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<document id="641A294ADF9D312D3B195294AADD78A4" ID-DOI="10.11646/zootaxa.3994.4.3" ID-GBIF-Dataset="04971140-5855-4d1c-9660-3a50a1f498d1" ID-ISSN="1175-5326" ID-Zenodo-Dep="243645" ID-ZooBank="83E7EB77-EA86-4720-9679-A8204992D786" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461276490491" checkinUser="plazi" docAuthor="Brancelj, Anton" docDate="2015" docId="03E987EBFFA9C43B69DAFCA3FD70FBEE" docLanguage="en" docName="zt03994p555.pdf" docOrigin="Zootaxa 3994 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parastenocaris joi Brancelj, 2015, sp. nov." docType="treatment" docVersion="11" lastPageNumber="553" masterDocId="FFD0FF93FFBBC42D694DFD11FFF3FFDA" masterDocTitle="Two new stygobiotic copepod species from the Tibesti area (Northern Chad) and a re-description of Pilocamptus schroederi (van Douwe, 1915)" masterLastPageNumber="555" masterPageNumber="531" pageNumber="549" updateTime="1698612941627" updateUser="plazi">
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<mods:title id="DF47435981F9E5B64A88800C7EA5E6F1">Two new stygobiotic copepod species from the Tibesti area (Northern Chad) and a re-description of Pilocamptus schroederi (van Douwe, 1915)</mods:title>
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<mods:namePart id="2EA3BAF31C19A6182BCEE8FB399D6D9D">Brancelj, Anton</mods:namePart>
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<mods:date id="758CE49C0E74DDB954AFFD3982B9B079">2015</mods:date>
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<treatment id="03E987EBFFA9C43B69DAFCA3FD70FBEE" ID-DOI="http://doi.org/10.5281/zenodo.5615419" ID-GBIF-Taxon="119639727" ID-Zenodo-Dep="5615419" LSID="urn:lsid:plazi:treatment:03E987EBFFA9C43B69DAFCA3FD70FBEE" httpUri="http://treatment.plazi.org/id/03E987EBFFA9C43B69DAFCA3FD70FBEE" lastPageId="22" lastPageNumber="553" pageId="18" pageNumber="549">
<subSubSection id="C35A6576FFA9C43F69DAFCA3FED3FE34" pageId="18" pageNumber="549" type="nomenclature">
<paragraph id="8BFF36FDFFA9C43F69DAFCA3FE24FE16" blockId="18.[151,471,434,494]" box="[151,471,434,460]" pageId="18" pageNumber="549">
<heading id="D0B78191FFA9C43F69DAFCA3FE24FE16" bold="true" box="[151,471,434,460]" fontSize="11" level="1" pageId="18" pageNumber="549" reason="1">
<emphasis id="B934EAEFFFA9C43F69DAFCA3FE24FE16" bold="true" box="[151,471,434,460]" pageId="18" pageNumber="549">
<taxonomicName id="4C404D7EFFA9C43F69DAFCA3FE81FE16" box="[151,370,434,460]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="549" phylum="Arthropoda" rank="species" species="joi" status="sp. nov.">
<emphasis id="B934EAEFFFA9C43F69DAFCA3FE81FE16" bold="true" box="[151,370,434,460]" italics="true" pageId="18" pageNumber="549">Parastenocaris joi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2075794FFA9C43F6834FCA3FE24FE16" box="[377,471,434,460]" pageId="18" pageNumber="549" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFF36FDFFA9C43F69DAFCC4FED3FE34" blockId="18.[151,471,434,494]" box="[151,288,469,494]" pageId="18" pageNumber="549">(Figs. 910)</paragraph>
</subSubSection>
<subSubSection id="C35A6576FFA9C43F69DAFF0CFDB7FCCB" pageId="18" pageNumber="549" type="materials_examined">
<paragraph id="8BFF36FDFFA9C43F69DAFF0CFD3AFD86" blockId="18.[151,1437,541,2009]" pageId="18" pageNumber="549">
<emphasis id="B934EAEFFFA9C43F69DAFF0CFEDEFDEC" bold="true" box="[151,301,541,566]" pageId="18" pageNumber="549">
<typeStatus id="54FB885FFFA9C43F69DAFF0CFF21FDEC" box="[151,210,541,566]" pageId="18" pageNumber="549">Type</typeStatus>
locality
</emphasis>
.
<collectingCountry id="F357766DFFA9C43F6877FF0CFE8BFDEC" box="[314,376,541,566]" name="Chad" pageId="18" pageNumber="549">Chad</collectingCountry>
, Tibetsi area; groundwater in dry channel of river Uet Duar near village Totous, northern
<collectingCountry id="F357766DFFA9C43F6C14FF0CFA6BFDEC" box="[1369,1432,541,566]" name="Chad" pageId="18" pageNumber="549">Chad</collectingCountry>
;
<geoCoordinate id="EE74503AFFA9C43F69DAFF55FED3FD86" box="[151,288,576,605]" direction="north" orientation="latitude" pageId="18" pageNumber="549" precision="15" value="19.439444">19 o26'22'' N</geoCoordinate>
<geoCoordinate id="EE74503AFFA9C43F6865FF55FE42FD86" box="[296,433,576,604]" direction="east" orientation="longitude" pageId="18" pageNumber="549" precision="15" value="17.529722">17o31'47'' E</geoCoordinate>
; altitude:
<quantity id="4CB89B18FFA9C43F6B6DFF55FD94FD86" box="[544,615,580,605]" metricMagnitude="2" metricUnit="m" metricValue="5.85" pageId="18" pageNumber="549" unit="m" value="585.0">585 m</quantity>
(
<figureCitation id="137B2A78FFA9C43F6B3BFF55FD44FD86" box="[630,695,580,605]" captionStart="FIGURE 1" captionStartId="3.[151,250,1922,1944]" captionTargetBox="[252,1315,194,1885]" captionTargetId="figure@3.[252,1316,194,1886]" captionTargetPageId="3" captionText="FIGURE 1. Locations for sampling of groundwater fauna in the Ounianga-Tibesti region (northern Chad). White arrows: shore of the lakes Ounianga Kebir and Bokou; white circles: the river bank near the village Yebi Bou, a spring near the village Gonoa and the river bed of the Uet Duar at the village Totous." httpUri="https://zenodo.org/record/243646/files/figure.png" pageId="18" pageNumber="549">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BFF36FDFFA9C43F698AFF79FDFFFD13" blockId="18.[151,1437,541,2009]" pageId="18" pageNumber="549">
<emphasis id="B934EAEFFFA9C43F698AFF79FE40FD5B" bold="true" box="[199,435,616,641]" pageId="18" pageNumber="549">Material examined.</emphasis>
<typeStatus id="54FB885FFFA9C43F68F0FF79FDD9FD5B" box="[445,554,616,641]" pageId="18" pageNumber="549" type="holotype">Holotype</typeStatus>
: adult female (length 390 µm), completely dissected, mounted on one slide; collected on
<date id="FFFE103DFFA9C43F6864FF9DFE24FD7E" box="[297,471,652,677]" pageId="18" pageNumber="549" value="2014-03-14">14 March 2014</date>
from ground water on the bottom of
<quantity id="4CB89B18FFA9C43F6AC9FF9DFC36FD7E" box="[900,965,652,677]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="549" unit="m" value="1.5">1.5 m</quantity>
deep Karaman-Chappuis pit; access No.: PMSL-Copepoda-Brancelj-1362.
</paragraph>
<paragraph id="8BFF36FDFFA9C43F698AFFC4FDB7FCCB" blockId="18.[151,1437,541,2009]" pageId="18" pageNumber="549">
<typeStatus id="54FB885FFFA9C43F698AFFC4FECAFD37" box="[199,313,725,749]" pageId="18" pageNumber="549" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D46FD74FFA9C43F680EFFC5FE62FD37" box="[323,401,724,749]" pageId="18" pageNumber="549" type="adult">4 adult</specimenCount>
females stored in 70% alcohol; collected on the same date and location as
<typeStatus id="54FB885FFFA9C43F6DAEFFC5FABBFD37" box="[1251,1352,724,749]" pageId="18" pageNumber="549" type="holotype">holotype</typeStatus>
; access No.: PMSL-Copepoda-Brancelj-1363.
</paragraph>
</subSubSection>
<subSubSection id="C35A6576FFA9C43F698AFE0DFF17FCA7" pageId="18" pageNumber="549" type="etymology">
<paragraph id="8BFF36FDFFA9C43F698AFE0DFF17FCA7" blockId="18.[151,1437,541,2009]" pageId="18" pageNumber="549">
<emphasis id="B934EAEFFFA9C43F698AFE0DFEBDFCEF" bold="true" box="[199,334,796,821]" pageId="18" pageNumber="549">Etymology.</emphasis>
The new species is named after Jo Vermeir, a member of the
<collectingCountry id="F357766DFFA9C43F6D66FE0DFB95FCEF" box="[1067,1126,796,821]" name="Chad" pageId="18" pageNumber="549">Chad</collectingCountry>
expedition. I dedicate this species to him in a honour of our long-lasting friendship and his reliability during many expeditions in remote places.
</paragraph>
</subSubSection>
<subSubSection id="C35A6576FFA9C438698AFE99FBAFFB7A" lastPageId="21" lastPageNumber="552" pageId="18" pageNumber="549" type="description">
<paragraph id="8BFF36FDFFA9C43F698AFE99FBF4FB1A" blockId="18.[151,1437,541,2009]" pageId="18" pageNumber="549">
<emphasis id="B934EAEFFFA9C43F698AFE99FEAAFC7B" bold="true" box="[199,345,904,929]" pageId="18" pageNumber="549">Description.</emphasis>
Female, body length, measured from tip of rostrum to posterior margin of caudal rami, 384409 µm (mean: 388 µm; n = 5); elongated, slightly compressed in dorsoventral direction; largest body width on fourth pedigerous somite; preserved specimens colourless (Figs. 9A, B). Naupliar eye not discernible. Rostrum very small, triangular; no sensilla observed. Integumental window not discernible on any somite. Posterior margins of prosomites and urosomites smooth dorsally (Figs. 9A, B). Genital double-somite as long as wide. Genital complex with single copulatory pore, poorly sclerotized, broadly rounded; seminal receptacles small, simple. Fused plate with reduced P6 positioned well above seminal receptacles (Fig. 10E). Anal segment on ventral side with 2 groups of minute spinules mediolaterally (Figs. 9 B, C). Anal operculum large; 2 sensilla positioned anteriorly; free margin almost straight, not reaching distal end of anal somite (Figs. 9A, B, C).
</paragraph>
<paragraph id="8BFF36FDFFA9C43F698AF9DDFDBCFA3A" blockId="18.[151,1437,541,2009]" pageId="18" pageNumber="549">Caudal rami slightly conical, slightly curved, diverging; each ramus about 3.5-times as long as wide, with few small spines at the distal inner corner (Fig. 9C). Anterolateral external accessory seta (I) slightly shorter than width of furcal ramus, inserted at about 2/3 of its length. Anterolateral external accessory seta (II) about 1.5-times longer than seta I, positioned close to it. Posterolateral seta (III) positioned close to seta II, about twice as long as seta I. Outer apical seta (IV), about 0.5 length of caudal ramus, spiniform, smooth. Inner apical seta (V) straight, with several minute spines on both sides, about 5-times as long as caudal ramus. Inner accessory seta (VI) as long as outer apical seta, soft, smooth. Dorsal seta (VII) slightly longer than caudal ramus, inserted on internal side of it, at about 2/3 of its length (Figs. 9A, B, C).</paragraph>
<paragraph id="8BFF36FDFFA9C43F698AF8FDFB67F9F2" blockId="18.[151,1437,541,2009]" pageId="18" pageNumber="549">Antennule (Fig. 9D) as long as cephalothorax, 7-segmented. Aesthetasc (ae) on fourth segment cylindrical, with rounded tip, overreaching tip of seventh segment. Setal formula: 0.4.3.2+ae.1.1. 8+ae.</paragraph>
<paragraph id="8BFF36FDFFA9C43F698AFB25FB30F94F" blockId="18.[151,1437,541,2009]" pageId="18" pageNumber="549">Antenna (Fig. 9E) with allobasis; small, 1-segmented Exp; robust 1-segmented Enp. Exp with one seta apically. 3 strong spines on outer margin of Enp, increasing in length distally; apical armature consisting of one short and one long robust spine on outermost locations; 3 geniculate setae between both spines.</paragraph>
<paragraph id="8BFF36FDFFA9C43F698AFBB1FBAAF907" blockId="18.[151,1437,541,2009]" pageId="18" pageNumber="549">Mandible (Fig. 9F) short, robust, with 3 strongly chitinised teeth on cutting edge. Dorsal seta near gnathobase. Mandibular palp relatively long, 1-segmented; apically 2 long setae, similar in length.</paragraph>
<paragraph id="8BFF36FDFFA9C43F698AFBF9FEC1F8FE" blockId="18.[151,1437,541,2009]" pageId="18" pageNumber="549">Maxillule (Fig. 9G) with 3 strong, long spines on praecoxal arthrite. Coxa with 2 setae unequal in length. Basis with one seta.</paragraph>
<paragraph id="8BFF36FDFFA9C43F698AFA21FCECF8B7" blockId="18.[151,1437,541,2009]" pageId="18" pageNumber="549">Maxilla (Fig. 9H) 2-segmented; syncoxa with 2 endites, each with 2 elements apically. Basis with strong beaklike outgrowth. Enp small, with 2 setae unequal in length.</paragraph>
<paragraph id="8BFF36FDFFA9C43F698AFA69FE34F802" blockId="18.[151,1437,541,2009]" pageId="18" pageNumber="549">
Maxilliped (Fig.
<date id="FFFE103DFFA9C43F68C8FA69FE6EF84A" box="[389,413,1912,1936]" pageId="18" pageNumber="549">9I</date>
) comprising syncoxa, basis, 1-segmented Enp. Syncoxa with no seta distally. Basis 5-times as long as wide. Enp drawn out into strong, acutely curved claw; about half of basis length; additional armature represented by a short seta.
</paragraph>
<paragraph id="8BFF36FDFFA8C43E69DAFA83FAABF81D" blockId="19.[151,1435,1935,1991]" pageId="19" pageNumber="550">
<emphasis id="B934EAEFFFA8C43E69DAFA83FEE1F87D" bold="true" box="[151,274,1938,1960]" pageId="19" pageNumber="550">FIGURE 9.</emphasis>
<taxonomicName id="4C404D7EFFA8C43E6855FA82FE20F872" box="[280,467,1939,1960]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="19" pageNumber="550" phylum="Arthropoda" rank="species" species="joi" status="sp. nov.">
<emphasis id="B934EAEFFFA8C43E6855FA82FE20F872" box="[280,467,1939,1960]" italics="true" pageId="19" pageNumber="550">Parastenocaris joi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2075794FFA8C43E6894FA83FDDAF87D" box="[473,553,1938,1959]" pageId="19" pageNumber="550" rank="species">
<emphasis id="B934EAEFFFA8C43E6894FA83FDDAF87D" bold="true" box="[473,553,1938,1959]" pageId="19" pageNumber="550">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<typeStatus id="54FB885FFFA8C43E6B78FA83FD7DF872" box="[565,654,1938,1960]" pageId="19" pageNumber="550" type="holotype">holotype</typeStatus>
female: A, habitus dorsal view; B, urosome, lateral view; C, 5th urosomal and anal somite with furcal rami, dorsal view; D, antennule; E, antenna; F, mandible; G, maxillule; H, maxilla; I, maxilliped.
</paragraph>
<paragraph id="8BFF36FDFFAFC43969DAFAA1FAC0F81C" blockId="20.[151,1331,1968,1990]" box="[151,1331,1968,1990]" pageId="20" pageNumber="551">
<emphasis id="B934EAEFFFAFC43969DAFAA1FEECF81F" bold="true" box="[151,287,1968,1990]" pageId="20" pageNumber="551">FIGURE 10.</emphasis>
<taxonomicName id="4C404D7EFFAFC4396868FAA0FE12F81C" box="[293,481,1969,1990]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="joi" status="sp. nov.">
<emphasis id="B934EAEFFFAFC4396868FAA0FE12F81C" box="[293,481,1969,1990]" italics="true" pageId="20" pageNumber="551">Parastenocaris joi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2075794FFAFC43968AAFAA1FDCBF81F" box="[487,568,1968,1989]" pageId="20" pageNumber="551" rank="species">
<emphasis id="B934EAEFFFAFC43968AAFAA1FDCBF81F" bold="true" box="[487,568,1968,1989]" pageId="20" pageNumber="551">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<typeStatus id="54FB885FFFAFC4396B08FAA1FD6DF81C" box="[581,670,1968,1990]" pageId="20" pageNumber="551" type="holotype">holotype</typeStatus>
female: A, P1; B, P2; C, P3; D, P4; E, P5 &amp; P6 and genital field.
</paragraph>
<paragraph id="8BFF36FDFFAEC438698AFD89FE72FE9A" blockId="21.[151,1437,151,1977]" pageId="21" pageNumber="552">P1 (Fig. 10A): basis with row of spinules on anterior margin distally. 3-segmented Exp, 2-segmented Enp; equal in length. Exp-1 with one strong outer spine; Exp-2 with no spine or seta; Exp-3 with one strong spine subapically; 3 setae apically, increasing in length, longer 2 genicualte. Longest geniculate seta longer than Exp. Enp-1 with no seta or spine; row of spines along outer margin. Enp-2 with 2 setae apically, longer one geniculate and longer than End.</paragraph>
<paragraph id="8BFF36FDFFAEC438698AFC5DFE9BFDE6" blockId="21.[151,1437,151,1977]" pageId="21" pageNumber="552">P2 (Fig. 10B): basis with one spiniform outer seta. 3-segmented Exp, 1-segmented Enp. Enp about 1/2 length of Exp-1. Exp-1 about 4-times as long as wide, with 2 spines along outer margin. Proximal spine smooth, at 1/4 of Exp-1 length; distal one with small spinules, at the distal corner. Exp-2 about 3-times as long as wide, with few spinules along outer margin. Exp-3 as long as Exp-2, with 2 outer spines; proximal very sort, distal about 1/3 longer than Exp-3; 2 spiniform setae apically; shorter one with spinules unilaterally, longer one bipinnate. Enp-1 4.5- times as long as wide; with 4 elements apically: 3 short spiniform setae, one long smooth seta, about 1.5-times longer than Enp-1.</paragraph>
<paragraph id="8BFF36FDFFAEC438698AFF59FD77FD16" blockId="21.[151,1437,151,1977]" pageId="21" pageNumber="552">P3 (Fig. 10C): basis with one long, thin, smooth seta on outer margin. 2-segmented Exp, 1-segmented Enp. Enp about 0.5 length of Exp-1. Exp-1 similar to Exp-1 of P2. Exp-2 with one long spine subapically, one long spiniform seta apically. Enp-1 about 6-times as long as wide; with 3 spines unequal in length apically; one very short, the longest about 0.5 length of Enp-1.</paragraph>
<paragraph id="8BFF36FDFFAEC4386982FFC9FF3FFC36" blockId="21.[151,1437,151,1977]" pageId="21" pageNumber="552">P4 (Fig. 10D): basis with one long, thin, smooth seta on outer margin. 3-segmented Exp, 1-segmented Enp. Enp about 1/2 length of Exp-1. Exp-1 with 3 spines, increasing in length; proximal shortest, distal longest; proximal spine at 1/3 of outer margin length; middle spine at 2/3 of margin length; distal one subapically. Exp-2 with 2 small spines; one at 2/3 of margin length, second one subapically. Exp-3 4.5-times as long as wide, with 2 relatively weak spines on outer margin; proximal at 3/4 of margin length, distal one subapically; two elements apically: one spiniform seta as long as Exp-3, one about 3-times longer than Exp-3. Enp-1 5-times as wide as long, about 0.5-times length of Exp-1. 3 elements apically: 2 very short spines, one spiniform seta about 1.5-length of Enp.</paragraph>
<paragraph id="8BFF36FDFFAEC438698AFEE6FC43FBEF" blockId="21.[151,1437,151,1977]" pageId="21" pageNumber="552">P5 (Fig. 10E) well separated, each half triangular in shape, with sharply pointed tip. Outer margin with 2 short spines distally, one long smooth seta proximally. Inner margin smooth.</paragraph>
<paragraph id="8BFF36FDFFAEC438698AF92EFC0CFB82" blockId="21.[151,1437,151,1977]" box="[199,1023,1087,1112]" pageId="21" pageNumber="552">P6 (Fig. 10E) reduced to small, fused plate, with pointed tip on each side.</paragraph>
<paragraph id="8BFF36FDFFAEC438698AF975FE60FBA7" blockId="21.[151,1437,151,1977]" box="[199,403,1124,1149]" pageId="21" pageNumber="552">Male. Not known.</paragraph>
<paragraph id="8BFF36FDFFAEC438698AF996FBAFFB7A" blockId="21.[151,1437,151,1977]" box="[199,1116,1159,1184]" pageId="21" pageNumber="552">
<emphasis id="B934EAEFFFAEC438698AF996FEBEFB7A" bold="true" box="[199,333,1159,1184]" pageId="21" pageNumber="552">Variability.</emphasis>
Except slight differences in body length, no variability was observed.
</paragraph>
</subSubSection>
<subSubSection id="C35A6576FFAEC43B698AF9BDFD70FBEE" lastPageId="22" lastPageNumber="553" pageId="21" pageNumber="552" type="discussion">
<paragraph id="8BFF36FDFFAEC438698AF9BDFCABFAEA" blockId="21.[151,1437,151,1977]" pageId="21" pageNumber="552">
<emphasis id="B934EAEFFFAEC438698AF9BDFEC8FB1F" bold="true" box="[199,315,1196,1221]" pageId="21" pageNumber="552">Remarks.</emphasis>
Since only females were collected, it is not possible to allocate the new species accurately to any of the many new genera recently described or resurrected (
<bibRefCitation id="EFD14B0CFFAEC4386A48F9DEFBFFFB32" author="Karanovic" box="[773,1036,1231,1256]" pageId="21" pageNumber="552" refString="Karanovic, T. &amp; Lee, W. (2012) A new species of Parastenocaris from Korea, with a redescription of the closely related P. biwae from Japan (Coppeoda: Harpacticoida: Parastenocarididae). Journal of Species Research, 1, 4 - 34. http: // dx. doi. org / 10.12651 / JSR. 2012.1.1.004" type="journal article" year="2012">Karanovic &amp; Lee 2012</bibRefCitation>
). For that reason this species should be provisionally included into the genus
<emphasis id="B934EAEFFFAEC4386B2CF9E4FCE4FAD6" box="[609,791,1269,1292]" italics="true" pageId="21" pageNumber="552">Parastenoscaris</emphasis>
Kessler, 1913,
<emphasis id="B934EAEFFFAEC4386A8BF9E4FBCBFAD6" box="[966,1080,1268,1292]" italics="true" pageId="21" pageNumber="552">sensu lato</emphasis>
, until males are found enabling precise allocation of the species into its definitive genus status.
</paragraph>
<paragraph id="8BFF36FDFFAEC438698AF82DFF2CFA3E" blockId="21.[151,1437,151,1977]" pageId="21" pageNumber="552">
So far only five species of parastenocaridids have been recorded from the Sahara (
<tableCitation id="C6C20346FFAEC4386DECF82DFB09FA8E" box="[1185,1274,1340,1365]" captionStart="TABLE 1" captionStartId="1.[151,239,690,713]" captionTargetBox="[151,1420,773,1886]" captionTargetPageId="1" captionText="TABLE 1. List of Copepoda species from the Sahara (after Botosaneanu 1986 and Juberthie &amp; Decu 2001); three species from Chad (this study) are included." httpUri="http://table.plazi.org/id/DF3F6675FFBAC42C69DAFFA3FDE9FD33" pageId="21" pageNumber="552" tableUuid="DF3F6675FFBAC42C69DAFFA3FDE9FD33">Table 1</tableCitation>
). From
<collectingCountry id="F357766DFFAEC4386C2DF82DFA6BFA8F" box="[1376,1432,1340,1365]" name="Serbia" pageId="21" pageNumber="552">Mali</collectingCountry>
,
<bibRefCitation id="EFD14B0CFFAEC43869DAF84EFEA0FAA2" author="Chappuis" box="[151,339,1375,1400]" pageId="21" pageNumber="552" refString="Chappuis, P. A. (1959) Deux Parastenocaris des sables du Niger. Bulletin de l'Institut Francais Afrique Noire, 21 A, 1211 - 1215." type="journal article" year="1959">Chappuis (1959)</bibRefCitation>
reported
<taxonomicName id="4C404D7EFFAEC438688CF871FC56FAA2" authority="Chappuis, 1959" authorityName="Chappuis" authorityYear="1959" box="[449,933,1375,1400]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="21" pageNumber="552" phylum="Arthropoda" rank="species" species="nigerianus">
<emphasis id="B934EAEFFFAEC438688CF871FD1FFAAD" box="[449,748,1376,1399]" italics="true" pageId="21" pageNumber="552">Parastenocaris nigerianus</emphasis>
<bibRefCitation id="EFD14B0CFFAEC4386BB9F84EFC56FAA2" author="Chappuis" box="[756,933,1375,1400]" pageId="21" pageNumber="552" refString="Chappuis, P. A. (1959) Deux Parastenocaris des sables du Niger. Bulletin de l'Institut Francais Afrique Noire, 21 A, 1211 - 1215." type="journal article" year="1959">Chappuis, 1959</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C404D7EFFAEC4386A93F871FA6BFAA2" authority="Chappuis, 1959" authorityName="Chappuis" authorityYear="1959" box="[990,1432,1375,1400]" class="Maxillopoda" family="Parastenocarididae" genus="Monodicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="21" pageNumber="552" phylum="Arthropoda" rank="species" species="monodi">
<emphasis id="B934EAEFFFAEC4386A93F871FB3AFAA2" box="[990,1225,1376,1400]" italics="true" pageId="21" pageNumber="552">Monodicaris monodi</emphasis>
(
<bibRefCitation id="EFD14B0CFFAEC4386D94F84EFA63FAA2" author="Chappuis" box="[1241,1424,1375,1400]" pageId="21" pageNumber="552" refString="Chappuis, P. A. (1959) Deux Parastenocaris des sables du Niger. Bulletin de l'Institut Francais Afrique Noire, 21 A, 1211 - 1215." type="journal article" year="1959">Chappuis, 1959</bibRefCitation>
)
</taxonomicName>
. A further three species have been reported from
<collectingCountry id="F357766DFFAEC4386B8EF895FCEFFA47" box="[707,796,1412,1437]" name="Algeria" pageId="21" pageNumber="552">Algeria</collectingCountry>
:
<taxonomicName id="4C404D7EFFAEC4386A65F894FB10FA47" authority="Chappuis, 1954" authorityName="Chappuis" authorityYear="1954" box="[808,1251,1412,1437]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="21" pageNumber="552" phylum="Arthropoda" rank="species" species="kabyla">
<emphasis id="B934EAEFFFAEC4386A65F894FBD4FA46" box="[808,1063,1412,1436]" italics="true" pageId="21" pageNumber="552">Parastenocaris kabyla</emphasis>
<bibRefCitation id="EFD14B0CFFAEC4386D62F895FB10FA47" author="Chappuis" box="[1071,1251,1412,1437]" pageId="21" pageNumber="552" refString="Chappuis, P. A. (1954) Harpacticides psammiques recoltes par Cl. Delamare Deboutteville en Mediterranee. Vie et Milieu, 4, 259 - 276." type="journal article" year="1954">Chappuis, 1954</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C404D7EFFAEC4386DBCF894FE5EFA1A" authority="Bozic, 1978" authorityName="Bozic" authorityYear="1978" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="21" pageNumber="552" phylum="Arthropoda" rank="species" species="ahaggarica">
<emphasis id="B934EAEFFFAEC4386DBCF894FEEBFA1A" italics="true" pageId="21" pageNumber="552">Parastenocaris ahaggarica</emphasis>
<bibRefCitation id="EFD14B0CFFAEC438686CF8B6FE5EFA1A" author="Bozic" box="[289,429,1447,1472]" pageId="21" pageNumber="552" refString="Bozic, B. (1978) Une nouvelle Parastenocaris du Hoggar (Crustacea, Copepoda, Harpacticoida). Bulletin du Museum National d'Histoire Naturelle, Paris, 514, 239 - 244." type="journal article" year="1978">Božić, 1978</bibRefCitation>
</taxonomicName>
; and
<emphasis id="B934EAEFFFAEC43868A0F8B9FD7AFA1A" box="[493,649,1448,1472]" italics="true" pageId="21" pageNumber="552">Cotarellicaris</emphasis>
<taxonomicName id="4C404D7EFFAEC4386BDFF8B9FC36FA1A" authority="(Rouch, 1987)" authorityName="Rouch" authorityYear="1987" baseAuthorityName="Rouch" baseAuthorityYear="1987" box="[658,965,1447,1472]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="21" pageNumber="552" phylum="Arthropoda" rank="species" species="numidiensis">
<emphasis id="B934EAEFFFAEC4386BDFF8B9FCEBFA1A" box="[658,792,1448,1472]" italics="true" pageId="21" pageNumber="552">numidiensis</emphasis>
(
<bibRefCitation id="EFD14B0CFFAEC4386A65F8B6FC4FFA1A" author="Rouch" box="[808,956,1447,1472]" pageId="21" pageNumber="552" refString="Rouch, R. (1987) Copepodes harpacticoides stygobies d'Algerie. Bijdragen tot der Dierkunde, 57, 71 - 86." type="journal article" year="1987">Rouch, 1987</bibRefCitation>
)
</taxonomicName>
(see
<bibRefCitation id="EFD14B0CFFAEC4386D49F8B6FB41FA1A" author="Chappuis" box="[1028,1202,1447,1472]" pageId="21" pageNumber="552" refString="Chappuis, P. A. (1954) Harpacticides psammiques recoltes par Cl. Delamare Deboutteville en Mediterranee. Vie et Milieu, 4, 259 - 276." type="journal article" year="1954">Chappuis 1954</bibRefCitation>
;
<bibRefCitation id="EFD14B0CFFAEC4386DF2F8B6FAB6FA1A" author="Bozic" box="[1215,1349,1447,1472]" pageId="21" pageNumber="552" refString="Bozic, B. (1978) Une nouvelle Parastenocaris du Hoggar (Crustacea, Copepoda, Harpacticoida). Bulletin du Museum National d'Histoire Naturelle, Paris, 514, 239 - 244." type="journal article" year="1978">Božić 1978</bibRefCitation>
;
<bibRefCitation id="EFD14B0CFFAEC4386C1EF8B6FF20FA3E" author="Rouch" pageId="21" pageNumber="552" refString="Rouch, R. (1987) Copepodes harpacticoides stygobies d'Algerie. Bijdragen tot der Dierkunde, 57, 71 - 86." type="journal article" year="1987">Rouch 1987</bibRefCitation>
).
</paragraph>
<paragraph id="8BFF36FDFFAEC438698AF8E0FCEAF942" blockId="21.[151,1437,151,1977]" pageId="21" pageNumber="552">
More detailed analysis of possible differences between those five species and the new species is difficult as the previous descriptions were frequently incomplete. In fact, there are no figures for
<taxonomicName id="4C404D7EFFAEC4386D73FB04FB55F9F6" box="[1086,1190,1556,1580]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="21" pageNumber="552" phylum="Arthropoda" rank="species" species="kabyla">
<emphasis id="B934EAEFFFAEC4386D73FB04FB55F9F6" box="[1086,1190,1556,1580]" italics="true" pageId="21" pageNumber="552">P. kabyla</emphasis>
</taxonomicName>
, which was collected close to the coast, and only males were found.
<bibRefCitation id="EFD14B0CFFAEC4386BD2FB26FCAAF98A" author="Chappuis" box="[671,857,1591,1616]" pageId="21" pageNumber="552" refString="Chappuis, P. A. (1954) Harpacticides psammiques recoltes par Cl. Delamare Deboutteville en Mediterranee. Vie et Milieu, 4, 259 - 276." type="journal article" year="1954">Chappuis (1954)</bibRefCitation>
reported that P4 and P5 of the male of
<taxonomicName id="4C404D7EFFAEC4386C40FB28FA80F98A" box="[1293,1395,1592,1616]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="21" pageNumber="552" phylum="Arthropoda" rank="species" species="kabyla">
<emphasis id="B934EAEFFFAEC4386C40FB28FA80F98A" box="[1293,1395,1592,1616]" italics="true" pageId="21" pageNumber="552">P. kabyla</emphasis>
</taxonomicName>
are similar to those in
<taxonomicName id="4C404D7EFFAEC4386828FB4CFCDAF9AF" authority="Schnitter &amp; Chappuis, 1915" authorityName="Schnitter &amp; Chappuis" authorityYear="1915" box="[357,809,1628,1653]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="21" pageNumber="552" phylum="Arthropoda" rank="species" species="fontinalis">
<emphasis id="B934EAEFFFAEC4386828FB4CFE19F9AE" box="[357,490,1628,1652]" italics="true" pageId="21" pageNumber="552">P. fontinalis</emphasis>
Schnitter &amp; Chappuis, 1915
</taxonomicName>
. As figured in Janetzky et al. (1996), P
<quantity id="4CB89B18FFAEC4386D97FB4DFAF0F9AF" box="[1242,1283,1628,1653]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="21" pageNumber="552" unit="in" value="5.0">5 in</quantity>
the female of
<taxonomicName id="4C404D7EFFAEC43869DAFB90FEEFF942" box="[151,284,1664,1688]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="21" pageNumber="552" phylum="Arthropoda" rank="species" species="fontinalis">
<emphasis id="B934EAEFFFAEC43869DAFB90FEEFF942" box="[151,284,1664,1688]" italics="true" pageId="21" pageNumber="552">P. fontinalis</emphasis>
</taxonomicName>
is quite different from those in
<taxonomicName id="4C404D7EFFAEC4386B32FB90FD4BF94D" box="[639,696,1664,1688]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="21" pageNumber="552" phylum="Arthropoda" rank="species" species="joi" status="sp. nov.">
<emphasis id="B934EAEFFFAEC4386B32FB90FD7EF942" box="[639,653,1665,1688]" italics="true" pageId="21" pageNumber="552">P</emphasis>
.
<emphasis id="B934EAEFFFAEC4386BD6FB91FD4BF94D" box="[667,696,1664,1687]" italics="true" pageId="21" pageNumber="552">joi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2075794FFAEC4386BF2FB6EFCEAF942" box="[703,793,1663,1688]" pageId="21" pageNumber="552" rank="species">
<emphasis id="B934EAEFFFAEC4386BF2FB6EFCEAF942" bold="true" box="[703,793,1663,1688]" pageId="21" pageNumber="552">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BFF36FDFFAEC438698AFBB4FCB9F862" blockId="21.[151,1437,151,1977]" pageId="21" pageNumber="552">
For
<taxonomicName id="4C404D7EFFAEC43869BBFBB4FE78F966" box="[246,395,1701,1724]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="21" pageNumber="552" phylum="Arthropoda" rank="species" species="nigerianus">
<emphasis id="B934EAEFFFAEC43869BBFBB4FE78F966" box="[246,395,1701,1724]" italics="true" pageId="21" pageNumber="552">P. nigerianus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C404D7EFFAEC4386889FBB4FDCCF966" box="[452,575,1700,1724]" class="Maxillopoda" family="Parastenocarididae" genus="Monodicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="21" pageNumber="552" phylum="Arthropoda" rank="species" species="monodi">
<emphasis id="B934EAEFFFAEC4386889FBB4FDCCF966" box="[452,575,1700,1724]" italics="true" pageId="21" pageNumber="552">M. monodi</emphasis>
</taxonomicName>
, only figures of males are presented and there are not enough details in the text to make any concise conclusions on taxonomic relations between those two species and the new one. However, the shape and armature of furcal ramus in males, and especially the position of lateral and dorsal setae, are quite different from those observed in the female of the new species. In
<taxonomicName id="4C404D7EFFAEC4386AE4FA00FBD5F8F2" box="[937,1062,1808,1832]" class="Maxillopoda" family="Parastenocarididae" genus="Monodicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="21" pageNumber="552" phylum="Arthropoda" rank="species" species="monodi">
<emphasis id="B934EAEFFFAEC4386AE4FA00FBD5F8F2" box="[937,1062,1808,1832]" italics="true" pageId="21" pageNumber="552">M. monodi</emphasis>
</taxonomicName>
furcal ramus is longer, straight, conical, with width/length ratio of about 1:6 (about 1:
<quantity id="4CB89B18FFAEC4386BBFFA25FCC2F897" box="[754,817,1844,1869]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" pageId="21" pageNumber="552" unit="in" value="3.5">3.5 in</quantity>
the new species) and with only one external lateral seta (anterolateral external accessory seta (II)?)
<emphasis id="B934EAEFFFAEC4386BC5FA48FD6CF8AA" box="[648,671,1881,1904]" italics="true" pageId="21" pageNumber="552">vs</emphasis>
. three relatively long setae (I-III) in the new species. Outer apical seta (IV) in
<taxonomicName id="4C404D7EFFAEC4386851FA6CFE66F84E" box="[284,405,1916,1940]" class="Maxillopoda" family="Parastenocarididae" genus="Monodicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="21" pageNumber="552" phylum="Arthropoda" rank="species" species="monodi">
<emphasis id="B934EAEFFFAEC4386851FA6CFE66F84E" box="[284,405,1916,1940]" italics="true" pageId="21" pageNumber="552">M. monodi</emphasis>
</taxonomicName>
looks long and soft, but short and spiniform in the new species. Only insertion of dorsal seta (VII) at 1/2 of inner margin length is similar to those in
<emphasis id="B934EAEFFFAEC4386A44FAB0FCB9F862" box="[777,842,1953,1976]" italics="true" pageId="21" pageNumber="552">
<taxonomicName id="4C404D7EFFAEC4386A44FAB0FCB6F862" box="[777,837,1953,1976]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="21" pageNumber="552" phylum="Arthropoda" rank="species" species="joi">P. joi</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BFF36FDFFADC43B698AFD89FE67FE52" blockId="22.[151,1437,151,1077]" pageId="22" pageNumber="553">
In
<taxonomicName id="4C404D7EFFADC43B69A4FD88FE8CFF75" box="[233,383,152,176]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="22" pageNumber="553" phylum="Arthropoda" rank="species" species="nigerianus">
<emphasis id="B934EAEFFFADC43B69A4FD88FE8CFF75" box="[233,383,152,176]" italics="true" pageId="22" pageNumber="553">P. nigerianus</emphasis>
</taxonomicName>
width/length of furca is about 1:4; divergent and slightly curved outward, like in the new species. In
<taxonomicName id="4C404D7EFFADC43B6859FDACFE45FF0E" box="[276,438,188,212]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="22" pageNumber="553" phylum="Arthropoda" rank="species" species="numidiensis">
<emphasis id="B934EAEFFFADC43B6859FDACFE45FF0E" box="[276,438,188,212]" italics="true" pageId="22" pageNumber="553">P. numidiensis</emphasis>
</taxonomicName>
one single external seta is figured (anterolateral external accessory seta (II)?) at about 1/ 3 of external margin length, compared to three setae in the new species: anterolateral external accessory seta (I), anterolateral external accessory seta (II) and posterolateral seta (III); all inserted at about 2/3 of external margin length (Fig. 9C). Shape and position of outer apical seta (IV) is similar in both species. Insertion of dorsal seta (VII) in
<taxonomicName id="4C404D7EFFADC43B69B8FC5CFE7AFEBE" box="[245,393,333,356]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="22" pageNumber="553" phylum="Arthropoda" rank="species" species="nigerianus">
<emphasis id="B934EAEFFFADC43B69B8FC5CFE7AFEBE" box="[245,393,333,356]" italics="true" pageId="22" pageNumber="553">P. nigerianus</emphasis>
</taxonomicName>
is at the distal middle part of furcal rami, while in the new species is at internal side of it, at about 2/3 of its length.
</paragraph>
<paragraph id="8BFF36FDFFADC43B698AFC84FC19FDE7" blockId="22.[151,1437,151,1077]" pageId="22" pageNumber="553">
For two species enough detailed description exists to make more firm conclusions on taxonomic relations between them and the new species. In
<taxonomicName id="4C404D7EFFADC43B6B1DFCA8FCCFFE0A" authority="End P" authorityName="End P" box="[592,828,439,464]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="22" pageNumber="553" phylum="Arthropoda" rank="species" species="ahaggarica">
<emphasis id="B934EAEFFFADC43B6B1DFCA8FD1DFE0A" box="[592,750,440,464]" italics="true" pageId="22" pageNumber="553">P. ahaggarica</emphasis>
End P
</taxonomicName>
2P4 of the female have one apical element each and thus are different from those in
<taxonomicName id="4C404D7EFFADC43B68B8FCCCFDDDFE2E" box="[501,558,477,500]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="22" pageNumber="553" phylum="Arthropoda" rank="species" species="joi">
<emphasis id="B934EAEFFFADC43B68B8FCCCFDF0FE2E" box="[501,515,477,500]" italics="true" pageId="22" pageNumber="553">P</emphasis>
.
<emphasis id="B934EAEFFFADC43B6B5CFCCCFDDDFE2E" box="[529,558,477,500]" italics="true" pageId="22" pageNumber="553">joi</emphasis>
</taxonomicName>
, with 3 or 4 elements of different lengths on each End. Also P
<quantity id="4CB89B18FFADC43B6DAEFCCDFAFEFE2F" box="[1251,1293,476,501]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="22" pageNumber="553" unit="in" value="5.0">5 in</quantity>
both species differ in shape and number of elements. P
<quantity id="4CB89B18FFADC43B6B38FCEEFD53FDC2" box="[629,672,511,536]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="22" pageNumber="553" unit="in" value="5.0">5 in</quantity>
<taxonomicName id="4C404D7EFFADC43B6BE5FF10FD11FDCD" box="[680,738,512,536]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="22" pageNumber="553" phylum="Arthropoda" rank="species" species="joi">
<emphasis id="B934EAEFFFADC43B6BE5FF10FD45FDC2" box="[680,694,513,536]" italics="true" pageId="22" pageNumber="553">P</emphasis>
.
<emphasis id="B934EAEFFFADC43B6B88FF11FD11FDCD" box="[709,738,512,535]" italics="true" pageId="22" pageNumber="553">joi</emphasis>
</taxonomicName>
has three elements laterally and a pointed distal end while
<taxonomicName id="4C404D7EFFADC43B6CCAFF10FEEBFDE6" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="22" pageNumber="553" phylum="Arthropoda" rank="species" species="ahaggarica">
<emphasis id="B934EAEFFFADC43B6CCAFF10FEEBFDE6" italics="true" pageId="22" pageNumber="553">P. ahaggarica</emphasis>
</taxonomicName>
has four elements; two of them apically and with no distinct tip.
</paragraph>
<paragraph id="8BFF36FDFFADC43B698AFF56FA65FC7F" blockId="22.[151,1437,151,1077]" pageId="22" pageNumber="553">
There are some similarities between
<taxonomicName id="4C404D7EFFADC43B6B2CFF58FCF2FDBA" box="[609,769,584,608]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="22" pageNumber="553" phylum="Arthropoda" rank="species" species="numidiensis">
<emphasis id="B934EAEFFFADC43B6B2CFF58FCF2FDBA" box="[609,769,584,608]" italics="true" pageId="22" pageNumber="553">P. numidiensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C404D7EFFADC43B6A75FF58FC81FD85" box="[824,882,584,608]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="22" pageNumber="553" phylum="Arthropoda" rank="species" species="joi">
<emphasis id="B934EAEFFFADC43B6A75FF58FCB5FDBA" box="[824,838,585,608]" italics="true" pageId="22" pageNumber="553">P</emphasis>
.
<emphasis id="B934EAEFFFADC43B6A19FF59FC81FD85" box="[852,882,584,607]" italics="true" pageId="22" pageNumber="553">joi</emphasis>
</taxonomicName>
in the armatures of P2 and P4, but not of P3. End P3 of
<taxonomicName id="4C404D7EFFADC43B6997FF7CFEE0FD5E" box="[218,275,621,644]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="22" pageNumber="553" phylum="Arthropoda" rank="species" species="joi">
<emphasis id="B934EAEFFFADC43B6997FF7CFF1BFD5E" box="[218,232,621,644]" italics="true" pageId="22" pageNumber="553">P</emphasis>
.
<emphasis id="B934EAEFFFADC43B69BBFF7CFEE0FD5E" box="[246,275,621,644]" italics="true" pageId="22" pageNumber="553">joi</emphasis>
</taxonomicName>
has three elements apically, while in
<taxonomicName id="4C404D7EFFADC43B6BF7FF7CFCA8FD5E" box="[698,859,620,644]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="22" pageNumber="553" phylum="Arthropoda" rank="species" species="numidiensis">
<emphasis id="B934EAEFFFADC43B6BF7FF7CFCA8FD5E" box="[698,859,620,644]" italics="true" pageId="22" pageNumber="553">P. numidiensis</emphasis>
</taxonomicName>
, the distal part of End ends with a sharp point. The distal end of P
<quantity id="4CB89B18FFADC43B6876FF9EFE95FD72" box="[315,358,655,680]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="22" pageNumber="553" unit="in" value="5.0">5 in</quantity>
both species ends with a sharp point, but on the outer margin there are 3 elements in
<taxonomicName id="4C404D7EFFADC43B6C7CFF80FA98FD7D" box="[1329,1387,656,680]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="22" pageNumber="553" phylum="Arthropoda" rank="species" species="joi">
<emphasis id="B934EAEFFFADC43B6C7CFF80FACCFD72" box="[1329,1343,657,680]" italics="true" pageId="22" pageNumber="553">P</emphasis>
.
<emphasis id="B934EAEFFFADC43B6C03FF81FA98FD7D" box="[1358,1387,656,679]" italics="true" pageId="22" pageNumber="553">joi</emphasis>
</taxonomicName>
and four in
<emphasis id="B934EAEFFFADC43B69A4FFA4FE62FD16" box="[233,401,692,716]" italics="true" pageId="22" pageNumber="553">
<taxonomicName id="4C404D7EFFADC43B69A4FFA4FE7EFD16" box="[233,397,692,716]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="22" pageNumber="553" phylum="Arthropoda" rank="species" species="numidiensis">P. numidiensis</taxonomicName>
.
</emphasis>
In
<taxonomicName id="4C404D7EFFADC43B68F8FFA4FDA5FD16" box="[437,598,692,716]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="22" pageNumber="553" phylum="Arthropoda" rank="species" species="numidiensis">
<emphasis id="B934EAEFFFADC43B68F8FFA4FDA5FD16" box="[437,598,692,716]" italics="true" pageId="22" pageNumber="553">P. numidiensis</emphasis>
</taxonomicName>
there is a very obvious incision in the middle of the outer margin, just next to the longest seta on P5, while in
<taxonomicName id="4C404D7EFFADC43B6B7FFFC8FD9DFD35" box="[562,622,728,752]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="22" pageNumber="553" phylum="Arthropoda" rank="species" species="joi">
<emphasis id="B934EAEFFFADC43B6B7FFFC8FDB3FD2A" box="[562,576,729,752]" italics="true" pageId="22" pageNumber="553">P</emphasis>
.
<emphasis id="B934EAEFFFADC43B6B1CFFC9FD9DFD35" box="[593,622,728,751]" italics="true" pageId="22" pageNumber="553">joi</emphasis>
</taxonomicName>
there is only slight depression. There are also differences on the anal operculum, which is short, and with straight free margin in
<taxonomicName id="4C404D7EFFADC43B6A64FFECFC92FCCE" box="[809,865,765,788]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="22" pageNumber="553" phylum="Arthropoda" rank="species" species="joi">
<emphasis id="B934EAEFFFADC43B6A64FFECFCC4FCCE" box="[809,823,765,788]" italics="true" pageId="22" pageNumber="553">P</emphasis>
.
<emphasis id="B934EAEFFFADC43B6A09FFECFC92FCCE" box="[836,865,765,788]" italics="true" pageId="22" pageNumber="553">joi</emphasis>
</taxonomicName>
, but longer and semicircular in
<emphasis id="B934EAEFFFADC43B6D8EFFECFA98FCCE" box="[1219,1387,764,788]" italics="true" pageId="22" pageNumber="553">
<taxonomicName id="4C404D7EFFADC43B6D8EFFECFA95FCCE" box="[1219,1382,764,788]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="22" pageNumber="553" phylum="Arthropoda" rank="species" species="numidiensis">P. numidiensis</taxonomicName>
.
</emphasis>
The two species also differ considerably in the shape of the furcal rami and their armature. In
<taxonomicName id="4C404D7EFFADC43B6DCAFE30FAD9FCE2" box="[1159,1322,800,824]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="22" pageNumber="553" phylum="Arthropoda" rank="species" species="numidiensis">
<emphasis id="B934EAEFFFADC43B6DCAFE30FAD9FCE2" box="[1159,1322,800,824]" italics="true" pageId="22" pageNumber="553">P. numidiensis</emphasis>
</taxonomicName>
the furcal ramus is curved inward, while in
<taxonomicName id="4C404D7EFFADC43B6B46FE54FDB7FC86" box="[523,580,837,860]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="22" pageNumber="553" phylum="Arthropoda" rank="species" species="joi">
<emphasis id="B934EAEFFFADC43B6B46FE54FDEAFC86" box="[523,537,837,860]" italics="true" pageId="22" pageNumber="553">P</emphasis>
.
<emphasis id="B934EAEFFFADC43B6B6AFE54FDB7FC86" box="[551,580,837,860]" italics="true" pageId="22" pageNumber="553">joi</emphasis>
</taxonomicName>
it is curved outward. The lateral setae and dorsal seta in
<taxonomicName id="4C404D7EFFADC43B6DF3FE54FB0BFC86" box="[1214,1272,837,860]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="22" pageNumber="553" phylum="Arthropoda" rank="species" species="joi">
<emphasis id="B934EAEFFFADC43B6DF3FE54FB3FFC86" box="[1214,1228,837,860]" italics="true" pageId="22" pageNumber="553">P</emphasis>
.
<emphasis id="B934EAEFFFADC43B6D97FE54FB0BFC86" box="[1242,1272,837,860]" italics="true" pageId="22" pageNumber="553">joi</emphasis>
</taxonomicName>
are positioned at 2/3 of caudal ramus length, while in
<taxonomicName id="4C404D7EFFADC43B6B1EFE78FD06FC5A" box="[595,757,872,896]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="22" pageNumber="553" phylum="Arthropoda" rank="species" species="numidiensis">
<emphasis id="B934EAEFFFADC43B6B1EFE78FD06FC5A" box="[595,757,872,896]" italics="true" pageId="22" pageNumber="553">P. numidiensis</emphasis>
</taxonomicName>
they are positioned at about 3/4 of the caudal ramus length. The inner apical seta (V) is swollen at the base in
<taxonomicName id="4C404D7EFFADC43B6B8DFE9CFC92FC7E" box="[704,865,908,932]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="22" pageNumber="553" phylum="Arthropoda" rank="species" species="numidiensis">
<emphasis id="B934EAEFFFADC43B6B8DFE9CFC92FC7E" box="[704,865,908,932]" italics="true" pageId="22" pageNumber="553">P. numidiensis</emphasis>
</taxonomicName>
, while in
<taxonomicName id="4C404D7EFFADC43B6A82FE9CFBF4FC7E" box="[975,1031,909,932]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="22" pageNumber="553" phylum="Arthropoda" rank="species" species="joi">
<emphasis id="B934EAEFFFADC43B6A82FE9CFC2EFC7E" box="[975,989,909,932]" italics="true" pageId="22" pageNumber="553">P</emphasis>
.
<emphasis id="B934EAEFFFADC43B6AA7FE9CFBF4FC7E" box="[1002,1031,909,932]" italics="true" pageId="22" pageNumber="553">joi</emphasis>
</taxonomicName>
it tapers evenly from its base to tip.
</paragraph>
<paragraph id="8BFF36FDFFADC43B698AFEA1FD70FBEE" blockId="22.[151,1437,151,1077]" pageId="22" pageNumber="553">
On consideration of the above listed differences between the six species from the Sahara it is evident that the collected specimens belong to the new species,
<taxonomicName id="4C404D7EFFADC43B6B85FEC4FCF6FC36" box="[712,773,981,1004]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="22" pageNumber="553" phylum="Arthropoda" rank="species" species="joi">
<emphasis id="B934EAEFFFADC43B6B85FEC4FD25FC36" box="[712,726,981,1004]" italics="true" pageId="22" pageNumber="553">P</emphasis>
.
<emphasis id="B934EAEFFFADC43B6BA5FEC4FCF6FC36" box="[744,773,981,1004]" italics="true" pageId="22" pageNumber="553">joi</emphasis>
</taxonomicName>
. At the same time the existence of several characters in common between
<taxonomicName id="4C404D7EFFADC43B6826FEE8FDE2FBCA" box="[363,529,1016,1040]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="22" pageNumber="553" phylum="Arthropoda" rank="species" species="numidiensis">
<emphasis id="B934EAEFFFADC43B6826FEE8FDE2FBCA" box="[363,529,1016,1040]" italics="true" pageId="22" pageNumber="553">P. numidiensis</emphasis>
</taxonomicName>
and the new species also indicates that the new species belongs to the genus
<taxonomicName id="4C404D7EFFADC43B69DAF90CFE16FBEE" authority="Kessler, 1913" authorityName="Kessler" authorityYear="1913" box="[151,485,1052,1077]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="22" pageNumber="553" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAEFFFADC43B69DAF90CFEB1FBEE" box="[151,322,1053,1076]" italics="true" pageId="22" pageNumber="553">Parastenocaris</emphasis>
Kessler, 1913
</taxonomicName>
,
<emphasis id="B934EAEFFFADC43B68BDF90CFD8EFBEE" box="[496,637,1053,1076]" italics="true" pageId="22" pageNumber="553">sensu stricto</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>