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<document ID-DOI="10.11646/zootaxa.5277.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7889695" ID-ZooBank="73ED69F3-3887-460D-86DE-C9F1302C8EC3" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1683099613832" checkinUser="plazi" docAuthor="Kluge, Nikita, Srinivasan, Pandiarajan, Sivaruban, T., Barathy, S. &amp; Isack, Rajasekaran" docDate="2023" docId="03F487D0A3704D49A2DBFF43FA24EBC1" docLanguage="en" docName="zootaxa.5277.2.1.pdf" docOrigin="Zootaxa 5277 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5277.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tenuibaetis Kang &amp; Yang 1994" docType="treatment" docVersion="3" lastPageNumber="208" masterDocId="FFCDFFA8A3764D4EA24CFFD4FFABE864" masterDocTitle="Contribution to the knowledge of the subgenus Tenuibaetis Kang &amp; Yang 1994 (Ephemeroptera, Baetidae, Baetis s. l.)" masterLastPageNumber="258" masterPageNumber="201" pageNumber="207" updateTime="1683138298321" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:title>Contribution to the knowledge of the subgenus Tenuibaetis Kang &amp; Yang 1994 (Ephemeroptera, Baetidae, Baetis s. l.)</mods:title>
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<mods:namePart>Kluge, Nikita</mods:namePart>
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Subgenus
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<emphasis bold="true" box="[279,418,151,178]" italics="true" pageId="6" pageNumber="207">Tenuibaetis</emphasis>
Kang &amp; Yang
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(in Kang, Chang &amp; Yang) 1994
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captionTargetPageId-4="9" captionTargetPageId-40="50" captionTargetPageId-41="51" captionTargetPageId-42="52" captionTargetPageId-43="53" captionTargetPageId-44="54" captionTargetPageId-45="55" captionTargetPageId-5="10" captionTargetPageId-6="11" captionTargetPageId-7="12" captionTargetPageId-8="13" captionTargetPageId-9="14" captionText-0="FIGURES 14. Setae on larval legs of Baetis (Tenuibaetis) bialatus sp. n.. 1, two-channel and hirudiform setae on outer side of middle tibia under scanning electron microscope; 23, hirudiform seta; 4, two-channel seta on anterior side of femur in transmitted light. Abbreviations: 2-can, two-channel seta; hirud, hirudiform seta." captionText-1="FIGURES 512. Pseudo-bifurcate setae of Baetis (Tenuibaetis) frequentus from India. 58, pseudo-bifurcate setae on outer side of tibia (7, with hirudiform seta); 910, pseudo-bifurcate setae on outer side of tibia (view in profile, in transmitted light); 1112, pseudo-bifurcate setae on anterior side of femur (surface view, in transmitted light)." captionText-10="FIGURES 5259. Baetis (Tenuibaetis) hissaricus. 5253, fore and middle legs of male imago; 5455, genitalia of males imago; 56, hind wing of male; 57, hind wing of female; 58, subimaginal exuviae of female fore leg; 59, subimaginal exuviae of half of mesonotum (57, 58, holotype)." captionText-11="FIGURES 6068. Baetis (Tenuibaetis) frequentus from Sri Lanka, larval exuviae. 60, abdomen; 61, fore leg; 62, hind leg; 63, half of pronotum and mesonotum; 6468, such details of other individuals." captionText-12="FIGURES 6977. Baetis (Tenuibaetis) frequentus, larval exuviae. 6975, from India (6974, at one magnification): 69, head; 7072, fore, middle and hind legs; 73, abdomen; 74, half of pronotum and mesonotum, 75, femur of dark individual. 7677, posterior margin of abdominal tergum VI of two individuals from Haputale (Sri Lanka)." captionText-13="FIGURES 7882. Baetis (Tenuibaetis) frequentus, larvae from India. 7879, left and right mandibles with mandibles of next instar developing inside; 8081, labrum of two individuals; 82, labium." captionText-14="FIGURES 8388. Baetis (Tenuibaetis) frequentus from Sri Lanka, setae on anterior side of femur. 83, pseudo-bifurcate setae and one long two-channel seta in individual from Sri Pada; 8485, fore and middle femora of another individual from Sri Pada, with all setae clavate and two-channel; 86, enlarged fragment of fore femur of the same individual; 8788, hind femur of individual from Haputale, with all setae short and pseudo-bifurcate." captionText-15="FIGURES 8991. Baetis (Tenuibaetis) frequentus from India. 8990, apex of femur and tibia of fore leg; 91, apex of femur and base of tibia of hind leg. Abbreviations: 2-can, two-channel seta; ant-ap, anterior-apical margin of femur; hirud, hirudiform seta; ps-bi, pseudo-bifurcate seta; sq, scale." captionText-16="FIGURES 92101. Baetis (Tenuibaetis) frequentus from Sri Lanka. 92, female imago; 9394, male imago from Sri Pada and its fore leg; 9598, fore leg, head, abdomen and genitalia of male imago from Haputale; 99101, male imago from Uma, its fore leg and genitalia." captionText-17="FIGURES 102111. Baetis (Tenuibaetis) frequentus from India. 102109, male imago and its subimaginal exuviae: 102, head and thorax; 103, abdomen; 104, exuviae of subimaginal half of mesonotum; 105106, fore and middle legs; 107108, their subimaginal exuviae; 109, hind wing. 110111, female imago and its hind wing." captionText-18="FIGURES 112115. Baetis (Tenuibaetis) frequentus from India, genitalia of male imago. Abbreviations: gv, gonovectus; m.gv, muscle of gonovectis; s.s, vestige of unpaired sterno-styligeral muscle." captionText-19="FIGURES 116124. Baetis (Tenuibaetis) frequentus (holotype of Indobaetis michaelohubbardi). 116, labrum; 117118, left and right mandibles; 119, maxilla; 120, hypopharynx with superlinguae; 121122, labium; 123, abdominal tergum with tergalius; 124, leg." captionText-2="FIGURES 1321. Imaginal hind wings and larval protoptera (under the same magnification). 1314, female Baetis (Tenuibaetis) frequentus from India; 1517, male B. (T.) octomaculatus sp. n.; 1819, female of same species; 2021, male B. (T.) bialatus sp. n. (13, 15, 20, metathorax of imago with hind wing; 14, 21, larval exuviae of metanotum; 16, 18, hind wing of imago; 17, 19, cuticle from metathorax of larva ready to molt to subimago, with subimaginal wing inside protopteron: crumpled on Fig. 17, non-crumpled on Fig. 19)." captionText-20="FIGURES 125128. Baetis (Tenuibaetis) fujitanii. 125127, larval exuviae: 125, head; 126, half of pronotum and mesonotum; 127, abdomen; 128, young larva in natural pose." captionText-21="FIGURES 129134. Baetis (Tenuibaetis) fujitanii, exuviae of larval legs. 129131, fore, middle and hind legs; 132, anterior surface of hind femur; 133134, middle tibia." captionText-22="FIGURES 135143. Baetis (Tenuibaetis) fujitanii. 135, male imago; 136, female imago; 137, subimaginal exuviae of half of mesonotum; 138, male hind wing; 139, female hind wing; 140, exuviae of male fore leg; 141, abdomen of male imago; 142143, middle and fore legs of male imago." captionText-23="FIGURES 144145. Baetis (Tenuibaetis) fujitanii, genitalia of male imagines." captionText-24="FIGURES 146153. Baetis (Tenuibaetis) kaltenbachi sp. n.. 146147, mature female and male larvae; 148153, larval exuviae: 148, abdomen; 149, half of pronotum and mesonotum; 150152, fore, middle and hind legs; 153, paraproct (148153, holotype)." captionText-25="FIGURES 154159. Baetis (Tenuibaetis) kaltenbachi sp. n., larval exuviae (holotype). 154155, labrum; 156158, labium; 159, posterior margin of abdominal tergum I." captionText-26="FIGURES 160168. Baetis (Tenuibaetis) kaltenbachi sp. n., larval exuviae (holotype). 160162, femora of fore, middle and hind legs; 163164, tibiae of fore and middle legs; 165, fragment of fore femur; 166167, fragments of mid femur; 168, claw." captionText-27="FIGURES 169180. Baetis (Tenuibaetis) kaltenbachi sp. n. 169170, male imago; 171, female imago; 172, subimaginal exuviae of abdomen; 173174, fore and middle legs of male imago; 175176, their subimaginal exuviae; 177, subimaginal exuviae of half of mesonotum; 178, hind wing of male imago; 179, the same, female; 180, blunt microlepides on fore tarsus of male subimago (169170, 172178, 180, holotype)." captionText-28="FIGURES 181183. Baetis (Tenuibaetis) kaltenbachi sp. n., genitalia of male imago (holotype)." captionText-29="FIGURES 184190. Baetis (Tenuibaetis) bialatus sp. n., larval exuviae (one and the same magnification). 184185, abdomen; 186, head with labrum; 187, half of pronotum and mesonotum; 188190, fore, middle and hind legs." captionText-3="FIGURES 2230. Baetis (Tenuibaetis) ursinus, larval exuviae (2429, one and the same magnification). 22, labrum; 23, maxilla and labium; 24, abdomen, 2527, middle and hind legs; 28, half of pronotum and mesonotum; 29, head; 30, paraproct." captionText-30="FIGURES 191194. Baetis (Tenuibaetis) bialatus sp. n., larval mouth parts. 191, labrum; 192, hypopharynx with superlinguae; 193, maxilla; 194, labium." captionText-31="FIGURES 195201. Baetis (Tenuibaetis) bialatus sp. n., larvae. 195196, left and right mandibles with developing mandibles of next instar inside; 197198, the same of another individual; 199, exuviae of abdominal tergum IX; 200, abdominal tergum II; 201, paraprocts. Abbreviations: hirud, hirudiform seta; sq, scale." captionText-32="FIGURES 202207. Baetis (Tenuibaetis) bialatus sp. n., larval legs. 202, middle femur (anterior side); 203, fore tibia (anterior side); 204, middle tibia (anterior side); 205, middle tibia (outer side); 206207, tibia and tarsus of fore and middle legs (anterior side) (202204, 206207, holotype). Abbreviations: 2-can, two-channel seta." captionText-33="FIGURES 208210. Baetis (Tenuibaetis) bialatus sp. n., proximal, middle and distal portions of larval femur (view from outer side).Abbreviations: 2-can, two-channel seta; ant-ap, anterior-apical margin of femur; hirud, hirudiform seta; post-ap, posteriorapical margin of femur." captionText-34="FIGURES211214. Baetis (Tenuibaetis) bialatus sp.n., larvaltibiae(view fromouter side).211213,fore tibia;214,middle tibia." captionText-35="FIGURES 215224. Baetis (Tenuibaetis) bialatus sp. n. 215, subimaginal exuviae of mesopleuron; 216, subimaginal exuviae of half of mesonotum; 217218, exuviae of fore and hind legs of male subimago; 219220, fore and middle legs of male imago; 221, male imago; 222, female imago; 223, subimaginal exuviae of abdominal segments VIIIIX; 224, egg (215220, holotype)." captionText-36="FIGURES 225229. Baetis (Tenuibaetis) bialatus sp. n., genitalia of male imago. 225, 227, ventral view; 226, 229, ventral view with focus on sterno-styligeral muscle; 228, enlarged fragment of sterno-styligeral muscle to show cross-striation (225226, holotype). Abbreviations: gv, gonovectis; s.s, sterno-styligeral muscle." captionText-37="FIGURES 230236. Baetis (Tenuibaetis) panhai, exuviae of larval legs. 230231, proximal-posterior portion of fore femur with villopore (230, anterior side; 231, posterior side); 232234, fore, middle and hind legs; 235236, fore and middle tibiae." captionText-38="FIGURES 237241. Baetis (Tenuibaetis) panhai. 237, abdomen of male subimago extracted from mature larva; 238, larval exuviae of half of pronotum and mesonotum; 239, larval exuviae of abdomen; 240, male subimago; 241, subimaginal gonostyli crumpled under larval cuticle." captionText-39="FIGURES 242248. Baetis (Tenuibaetis) octomaculatus sp. n. 242247, exuviae of female larvae: 242, 245, half of pronotum and mesonotum; 243, 246, head; 244, 247, abdomen; 248, young larva to show hypodermal coloration (242244, holotype)." captionText-4="FIGURES 3132. Baetis (Tenuibaetis) ursinus, larval exuviae of fore leg. 31, part of femur; 32, part of tibia. Abbreviation: 2-can, two-channel setae." captionText-40="FIGURES 249252. Baetis (Tenuibaetis) octomaculatus sp. n. 249, labrum (holotype); 250251, left and right mandibles; 252, maxilla and labium." captionText-41="FIGURES 253256. Baetis (Tenuibaetis) octomaculatus sp. n. (holotype). 253255, larval exuviae of fore, middle and hind legs; 256, base of femur with villopore." captionText-42="FIGURES 257264. Baetis (Tenuibaetis) octomaculatus sp. n. 257258, fore and hind femora; 259260, fore and middle tibiae; 261263, abdominal terga I, VII and X; 264, abdominal sternum IX and paraprocts (257260, holotype)." captionText-43="FIGURES 265274. Baetis (Tenuibaetis) octomaculatus sp. n. 265, subimaginal exuviae of half of mesonotum; 266, male imago; 267, female imago; 268269, fore legs of two female imagines (268, holotype); 270, female imago; 271, male subimago; 272, abdomen of male imago; 273, fore leg of male imago; 274, its subimaginal exuviae." captionText-44="FIGURES 275278. Baetis (Tenuibaetis) octomaculatus sp. n. 275, subimaginal gonostyli crumpled under larval cuticle; 276, imaginal genitalia; 277, imaginal genitalia of another individual, with focus on ventral side; 278, the same, with focus on gonovectes. Abbreviation: s.s, sterno-styligeral muscle." captionText-45="FIGURES 279284. Baetis (Tenuibaetis) sp. (Thailand). 279, fore femur; 280, portion of larval exuviae with left mandible, labrum and frons; 281, labium; 282, male subimaginal abdomen extracted from larva; 283, larval abdominal segments IXX and caudalii; 284, exuviae of larval abdominal segments IVIII." captionText-5="FIGURES 3341. Baetis (Tenuibaetis) ursinus. 33, subimaginal exuviae of half of mesonotum; 34, hind wing of male; 35, hind wing of female; 36, male imago and its spread abdomen; 3738, its fore and middle legs; 3940, their subimaginal exuviae (the same magnification); 41, female imago." captionText-6="FIGURES 4243. Baetis (Tenuibaetis) ursinus, genitalia of male imagines. Abbreviations: 1, 2, 3, segments of gonostylus; gd, gonoduct; gv, gonovectis; m.gs, gonostylar muscle; m.gv, muscle of gonovectis; s.s, sterno-styligeral muscle; us, unistyliger." captionText-7="FIGURES 4446. Baetis (Tenuibaetis) hissaricus, larval exuviae. 44, head; 45, labrum; 46, labium (44, 46, holotype)." captionText-8="FIGURES 4749. Baetis (Tenuibaetis) hissaricus, exuviae of larval fore, middle and hind femora. Abbreviation: 2-can, twochannel setae." captionText-9="FIGURES 5051. Baetis (Tenuibaetis) hissaricus, exuviae of larval hind tibia. 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)
</paragraph>
</subSubSection>
<subSubSection box="[151,878,256,280]" pageId="6" pageNumber="207" type="multiple">
<paragraph blockId="6.[151,980,256,341]" box="[151,878,256,280]" pageId="6" pageNumber="207">
<treatmentCitationGroup box="[151,878,256,280]" pageId="6" pageNumber="207">
Species
<treatmentCitation author="Novikova, E. A." box="[235,604,256,280]" page="793" pageId="6" pageNumber="207" year="1991">
<taxonomicName ID-CoL="KD3K" authority=": Novikova 1991: 793" authorityName=": Novikova" authorityPageNumber="793" authorityYear="1991" box="[235,604,256,280]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="ursinus">
<emphasis box="[235,380,256,279]" italics="true" pageId="6" pageNumber="207">Baetis ursinus</emphasis>
:
<bibRefCitation author="Novikova, E. A." box="[393,604,256,280]" pageId="6" pageNumber="207" pagination="793 - 795" refId="ref19156" refString="Novikova, E. A. (1991) [A new subspecies of mayflies Baetis ursinus hissaricus subsp. n. (Ephemeroptera, Baetidae) from Tadjikistan]. Entomologicheskoe Obozrenie, 70 (4), 793 - 795." type="journal article" year="1991">Novikova 1991: 793</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation author="Kluge, N. J." box="[614,797,256,280]" page="193" pageId="6" pageNumber="207" year="1997">
<bibRefCitation author="Kluge, N. J." box="[614,797,256,280]" pageId="6" pageNumber="207" refId="ref18180" refString="Kluge, N. J. (1997 b) Order Ephemeroptera. In: Tsalolikhin, S. J. (Ed.), Key to freshwater invertebrates of Russia and adjacent lands. Vol. 3. Nauka, S-Petersburg, pp. 176 - 220." type="book" year="1997">Kluge 1997b: 193</bibRefCitation>
</treatmentCitation>
(larva).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="207" type="reference_group">
<paragraph blockId="6.[151,980,256,341]" box="[151,980,286,311]" pageId="6" pageNumber="207">
Subgenus
<taxonomicName authority="Kang &amp; Yang" authorityName="Kang &amp; Yang" authorityYear="1994" box="[256,517,286,311]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="genus">
<emphasis box="[256,372,286,310]" italics="true" pageId="6" pageNumber="207">Tenuibaetis</emphasis>
Kang &amp; Yang
</taxonomicName>
(in Kang &amp; Chang &amp; Yang) 1994: 26 (larva).
</paragraph>
<paragraph blockId="6.[151,980,256,341]" box="[151,919,317,341]" pageId="6" pageNumber="207">
<treatmentCitationGroup box="[151,919,317,341]" pageId="6" pageNumber="207">
Genus
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<taxonomicName ID-CoL="63VC4" authority=": Fujitani, Hirowatari &amp; Tanida 2003: 126" authorityName=": Fujitani, Hirowatari &amp; Tanida" authorityPageNumber="126" authorityYear="2003" box="[222,762,317,341]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="genus">
<emphasis box="[222,338,317,341]" italics="true" pageId="6" pageNumber="207">Tenuibaetis</emphasis>
:
<bibRefCitation author="Fujitani, T. &amp; Hirowatari, T. &amp; Tanida, K." box="[352,762,317,341]" pageId="6" pageNumber="207" pagination="121 - 129" refId="ref17493" refString="Fujitani, T., Hirowatari, T. &amp; Tanida, K. (2003) Genera and species of Baetidae in Japan: Nigrobaetis, Alainites, Labiobaetis, and Tenuibaetis n. stat. (Ephemeroptera). Limnology, 4, 121 - 129. https: // doi. org / 10.1007 / s 10201 - 003 - 0105 - 2" type="journal article" year="2003">Fujitani, Hirowatari &amp; Tanida 2003: 126</bibRefCitation>
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(imago, larva).
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</paragraph>
</subSubSection>
<subSubSection box="[151,856,387,414]" pageId="6" pageNumber="207" type="type_taxon">
<paragraph blockId="6.[151,1437,387,2034]" box="[151,856,387,414]" pageId="6" pageNumber="207">
<emphasis bold="true" box="[151,309,387,413]" pageId="6" pageNumber="207">
<typeStatus box="[151,210,387,413]" pageId="6" pageNumber="207">Type</typeStatus>
species:
</emphasis>
<taxonomicName authority="Muller-Liebenau 1985" authorityName="Muller-Liebenau" authorityYear="1985" box="[316,851,387,414]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="pseudofrequentus">
<emphasis box="[316,589,387,413]" italics="true" pageId="6" pageNumber="207">Baetis pseudofrequentus</emphasis>
Müller-Liebenau 1985
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="207" type="diagnosis">
<paragraph blockId="6.[151,1437,387,2034]" pageId="6" pageNumber="207">
<emphasis bold="true" box="[199,322,423,449]" pageId="6" pageNumber="207">Diagnosis.</emphasis>
The subgenus
<taxonomicName authorityName=": Fujitani, Hirowatari &amp; Tanida" authorityYear="2003" box="[496,624,423,449]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="genus">
<emphasis box="[496,624,423,449]" italics="true" pageId="6" pageNumber="207">Tenuibaetis</emphasis>
</taxonomicName>
is characterized only by a combination of characters, each of which is found in some other taxa of Baetofemorata.
</paragraph>
<paragraph blockId="6.[151,1437,387,2034]" pageId="6" pageNumber="207">
<emphasis bold="true" box="[199,342,495,522]" pageId="6" pageNumber="207">
<emphasis bold="true" box="[199,333,495,522]" italics="true" pageId="6" pageNumber="207">Labial palp</emphasis>
:
</emphasis>
2nd segment is not significantly projected medially, 3rd segment is nearly symmetric, pointed, with both margins slightly convex (
<figureCitation box="[554,642,531,557]" captionStart="FIGURES 2230" captionStartId="8.[151,264,1852,1876]" captionTargetBox="[151,1436,181,1827]" captionTargetId="figure-36@8.[151,1436,181,1828]" captionTargetPageId="8" captionText="FIGURES 2230. Baetis (Tenuibaetis) ursinus, larval exuviae (2429, one and the same magnification). 22, labrum; 23, maxilla and labium; 24, abdomen, 2527, middle and hind legs; 28, half of pronotum and mesonotum; 29, head; 30, paraproct." figureDoi="http://doi.org/10.5281/zenodo.7889707" httpUri="https://zenodo.org/record/7889707/files/figure.png" pageId="6" pageNumber="207">Figs 23</figureCitation>
,
<figureCitation box="[656,687,531,557]" captionStart="FIGURES 4446" captionStartId="12.[151,264,1987,2011]" captionTargetBox="[169,1418,365,1963]" captionTargetId="figure-86@12.[169,1418,365,1963]" captionTargetPageId="12" captionText="FIGURES 4446. Baetis (Tenuibaetis) hissaricus, larval exuviae. 44, head; 45, labrum; 46, labium (44, 46, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889721" httpUri="https://zenodo.org/record/7889721/files/figure.png" pageId="6" pageNumber="207">46</figureCitation>
,
<figureCitation box="[699,729,531,557]" captionStart="FIGURES 7882" captionStartId="19.[151,264,1554,1578]" captionTargetBox="[158,1430,189,1531]" captionTargetId="figure-19@19.[158,1430,181,1531]" captionTargetPageId="19" captionText="FIGURES 7882. Baetis (Tenuibaetis) frequentus, larvae from India. 7879, left and right mandibles with mandibles of next instar developing inside; 8081, labrum of two individuals; 82, labium." figureDoi="http://doi.org/10.5281/zenodo.7889743" httpUri="https://zenodo.org/record/7889743/files/figure.png" pageId="6" pageNumber="207">82</figureCitation>
,
<figureCitation box="[742,786,531,558]" captionStart="FIGURES 154159" captionStartId="32.[151,264,1979,2003]" captionTargetBox="[151,1435,381,1956]" captionTargetId="figure-145@32.[151,1435,381,1956]" captionTargetPageId="32" captionText="FIGURES 154159. Baetis (Tenuibaetis) kaltenbachi sp. n., larval exuviae (holotype). 154155, labrum; 156158, labium; 159, posterior margin of abdominal tergum I." figureDoi="http://doi.org/10.5281/zenodo.7889773" httpUri="https://zenodo.org/record/7889773/files/figure.png" pageId="6" pageNumber="207">158</figureCitation>
,
<figureCitation box="[800,844,531,557]" captionStart="FIGURES 191194" captionStartId="38.[151,264,1802,1826]" captionTargetBox="[151,1430,181,1778]" captionTargetId="figure-36@38.[151,1436,181,1778]" captionTargetPageId="38" captionText="FIGURES 191194. Baetis (Tenuibaetis) bialatus sp. n., larval mouth parts. 191, labrum; 192, hypopharynx with superlinguae; 193, maxilla; 194, labium." figureDoi="http://doi.org/10.5281/zenodo.7889783" httpUri="https://zenodo.org/record/7889783/files/figure.png" pageId="6" pageNumber="207">194</figureCitation>
,
<figureCitation box="[857,902,531,558]" captionStart="FIGURES 249252" captionStartId="50.[151,264,1481,1505]" captionTargetBox="[151,1436,181,1457]" captionTargetId="figure-36@50.[151,1436,181,1457]" captionTargetPageId="50" captionText="FIGURES 249252. Baetis (Tenuibaetis) octomaculatus sp. n. 249, labrum (holotype); 250251, left and right mandibles; 252, maxilla and labium." figureDoi="http://doi.org/10.5281/zenodo.7889805" httpUri="https://zenodo.org/record/7889805/files/figure.png" pageId="6" pageNumber="207">252</figureCitation>
,
<figureCitation box="[914,960,531,557]" captionStart="FIGURES 279284" captionStartId="55.[151,264,1840,1864]" captionTargetBox="[151,1435,181,1816]" captionTargetId="figure-19@55.[151,1435,181,1816]" captionTargetPageId="55" captionText="FIGURES 279284. Baetis (Tenuibaetis) sp. (Thailand). 279, fore femur; 280, portion of larval exuviae with left mandible, labrum and frons; 281, labium; 282, male subimaginal abdomen extracted from larva; 283, larval abdominal segments IXX and caudalii; 284, exuviae of larval abdominal segments IVIII." figureDoi="http://doi.org/10.5281/zenodo.7889815" httpUri="https://zenodo.org/record/7889815/files/figure.png" pageId="6" pageNumber="207">281</figureCitation>
). The same in the group
<taxonomicName authorityName="Muller-Liebenau" authorityYear="1967" box="[1244,1319,531,557]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="lutheri">
<emphasis box="[1244,1319,531,557]" italics="true" pageId="6" pageNumber="207">lutheri</emphasis>
</taxonomicName>
of
<taxonomicName authorityName="Leach" authorityYear="1815" box="[1358,1428,532,557]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="genus">
<emphasis box="[1358,1428,532,557]" italics="true" pageId="6" pageNumber="207">Baetis</emphasis>
</taxonomicName>
; similar in
<taxonomicName authorityName="Ueno" authorityYear="1931" box="[265,366,567,593]" class="Insecta" family="Baetidae" genus="Baetiella" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="genus">
<emphasis box="[265,366,567,593]" italics="true" pageId="6" pageNumber="207">Baetiella</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="208" pageId="6" pageNumber="207" type="description">
<paragraph blockId="6.[151,1437,387,2034]" pageId="6" pageNumber="207">
<emphasis bold="true" box="[199,500,603,630]" pageId="6" pageNumber="207">
<emphasis bold="true" box="[199,491,603,630]" italics="true" pageId="6" pageNumber="207">Larval legs differentiated</emphasis>
:
</emphasis>
Femur of fore leg is wider than others, widest in proximal part; if cuticle of femur is pigmented, fore femur has large proximal blank lacking stout setae, while middle and hind femora have no such blank (
<figureCitation box="[230,362,675,702]" captionStart="FIGURES 2230" captionStartId="8.[151,264,1852,1876]" captionTargetBox="[151,1436,181,1827]" captionTargetId="figure-36@8.[151,1436,181,1828]" captionTargetPageId="8" captionText="FIGURES 2230. Baetis (Tenuibaetis) ursinus, larval exuviae (2429, one and the same magnification). 22, labrum; 23, maxilla and labium; 24, abdomen, 2527, middle and hind legs; 28, half of pronotum and mesonotum; 29, head; 30, paraproct." figureDoi="http://doi.org/10.5281/zenodo.7889707" httpUri="https://zenodo.org/record/7889707/files/figure.png" pageId="6" pageNumber="207">Figs 2527</figureCitation>
,
<figureCitation box="[375,448,675,701]" captionStart="FIGURES 4749" captionStartId="13.[151,264,1964,1988]" captionTargetBox="[151,1436,309,1941]" captionTargetId="figure-97@13.[151,1436,309,1941]" captionTargetPageId="13" captionText="FIGURES 4749. Baetis (Tenuibaetis) hissaricus, exuviae of larval fore, middle and hind femora. Abbreviation: 2-can, twochannel setae." figureDoi="http://doi.org/10.5281/zenodo.7889725" httpUri="https://zenodo.org/record/7889725/files/figure.png" pageId="6" pageNumber="207">4749</figureCitation>
,
<figureCitation box="[461,534,675,702]" captionStart="FIGURES 6977" captionStartId="18.[151,264,1844,1868]" captionTargetBox="[151,1425,181,1820]" captionTargetId="figure-36@18.[151,1436,181,1820]" captionTargetPageId="18" captionText="FIGURES 6977. Baetis (Tenuibaetis) frequentus, larval exuviae. 6975, from India (6974, at one magnification): 69, head; 7072, fore, middle and hind legs; 73, abdomen; 74, half of pronotum and mesonotum, 75, femur of dark individual. 7677, posterior margin of abdominal tergum VI of two individuals from Haputale (Sri Lanka)." figureDoi="http://doi.org/10.5281/zenodo.7889739" httpUri="https://zenodo.org/record/7889739/files/figure.png" pageId="6" pageNumber="207">7075</figureCitation>
,
<figureCitation box="[547,648,675,701]" captionStart="FIGURES 129134" captionStartId="28.[151,264,1712,1736]" captionTargetBox="[151,1436,185,1688]" captionTargetId="figure-36@28.[151,1436,181,1688]" captionTargetPageId="28" captionText="FIGURES 129134. Baetis (Tenuibaetis) fujitanii, exuviae of larval legs. 129131, fore, middle and hind legs; 132, anterior surface of hind femur; 133134, middle tibia." figureDoi="http://doi.org/10.5281/zenodo.7889763" httpUri="https://zenodo.org/record/7889763/files/figure.png" pageId="6" pageNumber="207">129131</figureCitation>
,
<figureCitation box="[661,763,675,701]" captionStart="FIGURES 160168" captionStartId="33.[151,264,1842,1866]" captionTargetBox="[151,1435,181,1818]" captionTargetId="figure-19@33.[151,1435,181,1818]" captionTargetPageId="33" captionText="FIGURES 160168. Baetis (Tenuibaetis) kaltenbachi sp. n., larval exuviae (holotype). 160162, femora of fore, middle and hind legs; 163164, tibiae of fore and middle legs; 165, fragment of fore femur; 166167, fragments of mid femur; 168, claw." figureDoi="http://doi.org/10.5281/zenodo.7889775" httpUri="https://zenodo.org/record/7889775/files/figure.png" pageId="6" pageNumber="207">160162</figureCitation>
,
<figureCitation box="[775,876,675,701]" captionStart="FIGURES 184190" captionStartId="37.[151,264,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-19@37.[151,1436,181,1721]" captionTargetPageId="37" captionText="FIGURES 184190. Baetis (Tenuibaetis) bialatus sp. n., larval exuviae (one and the same magnification). 184185, abdomen; 186, head with labrum; 187, half of pronotum and mesonotum; 188190, fore, middle and hind legs." figureDoi="http://doi.org/10.5281/zenodo.7889781" httpUri="https://zenodo.org/record/7889781/files/figure.png" pageId="6" pageNumber="207">188190</figureCitation>
,
<figureCitation box="[890,990,675,701]" captionStart="FIGURES 230236" captionStartId="46.[151,264,1840,1864]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-36@46.[151,1436,181,1817]" captionTargetPageId="46" captionText="FIGURES 230236. Baetis (Tenuibaetis) panhai, exuviae of larval legs. 230231, proximal-posterior portion of fore femur with villopore (230, anterior side; 231, posterior side); 232234, fore, middle and hind legs; 235236, fore and middle tibiae." figureDoi="http://doi.org/10.5281/zenodo.7889799" httpUri="https://zenodo.org/record/7889799/files/figure.png" pageId="6" pageNumber="207">232234</figureCitation>
,
<figureCitation box="[1004,1107,675,702]" captionStart="FIGURES 253256" captionStartId="51.[151,264,1450,1474]" captionTargetBox="[151,1436,181,1426]" captionTargetId="figure-19@51.[151,1436,181,1426]" captionTargetPageId="51" captionText="FIGURES 253256. Baetis (Tenuibaetis) octomaculatus sp. n. (holotype). 253255, larval exuviae of fore, middle and hind legs; 256, base of femur with villopore." figureDoi="http://doi.org/10.5281/zenodo.7889807" httpUri="https://zenodo.org/record/7889807/files/figure.png" pageId="6" pageNumber="207">253255</figureCitation>
). Among
<taxonomicName authorityName="Leach" authorityYear="1815" box="[1219,1289,676,701]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="genus">
<emphasis box="[1219,1289,676,701]" italics="true" pageId="6" pageNumber="207">Baetis</emphasis>
</taxonomicName>
, the same in the group
<taxonomicName authorityName="Muller-Liebenau" authorityYear="1967" box="[269,344,711,737]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="lutheri">
<emphasis box="[269,344,711,737]" italics="true" pageId="6" pageNumber="207">lutheri</emphasis>
</taxonomicName>
; among other mayflies, similar shape of femora and the proximal blank of fore femur occur in certain
<taxonomicName box="[235,417,747,773]" class="Insecta" family="Leptophlebiidae" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="family">Leptophlebiidae</taxonomicName>
(
<bibRefCitation author="Kluge, N. J." box="[433,569,747,773]" pageId="6" pageNumber="207" pagination="1 - 142" refId="ref18363" refString="Kluge, N. J. (2020) Systematic position of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae) and descriptions of new and little-known species. Zootaxa, 4756 (1), 1 - 142. https: // doi. org / 10.11646 / zootaxa. 4756.1.1" type="journal article" year="2020">Kluge 2020</bibRefCitation>
: fig. 9;
<bibRefCitation author="Kluge, N. &amp; Srinivasan, P. &amp; Vasanth, M. &amp; Sivaruban, T. &amp; Barathy, S. &amp; Isack, R." box="[654,986,747,774]" pageId="6" pageNumber="207" pagination="1 - 85" refId="ref18577" refString="Kluge, N., Srinivasan, P., Vasanth, M., Sivaruban, T., Barathy, S. &amp; Isack, R. (2022) Review of the subgenus Euthraulus (Ephemeroptera, Leptophlebiidae, genus Choroterpes) from the Western Ghats (India). Zootaxa, 5181 (1), 1 - 85. https: // doi. org / 10.11646 / zootaxa. 5181.1.1" type="journal article" year="2022">
Kluge, Srinivasan
<emphasis box="[862,912,747,773]" italics="true" pageId="6" pageNumber="207">et al</emphasis>
. 2022
</bibRefCitation>
: figs 6466;
<bibRefCitation author="Kluge, N. J. &amp; Vasanth, M. &amp; Balasubramanian, C. &amp; Sivaramakrishnan, K. G." box="[1131,1432,747,774]" pageId="6" pageNumber="207" pagination="1 - 140" refId="ref18653" refString="Kluge, N. J., Vasanth, M., Balasubramanian, C. &amp; Sivaramakrishnan, K. G. (2022) Review of the Kimminsula - complex (Ephemeroptera, Leptophlebiidae). Zootaxa, 5212 (1), 1 - 140. https: // doi. org / 10.11646 / zootaxa. 5212.1.1" type="journal article" year="2022">
Kluge, Vasanth
<emphasis box="[1309,1358,747,773]" italics="true" pageId="6" pageNumber="207">et al</emphasis>
. 2022
</bibRefCitation>
: figs 2729).
</paragraph>
<paragraph blockId="6.[151,1437,387,2034]" pageId="6" pageNumber="207">
<emphasis bold="true" box="[199,520,819,846]" italics="true" pageId="6" pageNumber="207">Stout setae on larval femora</emphasis>
(either two-channel, or pseudo-bifurcate) are located not only near outer and inner sides, but also on middle area of anterior (dorsal) side (
<figureCitation box="[778,865,855,881]" captionStart="FIGURES 3132" captionStartId="9.[151,264,1844,1868]" captionTargetBox="[151,1436,184,1821]" captionTargetId="figure-19@9.[151,1436,181,1821]" captionTargetPageId="9" captionText="FIGURES 3132. Baetis (Tenuibaetis) ursinus, larval exuviae of fore leg. 31, part of femur; 32, part of tibia. Abbreviation: 2-can, two-channel setae." figureDoi="http://doi.org/10.5281/zenodo.7889711" httpUri="https://zenodo.org/record/7889711/files/figure.png" pageId="6" pageNumber="207">Figs 31</figureCitation>
,
<figureCitation box="[878,951,855,881]" captionStart="FIGURES 4749" captionStartId="13.[151,264,1964,1988]" captionTargetBox="[151,1436,309,1941]" captionTargetId="figure-97@13.[151,1436,309,1941]" captionTargetPageId="13" captionText="FIGURES 4749. Baetis (Tenuibaetis) hissaricus, exuviae of larval fore, middle and hind femora. Abbreviation: 2-can, twochannel setae." figureDoi="http://doi.org/10.5281/zenodo.7889725" httpUri="https://zenodo.org/record/7889725/files/figure.png" pageId="6" pageNumber="207">4749</figureCitation>
,
<figureCitation box="[963,1035,855,881]" captionStart="FIGURES 8388" captionStartId="20.[151,264,1838,1862]" captionTargetBox="[151,1436,181,1815]" captionTargetId="figure-36@20.[151,1436,181,1815]" captionTargetPageId="20" captionText="FIGURES 8388. Baetis (Tenuibaetis) frequentus from Sri Lanka, setae on anterior side of femur. 83, pseudo-bifurcate setae and one long two-channel seta in individual from Sri Pada; 8485, fore and middle femora of another individual from Sri Pada, with all setae clavate and two-channel; 86, enlarged fragment of fore femur of the same individual; 8788, hind femur of individual from Haputale, with all setae short and pseudo-bifurcate." figureDoi="http://doi.org/10.5281/zenodo.7889747" httpUri="https://zenodo.org/record/7889747/files/figure.png" pageId="6" pageNumber="207">8388</figureCitation>
,
<figureCitation box="[1048,1149,855,881]" captionStart="FIGURES 129134" captionStartId="28.[151,264,1712,1736]" captionTargetBox="[151,1436,185,1688]" captionTargetId="figure-36@28.[151,1436,181,1688]" captionTargetPageId="28" captionText="FIGURES 129134. Baetis (Tenuibaetis) fujitanii, exuviae of larval legs. 129131, fore, middle and hind legs; 132, anterior surface of hind femur; 133134, middle tibia." figureDoi="http://doi.org/10.5281/zenodo.7889763" httpUri="https://zenodo.org/record/7889763/files/figure.png" pageId="6" pageNumber="207">129132</figureCitation>
,
<figureCitation box="[1161,1262,855,881]" captionStart="FIGURES 160168" captionStartId="33.[151,264,1842,1866]" captionTargetBox="[151,1435,181,1818]" captionTargetId="figure-19@33.[151,1435,181,1818]" captionTargetPageId="33" captionText="FIGURES 160168. Baetis (Tenuibaetis) kaltenbachi sp. n., larval exuviae (holotype). 160162, femora of fore, middle and hind legs; 163164, tibiae of fore and middle legs; 165, fragment of fore femur; 166167, fragments of mid femur; 168, claw." figureDoi="http://doi.org/10.5281/zenodo.7889775" httpUri="https://zenodo.org/record/7889775/files/figure.png" pageId="6" pageNumber="207">160162</figureCitation>
,
<figureCitation box="[1274,1319,855,881]" captionStart="FIGURES 202207" captionStartId="40.[151,264,1840,1864]" captionTargetBox="[151,1396,181,1811]" captionTargetId="figure-36@40.[151,1436,181,1816]" captionTargetPageId="40" captionText="FIGURES 202207. Baetis (Tenuibaetis) bialatus sp. n., larval legs. 202, middle femur (anterior side); 203, fore tibia (anterior side); 204, middle tibia (anterior side); 205, middle tibia (outer side); 206207, tibia and tarsus of fore and middle legs (anterior side) (202204, 206207, holotype). Abbreviations: 2-can, two-channel seta." figureDoi="http://doi.org/10.5281/zenodo.7889787" httpUri="https://zenodo.org/record/7889787/files/figure.png" pageId="6" pageNumber="207">202</figureCitation>
,
<figureCitation box="[1331,1432,855,881]" captionStart="FIGURES 230236" captionStartId="46.[151,264,1840,1864]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-36@46.[151,1436,181,1817]" captionTargetPageId="46" captionText="FIGURES 230236. Baetis (Tenuibaetis) panhai, exuviae of larval legs. 230231, proximal-posterior portion of fore femur with villopore (230, anterior side; 231, posterior side); 232234, fore, middle and hind legs; 235236, fore and middle tibiae." figureDoi="http://doi.org/10.5281/zenodo.7889799" httpUri="https://zenodo.org/record/7889799/files/figure.png" pageId="6" pageNumber="207">232234</figureCitation>
,
<figureCitation box="[151,252,891,918]" captionStart-0="FIGURES 253256" captionStart-1="FIGURES 257264" captionStart-2="FIGURES 265274" captionStartId-0="51.[151,264,1450,1474]" captionStartId-1="52.[151,264,1879,1903]" captionStartId-2="53.[151,264,1843,1867]" captionTargetBox-0="[151,1436,181,1426]" captionTargetBox-1="[151,1435,181,1855]" captionTargetBox-2="[151,1435,181,1819]" captionTargetId-0="figure-19@51.[151,1436,181,1426]" captionTargetId-1="figure-36@52.[151,1435,181,1855]" captionTargetId-2="figure-19@53.[151,1435,181,1819]" captionTargetPageId-0="51" captionTargetPageId-1="52" captionTargetPageId-2="53" captionText-0="FIGURES 253256. Baetis (Tenuibaetis) octomaculatus sp. n. (holotype). 253255, larval exuviae of fore, middle and hind legs; 256, base of femur with villopore." captionText-1="FIGURES 257264. Baetis (Tenuibaetis) octomaculatus sp. n. 257258, fore and hind femora; 259260, fore and middle tibiae; 261263, abdominal terga I, VII and X; 264, abdominal sternum IX and paraprocts (257260, holotype)." captionText-2="FIGURES 265274. Baetis (Tenuibaetis) octomaculatus sp. n. 265, subimaginal exuviae of half of mesonotum; 266, male imago; 267, female imago; 268269, fore legs of two female imagines (268, holotype); 270, female imago; 271, male subimago; 272, abdomen of male imago; 273, fore leg of male imago; 274, its subimaginal exuviae." figureDoi-0="http://doi.org/10.5281/zenodo.7889807" figureDoi-1="http://doi.org/10.5281/zenodo.7889809" figureDoi-2="http://doi.org/10.5281/zenodo.7889811" httpUri-0="https://zenodo.org/record/7889807/files/figure.png" httpUri-1="https://zenodo.org/record/7889809/files/figure.png" httpUri-2="https://zenodo.org/record/7889811/files/figure.png" pageId="6" pageNumber="207">253266</figureCitation>
,
<figureCitation box="[266,312,891,917]" captionStart="FIGURES 279284" captionStartId="55.[151,264,1840,1864]" captionTargetBox="[151,1435,181,1816]" captionTargetId="figure-19@55.[151,1435,181,1816]" captionTargetPageId="55" captionText="FIGURES 279284. Baetis (Tenuibaetis) sp. (Thailand). 279, fore femur; 280, portion of larval exuviae with left mandible, labrum and frons; 281, labium; 282, male subimaginal abdomen extracted from larva; 283, larval abdominal segments IXX and caudalii; 284, exuviae of larval abdominal segments IVIII." figureDoi="http://doi.org/10.5281/zenodo.7889815" httpUri="https://zenodo.org/record/7889815/files/figure.png" pageId="6" pageNumber="207">279</figureCitation>
). The same in many other mayflies, including certain species of
<taxonomicName authorityName="Leach" authorityYear="1815" box="[1062,1132,892,917]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="genus">
<emphasis box="[1062,1132,892,917]" italics="true" pageId="6" pageNumber="207">Baetis</emphasis>
</taxonomicName>
groups
<emphasis box="[1227,1302,892,917]" italics="true" pageId="6" pageNumber="207">vernus</emphasis>
,
<emphasis box="[1319,1429,891,917]" italics="true" pageId="6" pageNumber="207">buceratus</emphasis>
, subgenus
<taxonomicName box="[262,402,927,953]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="subGenus" subGenus="Rhodobaetis">
<emphasis box="[262,402,927,953]" italics="true" pageId="6" pageNumber="207">Rhodobaetis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph blockId="6.[151,1437,387,2034]" pageId="6" pageNumber="207">
<emphasis bold="true" box="[199,601,963,990]" pageId="6" pageNumber="207">
<emphasis bold="true" box="[199,592,963,990]" italics="true" pageId="6" pageNumber="207">Scales on larval abdominal terga</emphasis>
:
</emphasis>
Wide, in semicircular sockets (
<figureCitation box="[969,1131,963,989]" captionStart="FIGURES 195201" captionStartId="39.[151,264,1840,1864]" captionTargetBox="[151,1436,190,1817]" captionTargetId="figure-19@39.[151,1436,181,1817]" captionTargetPageId="39" captionText="FIGURES 195201. Baetis (Tenuibaetis) bialatus sp. n., larvae. 195196, left and right mandibles with developing mandibles of next instar inside; 197198, the same of another individual; 199, exuviae of abdominal tergum IX; 200, abdominal tergum II; 201, paraprocts. Abbreviations: hirud, hirudiform seta; sq, scale." figureDoi="http://doi.org/10.5281/zenodo.7889785" httpUri="https://zenodo.org/record/7889785/files/figure.png" pageId="6" pageNumber="207">Figs 199200</figureCitation>
). The same in many other
<taxonomicName authorityName="Leach" authorityYear="1815" box="[151,221,1000,1025]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="genus">
<emphasis box="[151,221,1000,1025]" italics="true" pageId="6" pageNumber="207">Baetis</emphasis>
</taxonomicName>
, but not in the group
<taxonomicName authorityName="Muller-Liebenau" authorityYear="1967" box="[463,538,999,1025]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="lutheri">
<emphasis box="[463,538,999,1025]" italics="true" pageId="6" pageNumber="207">lutheri</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph blockId="6.[151,1437,387,2034]" pageId="6" pageNumber="207">
<emphasis bold="true" box="[199,297,1035,1062]" pageId="6" pageNumber="207">
<emphasis bold="true" box="[199,288,1035,1062]" italics="true" pageId="6" pageNumber="207">Tergalii</emphasis>
:
</emphasis>
All seven pairs present, oval; tergalius I often much smaller than others (
<figureCitation box="[1111,1197,1035,1061]" captionStart="FIGURES 6977" captionStartId="18.[151,264,1844,1868]" captionTargetBox="[151,1425,181,1820]" captionTargetId="figure-36@18.[151,1436,181,1820]" captionTargetPageId="18" captionText="FIGURES 6977. Baetis (Tenuibaetis) frequentus, larval exuviae. 6975, from India (6974, at one magnification): 69, head; 7072, fore, middle and hind legs; 73, abdomen; 74, half of pronotum and mesonotum, 75, femur of dark individual. 7677, posterior margin of abdominal tergum VI of two individuals from Haputale (Sri Lanka)." figureDoi="http://doi.org/10.5281/zenodo.7889739" httpUri="https://zenodo.org/record/7889739/files/figure.png" pageId="6" pageNumber="207">Figs 73</figureCitation>
,
<figureCitation box="[1209,1253,1035,1061]" captionStart="FIGURES 184190" captionStartId="37.[151,264,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-19@37.[151,1436,181,1721]" captionTargetPageId="37" captionText="FIGURES 184190. Baetis (Tenuibaetis) bialatus sp. n., larval exuviae (one and the same magnification). 184185, abdomen; 186, head with labrum; 187, half of pronotum and mesonotum; 188190, fore, middle and hind legs." figureDoi="http://doi.org/10.5281/zenodo.7889781" httpUri="https://zenodo.org/record/7889781/files/figure.png" pageId="6" pageNumber="207">184</figureCitation>
,
<figureCitation box="[1264,1310,1035,1061]" captionStart="FIGURES 237241" captionStartId="47.[151,264,1838,1862]" captionTargetBox="[152,1435,181,1814]" captionTargetId="figure-19@47.[151,1435,181,1814]" captionTargetPageId="47" captionText="FIGURES 237241. Baetis (Tenuibaetis) panhai. 237, abdomen of male subimago extracted from mature larva; 238, larval exuviae of half of pronotum and mesonotum; 239, larval exuviae of abdomen; 240, male subimago; 241, subimaginal gonostyli crumpled under larval cuticle." figureDoi="http://doi.org/10.5281/zenodo.7889801" httpUri="https://zenodo.org/record/7889801/files/figure.png" pageId="6" pageNumber="207">239</figureCitation>
). The same in most other
<taxonomicName authorityName="Leach" authorityYear="1815" box="[306,376,1072,1097]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis box="[306,376,1072,1097]" italics="true" pageId="6" pageNumber="207">Baetis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[383,423,1071,1098]" pageId="6" pageNumber="207" sensu="lato">s. l.</taxonomicNameLabel>
In
<taxonomicName box="[460,683,1071,1097]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="subGenus" subGenus="Tenuibaetis">
<emphasis box="[460,530,1072,1097]" italics="true" pageId="6" pageNumber="207">Baetis</emphasis>
(
<emphasis box="[546,674,1071,1097]" italics="true" pageId="6" pageNumber="207">Tenuibaetis</emphasis>
)
</taxonomicName>
sp. from
<collectingCountry box="[790,892,1071,1097]" name="Thailand" pageId="6" pageNumber="207">Thailand</collectingCountry>
, the first tergalius is relatively large (
<figureCitation box="[1309,1409,1071,1097]" captionStart="FIGURES 279284" captionStartId="55.[151,264,1840,1864]" captionTargetBox="[151,1435,181,1816]" captionTargetId="figure-19@55.[151,1435,181,1816]" captionTargetPageId="55" captionText="FIGURES 279284. Baetis (Tenuibaetis) sp. (Thailand). 279, fore femur; 280, portion of larval exuviae with left mandible, labrum and frons; 281, labium; 282, male subimaginal abdomen extracted from larva; 283, larval abdominal segments IXX and caudalii; 284, exuviae of larval abdominal segments IVIII." figureDoi="http://doi.org/10.5281/zenodo.7889815" httpUri="https://zenodo.org/record/7889815/files/figure.png" pageId="6" pageNumber="207">Fig. 284</figureCitation>
).
</paragraph>
<paragraph blockId="6.[151,1437,387,2034]" pageId="6" pageNumber="207">
<emphasis bold="true" box="[199,339,1107,1134]" pageId="6" pageNumber="207">
<emphasis bold="true" box="[199,332,1107,1134]" italics="true" pageId="6" pageNumber="207">Larval pose</emphasis>
.
</emphasis>
When swimming, larva does not stretch its legs along the body, but after short swimming passively falls down with its legs bent down and abdomen often bent on back (
<figureCitation box="[928,1027,1143,1169]" captionStart="FIGURES 125128" captionStartId="27.[151,264,1955,1979]" captionTargetBox="[151,1436,345,1928]" captionTargetId="figure-113@27.[151,1436,345,1932]" captionTargetPageId="27" captionText="FIGURES 125128. Baetis (Tenuibaetis) fujitanii. 125127, larval exuviae: 125, head; 126, half of pronotum and mesonotum; 127, abdomen; 128, young larva in natural pose." figureDoi="http://doi.org/10.5281/zenodo.7889761" httpUri="https://zenodo.org/record/7889761/files/figure.png" pageId="6" pageNumber="207">Fig. 128</figureCitation>
); larva of
<taxonomicName authorityName="Novikova" authorityYear="1991" box="[1142,1336,1143,1170]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="217" phylum="Arthropoda" rank="species" species="hissaricus" subGenus="Tenuibaetis">
<emphasis box="[1142,1159,1144,1169]" italics="true" pageId="6" pageNumber="207">B</emphasis>
. (
<emphasis box="[1183,1199,1144,1169]" italics="true" pageId="6" pageNumber="207">T</emphasis>
.)
<emphasis box="[1222,1336,1143,1169]" italics="true" pageId="6" pageNumber="207">hissaricus</emphasis>
</taxonomicName>
bents its legs, but abdomen retains straight (
<bibRefCitation author="Kluge, N. J." box="[546,680,1179,1205]" pageId="6" pageNumber="207" pagination="53 - 68" refId="ref18000" refString="Kluge, N. J. (1983) New and little known mayflies of the family Baetidae (Ephemeroptera) from Primorye Territory. Entomologicheskoe Obozrenie = Revue d'Entomologie de l'URSS, 61 (1), 65 - 79. [in Russian, Entomological Review, 62 (1), 53 - 68, English translation]" type="journal article" year="1983">Kluge 1983</bibRefCitation>
,
<bibRefCitation author="Novikova, E. A." box="[692,869,1179,1205]" pageId="6" pageNumber="207" pagination="793 - 795" refId="ref19156" refString="Novikova, E. A. (1991) [A new subspecies of mayflies Baetis ursinus hissaricus subsp. n. (Ephemeroptera, Baetidae) from Tadjikistan]. Entomologicheskoe Obozrenie, 70 (4), 793 - 795." type="journal article" year="1991">Novikova 1991</bibRefCitation>
). The same in
<taxonomicName authorityName="Bengtsson" authorityYear="1912" box="[1033,1149,1179,1205]" class="Insecta" family="Baetidae" genus="Acentrella" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="genus">
<emphasis box="[1033,1149,1179,1205]" italics="true" pageId="6" pageNumber="207">Acentrella</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Ueno" authorityYear="1931" box="[1163,1264,1179,1205]" class="Insecta" family="Baetidae" genus="Baetiella" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="genus">
<emphasis box="[1163,1264,1179,1205]" italics="true" pageId="6" pageNumber="207">Baetiella</emphasis>
</taxonomicName>
and some nonrelated mayflies (e.g.
<taxonomicName authorityName="Klapalek" authorityYear="1909" box="[393,574,1215,1241]" class="Insecta" family="Ephemerellidae" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="family">Ephemerellidae</taxonomicName>
), in contrast to majority of
<taxonomicName authorityName="Leach" authorityYear="1815" box="[873,943,1216,1241]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="genus">
<emphasis box="[873,943,1216,1241]" italics="true" pageId="6" pageNumber="207">Baetis</emphasis>
</taxonomicName>
, which retain ability to swim stretching legs backward along the body.
</paragraph>
<paragraph blockId="6.[151,1437,387,2034]" pageId="6" pageNumber="207">
<emphasis bold="true" box="[199,519,1287,1314]" pageId="6" pageNumber="207">
<emphasis bold="true" box="[199,510,1287,1314]" italics="true" pageId="6" pageNumber="207">Larval cuticular coloration</emphasis>
:
</emphasis>
Varies from nearly uniformly brown (
<figureCitation box="[940,1039,1287,1313]" captionStart="FIGURES 184190" captionStartId="37.[151,264,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-19@37.[151,1436,181,1721]" captionTargetPageId="37" captionText="FIGURES 184190. Baetis (Tenuibaetis) bialatus sp. n., larval exuviae (one and the same magnification). 184185, abdomen; 186, head with labrum; 187, half of pronotum and mesonotum; 188190, fore, middle and hind legs." figureDoi="http://doi.org/10.5281/zenodo.7889781" httpUri="https://zenodo.org/record/7889781/files/figure.png" pageId="6" pageNumber="207">Figs 184</figureCitation>
) or diffusively maculated, to brown with contrasting V-shaped or transverse blank on mesonotum (
<figureCitation box="[855,937,1323,1349]" captionStart="FIGURES 2230" captionStartId="8.[151,264,1852,1876]" captionTargetBox="[151,1436,181,1827]" captionTargetId="figure-36@8.[151,1436,181,1828]" captionTargetPageId="8" captionText="FIGURES 2230. Baetis (Tenuibaetis) ursinus, larval exuviae (2429, one and the same magnification). 22, labrum; 23, maxilla and labium; 24, abdomen, 2527, middle and hind legs; 28, half of pronotum and mesonotum; 29, head; 30, paraproct." figureDoi="http://doi.org/10.5281/zenodo.7889707" httpUri="https://zenodo.org/record/7889707/files/figure.png" pageId="6" pageNumber="207">Fig. 28</figureCitation>
) and contrasting blanks on abdomen; in this case abdominal tergum IV is lighter then neighboring ones (
<figureCitation box="[812,895,1359,1385]" captionStart="FIGURES 2230" captionStartId="8.[151,264,1852,1876]" captionTargetBox="[151,1436,181,1827]" captionTargetId="figure-36@8.[151,1436,181,1828]" captionTargetPageId="8" captionText="FIGURES 2230. Baetis (Tenuibaetis) ursinus, larval exuviae (2429, one and the same magnification). 22, labrum; 23, maxilla and labium; 24, abdomen, 2527, middle and hind legs; 28, half of pronotum and mesonotum; 29, head; 30, paraproct." figureDoi="http://doi.org/10.5281/zenodo.7889707" httpUri="https://zenodo.org/record/7889707/files/figure.png" pageId="6" pageNumber="207">Fig. 24</figureCitation>
). The species name of the first described species,
<taxonomicName authorityName=": Novikova" authorityYear="1991" box="[151,311,1396,1421]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="ursinus">
<emphasis box="[151,311,1396,1421]" italics="true" pageId="6" pageNumber="207">Baetis ursinus</emphasis>
</taxonomicName>
, was given as an allusion to the similarity of its dorsal V-shaped blank to the ventral white marking of the Asian bear (
<taxonomicName authority="Cuv." authorityName="Cuv." box="[358,609,1431,1457]" class="Mammalia" family="Ursidae" genus="Ursus" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="207" phylum="Chordata" rank="species" species="thibetanus">
<emphasis box="[358,549,1431,1457]" italics="true" pageId="6" pageNumber="207">Ursus thibetanus</emphasis>
Cuv.
</taxonomicName>
)
</paragraph>
<paragraph blockId="6.[151,1437,387,2034]" pageId="6" pageNumber="207">
<emphasis bold="true" box="[199,316,1467,1494]" italics="true" pageId="6" pageNumber="207">Gonostyli.</emphasis>
Male imaginal gonostylus with terminal (3rd) segment short (
<figureCitation box="[1011,1141,1467,1493]" captionStart="FIGURES 4243" captionStartId="11.[151,264,1983,2007]" captionTargetBox="[151,1436,1065,1959]" captionTargetId="figure-414@11.[151,1436,1065,1959]" captionTargetPageId="11" captionText="FIGURES 4243. Baetis (Tenuibaetis) ursinus, genitalia of male imagines. Abbreviations: 1, 2, 3, segments of gonostylus; gd, gonoduct; gv, gonovectis; m.gs, gonostylar muscle; m.gv, muscle of gonovectis; s.s, sterno-styligeral muscle; us, unistyliger." figureDoi="http://doi.org/10.5281/zenodo.7889719" httpUri="https://zenodo.org/record/7889719/files/figure.png" pageId="6" pageNumber="207">Figs 4243</figureCitation>
,
<figureCitation box="[1152,1224,1467,1494]" captionStart="FIGURES 5259" captionStartId="15.[151,264,1834,1858]" captionTargetBox="[151,1435,181,1810]" captionTargetId="figure-19@15.[151,1435,181,1810]" captionTargetPageId="15" captionText="FIGURES 5259. Baetis (Tenuibaetis) hissaricus. 5253, fore and middle legs of male imago; 5455, genitalia of males imago; 56, hind wing of male; 57, hind wing of female; 58, subimaginal exuviae of female fore leg; 59, subimaginal exuviae of half of mesonotum (57, 58, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889731" httpUri="https://zenodo.org/record/7889731/files/figure.png" pageId="6" pageNumber="207">5455</figureCitation>
,
<figureCitation box="[1236,1321,1467,1493]" captionStart="FIGURES 92101" captionStartId="22.[151,264,1946,1970]" captionTargetBox="[162,1428,309,1923]" captionTargetId="figure-90@22.[159,1428,309,1923]" captionTargetPageId="22" captionText="FIGURES 92101. Baetis (Tenuibaetis) frequentus from Sri Lanka. 92, female imago; 9394, male imago from Sri Pada and its fore leg; 9598, fore leg, head, abdomen and genitalia of male imago from Haputale; 99101, male imago from Uma, its fore leg and genitalia." figureDoi="http://doi.org/10.5281/zenodo.7889751" httpUri="https://zenodo.org/record/7889751/files/figure.png" pageId="6" pageNumber="207">98, 101</figureCitation>
,
<figureCitation box="[1333,1432,1467,1493]" captionStart="FIGURES 112115" captionStartId="24.[151,264,1718,1742]" captionTargetBox="[151,1436,705,1694]" captionTargetId="figure-271@24.[151,1436,705,1695]" captionTargetPageId="24" captionText="FIGURES 112115. Baetis (Tenuibaetis) frequentus from India, genitalia of male imago. Abbreviations: gv, gonovectus; m.gv, muscle of gonovectis; s.s, vestige of unpaired sterno-styligeral muscle." figureDoi="http://doi.org/10.5281/zenodo.7889757" httpUri="https://zenodo.org/record/7889757/files/figure.png" pageId="6" pageNumber="207">112114</figureCitation>
,
<figureCitation box="[151,252,1503,1530]" captionStart="FIGURES 144145" captionStartId="30.[151,264,1131,1155]" captionTargetBox="[151,1436,181,1107]" captionTargetId="figure-36@30.[151,1436,181,1107]" captionTargetPageId="30" captionText="FIGURES 144145. Baetis (Tenuibaetis) fujitanii, genitalia of male imagines." figureDoi="http://doi.org/10.5281/zenodo.7889767" httpUri="https://zenodo.org/record/7889767/files/figure.png" pageId="6" pageNumber="207">144145</figureCitation>
,
<figureCitation box="[263,364,1503,1529]" captionStart="FIGURES 181183" captionStartId="35.[151,264,2000,2024]" captionTargetBox="[151,1436,777,1976]" captionTargetId="figure-290@35.[151,1436,777,1976]" captionTargetPageId="35" captionText="FIGURES 181183. Baetis (Tenuibaetis) kaltenbachi sp. n., genitalia of male imago (holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889779" httpUri="https://zenodo.org/record/7889779/files/figure.png" pageId="6" pageNumber="207">181183</figureCitation>
,
<figureCitation box="[375,476,1503,1530]" captionStart="FIGURES 225229" captionStartId="44.[151,264,1840,1864]" captionTargetBox="[151,1436,185,1817]" captionTargetId="figure-36@44.[151,1436,181,1817]" captionTargetPageId="44" captionText="FIGURES 225229. Baetis (Tenuibaetis) bialatus sp. n., genitalia of male imago. 225, 227, ventral view; 226, 229, ventral view with focus on sterno-styligeral muscle; 228, enlarged fragment of sterno-styligeral muscle to show cross-striation (225226, holotype). Abbreviations: gv, gonovectis; s.s, sterno-styligeral muscle." figureDoi="http://doi.org/10.5281/zenodo.7889797" httpUri="https://zenodo.org/record/7889797/files/figure.png" pageId="6" pageNumber="207">225, 227</figureCitation>
,
<figureCitation box="[487,591,1503,1529]" captionStart="FIGURES 275278" captionStartId="54.[151,264,1584,1608]" captionTargetBox="[151,1436,181,1560]" captionTargetId="figure-36@54.[151,1436,181,1560]" captionTargetPageId="54" captionText="FIGURES 275278. Baetis (Tenuibaetis) octomaculatus sp. n. 275, subimaginal gonostyli crumpled under larval cuticle; 276, imaginal genitalia; 277, imaginal genitalia of another individual, with focus on ventral side; 278, the same, with focus on gonovectes. Abbreviation: s.s, sterno-styligeral muscle." figureDoi="http://doi.org/10.5281/zenodo.7889813" httpUri="https://zenodo.org/record/7889813/files/figure.png" pageId="6" pageNumber="207">276277</figureCitation>
).
</paragraph>
<paragraph blockId="6.[151,1437,387,2034]" pageId="6" pageNumber="207">
<emphasis bold="true" box="[199,488,1539,1566]" pageId="6" pageNumber="207">
<emphasis bold="true" box="[199,479,1539,1566]" italics="true" pageId="6" pageNumber="207">Sterno-styligeral muscle</emphasis>
:
</emphasis>
Varies from vestigial with remnants of cross striation (
<figureCitation box="[1110,1267,1539,1565]" captionStart="FIGURES 225229" captionStartId="44.[151,264,1840,1864]" captionTargetBox="[151,1436,185,1817]" captionTargetId="figure-36@44.[151,1436,181,1817]" captionTargetPageId="44" captionText="FIGURES 225229. Baetis (Tenuibaetis) bialatus sp. n., genitalia of male imago. 225, 227, ventral view; 226, 229, ventral view with focus on sterno-styligeral muscle; 228, enlarged fragment of sterno-styligeral muscle to show cross-striation (225226, holotype). Abbreviations: gv, gonovectis; s.s, sterno-styligeral muscle." figureDoi="http://doi.org/10.5281/zenodo.7889797" httpUri="https://zenodo.org/record/7889797/files/figure.png" pageId="6" pageNumber="207">Figs 228229</figureCitation>
) to completely absent (
<figureCitation box="[238,336,1575,1602]" captionStart="FIGURES 144145" captionStartId="30.[151,264,1131,1155]" captionTargetBox="[151,1436,181,1107]" captionTargetId="figure-36@30.[151,1436,181,1107]" captionTargetPageId="30" captionText="FIGURES 144145. Baetis (Tenuibaetis) fujitanii, genitalia of male imagines." figureDoi="http://doi.org/10.5281/zenodo.7889767" httpUri="https://zenodo.org/record/7889767/files/figure.png" pageId="6" pageNumber="207">Fig. 145</figureCitation>
).
</paragraph>
<paragraph blockId="6.[151,1437,387,2034]" pageId="6" pageNumber="207">
<emphasis bold="true" box="[199,327,1611,1638]" pageId="6" pageNumber="207">
<emphasis bold="true" box="[199,320,1611,1638]" italics="true" pageId="6" pageNumber="207">Hind wing</emphasis>
.
</emphasis>
Hind wing, if present, with two longitudinal veins and short costal projection (
<figureCitation box="[1200,1282,1611,1637]" captionStart="FIGURES 1321" captionStartId="5.[151,264,1840,1864]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure-19@5.[151,1436,181,1816]" captionTargetPageId="5" captionText="FIGURES 1321. Imaginal hind wings and larval protoptera (under the same magnification). 1314, female Baetis (Tenuibaetis) frequentus from India; 1517, male B. (T.) octomaculatus sp. n.; 1819, female of same species; 2021, male B. (T.) bialatus sp. n. (13, 15, 20, metathorax of imago with hind wing; 14, 21, larval exuviae of metanotum; 16, 18, hind wing of imago; 17, 19, cuticle from metathorax of larva ready to molt to subimago, with subimaginal wing inside protopteron: crumpled on Fig. 17, non-crumpled on Fig. 19)." figureDoi="http://doi.org/10.5281/zenodo.7889705" httpUri="https://zenodo.org/record/7889705/files/figure.png" pageId="6" pageNumber="207">Fig. 13</figureCitation>
). The same in many other
<taxonomicName authorityName="Leach" authorityYear="1815" box="[285,355,1648,1673]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="genus">
<emphasis box="[285,355,1648,1673]" italics="true" pageId="6" pageNumber="207">Baetis</emphasis>
</taxonomicName>
, in contrast to representatives having three veins or lacking costal projection.
</paragraph>
<paragraph blockId="6.[151,1437,387,2034]" pageId="6" pageNumber="207">
<emphasis bold="true" box="[199,507,1683,1709]" pageId="6" pageNumber="207">Variability of hind wings.</emphasis>
Most species of
<taxonomicName authorityName=": Fujitani, Hirowatari &amp; Tanida" authorityYear="2003" box="[703,831,1683,1709]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="genus">
<emphasis box="[703,831,1683,1709]" italics="true" pageId="6" pageNumber="207">Tenuibaetis</emphasis>
</taxonomicName>
have hind wings as large as in most other species of
<taxonomicName authorityName="Leach" authorityYear="1815" box="[151,221,1720,1745]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis box="[151,221,1720,1745]" italics="true" pageId="6" pageNumber="207">Baetis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[231,275,1719,1746]" pageId="6" pageNumber="207" sensu="lato">s. l.</taxonomicNameLabel>
; in this case larval protoptera are also well-developed (
<figureCitation box="[917,1050,1719,1745]" captionStart="FIGURES 1321" captionStartId="5.[151,264,1840,1864]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure-19@5.[151,1436,181,1816]" captionTargetPageId="5" captionText="FIGURES 1321. Imaginal hind wings and larval protoptera (under the same magnification). 1314, female Baetis (Tenuibaetis) frequentus from India; 1517, male B. (T.) octomaculatus sp. n.; 1819, female of same species; 2021, male B. (T.) bialatus sp. n. (13, 15, 20, metathorax of imago with hind wing; 14, 21, larval exuviae of metanotum; 16, 18, hind wing of imago; 17, 19, cuticle from metathorax of larva ready to molt to subimago, with subimaginal wing inside protopteron: crumpled on Fig. 17, non-crumpled on Fig. 19)." figureDoi="http://doi.org/10.5281/zenodo.7889705" httpUri="https://zenodo.org/record/7889705/files/figure.png" pageId="6" pageNumber="207">Figs 1314</figureCitation>
). In female, hind wings are often somewhat smaller or at least narrower than in male (
<figureCitation box="[738,868,1755,1782]" captionStart="FIGURES 3341" captionStartId="10.[151,264,1788,1812]" captionTargetBox="[151,1435,181,1764]" captionTargetId="figure-36@10.[151,1435,181,1764]" captionTargetPageId="10" captionText="FIGURES 3341. Baetis (Tenuibaetis) ursinus. 33, subimaginal exuviae of half of mesonotum; 34, hind wing of male; 35, hind wing of female; 36, male imago and its spread abdomen; 3738, its fore and middle legs; 3940, their subimaginal exuviae (the same magnification); 41, female imago." figureDoi="http://doi.org/10.5281/zenodo.7889715" httpUri="https://zenodo.org/record/7889715/files/figure.png" pageId="6" pageNumber="207">Figs 3435</figureCitation>
,
<figureCitation box="[879,951,1755,1782]" captionStart="FIGURES 5259" captionStartId="15.[151,264,1834,1858]" captionTargetBox="[151,1435,181,1810]" captionTargetId="figure-19@15.[151,1435,181,1810]" captionTargetPageId="15" captionText="FIGURES 5259. Baetis (Tenuibaetis) hissaricus. 5253, fore and middle legs of male imago; 5455, genitalia of males imago; 56, hind wing of male; 57, hind wing of female; 58, subimaginal exuviae of female fore leg; 59, subimaginal exuviae of half of mesonotum (57, 58, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889731" httpUri="https://zenodo.org/record/7889731/files/figure.png" pageId="6" pageNumber="207">5657</figureCitation>
,
<figureCitation box="[963,1061,1755,1781]" captionStart="FIGURES 102111" captionStartId="23.[151,264,1841,1865]" captionTargetBox="[151,1431,185,1816]" captionTargetId="figure-19@23.[151,1435,181,1817]" captionTargetPageId="23" captionText="FIGURES 102111. Baetis (Tenuibaetis) frequentus from India. 102109, male imago and its subimaginal exuviae: 102, head and thorax; 103, abdomen; 104, exuviae of subimaginal half of mesonotum; 105106, fore and middle legs; 107108, their subimaginal exuviae; 109, hind wing. 110111, female imago and its hind wing." figureDoi="http://doi.org/10.5281/zenodo.7889755" httpUri="https://zenodo.org/record/7889755/files/figure.png" pageId="6" pageNumber="207">109111</figureCitation>
,
<figureCitation box="[1073,1174,1755,1781]" captionStart="FIGURES 135143" captionStartId="29.[151,264,1766,1790]" captionTargetBox="[151,1435,181,1742]" captionTargetId="figure-19@29.[151,1435,181,1742]" captionTargetPageId="29" captionText="FIGURES 135143. Baetis (Tenuibaetis) fujitanii. 135, male imago; 136, female imago; 137, subimaginal exuviae of half of mesonotum; 138, male hind wing; 139, female hind wing; 140, exuviae of male fore leg; 141, abdomen of male imago; 142143, middle and fore legs of male imago." figureDoi="http://doi.org/10.5281/zenodo.7889765" httpUri="https://zenodo.org/record/7889765/files/figure.png" pageId="6" pageNumber="207">138139</figureCitation>
,
<figureCitation box="[1185,1287,1755,1781]" captionStart="FIGURES 169180" captionStartId="34.[151,264,1880,1904]" captionTargetBox="[151,1436,181,1856]" captionTargetId="figure-36@34.[151,1436,181,1856]" captionTargetPageId="34" captionText="FIGURES 169180. Baetis (Tenuibaetis) kaltenbachi sp. n. 169170, male imago; 171, female imago; 172, subimaginal exuviae of abdomen; 173174, fore and middle legs of male imago; 175176, their subimaginal exuviae; 177, subimaginal exuviae of half of mesonotum; 178, hind wing of male imago; 179, the same, female; 180, blunt microlepides on fore tarsus of male subimago (169170, 172178, 180, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889777" httpUri="https://zenodo.org/record/7889777/files/figure.png" pageId="6" pageNumber="207">178179</figureCitation>
).
</paragraph>
<paragraph blockId="6.[151,1437,387,2034]" pageId="6" pageNumber="207">
In
<emphasis box="[231,248,1792,1817]" italics="true" pageId="6" pageNumber="207">B</emphasis>
.
<taxonomicName authority="(Fujitani et al. 2011)" authorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" authorityYear="2023" baseAuthorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" baseAuthorityYear="2011" box="[263,685,1791,1818]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="parvipterus" subGenus="Tenuibaetis">
(
<emphasis box="[272,288,1792,1817]" italics="true" pageId="6" pageNumber="207">T</emphasis>
.)
<emphasis box="[313,442,1792,1817]" italics="true" pageId="6" pageNumber="207">parvipterus</emphasis>
(
<bibRefCitation author="Fujitani, T. &amp; Kobayashi, N. &amp; Hirowatari, T. &amp; Tanida, K." box="[457,677,1791,1818]" pageId="6" pageNumber="207" pagination="213 - 223" refId="ref17559" refString="Fujitani, T., Kobayashi, N., Hirowatari, T. &amp; Tanida, K. (2011) Three species of a genus Tenuibaetis (Ephemeroptera: Baetidae) from Japan, with description of a new species. Limnology, 12, 213 - 223. https: // doi. org / 10.1007 / s 10201 - 010 - 0342 - 0" type="journal article" year="2011">
Fujitani
<emphasis box="[555,605,1791,1817]" italics="true" pageId="6" pageNumber="207">et al</emphasis>
. 2011
</bibRefCitation>
)
</taxonomicName>
hind wings of female are greatly diminished (
<bibRefCitation author="Fujitani, T. &amp; Kobayashi, N. &amp; Hirowatari, T. &amp; Tanida, K." box="[1211,1432,1791,1818]" pageId="6" pageNumber="207" pagination="213 - 223" refId="ref17559" refString="Fujitani, T., Kobayashi, N., Hirowatari, T. &amp; Tanida, K. (2011) Three species of a genus Tenuibaetis (Ephemeroptera: Baetidae) from Japan, with description of a new species. Limnology, 12, 213 - 223. https: // doi. org / 10.1007 / s 10201 - 010 - 0342 - 0" type="journal article" year="2011">
Fujitani
<emphasis box="[1308,1358,1791,1817]" italics="true" pageId="6" pageNumber="207">et al</emphasis>
. 2011
</bibRefCitation>
: fig. 5b). In
<taxonomicName authority="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack, 2023" authorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" authorityYear="2023" box="[278,520,1827,1854]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="206" phylum="Arthropoda" rank="species" species="octomaculatus" status="sp. n." subGenus="Tenuibaetis">
<emphasis box="[278,295,1828,1853]" italics="true" pageId="6" pageNumber="207">B</emphasis>
. (
<emphasis box="[318,334,1828,1853]" italics="true" pageId="6" pageNumber="207">T</emphasis>
.)
<emphasis box="[357,520,1827,1853]" italics="true" pageId="6" pageNumber="207">octomaculatus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[527,591,1827,1853]" pageId="6" pageNumber="207">
<taxonomicNameLabel box="[527,591,1827,1853]" pageId="6" pageNumber="207" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
hind wings of both sexes are diminished (
<figureCitation box="[1063,1145,1827,1854]" captionStart="FIGURES 1321" captionStartId="5.[151,264,1840,1864]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure-19@5.[151,1436,181,1816]" captionTargetPageId="5" captionText="FIGURES 1321. Imaginal hind wings and larval protoptera (under the same magnification). 1314, female Baetis (Tenuibaetis) frequentus from India; 1517, male B. (T.) octomaculatus sp. n.; 1819, female of same species; 2021, male B. (T.) bialatus sp. n. (13, 15, 20, metathorax of imago with hind wing; 14, 21, larval exuviae of metanotum; 16, 18, hind wing of imago; 17, 19, cuticle from metathorax of larva ready to molt to subimago, with subimaginal wing inside protopteron: crumpled on Fig. 17, non-crumpled on Fig. 19)." figureDoi="http://doi.org/10.5281/zenodo.7889705" httpUri="https://zenodo.org/record/7889705/files/figure.png" pageId="6" pageNumber="207">Fig. 15</figureCitation>
), but larval protoptera are nearly as large as in the species with full-sized hind wings (
<figureCitation box="[821,952,1863,1889]" captionStart="FIGURES 1321" captionStartId="5.[151,264,1840,1864]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure-19@5.[151,1436,181,1816]" captionTargetPageId="5" captionText="FIGURES 1321. Imaginal hind wings and larval protoptera (under the same magnification). 1314, female Baetis (Tenuibaetis) frequentus from India; 1517, male B. (T.) octomaculatus sp. n.; 1819, female of same species; 2021, male B. (T.) bialatus sp. n. (13, 15, 20, metathorax of imago with hind wing; 14, 21, larval exuviae of metanotum; 16, 18, hind wing of imago; 17, 19, cuticle from metathorax of larva ready to molt to subimago, with subimaginal wing inside protopteron: crumpled on Fig. 17, non-crumpled on Fig. 19)." figureDoi="http://doi.org/10.5281/zenodo.7889705" httpUri="https://zenodo.org/record/7889705/files/figure.png" pageId="6" pageNumber="207">Figs 17, 19</figureCitation>
); in female, imaginal hind wing is as small as its larval protopteron, so that it develops under larval cuticle without crumpling (
<figureCitation box="[1103,1235,1899,1925]" captionStart="FIGURES 1321" captionStartId="5.[151,264,1840,1864]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure-19@5.[151,1436,181,1816]" captionTargetPageId="5" captionText="FIGURES 1321. Imaginal hind wings and larval protoptera (under the same magnification). 1314, female Baetis (Tenuibaetis) frequentus from India; 1517, male B. (T.) octomaculatus sp. n.; 1819, female of same species; 2021, male B. (T.) bialatus sp. n. (13, 15, 20, metathorax of imago with hind wing; 14, 21, larval exuviae of metanotum; 16, 18, hind wing of imago; 17, 19, cuticle from metathorax of larva ready to molt to subimago, with subimaginal wing inside protopteron: crumpled on Fig. 17, non-crumpled on Fig. 19)." figureDoi="http://doi.org/10.5281/zenodo.7889705" httpUri="https://zenodo.org/record/7889705/files/figure.png" pageId="6" pageNumber="207">Figs 1819</figureCitation>
); in contrast to it, male hind wing is larger than protopteron and is crumpled before molt (
<figureCitation box="[956,1086,1935,1961]" captionStart="FIGURES 1321" captionStartId="5.[151,264,1840,1864]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure-19@5.[151,1436,181,1816]" captionTargetPageId="5" captionText="FIGURES 1321. Imaginal hind wings and larval protoptera (under the same magnification). 1314, female Baetis (Tenuibaetis) frequentus from India; 1517, male B. (T.) octomaculatus sp. n.; 1819, female of same species; 2021, male B. (T.) bialatus sp. n. (13, 15, 20, metathorax of imago with hind wing; 14, 21, larval exuviae of metanotum; 16, 18, hind wing of imago; 17, 19, cuticle from metathorax of larva ready to molt to subimago, with subimaginal wing inside protopteron: crumpled on Fig. 17, non-crumpled on Fig. 19)." figureDoi="http://doi.org/10.5281/zenodo.7889705" httpUri="https://zenodo.org/record/7889705/files/figure.png" pageId="6" pageNumber="207">Figs 1617</figureCitation>
), as in most other mayflies and most other insects. Thus, in contrast to most other mayflies, in
<taxonomicName authorityName=": Fujitani, Hirowatari &amp; Tanida" authorityYear="2003" box="[846,974,1971,1997]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="genus">
<emphasis box="[846,974,1971,1997]" italics="true" pageId="6" pageNumber="207">Tenuibaetis</emphasis>
</taxonomicName>
size of hind protopteron of larva does not allow to make conclusion about size of its imaginal hind wing.
</paragraph>
<paragraph blockId="7.[151,1437,151,933]" pageId="7" pageNumber="208">
In
<taxonomicName authority="Suttinun et al. 2022" authorityName="Suttinun" authorityYear="2022" box="[229,608,151,177]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="panhai" subGenus="Tenuibaetis">
<emphasis box="[229,253,151,176]" italics="true" pageId="7" pageNumber="208">B.</emphasis>
(
<emphasis box="[269,290,151,176]" italics="true" pageId="7" pageNumber="208">T.</emphasis>
)
<emphasis box="[305,383,151,177]" italics="true" pageId="7" pageNumber="208">panhai</emphasis>
<bibRefCitation author="Suttinun, C. &amp; Gattolliat, J. - L. &amp; Boonsoong, B." box="[389,608,151,177]" pageId="7" pageNumber="208" pagination="165 - 182" refId="ref19443" refString="Suttinun, C. &amp; Gattolliat, J. - L. &amp; Boonsoong, B. (2022) First report of the genus Tenuibaetis (Ephemeroptera, Baetidae) from Thailand revealing a complex of cryptic species. ZooKeys, 1084, 165 - 182. https: // doi. org / 10.3897 / zookeys. 1084.78405" type="journal article" year="2022">
Suttinun
<emphasis box="[491,539,151,177]" italics="true" pageId="7" pageNumber="208">et al</emphasis>
. 2022
</bibRefCitation>
</taxonomicName>
and
<taxonomicName authority="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack, 2023" authorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" authorityYear="2023" box="[662,830,151,177]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="246" phylum="Arthropoda" rank="species" species="bialatus" status="sp. n." subGenus="Tenuibaetis">
<emphasis box="[662,679,151,176]" italics="true" pageId="7" pageNumber="208">B</emphasis>
. (
<emphasis box="[701,717,151,176]" italics="true" pageId="7" pageNumber="208">T</emphasis>
.)
<emphasis box="[740,830,151,177]" italics="true" pageId="7" pageNumber="208">bialatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[836,898,151,177]" pageId="7" pageNumber="208" rank="species">
<emphasis bold="true" box="[836,892,151,177]" pageId="7" pageNumber="208">sp. n</emphasis>
.
</taxonomicNameLabel>
, hind wings are completely absent in both sexes (
<figureCitation box="[159,244,187,213]" captionStart="FIGURES 1321" captionStartId="5.[151,264,1840,1864]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure-19@5.[151,1436,181,1816]" captionTargetPageId="5" captionText="FIGURES 1321. Imaginal hind wings and larval protoptera (under the same magnification). 1314, female Baetis (Tenuibaetis) frequentus from India; 1517, male B. (T.) octomaculatus sp. n.; 1819, female of same species; 2021, male B. (T.) bialatus sp. n. (13, 15, 20, metathorax of imago with hind wing; 14, 21, larval exuviae of metanotum; 16, 18, hind wing of imago; 17, 19, cuticle from metathorax of larva ready to molt to subimago, with subimaginal wing inside protopteron: crumpled on Fig. 17, non-crumpled on Fig. 19)." figureDoi="http://doi.org/10.5281/zenodo.7889705" httpUri="https://zenodo.org/record/7889705/files/figure.png" pageId="7" pageNumber="208">Fig. 20</figureCitation>
); larvae of these species have no any vestiges of hind protoptera (
<figureCitation box="[979,1062,187,213]" captionStart="FIGURES 1321" captionStartId="5.[151,264,1840,1864]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure-19@5.[151,1436,181,1816]" captionTargetPageId="5" captionText="FIGURES 1321. Imaginal hind wings and larval protoptera (under the same magnification). 1314, female Baetis (Tenuibaetis) frequentus from India; 1517, male B. (T.) octomaculatus sp. n.; 1819, female of same species; 2021, male B. (T.) bialatus sp. n. (13, 15, 20, metathorax of imago with hind wing; 14, 21, larval exuviae of metanotum; 16, 18, hind wing of imago; 17, 19, cuticle from metathorax of larva ready to molt to subimago, with subimaginal wing inside protopteron: crumpled on Fig. 17, non-crumpled on Fig. 19)." figureDoi="http://doi.org/10.5281/zenodo.7889705" httpUri="https://zenodo.org/record/7889705/files/figure.png" pageId="7" pageNumber="208">Fig. 21</figureCitation>
).
</paragraph>
<paragraph blockId="7.[151,1437,151,933]" box="[199,658,222,249]" pageId="7" pageNumber="208">
<emphasis bold="true" box="[199,658,222,249]" pageId="7" pageNumber="208">
Characters common with other
<taxonomicName authorityName="Leach" authorityYear="1815" box="[580,652,223,249]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[580,652,223,249]" italics="true" pageId="7" pageNumber="208">Baetis</emphasis>
</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph blockId="7.[151,1437,151,933]" box="[199,1211,258,285]" pageId="7" pageNumber="208">
<emphasis bold="true" box="[199,503,258,285]" italics="true" pageId="7" pageNumber="208">Femoral patch (villopore).</emphasis>
Equally developed on all femora of larva (
<figureCitation box="[984,1142,259,285]" captionStart="FIGURES 230236" captionStartId="46.[151,264,1840,1864]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-36@46.[151,1436,181,1817]" captionTargetPageId="46" captionText="FIGURES 230236. Baetis (Tenuibaetis) panhai, exuviae of larval legs. 230231, proximal-posterior portion of fore femur with villopore (230, anterior side; 231, posterior side); 232234, fore, middle and hind legs; 235236, fore and middle tibiae." figureDoi="http://doi.org/10.5281/zenodo.7889799" httpUri="https://zenodo.org/record/7889799/files/figure.png" pageId="7" pageNumber="208">Figs 230231</figureCitation>
,
<figureCitation box="[1153,1199,258,285]" captionStart="FIGURES 253256" captionStartId="51.[151,264,1450,1474]" captionTargetBox="[151,1436,181,1426]" captionTargetId="figure-19@51.[151,1436,181,1426]" captionTargetPageId="51" captionText="FIGURES 253256. Baetis (Tenuibaetis) octomaculatus sp. n. (holotype). 253255, larval exuviae of fore, middle and hind legs; 256, base of femur with villopore." figureDoi="http://doi.org/10.5281/zenodo.7889807" httpUri="https://zenodo.org/record/7889807/files/figure.png" pageId="7" pageNumber="208">256</figureCitation>
).
</paragraph>
<paragraph blockId="7.[151,1437,151,933]" pageId="7" pageNumber="208">
<emphasis bold="true" box="[199,798,294,321]" italics="true" pageId="7" pageNumber="208">Subimaginal gonostyli folded under larval exuviae.</emphasis>
Pose of the
<taxonomicName authorityName="Leach" authorityYear="1815" box="[945,1029,295,321]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="genus">
<emphasis box="[945,1029,295,321]" italics="true" pageId="7" pageNumber="208">́Baetis</emphasis>
</taxonomicName>
-
<typeStatus box="[1037,1088,295,321]" pageId="7" pageNumber="208">type</typeStatus>
» or the
<taxonomicName authorityName="Bengtsson" authorityYear="1912" box="[1185,1315,295,321]" class="Insecta" family="Baetidae" genus="Acentrella" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="genus">
<emphasis box="[1185,1315,295,321]" italics="true" pageId="7" pageNumber="208">́Acentrella</emphasis>
</taxonomicName>
-
<typeStatus box="[1322,1375,295,321]" pageId="7" pageNumber="208">type</typeStatus>
», i.e. with the 2nd and 3rd segments directed medially (
<bibRefCitation author="Kluge, N. J. &amp; Novikova, E. A." box="[713,990,331,357]" pageId="7" pageNumber="208" pagination="1 - 56" refId="ref18456" refString="Kluge, N. J. &amp; Novikova, E. A. (2011) Systematics of the mayfly taxon Acentrella (Ephemeroptera, Baetidae), with description of new Asian and African species. Russian Entomological Journal, 20 (1), 1 - 56. https: // doi. org / 10.15298 / rusentj. 20.1.01" type="journal article" year="2011">Kluge &amp; Novikova 2011</bibRefCitation>
) (
<figureCitation box="[1013,1114,331,357]" captionStart="FIGURES 237241" captionStartId="47.[151,264,1838,1862]" captionTargetBox="[152,1435,181,1814]" captionTargetId="figure-19@47.[151,1435,181,1814]" captionTargetPageId="47" captionText="FIGURES 237241. Baetis (Tenuibaetis) panhai. 237, abdomen of male subimago extracted from mature larva; 238, larval exuviae of half of pronotum and mesonotum; 239, larval exuviae of abdomen; 240, male subimago; 241, subimaginal gonostyli crumpled under larval cuticle." figureDoi="http://doi.org/10.5281/zenodo.7889801" httpUri="https://zenodo.org/record/7889801/files/figure.png" pageId="7" pageNumber="208">Figs 241</figureCitation>
,
<figureCitation box="[1126,1172,330,357]" captionStart="FIGURES 275278" captionStartId="54.[151,264,1584,1608]" captionTargetBox="[151,1436,181,1560]" captionTargetId="figure-36@54.[151,1436,181,1560]" captionTargetPageId="54" captionText="FIGURES 275278. Baetis (Tenuibaetis) octomaculatus sp. n. 275, subimaginal gonostyli crumpled under larval cuticle; 276, imaginal genitalia; 277, imaginal genitalia of another individual, with focus on ventral side; 278, the same, with focus on gonovectes. Abbreviation: s.s, sterno-styligeral muscle." figureDoi="http://doi.org/10.5281/zenodo.7889813" httpUri="https://zenodo.org/record/7889813/files/figure.png" pageId="7" pageNumber="208">275</figureCitation>
).
</paragraph>
<paragraph blockId="7.[151,1437,151,933]" pageId="7" pageNumber="208">
<emphasis bold="true" box="[199,530,366,393]" italics="true" pageId="7" pageNumber="208">Texture of subimaginal tarsi.</emphasis>
All tarsomeres of all leg pairs covered mostly with blunt microlepides (
<figureCitation box="[1326,1424,367,393]" captionStart="FIGURES 169180" captionStartId="34.[151,264,1880,1904]" captionTargetBox="[151,1436,181,1856]" captionTargetId="figure-36@34.[151,1436,181,1856]" captionTargetPageId="34" captionText="FIGURES 169180. Baetis (Tenuibaetis) kaltenbachi sp. n. 169170, male imago; 171, female imago; 172, subimaginal exuviae of abdomen; 173174, fore and middle legs of male imago; 175176, their subimaginal exuviae; 177, subimaginal exuviae of half of mesonotum; 178, hind wing of male imago; 179, the same, female; 180, blunt microlepides on fore tarsus of male subimago (169170, 172178, 180, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889777" httpUri="https://zenodo.org/record/7889777/files/figure.png" pageId="7" pageNumber="208">Fig. 180</figureCitation>
). The same in all other Baetofemorata (Kluge 2022).
</paragraph>
<paragraph blockId="7.[151,1437,151,933]" box="[199,874,439,465]" pageId="7" pageNumber="208">
<emphasis bold="true" box="[199,253,439,465]" pageId="7" pageNumber="208">
<emphasis bold="true" box="[199,246,439,465]" italics="true" pageId="7" pageNumber="208">Egg</emphasis>
.
</emphasis>
Oval, chorion rugose, without regular relief (
<figureCitation box="[763,862,439,465]" captionStart="FIGURES 215224" captionStartId="43.[151,264,1770,1794]" captionTargetBox="[151,1435,181,1746]" captionTargetId="figure-19@43.[151,1435,181,1746]" captionTargetPageId="43" captionText="FIGURES 215224. Baetis (Tenuibaetis) bialatus sp. n. 215, subimaginal exuviae of mesopleuron; 216, subimaginal exuviae of half of mesonotum; 217218, exuviae of fore and hind legs of male subimago; 219220, fore and middle legs of male imago; 221, male imago; 222, female imago; 223, subimaginal exuviae of abdominal segments VIIIIX; 224, egg (215220, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889793" httpUri="https://zenodo.org/record/7889793/files/figure.png" pageId="7" pageNumber="208">Fig. 224</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection box="[199,1113,474,501]" pageId="7" pageNumber="208" type="distribution">
<paragraph blockId="7.[151,1437,151,933]" box="[199,1113,474,501]" pageId="7" pageNumber="208">
<emphasis bold="true" box="[199,353,474,500]" pageId="7" pageNumber="208">Distribution.</emphasis>
Central and Eastern Asia: East Palaearctic and the Oriental Region.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="208" type="taxon_list">
<paragraph blockId="7.[151,1437,151,933]" pageId="7" pageNumber="208">
<emphasis bold="true" box="[199,447,510,536]" pageId="7" pageNumber="208">Species composition.</emphasis>
SIBERIA and RUSSIAN FAR EAST:
<taxonomicName authority="Kazlauskas 1963" authorityName="Kazlauskas" authorityYear="1963" box="[910,1431,511,537]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="ursinus" subGenus="Tenuibaetis">
<emphasis box="[910,980,511,536]" italics="true" pageId="7" pageNumber="208">Baetis</emphasis>
(
<emphasis box="[998,1126,511,537]" italics="true" pageId="7" pageNumber="208">Tenuibaetis</emphasis>
)
<emphasis box="[1145,1227,511,536]" italics="true" pageId="7" pageNumber="208">ursinus</emphasis>
<bibRefCitation author="Kazlauskas, R. S." box="[1236,1431,511,537]" pageId="7" pageNumber="208" pagination="582 - 592" refId="ref17956" refString="Kazlauskas, R. S. (1963) [New and little-known may flies (Ephemeroptera) from the USSR]. Entomologicheskoe Obozrenie = Revue d'Entomologie de l'URSS, 42 (3), 582 - 592. [in Russian]" type="journal article" year="1963">Kazlauskas 1963</bibRefCitation>
</taxonomicName>
; CENTRAL ASIA:
<taxonomicName authority="Novikova 1991" authorityName="Novikova" authorityYear="1991" box="[374,901,547,573]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="hissaricus" subGenus="Tenuibaetis">
<emphasis box="[374,444,547,572]" italics="true" pageId="7" pageNumber="208">Baetis</emphasis>
(
<emphasis box="[460,588,547,573]" italics="true" pageId="7" pageNumber="208">Tenuibaetis</emphasis>
)
<emphasis box="[605,719,547,573]" italics="true" pageId="7" pageNumber="208">hissaricus</emphasis>
<bibRefCitation author="Novikova, E. A." box="[725,901,547,573]" pageId="7" pageNumber="208" pagination="793 - 795" refId="ref19156" refString="Novikova, E. A. (1991) [A new subspecies of mayflies Baetis ursinus hissaricus subsp. n. (Ephemeroptera, Baetidae) from Tadjikistan]. Entomologicheskoe Obozrenie, 70 (4), 793 - 795." type="journal article" year="1991">Novikova 1991</bibRefCitation>
</taxonomicName>
.
<collectingCountry box="[912,1004,547,573]" name="Japan" pageId="7" pageNumber="208">JAPAN</collectingCountry>
:
<taxonomicName authority="Gose 1980" authorityName="Gose" authorityYear="1980" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="flexifemora" subGenus="Tenuibaetis">
<emphasis box="[1014,1084,547,572]" italics="true" pageId="7" pageNumber="208">Baetis</emphasis>
(
<emphasis box="[1100,1228,547,573]" italics="true" pageId="7" pageNumber="208">Tenuibaetis</emphasis>
)
<emphasis box="[1244,1372,547,573]" italics="true" pageId="7" pageNumber="208">flexifemora</emphasis>
<bibRefCitation author="Gose, K." pageId="7" pageNumber="208" pagination="122 - 123" refId="ref17790" refString="Gose, K. (1980) The mayflies of Japanese 7 (in Japanese). Aquabiology, 2, 122 - 123." type="journal article" year="1980">Gose 1980</bibRefCitation>
</taxonomicName>
;
<taxonomicName authority="(Fujitani et al. 2011)" authorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" authorityYear="2023" baseAuthorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" baseAuthorityYear="2011" box="[218,793,583,609]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="parvipterus" status="comb. n." subGenus="Tenuibaetis">
<emphasis box="[218,288,583,608]" italics="true" pageId="7" pageNumber="208">Baetis</emphasis>
(
<emphasis box="[301,429,583,609]" italics="true" pageId="7" pageNumber="208">Tenuibaetis</emphasis>
)
<emphasis box="[441,570,583,608]" italics="true" pageId="7" pageNumber="208">parvipterus</emphasis>
(
<bibRefCitation author="Fujitani, T. &amp; Kobayashi, N. &amp; Hirowatari, T. &amp; Tanida, K." box="[581,786,583,609]" pageId="7" pageNumber="208" pagination="213 - 223" refId="ref17559" refString="Fujitani, T., Kobayashi, N., Hirowatari, T. &amp; Tanida, K. (2011) Three species of a genus Tenuibaetis (Ephemeroptera: Baetidae) from Japan, with description of a new species. Limnology, 12, 213 - 223. https: // doi. org / 10.1007 / s 10201 - 010 - 0342 - 0" type="journal article" year="2011">
Fujitani
<emphasis box="[673,726,583,609]" italics="true" pageId="7" pageNumber="208">et al.</emphasis>
2011
</bibRefCitation>
)
</taxonomicName>
<emphasis bold="true" box="[796,895,583,609]" pageId="7" pageNumber="208">
<taxonomicNameLabel box="[796,895,583,609]" pageId="7" pageNumber="208" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
<collectingCountry box="[897,1010,583,609]" name="Taiwan" pageId="7" pageNumber="208">TAIWAN</collectingCountry>
:
<taxonomicName authority="Muller-Liebenau 1985" authorityName="Muller-Liebenau" authorityYear="1985" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="pseudofrequentus" subGenus="Tenuibaetis">
<emphasis box="[1017,1087,583,608]" italics="true" pageId="7" pageNumber="208">Baetis</emphasis>
(
<emphasis box="[1099,1227,583,609]" italics="true" pageId="7" pageNumber="208">Tenuibaetis</emphasis>
)
<emphasis box="[1240,1437,583,609]" italics="true" pageId="7" pageNumber="208">pseudofrequentus</emphasis>
Müller-Liebenau 1985
</taxonomicName>
;
<taxonomicName authority="Kang &amp; Yang" authorityName="Kang &amp; Yang" box="[428,921,619,645]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="arduus" subGenus="Tenuibaetis">
<emphasis box="[428,498,619,644]" italics="true" pageId="7" pageNumber="208">Baetis</emphasis>
(
<emphasis box="[520,648,619,645]" italics="true" pageId="7" pageNumber="208">Tenuibaetis</emphasis>
)
<emphasis box="[669,746,619,645]" italics="true" pageId="7" pageNumber="208">arduus</emphasis>
Kang &amp; Yang
</taxonomicName>
(in Kang
<emphasis box="[1049,1111,619,645]" italics="true" pageId="7" pageNumber="208">et al.</emphasis>
) 1994;
<taxonomicName authority="Kang &amp; Yang" authorityName="Kang &amp; Yang" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="inornatus" subGenus="Tenuibaetis">
<emphasis box="[1207,1277,619,644]" italics="true" pageId="7" pageNumber="208">Baetis</emphasis>
(
<emphasis box="[1299,1427,619,645]" italics="true" pageId="7" pageNumber="208">Tenuibaetis</emphasis>
)
<emphasis box="[151,258,655,680]" italics="true" pageId="7" pageNumber="208">inornatus</emphasis>
Kang &amp; Yang
</taxonomicName>
(in Kang
<emphasis box="[550,609,655,681]" italics="true" pageId="7" pageNumber="208">et al.</emphasis>
) 1994.
<collectingCountry box="[701,850,655,681]" name="Thailand" pageId="7" pageNumber="208">THAILAND</collectingCountry>
:
<taxonomicName authority="(Suttinun et al. 2022)" authorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" authorityYear="2023" baseAuthorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" baseAuthorityYear="2022" box="[863,1437,655,681]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="panhai" status="comb. n." subGenus="Tenuibaetis">
<emphasis box="[863,933,655,680]" italics="true" pageId="7" pageNumber="208">Baetis</emphasis>
(
<emphasis box="[952,1080,655,681]" italics="true" pageId="7" pageNumber="208">Tenuibaetis</emphasis>
)
<emphasis box="[1100,1178,655,681]" italics="true" pageId="7" pageNumber="208">panhai</emphasis>
(
<bibRefCitation author="Suttinun, C. &amp; Gattolliat, J. - L. &amp; Boonsoong, B." box="[1195,1429,655,681]" pageId="7" pageNumber="208" pagination="165 - 182" refId="ref19443" refString="Suttinun, C. &amp; Gattolliat, J. - L. &amp; Boonsoong, B. (2022) First report of the genus Tenuibaetis (Ephemeroptera, Baetidae) from Thailand revealing a complex of cryptic species. ZooKeys, 1084, 165 - 182. https: // doi. org / 10.3897 / zookeys. 1084.78405" type="journal article" year="2022">
Suttinun
<emphasis box="[1302,1361,655,681]" italics="true" pageId="7" pageNumber="208">et al.</emphasis>
2022
</bibRefCitation>
)
</taxonomicName>
<emphasis bold="true" box="[151,252,691,717]" pageId="7" pageNumber="208">
<taxonomicNameLabel box="[151,252,691,717]" pageId="7" pageNumber="208" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
and some other species, among which
<taxonomicName authority="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack, 2023" authorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" authorityYear="2023" box="[681,1071,691,717]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="octomaculatus" status="sp. n." subGenus="Tenuibaetis">
<emphasis box="[681,751,691,716]" italics="true" pageId="7" pageNumber="208">Baetis</emphasis>
(
<emphasis box="[765,893,691,717]" italics="true" pageId="7" pageNumber="208">Tenuibaetis</emphasis>
)
<emphasis box="[908,1071,691,717]" italics="true" pageId="7" pageNumber="208">octomaculatus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1077,1139,691,717]" pageId="7" pageNumber="208">
<taxonomicNameLabel box="[1077,1139,691,717]" pageId="7" pageNumber="208" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is described below. WEST
<collectingCountry box="[151,296,727,753]" name="Malaysia" pageId="7" pageNumber="208">MALAYSIA</collectingCountry>
:
<taxonomicName box="[313,538,727,753]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="subGenus" subGenus="Tenuibaetis">
<emphasis box="[313,383,727,752]" italics="true" pageId="7" pageNumber="208">Baetis</emphasis>
(
<emphasis box="[401,529,727,753]" italics="true" pageId="7" pageNumber="208">Tenuibaetis</emphasis>
)
</taxonomicName>
<taxonomicName authority="Muller-Liebenau 1984" authorityName="Muller-Liebenau" authorityYear="1984" box="[548,899,727,753]" family="Coelophysidae" genus="Lepidus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis box="[548,637,727,753]" italics="true" pageId="7" pageNumber="208">Lepidus</emphasis>
Müller-Liebenau 1984
</taxonomicName>
(Kaltenbach, personal communication). JAVA:
<taxonomicName authority="(Kaltenbach &amp; Gattolliat 2019)" authorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" authorityYear="2023" baseAuthorityName="Kaltenbach &amp; Gattolliat" baseAuthorityYear="2019" box="[151,829,763,789]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="fujitanii" status="comb. n." subGenus="Tenuibaetis">
<emphasis box="[151,221,763,788]" italics="true" pageId="7" pageNumber="208">Baetis</emphasis>
(
<emphasis box="[238,366,763,789]" italics="true" pageId="7" pageNumber="208">Tenuibaetis</emphasis>
)
<emphasis box="[382,471,763,789]" italics="true" pageId="7" pageNumber="208">fujitanii</emphasis>
(
<bibRefCitation author="Kaltenbach, T. &amp; Gattolliat, J. - L." box="[487,821,763,789]" pageId="7" pageNumber="208" pagination="13 - 23" refId="ref17903" refString="Kaltenbach, T. &amp; Gattolliat, J. - L. (2019) A new species of Tenuibaetis Kang &amp; Yang, 1994 from Indonesia (Ephemeroptera, Baetidae). ZooKeys, 820, 13 - 23. https: // doi. org / 10.3897 / zookeys. 820.31487" type="journal article" year="2019">Kaltenbach &amp; Gattolliat 2019</bibRefCitation>
)
</taxonomicName>
<emphasis bold="true" box="[835,938,763,789]" pageId="7" pageNumber="208">
<taxonomicNameLabel box="[835,938,763,789]" pageId="7" pageNumber="208" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
INDIAN HIMALAYA:
<taxonomicName class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="himani" subGenus="Tenuibaetis">
<emphasis box="[1212,1282,763,788]" italics="true" pageId="7" pageNumber="208">Baetis</emphasis>
(
<emphasis box="[1299,1427,763,789]" italics="true" pageId="7" pageNumber="208">Tenuibaetis</emphasis>
)
<emphasis box="[151,229,799,825]" italics="true" pageId="7" pageNumber="208">himani</emphasis>
</taxonomicName>
(Kubendran
<emphasis box="[384,443,799,825]" italics="true" pageId="7" pageNumber="208">et al.</emphasis>
in
<bibRefCitation author="Kubendran, T. &amp; Vasanth, M. &amp; Subramanian, K. A. &amp; Gattolliat, J. - L. &amp; Selvakumar, C. &amp; Jabeen, F. &amp; Sinha, B." box="[486,749,799,825]" pageId="7" pageNumber="208" pagination="511 - 534" refId="ref18803" refString="Kubendran, T., Vasanth, M., Subramanian, K. A., Gattolliat, J. - L., Selvakumar, C., Jabeen, F. &amp; Sinha, B. (2022) Two new species and two new records of the genus Tenuibaetis Kang &amp; Yang, 1994 (Ephemeroptera: Baetidae) from Indian Himalaya. Zootaxa, 5196 (4), 511 - 534. https: // doi. org / 10.11646 / zootaxa. 5196.4.3" type="journal article" year="2022">
Kubendran
<emphasis box="[621,680,799,825]" italics="true" pageId="7" pageNumber="208">et al.</emphasis>
2022
</bibRefCitation>
)
<emphasis bold="true" box="[766,872,799,825]" pageId="7" pageNumber="208">comb. n.</emphasis>
;
<taxonomicName box="[890,1190,799,825]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="kangi" subGenus="Tenuibaetis">
<emphasis box="[890,960,799,824]" italics="true" pageId="7" pageNumber="208">Baetis</emphasis>
(
<emphasis box="[980,1108,799,825]" italics="true" pageId="7" pageNumber="208">Tenuibaetis</emphasis>
)
<emphasis box="[1128,1190,799,825]" italics="true" pageId="7" pageNumber="208">kangi</emphasis>
</taxonomicName>
(Kubendran
<emphasis box="[1345,1404,799,825]" italics="true" pageId="7" pageNumber="208">et al.</emphasis>
in
<bibRefCitation author="Kubendran, T. &amp; Vasanth, M. &amp; Subramanian, K. A. &amp; Gattolliat, J. - L. &amp; Selvakumar, C. &amp; Jabeen, F. &amp; Sinha, B." box="[151,406,835,861]" pageId="7" pageNumber="208" pagination="511 - 534" refId="ref18803" refString="Kubendran, T., Vasanth, M., Subramanian, K. A., Gattolliat, J. - L., Selvakumar, C., Jabeen, F. &amp; Sinha, B. (2022) Two new species and two new records of the genus Tenuibaetis Kang &amp; Yang, 1994 (Ephemeroptera: Baetidae) from Indian Himalaya. Zootaxa, 5196 (4), 511 - 534. https: // doi. org / 10.11646 / zootaxa. 5196.4.3" type="journal article" year="2022">
Kubendran
<emphasis box="[283,340,835,861]" italics="true" pageId="7" pageNumber="208">et al.</emphasis>
2022
</bibRefCitation>
)
<emphasis bold="true" box="[421,524,835,861]" pageId="7" pageNumber="208">comb. n.</emphasis>
SOUTHERN
<collectingCountry box="[688,771,835,861]" name="India" pageId="7" pageNumber="208">INDIA</collectingCountry>
and
<collectingCountry box="[826,978,835,861]" name="Sri Lanka" pageId="7" pageNumber="208">SRI LANKA</collectingCountry>
:
<taxonomicName authority="Muller-Liebenau &amp; Hubbard 1985" authorityName="Muller-Liebenau &amp; Hubbard" authorityYear="1985" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="frequentus" subGenus="Tenuibaetis">
<emphasis box="[994,1064,835,860]" italics="true" pageId="7" pageNumber="208">Baetis</emphasis>
(
<emphasis box="[1081,1209,835,861]" italics="true" pageId="7" pageNumber="208">Tenuibaetis</emphasis>
)
<emphasis box="[1226,1343,835,861]" italics="true" pageId="7" pageNumber="208">frequentus</emphasis>
<bibRefCitation author="Muller-Liebenau, I. &amp; Hubbard, M. D." pageId="7" pageNumber="208" pagination="537 - 561" refId="ref19053" refString="Muller-Liebenau, I. &amp; Hubbard, M. D. (1985) Baetidae from Sri Lanka with some general remarks on the Baetidae of the Oriental Region (Insecta: Ephemeroptera). Florida Entomologist, 68 (4), 537 - 561. https: // doi. org / 10.2307 / 3494855" type="journal article" year="1985">Müller-Liebenau &amp; Hubbard 1985</bibRefCitation>
</taxonomicName>
(=
<taxonomicName authorityName=": Kluge &amp; Novikova" authorityYear="2014" box="[492,773,871,897]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="michaelohubbardi">
<emphasis box="[492,773,871,897]" italics="true" pageId="7" pageNumber="208">Baetis michaelohubbardi</emphasis>
</taxonomicName>
(Selva-kumar
<emphasis box="[940,997,871,897]" italics="true" pageId="7" pageNumber="208">et al.</emphasis>
2012)
<emphasis bold="true" box="[1076,1154,871,897]" pageId="7" pageNumber="208">syn. n.</emphasis>
) and some other species, among which
<taxonomicName authority="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack, 2023" authorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" authorityYear="2023" box="[310,672,907,933]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="kaltenbachi" status="sp. n." subGenus="Tenuibaetis">
<emphasis box="[310,380,907,932]" italics="true" pageId="7" pageNumber="208">Baetis</emphasis>
(
<emphasis box="[396,524,907,933]" italics="true" pageId="7" pageNumber="208">Tenuibaetis</emphasis>
)
<emphasis box="[541,672,907,933]" italics="true" pageId="7" pageNumber="208">kaltenbachi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[678,742,907,933]" pageId="7" pageNumber="208">
<taxonomicNameLabel box="[678,742,907,933]" pageId="7" pageNumber="208" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName authority="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack, 2023" authorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" authorityYear="2023" box="[796,1116,907,933]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="bialatus" status="sp. n." subGenus="Tenuibaetis">
<emphasis box="[796,866,907,932]" italics="true" pageId="7" pageNumber="208">Baetis</emphasis>
(
<emphasis box="[882,1010,907,933]" italics="true" pageId="7" pageNumber="208">Tenuibaetis</emphasis>
)
<emphasis box="[1026,1116,907,933]" italics="true" pageId="7" pageNumber="208">bialatus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1124,1187,907,933]" pageId="7" pageNumber="208">
<taxonomicNameLabel box="[1124,1187,907,933]" pageId="7" pageNumber="208" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
are described below.
</paragraph>
</subSubSection>
</treatment>
</document>