9C330BB956" box="[762,820,587,609]" captionStart="FIGURE 7" captionStartId="13.[151,250,871,893]" captionTargetBox="[135,1418,201,858]" captionTargetId="figure@13.[133,1418,201,858]" captionTargetPageId="13" captionText="FIGURE 7. CompaRISoN of doRSal coloUR pattERN dIffERENcES amoNg thE thREE SUbSpEcIES of Zamenis hohenackeri: (a) Z. h. lyciensis ssp. nov.; (b) Z. h. tauricus; (c) Z. h. hohenackeri." httpUri="https://zenodo.org/record/1454485/files/figure.png" pageId="10" pageNumber="147">Fig. 7</figureCitation>
), usually&gt;70 blotches that are indistinctly black bordered with scales often exhibiting black edges or tips throughout the blotch................................................. 3
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</paragraph>
<paragraph id="8BE236C6FFC5A54F2CC4FD5235A3B9E0" blockId="10.[151,1436,293,961]" pageId="10" pageNumber="147">
<keyLead id="30AC95F3FFC5A54F2CC4FD5235A3B9E0" pageId="10" pageNumber="147">
2b Dorsal blotches light brown or reddish brown in adults (similar to
<taxonomicName id="4C5D4D45FFC5A54F2F3DFD513381B9AD" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[878,958,646,667]" class="Reptilia" family="Colubridae" genus="Zamenis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="situla">
<emphasis id="B929EAD4FFC5A54F2F3DFD513381B9AD" box="[878,958,646,667]" italics="true" pageId="10" pageNumber="147">Z. situla</emphasis>
</taxonomicName>
,
<figureCitation id="13662A43FFC5A54F2F9FFD523438B9AD" box="[972,1031,645,667]" captionStart="FIGURE 7" captionStartId="13.[151,250,871,893]" captionTargetBox="[135,1418,201,858]" captionTargetId="figure@13.[133,1418,201,858]" captionTargetPageId="13" captionText="FIGURE 7. CompaRISoN of doRSal coloUR pattERN dIffERENcES amoNg thE thREE SUbSpEcIES of Zamenis hohenackeri: (a) Z. h. lyciensis ssp. nov.; (b) Z. h. tauricus; (c) Z. h. hohenackeri." httpUri="https://zenodo.org/record/1454485/files/figure.png" pageId="10" pageNumber="147">Fig. 7</figureCitation>
), 35 dorsals in width (along body axis), rounded, usually &lt;70 blotches that are distinctly black bordered and no or few scales within the blotch with black edges or tips, brown blotches in juveniles; 214244 ventrals...............................
<taxonomicName id="4C5D4D45FFC5A54F2846FD163503B9E0" authorityName="Hofmann &amp; Mebert &amp; Schulz &amp; Helfenberger &amp; Göçmen &amp; Böhme" authorityYear="2018" box="[1045,1340,705,726]" class="Reptilia" family="Colubridae" genus="Zamenis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="147" phylum="Chordata" rank="subSpecies" species="hohenackeri" subSpecies="lyciensis">
<emphasis id="B929EAD4FFC5A54F2846FD163503B9E0" box="[1045,1340,705,726]" italics="true" pageId="10" pageNumber="147">Zamenis hohenackeri lyciensis</emphasis>
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<emphasis id="B929EAD4FFC5A54F2911FD1735A3B9E0" bold="true" box="[1346,1436,704,726]" pageId="10" pageNumber="147">spp. nov.</emphasis>
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<keyStep id="30A92E63FFC5A54F2CC4FD0A35A4B8F7" pageId="10" pageNumber="147">
<paragraph id="8BE236C6FFC5A54F2CC4FD0A35A3B87D" blockId="10.[151,1436,293,961]" pageId="10" pageNumber="147">
<keyLead id="30AC95F3FFC5A54F2CC4FD0A35A3B87D" pageId="10" pageNumber="147">
3a Dorsal blotches 12 dorsals in width (along body axis,
<figureCitation id="13662A43FFC5A54F2EACFD0A3306B9C5" box="[767,825,733,755]" captionStart="FIGURE 7" captionStartId="13.[151,250,871,893]" captionTargetBox="[135,1418,201,858]" captionTargetId="figure@13.[133,1418,201,858]" captionTargetPageId="13" captionText="FIGURE 7. CompaRISoN of doRSal coloUR pattERN dIffERENcES amoNg thE thREE SUbSpEcIES of Zamenis hohenackeri: (a) Z. h. lyciensis ssp. nov.; (b) Z. h. tauricus; (c) Z. h. hohenackeri." httpUri="https://zenodo.org/record/1454485/files/figure.png" pageId="10" pageNumber="147">Fig. 7</figureCitation>
), commonly dorsal blotches arranged in 2 rows (twin-spotted) and separated by a light coloured vertebral line, whereas some of the first 20 dorsal blotches posterior the nuchal band might be formed as translateral bars, but never all 20; colour of blotches brown in juveniles and adults; 200225 ventrals.....................................................................................
<taxonomicName id="4C5D4D45FFC5A54F2802FCE135A3B87D" authority="hohenackeri" box="[1105,1436,822,843]" class="Reptilia" family="Colubridae" genus="Zamenis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="147" phylum="Chordata" rank="subSpecies" species="hohenackeri" subSpecies="hohenackeri">
<emphasis id="B929EAD4FFC5A54F2802FCE135A3B87D" box="[1105,1436,822,843]" italics="true" pageId="10" pageNumber="147">Zamenis hohenackeri hohenackeri</emphasis>
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</paragraph>
<paragraph id="8BE236C6FFC5A54F2CC4FC8435A4B8F7" blockId="10.[151,1436,293,961]" pageId="10" pageNumber="147">
<keyLead id="30AC95F3FFC5A54F2CC4FC8435A4B8F7" pageId="10" pageNumber="147">
3b Dorsal blotches 23 dorsals in width (along body axis,
<figureCitation id="13662A43FFC5A54F2EA8FC84330AB85F" box="[763,821,851,873]" captionStart="FIGURE 7" captionStartId="13.[151,250,871,893]" captionTargetBox="[135,1418,201,858]" captionTargetId="figure@13.[133,1418,201,858]" captionTargetPageId="13" captionText="FIGURE 7. CompaRISoN of doRSal coloUR pattERN dIffERENcES amoNg thE thREE SUbSpEcIES of Zamenis hohenackeri: (a) Z. h. lyciensis ssp. nov.; (b) Z. h. tauricus; (c) Z. h. hohenackeri." httpUri="https://zenodo.org/record/1454485/files/figure.png" pageId="10" pageNumber="147">Fig. 7</figureCitation>
), usually all of the first 20 blotches posterior the nuchal blotch formed as elongated (oblong to oval) translateral bars (and&gt;50% over the entire trunk), with some dorsal blotches separated into twin spots more posteriorly; colour of blotches brown to olive in juveniles and adults; 196224 ventrals...........................................................................................
<taxonomicName id="4C5D4D45FFC5A54F2826FC7B35A4B8F7" baseAuthorityName="Werner" baseAuthorityYear="1898" box="[1141,1435,940,961]" class="Reptilia" family="Colubridae" genus="Zamenis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="147" phylum="Chordata" rank="subSpecies" species="hohenackeri" subSpecies="tauricus">
<emphasis id="B929EAD4FFC5A54F2826FC7B35A4B8F7" box="[1141,1435,940,961]" italics="true" pageId="10" pageNumber="147">Zamenis hohenackeri tauricus</emphasis>
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