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<document id="2C446E809E2B468775A81F8615547F24" ID-CLB-Dataset="3948" ID-DOI="10.5852/ejt.2021.775.1551" ID-GBIF-Dataset="cc10c087-bec3-4947-81d7-b3e5580a5c94" ID-ISSN="2118-9773" ID-Zenodo-Dep="5589689" ID-ZooBank="976D7020-5904-4951-97CE-B4FE58DA12A8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1634822039012" checkinUser="felipe" docAuthor="González-Alvarado, Arturo &amp; Vaz-De, Fernando Z." docDate="2021" docId="03F48795DC4CFFA1A4E7A106FABAFA9A" docLanguage="en" docName="ejt-775_86-106.pdf" docOrigin="European Journal of Taxonomy 775" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Deltochilum gilli Gonzalez-Alvarado &amp; Vaz-de-Mello 2021" docType="key" docVersion="5" lastPageNumber="104" masterDocId="FFCDFFEDDC5DFFB3A45AA753FFCDFFC7" masterDocTitle="Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum)" masterLastPageNumber="106" masterPageNumber="86" pageNumber="103" updateTime="1699464201688" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="5960425FB63DACDD4303905F6C44C16C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DEBBA07651C0357373BA96F5B6CA980C">
<mods:title id="FB07223085E2D6C4A0AEC52CAFFE96D6">Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum)</mods:title>
</mods:titleInfo>
<mods:name id="D1BA0FC81869BAC1E8F5FF2D6600ECBE" type="personal">
<mods:role id="8E27D2854B2BFA1FA94FF4912433A4A9">
<mods:roleTerm id="838293F9253EB09A13F05724E1D8B7CF">Author</mods:roleTerm>
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<mods:namePart id="2F3F8E9370353738244AFF113E73633C">González-Alvarado, Arturo</mods:namePart>
<mods:nameIdentifier id="7199052057FC44977CB171D011B41935" type="LSID">2D58A702-73A9-4C66-8CED-6B8B9A2AB10B</mods:nameIdentifier>
<mods:affiliation id="464030201C27CECED5FEEE44D8CCF68D">Programa de Pós-graduação em Ecologia e Conservação da Biodiversidade, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
<mods:nameIdentifier id="C56DB0F889C0CD6CCB9C35B9D76FDB38" type="email">fagakorn@gmail.com</mods:nameIdentifier>
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<mods:name id="3BE4446304EF659CF88CC6D496E224A1" type="personal">
<mods:role id="B6A07EC8C2059D838C57249B89D78F8F">
<mods:roleTerm id="752D56326DC46F4DB905110F0005F6A5">Author</mods:roleTerm>
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<mods:namePart id="53F51F23ECEEF78E9D81582540A679BE">Vaz-De, Fernando Z.</mods:namePart>
<mods:nameIdentifier id="66006A8B0B7285E799FFE504975719BE" type="LSID">2FF2B7D6-1A6B-43C1-9966-A1A949FB2B05</mods:nameIdentifier>
<mods:affiliation id="A3E4E479EBCB8A0E4E5C60C55B07FB97">Instituto de Biociências, Departamento de Biologia e Zoologia, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
<mods:nameIdentifier id="CBF0EC71C75DB2AFDC8487417AEB548D" type="email">vazdemello@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="4DCE217618E516C96B4EB389E99E7903">text</mods:typeOfResource>
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<mods:titleInfo id="3CD72A4197912B3ABA529413E50BE7E9">
<mods:title id="9F509D6BF82057632D256F9999DAE53C">European Journal of Taxonomy</mods:title>
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<mods:part id="ED66558909840EF3A841A7A859FC815F">
<mods:date id="4F1285DC84E3F632D3C04BCE8692EF46">2021</mods:date>
<mods:detail id="56469A5F82684CED280A9E4BF3BF48FA" type="pubDate">
<mods:number id="171781970D29F4DD9B869FFF21ABC54D">2021-10-20</mods:number>
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<mods:number id="39AC31FDF9CAAEC309DD902A4E9622B9">775</mods:number>
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<mods:start id="26551F76B88D0A4016924D41D1E4684B">86</mods:start>
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<mods:classification id="A98C5C99D290E7B55D8E11E69D7FEE8C">journal article</mods:classification>
<mods:identifier id="79A2BF907974596113062482A7DD29AC" type="CLB-Dataset">3948</mods:identifier>
<mods:identifier id="CCA580A8A970AA095EE917CAEBF114F7" type="DOI">10.5852/ejt.2021.775.1551</mods:identifier>
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<mods:identifier id="B6D9765B14371D9F2E62D293F3AAA2CA" type="ISSN">2118-9773</mods:identifier>
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<subSection id="E2D22F68DC4CFFA1A4E7A106FABAFA9A" lastPageId="18" lastPageNumber="104" pageId="17" pageNumber="103" type="multiple">
<treatment id="03F48795DC4CFFA1A4E7A106FABAFA9A" ID-DOI="http://doi.org/10.5281/zenodo.5590321" ID-GBIF-Taxon="189151493" ID-Zenodo-Dep="5590321" LSID="urn:lsid:plazi:treatment:03F48795DC4CFFA1A4E7A106FABAFA9A" httpUri="http://treatment.plazi.org/id/03F48795DC4CFFA1A4E7A106FABAFA9A" lastPageId="18" lastPageNumber="104" pageId="17" pageNumber="103">
<subSubSection id="C3476508DC4CFFA2A4E7A106FC0CF9B3" box="[189,961,1620,1652]" pageId="17" pageNumber="103" type="nomenclature">
<paragraph id="8BE23683DC4CFFA2A4E7A106FC0CF9B3" blockId="17.[189,961,1620,1652]" box="[189,961,1620,1652]" pageId="17" pageNumber="103">
<heading id="D0AA81EFDC4CFFA2A4E7A106FC0CF9B3" bold="true" box="[189,961,1620,1652]" fontSize="13" level="1" pageId="17" pageNumber="103" reason="1">
<emphasis id="B929EA91DC4CFFA2A4E7A106FC0CF9B3" bold="true" box="[189,961,1620,1652]" pageId="17" pageNumber="103">
Identification key to species of the
<taxonomicName id="4C5D4D00DC4CFFA2A6E3A107FD3CF9B4" ID-CoL="7VN7Q" authorityName="Gonzalez-Alvarado &amp; Vaz-de-Mello" authorityYear="2021" box="[697,753,1620,1651]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="103" phylum="Arthropoda" rank="species" species="gilli">
<emphasis id="B929EA91DC4CFFA2A6E3A107FD3CF9B4" bold="true" box="[697,753,1620,1651]" italics="true" pageId="17" pageNumber="103">gilli</emphasis>
</taxonomicName>
species group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3476508DC4CFFA1A4E7A1DFFABAFA9A" lastPageId="18" lastPageNumber="104" pageId="17" pageNumber="103" type="key">
<key id="FFCC61B2DC4CFFA1A4E7A1DFFABAFA9A" lastPageId="18" lastPageNumber="104" pageId="17" pageNumber="103">
<keyStep id="30A92E26DC4CFFA2A4E7A1DFFABAF928" pageId="17" pageNumber="103">
<paragraph id="8BE23683DC4CFFA2A4E7A1DFFABAF90C" blockId="17.[189,1399,1676,2025]" pageId="17" pageNumber="103">
<keyLead id="30AC95B6DC4CFFA2A4E7A1DFFABAF90C" pageId="17" pageNumber="103">
1. Striae I–VII inconspicuous, including apically, only in some parts striae slightly visible and very narrow (
<figureCitation id="13662A06DC4CFFA2A517A1E2FE07F90C" box="[333,458,1713,1739]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[219,1369,314,1798]" captionTargetId="figure-77@6.[219,1369,314,1798]" captionTargetPageId="6" captionText="Fig. 4. Elytra of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589701" httpUri="https://zenodo.org/record/5589701/files/figure.png" pageId="17" pageNumber="103">Fig. 4B–C</figureCitation>
)............................................................................................................................ 2
</keyLead>
</paragraph>
<paragraph id="8BE23683DC4CFFA2A49BA185FABAF928" blockId="17.[189,1399,1676,2025]" box="[193,1399,1749,1776]" pageId="17" pageNumber="103">
<keyLead id="30AC95B6DC4CFFA2A49BA185FABAF928" box="[193,1399,1749,1776]" pageId="17" pageNumber="103">
– Striae I–VII conspicuous (
<figureCitation id="13662A06DC4CFFA2A67AA186FD0DF937" box="[544,704,1749,1776]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[219,1369,314,1798]" captionTargetId="figure-77@6.[219,1369,314,1798]" captionTargetPageId="6" captionText="Fig. 4. Elytra of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589701" httpUri="https://zenodo.org/record/5589701/files/figure.png" pageId="17" pageNumber="103">Fig. 4A, D–E</figureCitation>
) .......................................................................................... 3
</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BE23683DC4CFFA1A4E7A04BFABAFE14" blockId="17.[189,1399,1676,2025]" lastBlockId="18.[189,1399,265,1373]" lastPageId="18" lastPageNumber="104" pageId="17" pageNumber="103">
<keyStep id="30A92E26DC4CFFA1A4E7A04BFABAFE14" lastPageId="18" lastPageNumber="104" pageId="17" pageNumber="103">
<keyLead id="30AC95B6DC4CFFA1A4E7A04BFABAFE14" lastPageId="18" lastPageNumber="104" pageId="17" pageNumber="103">
2. Head punctures on frons separated by less than one diameter of a puncture (
<figureCitation id="13662A06DC4CFFA2A0C3A04BFB37F8F5" box="[1177,1274,1816,1842]" captionStart="Fig" captionStartId="4.[189,232,1892,1918]" captionTargetBox="[237,1351,525,1826]" captionTargetId="figure-106@4.[237,1351,525,1826]" captionTargetPageId="4" captionText="Fig. 2. Head of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589695" httpUri="https://zenodo.org/record/5589695/files/figure.png" pageId="17" pageNumber="103">Fig. 2B</figureCitation>
). Pronotal disc punctures separated by less than one diameter (
<figureCitation id="13662A06DC4CFFA2A73EA06FFC2BF890" box="[868,998,1852,1879]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[217,1370,471,1774]" captionTargetId="figure-147@5.[217,1370,471,1774]" captionTargetPageId="5" captionText="Fig. 3. Pronotum of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744 E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745. F. Miscrosculpture of pronotal disc, Holotype of D. jocelynae sp. nov. G. Miscrosculpture pronotal disc, Holotype of D. nonstriatum sp. nov. White arrows show the shiny points." figureDoi="http://doi.org/10.5281/zenodo.5589697" httpUri="https://zenodo.org/record/5589697/files/figure.png" pageId="17" pageNumber="103">Fig. 3B, F</figureCitation>
). Pronotal disc with well-defined shiny points (
<figureCitation id="13662A06DC4CFFA2A5D3A032FE12F8BC" box="[393,479,1889,1915]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[217,1370,471,1774]" captionTargetId="figure-147@5.[217,1370,471,1774]" captionTargetPageId="5" captionText="Fig. 3. Pronotum of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744 E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745. F. Miscrosculpture of pronotal disc, Holotype of D. jocelynae sp. nov. G. Miscrosculpture pronotal disc, Holotype of D. nonstriatum sp. nov. White arrows show the shiny points." figureDoi="http://doi.org/10.5281/zenodo.5589697" httpUri="https://zenodo.org/record/5589697/files/figure.png" pageId="17" pageNumber="103">Fig. 3F</figureCitation>
, white arrows). Punctures of third interstria on disc occupying about 1/14
<superScript id="7C289BCBDC4CFFA2A164A033FA86F8A9" attach="left" box="[1342,1355,1888,1902]" fontSize="6" pageId="17" pageNumber="103">th</superScript>
the distance between striae II and III (
<figureCitation id="13662A06DC4CFFA2A6D2A0D6FD2AF858" box="[648,743,1925,1951]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[219,1369,314,1798]" captionTargetId="figure-77@6.[219,1369,314,1798]" captionTargetPageId="6" captionText="Fig. 4. Elytra of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589701" httpUri="https://zenodo.org/record/5589701/files/figure.png" pageId="17" pageNumber="103">Fig. 4B</figureCitation>
). Width of expansion of first ventrite of male, on third ventrite, subequal to distance between clypeal teeth (
<figureCitation id="13662A06DC4CFFA2A710A0F9FC68F802" box="[842,933,1962,1989]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[230,1363,815,1803]" captionTargetId="figure-152@8.[220,1368,814,1813]" captionTargetPageId="8" captionText="Fig. 6. Ventral view of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589707" httpUri="https://zenodo.org/record/5589707/files/figure.png" pageId="17" pageNumber="103">Fig. 6B</figureCitation>
).
<collectingCountry id="F34A7613DC4CFFA2A7E3A0F9FBDAF803" box="[953,1047,1962,1988]" name="Guyana" pageId="17" pageNumber="103">Guyana</collectingCountry>
, District 8: Mount Wokomung (
<figureCitation id="13662A06DC4CFFA2A4A9A09DFE87F82F" box="[243,330,1998,2024]" captionStart="Fig" captionStartId="14.[189,232,1928,1954]" captionTargetBox="[336,1251,578,1870]" captionTargetId="figure-135@14.[336,1251,578,1870]" captionTargetPageId="14" captionText="Fig. 10. Distribution map of thegilli species group. Blue circle =Deltochilum gilli González-Alvarado &amp; Vaz-de-Mello, 2021; yellow square = D. jocelynae sp. nov.; red star = D. nonstriatum sp. nov.; green triangle = D. quasistriatum sp. nov.; white eight-point star = D. tenuistriatum sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5589717" httpUri="https://zenodo.org/record/5589717/files/figure.png" pageId="17" pageNumber="103">Fig. 10</figureCitation>
, yellow square) .....................................................................
<taxonomicName id="4C5D4D00DC4CFFA2A05CA09DFAD8F82E" ID-CoL="8SSPY" authority="González-Alvarado &amp; Vaz-De, 2021" authorityName="González-Alvarado &amp; Vaz-De" authorityYear="2021" box="[1030,1301,1998,2025]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="103" phylum="Arthropoda" rank="species" species="jocelynae" status="sp. nov.">
<emphasis id="B929EA91DC4CFFA2A05CA09DFAD8F82E" bold="true" box="[1030,1301,1998,2025]" italics="true" pageId="17" pageNumber="103">Deltochilum jocelynae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A57EADC4CFFA2A146A09CFABAF82E" box="[1308,1399,1999,2025]" pageId="17" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
– Head punctures on frons almost contiguous (
<figureCitation id="13662A06DC4FFFA1A756A65AFCA1FEE4" box="[780,876,265,291]" captionStart="Fig" captionStartId="4.[189,232,1892,1918]" captionTargetBox="[237,1351,525,1826]" captionTargetId="figure-106@4.[237,1351,525,1826]" captionTargetPageId="4" captionText="Fig. 2. Head of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589695" httpUri="https://zenodo.org/record/5589695/files/figure.png" pageId="18" pageNumber="104">Fig. 2C</figureCitation>
). Pronotal disc punctures separated by less than one diameter, almost contiguous (
<figureCitation id="13662A06DC4FFFA1A689A67FFCACFE81" box="[723,865,300,326]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[217,1370,471,1774]" captionTargetId="figure-147@5.[217,1370,471,1774]" captionTargetPageId="5" captionText="Fig. 3. Pronotum of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744 E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745. F. Miscrosculpture of pronotal disc, Holotype of D. jocelynae sp. nov. G. Miscrosculpture pronotal disc, Holotype of D. nonstriatum sp. nov. White arrows show the shiny points." figureDoi="http://doi.org/10.5281/zenodo.5589697" httpUri="https://zenodo.org/record/5589697/files/figure.png" pageId="18" pageNumber="104">Fig. 3C, G</figureCitation>
). Pronotal disc with irregular shiny points (
<figureCitation id="13662A06DC4FFFA1A4A9A61CFE99FEAE" box="[243,340,335,361]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[217,1370,471,1774]" captionTargetId="figure-147@5.[217,1370,471,1774]" captionTargetPageId="5" captionText="Fig. 3. Pronotum of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744 E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745. F. Miscrosculpture of pronotal disc, Holotype of D. jocelynae sp. nov. G. Miscrosculpture pronotal disc, Holotype of D. nonstriatum sp. nov. White arrows show the shiny points." figureDoi="http://doi.org/10.5281/zenodo.5589697" httpUri="https://zenodo.org/record/5589697/files/figure.png" pageId="18" pageNumber="104">Fig. 3G</figureCitation>
, white arrows). Punctures of third interstria on disc occupying about
<emphasis id="B929EA91DC4FFFA1A0F6A61CFB03FEAE" bold="true" box="[1196,1230,335,361]" pageId="18" pageNumber="104">1/10</emphasis>
<superScript id="7C289BCBDC4FFFA1A094A61CFB16FE9A" attach="left" box="[1230,1243,335,349]" fontSize="6" pageId="18" pageNumber="104">th</superScript>
the distance between striae II and III (
<figureCitation id="13662A06DC4FFFA1A644A621FDB3FE4A" box="[542,638,370,397]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[219,1369,314,1798]" captionTargetId="figure-77@6.[219,1369,314,1798]" captionTargetPageId="6" captionText="Fig. 4. Elytra of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589701" httpUri="https://zenodo.org/record/5589701/files/figure.png" pageId="18" pageNumber="104">Fig. 4C</figureCitation>
). Width of expansion of first ventrite of male, on third ventrite, narrower than distance between clypeal teeth (
<figureCitation id="13662A06DC4FFFA1A716A6C5FC79FE76" box="[844,948,406,433]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[230,1363,815,1803]" captionTargetId="figure-152@8.[220,1368,814,1813]" captionTargetPageId="8" captionText="Fig. 6. Ventral view of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589707" httpUri="https://zenodo.org/record/5589707/files/figure.png" pageId="18" pageNumber="104">Fig. 6C</figureCitation>
).
<collectingCountry id="F34A7613DC4FFFA1A789A6C5FB82FE77" box="[979,1103,406,432]" name="Venezuela" pageId="18" pageNumber="104">Venezuela</collectingCountry>
,
<collectingRegion id="4999F861DC4FFFA1A03FA6C5FB70FE77" box="[1125,1213,406,432]" country="Venezuela" name="Bolivar" pageId="18" pageNumber="104">Bolivar</collectingRegion>
(
<figureCitation id="13662A06DC4FFFA1A082A6C5FAF7FE77" box="[1240,1338,406,432]" captionStart="Fig" captionStartId="14.[189,232,1928,1954]" captionTargetBox="[336,1251,578,1870]" captionTargetId="figure-135@14.[336,1251,578,1870]" captionTargetPageId="14" captionText="Fig. 10. Distribution map of thegilli species group. Blue circle =Deltochilum gilli González-Alvarado &amp; Vaz-de-Mello, 2021; yellow square = D. jocelynae sp. nov.; red star = D. nonstriatum sp. nov.; green triangle = D. quasistriatum sp. nov.; white eight-point star = D. tenuistriatum sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5589717" httpUri="https://zenodo.org/record/5589717/files/figure.png" pageId="18" pageNumber="104">Fig. 10</figureCitation>
, red star) ...............................................................................................
<taxonomicName id="4C5D4D00DC4FFFA1A7B9A6EBFAD8FE14" ID-CoL="8SSP2" authority="González-Alvarado &amp; Vaz-De, 2021" authorityName="González-Alvarado &amp; Vaz-De" authorityYear="2021" box="[995,1301,440,467]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="104" phylum="Arthropoda" rank="species" species="nonstriatum" status="sp. nov.">
<emphasis id="B929EA91DC4FFFA1A7B9A6EBFAD8FE14" bold="true" box="[995,1301,440,467]" italics="true" pageId="18" pageNumber="104">Deltochilum nonstriatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A57EADC4FFFA1A146A6EAFABAFE14" box="[1308,1399,441,467]" pageId="18" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</keyStep>
</paragraph>
<keyStep id="30A92E26DC4FFFA1A4E7A6A8FABAFC19" pageId="18" pageNumber="104">
<paragraph id="8BE23683DC4FFFA1A4E7A6A8FABAFD2F" blockId="18.[189,1399,265,1373]" pageId="18" pageNumber="104">
<keyLead id="30AC95B6DC4FFFA1A4E7A6A8FABAFD2F" pageId="18" pageNumber="104">
3. Elytral apical tubercles on interstriae III–VII (
<figureCitation id="13662A06DC4FFFA1A762A6A9FC52FDD2" box="[824,927,506,533]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[201,1386,717,1836]" captionTargetId="figure-170@7.[201,1386,717,1836]" captionTargetPageId="7" captionText="Fig. 5. Caudal view of mmbers of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589705" httpUri="https://zenodo.org/record/5589705/files/figure.png" pageId="18" pageNumber="104">Fig. 5A</figureCitation>
). First stria almost twice wider than second stria (
<figureCitation id="13662A06DC4FFFA1A5D0A54DFE26FDFF" box="[394,491,542,568]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[219,1369,314,1798]" captionTargetId="figure-77@6.[219,1369,314,1798]" captionTargetPageId="6" captionText="Fig. 4. Elytra of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589701" httpUri="https://zenodo.org/record/5589701/files/figure.png" pageId="18" pageNumber="104">Fig. 4A</figureCitation>
). Striae III–VII subequal in width and conspicuous (
<figureCitation id="13662A06DC4FFFA1A039A54DFB09FDFF" box="[1123,1220,542,568]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[219,1369,314,1798]" captionTargetId="figure-77@6.[219,1369,314,1798]" captionTargetPageId="6" captionText="Fig. 4. Elytra of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589701" httpUri="https://zenodo.org/record/5589701/files/figure.png" pageId="18" pageNumber="104">Fig. 4A</figureCitation>
). Pronotal disc with shiny points well-defined and separated from punctures (
<figureCitation id="13662A06DC4FFFA1A7A0A512FB92FD9C" box="[1018,1119,577,603]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[217,1370,471,1774]" captionTargetId="figure-147@5.[217,1370,471,1774]" captionTargetPageId="5" captionText="Fig. 3. Pronotum of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744 E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745. F. Miscrosculpture of pronotal disc, Holotype of D. jocelynae sp. nov. G. Miscrosculpture pronotal disc, Holotype of D. nonstriatum sp. nov. White arrows show the shiny points." figureDoi="http://doi.org/10.5281/zenodo.5589697" httpUri="https://zenodo.org/record/5589697/files/figure.png" pageId="18" pageNumber="104">Fig. 3A</figureCitation>
). Male metatrochanter with expansion on distal third. Width of expansion of first ventrite of male, on third ventrite, wider than distance between clypeal teeth (
<figureCitation id="13662A06DC4FFFA1A746A5D4FC49FD65" box="[796,900,647,674]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[230,1363,815,1803]" captionTargetId="figure-152@8.[220,1368,814,1813]" captionTargetPageId="8" captionText="Fig. 6. Ventral view of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589707" httpUri="https://zenodo.org/record/5589707/files/figure.png" pageId="18" pageNumber="104">Fig. 6A</figureCitation>
). Sub-medial area of endophallus with elongate scales (
<figureCitation id="13662A06DC4FFFA1A59FA5F9FDE2FD02" box="[453,559,682,709]" captionStart="Fig" captionStartId="10.[189,232,1857,1883]" captionTargetBox="[233,1362,650,1777]" captionTargetId="figure-181@10.[218,1370,644,1787]" captionTargetPageId="10" captionText="Fig. 8. Endophallus of members of thegilli species group.Arrows show the scales on the sub-medial area of the endophallus. A. Elongate scales. C. Regular scales. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589713" httpUri="https://zenodo.org/record/5589713/files/figure.png" pageId="18" pageNumber="104">Fig. 8A</figureCitation>
, arrow).
<collectingCountry id="F34A7613DC4FFFA1A6E8A5F8FCDDFD02" box="[690,784,683,709]" name="Guyana" pageId="18" pageNumber="104">Guyana</collectingCountry>
, District 8: Mount Wokomung (
<figureCitation id="13662A06DC4FFFA1A09DA5F9FAE6FD02" box="[1223,1323,682,709]" captionStart="Fig" captionStartId="14.[189,232,1928,1954]" captionTargetBox="[336,1251,578,1870]" captionTargetId="figure-135@14.[336,1251,578,1870]" captionTargetPageId="14" captionText="Fig. 10. Distribution map of thegilli species group. Blue circle =Deltochilum gilli González-Alvarado &amp; Vaz-de-Mello, 2021; yellow square = D. jocelynae sp. nov.; red star = D. nonstriatum sp. nov.; green triangle = D. quasistriatum sp. nov.; white eight-point star = D. tenuistriatum sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5589717" httpUri="https://zenodo.org/record/5589717/files/figure.png" pageId="18" pageNumber="104">Fig. 10</figureCitation>
, blue circle) ..................................................
<taxonomicName id="4C5D4D00DC4FFFA1A6F7A59EFABAFD2F" ID-CoL="7VN7Q" authority="Gonzalez-Alvarado &amp; Vaz-de-Mello, 2021" authorityName="Gonzalez-Alvarado &amp; Vaz-de-Mello" authorityYear="2021" box="[685,1399,717,744]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="104" phylum="Arthropoda" rank="species" species="gilli">
<emphasis id="B929EA91DC4FFFA1A6F7A59EFCB7FD2F" bold="true" box="[685,890,717,744]" italics="true" pageId="18" pageNumber="104">Deltochilum gilli</emphasis>
<bibRefCitation id="EFCC4B72DC4FFFA1A7DBA59DFABAFD2F" author="Gonzalez-Alvarado A. &amp; Vaz-de-Mello F. Z." box="[897,1399,718,744]" pageId="18" pageNumber="104" pagination="0244657" refId="ref7969" refString="Gonzalez-Alvarado A. &amp; Vaz-de-Mello F. Z. 2021. Towards a comprehensive taxonomic revision of the Neotropical dung beetle subgenus Deltochilum (Deltohyboma) Lane, 1946 (Coleoptera: Scarabaeidae: Scarabaeinae): Division into species groups. PLOS ONE 16: e 0244657. https: // doi. org / 10.1371 / journal. pone. 0244657" type="journal article" year="2021">González-Alvarado &amp; Vaz-de-Mello, 2021</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE23683DC4FFFA1A49BA5A2FABAFC19" blockId="18.[189,1399,265,1373]" pageId="18" pageNumber="104">
<keyLead id="30AC95B6DC4FFFA1A49BA5A2FABAFC19" pageId="18" pageNumber="104">
– Elytral apical tubercles on interstriae III, V–VII (
<figureCitation id="13662A06DC4FFFA1A769A5A2FC79FCCB" box="[819,948,753,780]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[201,1386,717,1836]" captionTargetId="figure-170@7.[201,1386,717,1836]" captionTargetPageId="7" captionText="Fig. 5. Caudal view of mmbers of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589705" httpUri="https://zenodo.org/record/5589705/files/figure.png" pageId="18" pageNumber="104">Fig. 5D–E</figureCitation>
). First stria subequal to slightly wider than second stria (
<figureCitation id="13662A06DC4FFFA1A59DA447FD84FCE9" box="[455,585,788,814]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[219,1369,314,1798]" captionTargetId="figure-77@6.[219,1369,314,1798]" captionTargetPageId="6" captionText="Fig. 4. Elytra of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589701" httpUri="https://zenodo.org/record/5589701/files/figure.png" pageId="18" pageNumber="104">Fig. 4D–E</figureCitation>
). Striae III–VII successively thinner and more ill-defined (
<figureCitation id="13662A06DC4FFFA1A151A447FECDFC95" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[219,1369,314,1798]" captionTargetId="figure-77@6.[219,1369,314,1798]" captionTargetPageId="6" captionText="Fig. 4. Elytra of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589701" httpUri="https://zenodo.org/record/5589701/files/figure.png" pageId="18" pageNumber="104">Fig. 4D– E</figureCitation>
). Pronotal disc with shiny points irregular and contiguous to punctures (
<figureCitation id="13662A06DC4FFFA1A0C3A464FAEDFC96" box="[1177,1312,823,849]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[217,1370,471,1774]" captionTargetId="figure-147@5.[217,1370,471,1774]" captionTargetPageId="5" captionText="Fig. 3. Pronotum of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744 E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745. F. Miscrosculpture of pronotal disc, Holotype of D. jocelynae sp. nov. G. Miscrosculpture pronotal disc, Holotype of D. nonstriatum sp. nov. White arrows show the shiny points." figureDoi="http://doi.org/10.5281/zenodo.5589697" httpUri="https://zenodo.org/record/5589697/files/figure.png" pageId="18" pageNumber="104">Fig. 3D–E</figureCitation>
). Male metatrochanter not modified. Width of expansion of first ventrite of male, on third ventrite, subequal to distance between clypeal teeth (
<figureCitation id="13662A06DC4FFFA1A6DAA42DFCCCFC5F" box="[640,769,893,920]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[230,1363,815,1803]" captionTargetId="figure-152@8.[220,1368,814,1813]" captionTargetPageId="8" captionText="Fig. 6. Ventral view of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589707" httpUri="https://zenodo.org/record/5589707/files/figure.png" pageId="18" pageNumber="104">Fig. 6D–E</figureCitation>
). Sub-medial area of endophallus with scales, but not elongate (
<figureCitation id="13662A06DC4FFFA1A53AA4F2FE12FC7C" box="[352,479,929,955]" captionStart="Fig" captionStartId="10.[189,232,1857,1883]" captionTargetBox="[233,1362,650,1777]" captionTargetId="figure-181@10.[218,1370,644,1787]" captionTargetPageId="10" captionText="Fig. 8. Endophallus of members of thegilli species group.Arrows show the scales on the sub-medial area of the endophallus. A. Elongate scales. C. Regular scales. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589713" httpUri="https://zenodo.org/record/5589713/files/figure.png" pageId="18" pageNumber="104">Fig. 8D–E</figureCitation>
)
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, Mazaruni-Potaro District.
<collectingCountry id="F34A7613DC4FFFA1A7CFA4F2FBC3FC7C" box="[917,1038,929,955]" name="Venezuela" pageId="18" pageNumber="104">Venezuela</collectingCountry>
<collectingRegion id="4999F861DC4FFFA1A04DA4F2FBA2FC7C" box="[1047,1135,929,955]" country="Venezuela" name="Bolivar" pageId="18" pageNumber="104">Bolívar</collectingRegion>
(
<figureCitation id="13662A06DC4FFFA1A0DBA4F2FB17FC7C" box="[1153,1242,929,955]" captionStart="Fig" captionStartId="14.[189,232,1928,1954]" captionTargetBox="[336,1251,578,1870]" captionTargetId="figure-135@14.[336,1251,578,1870]" captionTargetPageId="14" captionText="Fig. 10. Distribution map of thegilli species group. Blue circle =Deltochilum gilli González-Alvarado &amp; Vaz-de-Mello, 2021; yellow square = D. jocelynae sp. nov.; red star = D. nonstriatum sp. nov.; green triangle = D. quasistriatum sp. nov.; white eight-point star = D. tenuistriatum sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5589717" httpUri="https://zenodo.org/record/5589717/files/figure.png" pageId="18" pageNumber="104">Fig. 10</figureCitation>
, white eightpoint star and green triangle) ............................................................................................................. 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A92E26DC4FFFA1A4E7A355FABAFA9A" pageId="18" pageNumber="104">
<paragraph id="8BE23683DC4FFFA1A4E7A355FABAFB6A" blockId="18.[189,1399,265,1373]" pageId="18" pageNumber="104">
<keyLead id="30AC95B6DC4FFFA1A4E7A355FABAFB6A" pageId="18" pageNumber="104">
4. First stria wider than second stria (
<figureCitation id="13662A06DC4FFFA1A6F6A356FCDFFBE7" box="[684,786,1029,1056]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[219,1369,314,1798]" captionTargetId="figure-77@6.[219,1369,314,1798]" captionTargetPageId="6" captionText="Fig. 4. Elytra of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589701" httpUri="https://zenodo.org/record/5589701/files/figure.png" pageId="18" pageNumber="104">Fig. 4D</figureCitation>
). Punctures of third interstria on disc occupying about 1/10
<superScript id="7C289BCBDC4FFFA1A537A37BFEB7FBF1" attach="left" box="[365,378,1064,1078]" fontSize="6" pageId="18" pageNumber="104">th</superScript>
the distance between striae II and III (
<figureCitation id="13662A06DC4FFFA1A72BA37AFC1BFB84" box="[881,982,1065,1091]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[219,1369,314,1798]" captionTargetId="figure-77@6.[219,1369,314,1798]" captionTargetPageId="6" captionText="Fig. 4. Elytra of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589701" httpUri="https://zenodo.org/record/5589701/files/figure.png" pageId="18" pageNumber="104">Fig. 4D</figureCitation>
). Pygidium with discal punctures occupying approximately
<emphasis id="B929EA91DC4FFFA1A676A31FFD83FBA1" bold="true" box="[556,590,1100,1126]" pageId="18" pageNumber="104">1/29</emphasis>
<superScript id="7C289BCBDC4FFFA1A614A318FD96FB9E" attach="left" box="[590,603,1099,1113]" fontSize="6" pageId="18" pageNumber="104">th</superScript>
the width on middle of pygidium and basal punctures almost contiguous (
<figureCitation id="13662A06DC4FFFA1A5DFA33CFE20FB4D" box="[389,493,1135,1162]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[201,1386,717,1836]" captionTargetId="figure-170@7.[201,1386,717,1836]" captionTargetPageId="7" captionText="Fig. 5. Caudal view of mmbers of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589705" httpUri="https://zenodo.org/record/5589705/files/figure.png" pageId="18" pageNumber="104">Fig. 5D</figureCitation>
).
<collectingCountry id="F34A7613DC4FFFA1A651A33CFDA4FB4E" box="[523,617,1135,1161]" name="Guyana" pageId="18" pageNumber="104">Guyana</collectingCountry>
, Mazaruni-Potaro District: Takutu Mountains (
<figureCitation id="13662A06DC4FFFA1A0E5A33CFAEDFB4E" box="[1215,1312,1135,1161]" captionStart="Fig" captionStartId="14.[189,232,1928,1954]" captionTargetBox="[336,1251,578,1870]" captionTargetId="figure-135@14.[336,1251,578,1870]" captionTargetPageId="14" captionText="Fig. 10. Distribution map of thegilli species group. Blue circle =Deltochilum gilli González-Alvarado &amp; Vaz-de-Mello, 2021; yellow square = D. jocelynae sp. nov.; red star = D. nonstriatum sp. nov.; green triangle = D. quasistriatum sp. nov.; white eight-point star = D. tenuistriatum sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5589717" httpUri="https://zenodo.org/record/5589717/files/figure.png" pageId="18" pageNumber="104">Fig. 10</figureCitation>
, green triangle).......................................................................................
<taxonomicName id="4C5D4D00DC4FFFA1A78BA3C1FAD8FB6B" ID-CoL="8SSNC" authority="González-Alvarado &amp; Vaz-De, 2021" authorityName="González-Alvarado &amp; Vaz-De" authorityYear="2021" box="[977,1301,1170,1197]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="104" phylum="Arthropoda" rank="species" species="quasistriatum" status="sp. nov.">
<emphasis id="B929EA91DC4FFFA1A78BA3C1FAD8FB6B" bold="true" box="[977,1301,1170,1197]" italics="true" pageId="18" pageNumber="104">Deltochilum quasistriatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A57EADC4FFFA1A146A3C0FABAFB6A" box="[1308,1399,1171,1197]" pageId="18" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BE23683DC4FFFA1A49BA3E5FABAFA9A" blockId="18.[189,1399,265,1373]" pageId="18" pageNumber="104">
<keyLead id="30AC95B6DC4FFFA1A49BA3E5FABAFA9A" pageId="18" pageNumber="104">
– First stria subequal in width to second stria (
<figureCitation id="13662A06DC4FFFA1A6B7A3E6FC84FB17" box="[749,841,1205,1232]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[219,1369,314,1798]" captionTargetId="figure-77@6.[219,1369,314,1798]" captionTargetPageId="6" captionText="Fig. 4. Elytra of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589701" httpUri="https://zenodo.org/record/5589701/files/figure.png" pageId="18" pageNumber="104">Fig. 4E</figureCitation>
). Punctures of third interstria on disc occupying about
<emphasis id="B929EA91DC4FFFA1A56CA38BFE94FB35" bold="true" box="[310,345,1240,1266]" pageId="18" pageNumber="104">1/14</emphasis>
<superScript id="7C289BCBDC4FFFA1A503A38BFEABFB21" attach="left" box="[345,358,1240,1254]" fontSize="6" pageId="18" pageNumber="104">th</superScript>
the distance between II and III (
<figureCitation id="13662A06DC4FFFA1A6A7A38AFC91FB34" box="[765,860,1241,1267]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[219,1369,314,1798]" captionTargetId="figure-77@6.[219,1369,314,1798]" captionTargetPageId="6" captionText="Fig. 4. Elytra of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589701" httpUri="https://zenodo.org/record/5589701/files/figure.png" pageId="18" pageNumber="104">Fig. 4E</figureCitation>
). Pygidium with discal punctures occupying approximately
<emphasis id="B929EA91DC4FFFA1A5C4A3AFFE0CFAD1" bold="true" box="[414,449,1276,1302]" pageId="18" pageNumber="104">1/38</emphasis>
<superScript id="7C289BCBDC4FFFA1A59BA3A8FE03FACE" attach="left" box="[449,462,1275,1289]" fontSize="6" pageId="18" pageNumber="104">th</superScript>
the width on middle of pygidium basal punctures separated by less than one diameter but not almost contiguous (
<figureCitation id="13662A06DC4FFFA1A6CAA24CFD24FAFD" box="[656,745,1311,1338]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[201,1386,717,1836]" captionTargetId="figure-170@7.[201,1386,717,1836]" captionTargetPageId="7" captionText="Fig. 5. Caudal view of mmbers of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589705" httpUri="https://zenodo.org/record/5589705/files/figure.png" pageId="18" pageNumber="104">Fig. 5E</figureCitation>
).
<collectingCountry id="F34A7613DC4FFFA1A6A6A24CFCB5FAFE" box="[764,888,1311,1337]" name="Venezuela" pageId="18" pageNumber="104">Venezuela</collectingCountry>
,
<collectingRegion id="4999F861DC4FFFA1A7DBA24CFC13FAFE" box="[897,990,1311,1337]" country="Venezuela" name="Bolivar" pageId="18" pageNumber="104">Bolívar</collectingRegion>
:
<quantity id="4CA59B66DC4FFFA1A7BDA24CFBE2FAFE" box="[999,1071,1311,1337]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="104" unit="km" value="10.0">10 km</quantity>
E of San Francisco (
<figureCitation id="13662A06DC4FFFA1A147A24CFABFFAFE" box="[1309,1394,1311,1337]" captionStart="Fig" captionStartId="14.[189,232,1928,1954]" captionTargetBox="[336,1251,578,1870]" captionTargetId="figure-135@14.[336,1251,578,1870]" captionTargetPageId="14" captionText="Fig. 10. Distribution map of thegilli species group. Blue circle =Deltochilum gilli González-Alvarado &amp; Vaz-de-Mello, 2021; yellow square = D. jocelynae sp. nov.; red star = D. nonstriatum sp. nov.; green triangle = D. quasistriatum sp. nov.; white eight-point star = D. tenuistriatum sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5589717" httpUri="https://zenodo.org/record/5589717/files/figure.png" pageId="18" pageNumber="104">Fig. 10</figureCitation>
, white eight-point star) .................................................................
<taxonomicName id="4C5D4D00DC4FFFA1A78FA211FAD8FA9B" ID-CoL="8SSQ6" authority="González-Alvarado &amp; Vaz-De, 2021" authorityName="González-Alvarado &amp; Vaz-De" authorityYear="2021" box="[981,1301,1346,1373]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="104" phylum="Arthropoda" rank="species" species="tenuistriatum" status="sp. nov.">
<emphasis id="B929EA91DC4FFFA1A78FA211FAD8FA9B" bold="true" box="[981,1301,1346,1373]" italics="true" pageId="18" pageNumber="104">Deltochilum tenuistriatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A57EADC4FFFA1A146A210FABAFA9A" box="[1308,1399,1347,1373]" pageId="18" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</subSection>
</document>
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