167 lines
19 KiB
XML
167 lines
19 KiB
XML
<document id="C267D57CE04DFBD338AAFC754EDC3D94" ID-DOI="10.5281/zenodo.178756" ID-GBIF-Dataset="f9ea8775-9b2d-4364-b20a-4738aab6a54c" ID-ISSN="1175-5326" ID-Zenodo-Dep="178756" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459920987951" checkinUser="plazi" docAuthor="Kataev, Boris M. & Liang, Hongbin" docDate="2007" docId="03E987D1FFB2FFFCFF178E863435FCD9" docLanguage="en" docName="zt01604p020.pdf" docOrigin="Zootaxa 1604" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Harpalus viridanus subsp. staudingerianus Schauberger 1932" docType="treatment" docVersion="9" lastPageNumber="9" masterDocId="FFD0FFA9FFB5FFF4FF8088153517FFD3" masterDocTitle="Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini)" masterLastPageNumber="20" masterPageNumber="1" pageNumber="8" updateTime="1698229235843" updateUser="plazi">
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<mods:title id="79ED06DE5D2529F77CCD518BF1544AE9">Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini)</mods:title>
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<mods:name id="03352E5932106D235901583F1269B790" type="personal">
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<mods:namePart id="79F81B565B98FBF9EBD37203330BE419">Kataev, Boris M.</mods:namePart>
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<mods:name id="4629942FD4B0F10A5EFE98DE059878FB" type="personal">
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<mods:roleTerm id="67F34FC9FD1FB5C3AD9A70EDF2E36407">Author</mods:roleTerm>
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<mods:namePart id="DB44E6D80917EF58B053DBB13FEBCF02">Liang, Hongbin</mods:namePart>
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<mods:typeOfResource id="073C4CA3B8513892FFB7EA66236BABAC">text</mods:typeOfResource>
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<mods:title id="2E67A3E8BAF7E689A03C4B62B4E602E6">Zootaxa</mods:title>
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<mods:part id="667CCDA5A5C61CE122EF763D8F4C361A">
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<mods:date id="F78E131CC5570130CD8CDCDC6C86B9D0">2007</mods:date>
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<mods:detail id="9D8EBF57538A85BEDB90ADE5B2EE2A7B" type="volume">
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<mods:number id="29C008568196E85E05108117D7A04283">1604</mods:number>
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<mods:start id="95AA21CF3CDE79EAD5D241F506155BAE">1</mods:start>
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<mods:classification id="8A0D42D44B82D4E8AC2044B5823FFD12">journal article</mods:classification>
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<mods:identifier id="066E1061C25B66BC653A65D4CA11A9B7" type="DOI">10.5281/zenodo.178756</mods:identifier>
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<mods:identifier id="031855CE91028CCDB729853D6F8D0F22" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03E987D1FFB2FFFCFF178E863435FCD9" ID-DOI="http://doi.org/10.5281/zenodo.6250234" ID-GBIF-Taxon="119373723" ID-Zenodo-Dep="6250234" LSID="urn:lsid:plazi:treatment:03E987D1FFB2FFFCFF178E863435FCD9" httpUri="http://treatment.plazi.org/id/03E987D1FFB2FFFCFF178E863435FCD9" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
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<subSubSection id="C35A654CFFB2FFF3FF178E86345CF903" pageId="7" pageNumber="8" type="nomenclature">
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<paragraph id="8BFF36C7FFB2FFF3FF178E8636ADF97E" blockId="7.[151,954,1683,1744]" box="[151,954,1683,1709]" pageId="7" pageNumber="8">
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<heading id="D0B781ABFFB2FFF3FF178E8636ADF97E" bold="true" box="[151,954,1683,1709]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
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<emphasis id="B934EAD5FFB2FFF3FF178E8636ADF97E" bold="true" box="[151,954,1683,1709]" pageId="7" pageNumber="8">
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<taxonomicName id="4C404D44FFB2FFF3FF178E86362CF97E" authority="Schauberger, 1932" authorityName="Schauberger" authorityYear="1932" box="[151,827,1683,1709]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="subSpecies" species="viridanus" subSpecies="staudingerianus">
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<emphasis id="B934EAD5FFB2FFF3FF178E86375BF97E" bold="true" box="[151,588,1683,1709]" italics="true" pageId="7" pageNumber="8">Harpalus viridanus staudingerianus</emphasis>
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<bibRefCitation id="EFD14B36FFB2FFF3FDD38E86362CF97E" author="Schauberger" box="[595,827,1683,1709]" pageId="7" pageNumber="20" refString="Schauberger, E. (1932) Zur Kenntnis der palaarktischen Harpalinen (X. Beitrag). Wiener Entomologische Zeitung, 49 (1), 23 - 44." type="journal article" year="1932">Schauberger, 1932</bibRefCitation>
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</taxonomicName>
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, stat. rest.
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BFF36C7FFB2FFF3FF178EA3345CF903" blockId="7.[151,954,1683,1744]" box="[151,331,1718,1744]" pageId="7" pageNumber="8">
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(
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<figureCitation id="137B2A42FFB2FFF3FF1F8EA335FCF903" box="[159,235,1718,1744]" captionStart="FIGURES 7 – 10" captionStartId="4.[151,269,1152,1176]" captionTargetBox="[212,1307,422,1124]" captionTargetId="figure@4.[184,1422,416,1147]" captionTargetPageId="4" captionText="FIGURES 7 – 10. Harapalus viridanus, pronotum. 7 – 8, H. v. angustibasis ssp. n. (7, Qinghai, Yushu Co., paratype; 8, Sichuan, Garzê Co., holotype); 9, H. v. staudingerianus (Minxian, paratype); 10, H. v. viridanus (Russia, Irkutsk env.). Scale = 1.0 mm." httpUri="https://zenodo.org/record/178758/files/figure.png" pageId="7" pageNumber="8">Figs 9</figureCitation>
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,
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<figureCitation id="137B2A42FFB2FFF3FF788EA33455F903" box="[248,322,1718,1744]" captionStart="FIGURES 11 – 16" captionStartId="7.[151,269,1499,1523]" captionTargetBox="[351,1280,718,1455]" captionTargetId="figure@7.[264,1323,696,1493]" captionTargetPageId="7" captionText="FIGURES 11 – 16. Harpalus viridanus, median lobe of aedeagus (13 – 14, terminal lamella). 11 – 13, H. v. angustibasis ssp. n. (Qinghai, Yushu Co., paratype); 14 – 16, H. v. staudingerianus (Minxian, paratype). 11, 13 – 16, dorsal aspect; 12, 16, lateral aspect. Scales: A = 0.5 mm (Figs 13 – 14), B = 1.0 mm (Figs 11 – 12, 15 – 16)." httpUri="https://zenodo.org/record/178759/files/figure.png" pageId="7" pageNumber="8">14–16</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C35A654CFFB2FFF3FF178F133048F8CE" box="[151,1375,1798,1821]" pageId="7" pageNumber="8" type="reference_group">
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<paragraph id="8BFF36C7FFB2FFF3FF178F133048F8CE" blockId="7.[151,1375,1798,1821]" box="[151,1375,1798,1821]" pageId="7" pageNumber="8">
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<treatmentCitationGroup id="AB5011E9FFB2FFF3FF178F133048F8CE" box="[151,1375,1798,1821]" pageId="7" pageNumber="8">
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<taxonomicName id="4C404D44FFB2FFF3FF178F133605F8CE" authority="Schauberger, 1932: 42" authorityName="Schauberger" authorityPageNumber="42" authorityYear="1932" box="[151,786,1798,1821]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="subSpecies" species="viridanus" subSpecies="staudingerianus">
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<emphasis id="B934EAD5FFB2FFF3FF178F13370EF8CE" box="[151,537,1798,1821]" italics="true" pageId="7" pageNumber="8">Harpalus viridanus staudingerianus</emphasis>
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<treatmentCitation id="0AE110D6FFB2FFF3FDA08F133605F8CE" author="Schauberger" box="[544,786,1798,1821]" page="42" pageId="7" pageNumber="8" year="1932">
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<bibRefCitation id="EFD14B36FFB2FFF3FDA08F1337FDF8CE" author="Schauberger" box="[544,746,1798,1821]" pageId="7" pageNumber="20" refString="Schauberger, E. (1932) Zur Kenntnis der palaarktischen Harpalinen (X. Beitrag). Wiener Entomologische Zeitung, 49 (1), 23 - 44." type="journal article" year="1932">Schauberger, 1932</bibRefCitation>
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: 42
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</treatmentCitation>
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</taxonomicName>
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.
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<typeStatus id="54FB8865FFB2FFF3FC9C8F133646F8CE" box="[796,849,1798,1821]" pageId="7" pageNumber="8">Type</typeStatus>
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locality: Min-Tschou (= Minxian), Gansu,
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<collectingCountry id="F3577657FFB2FFF3FA9F8F133048F8CE" box="[1311,1375,1798,1821]" name="China" pageId="7" pageNumber="8">China</collectingCountry>
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35A654CFFB2FFFCFF178F5831E0FF09" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="materials_examined">
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<paragraph id="8BFF36C7FFB2FFF3FF178F583056F843" blockId="7.[151,1436,1869,2016]" pageId="7" pageNumber="8">
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<emphasis id="B934EAD5FFB2FFF3FF178F583453F8B4" bold="true" box="[151,324,1869,1895]" pageId="7" pageNumber="8">
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<typeStatus id="54FB8865FFB2FFF3FF178F5835C5F8B4" box="[151,210,1869,1895]" pageId="7" pageNumber="8">Type</typeStatus>
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material
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</emphasis>
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.
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<typeStatus id="54FB8865FFB2FFF3FED48F5B34D3F8BB" box="[340,452,1870,1896]" pageId="7" pageNumber="8" type="syntype">Syntypes</typeStatus>
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: 14 ɗ, 17 Ψ labelled "Minchow,
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<collectingCountry id="F3577657FFB2FFF3FCC28F5B3699F8BB" box="[834,910,1870,1896]" name="China" pageId="7" pageNumber="8">China</collectingCountry>
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", "Cotype", "
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<taxonomicName id="4C404D44FFB2FFF3FBAA8F5B31F0F8BB" authority="Mots." authorityName="Mots." box="[1066,1255,1870,1896]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="viridanus">
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<emphasis id="B934EAD5FFB2FFF3FBAA8F5B318EF8B4" box="[1066,1177,1870,1895]" italics="true" pageId="7" pageNumber="8">viridanus</emphasis>
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Mots.
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</taxonomicName>
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, s.
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<emphasis id="B934EAD5FFB2FFF3FA8C8F5B35D8F85C" italics="true" pageId="7" pageNumber="8">staudingerianus</emphasis>
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Schaub.", "W. H. Muche, Radeberg, Ankauf", "Staatl. Museum für Tierkunde, Dresden" (
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<collectionCode id="ED51AE02FFB2FFF3FB6D8F633023F843" box="[1261,1332,1910,1936]" httpUri="http://grbio.org/cool/4j3t-w40g" name="Museum of Zoology Senckenberg Dresden" pageId="7" pageNumber="8">MTD</collectionCode>
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).
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</paragraph>
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<paragraph id="8BFF36C7FFB2FFFCFF468F8831E0FF09" blockId="7.[151,1436,1869,2016]" lastBlockId="8.[151,1437,151,778]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
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<emphasis id="B934EAD5FFB2FFF3FF468F88372AF864" bold="true" box="[198,573,1949,1975]" pageId="7" pageNumber="8">Additional material examined</emphasis>
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.
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<collectingCountry id="F3577657FFB2FFF3FDCC8F8B3781F86B" box="[588,662,1950,1976]" name="China" pageId="7" pageNumber="8">China</collectingCountry>
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.
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<collectionCode id="ED51AE02FFB2FFF3FD218F8B361CF86B" box="[673,779,1950,1976]" pageId="7" pageNumber="8">GANSU</collectionCode>
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: 1 ɗ, 2 Ψ, Minxian Co.,
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<date id="FFFE1007FFB2FFF3FBAA8F8B31ABF86B" box="[1066,1212,1950,1976]" pageId="7" pageNumber="8" value="1987-07-20">20.VII.1987</date>
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, Zhong Tiesen leg. (
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<collectionCode id="ED51AE02FFB2FFF3FF208FD335C3F833" box="[160,212,1990,2016]" pageId="7" pageNumber="8">IZB</collectionCode>
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); 1 ɗ, "Kan–ssu ["Pag. Sche-pu et fl. Po-i-cho,
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<quantity id="4CB89B22FFB2FFF3FC9D8FD3366AF833" box="[797,893,1990,2016]" metricMagnitude="3" metricUnit="m" metricValue="1.9369999999999998" pageId="7" pageNumber="8" unit="m" value="1937.0">1937 m</quantity>
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", NE of Nanping near border with Sichuan], 1885, G. Patanin/
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<date id="FFFE1007FFBDFFFCFEE9888D34AFFF61" box="[361,440,152,178]" pageId="8" pageNumber="9">13.VII</date>
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" (
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<collectionCode id="ED51AE02FFBDFFFCFE52888D3703FF61" box="[466,532,152,178]" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="8" pageNumber="9">ZISP</collectionCode>
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); 1 ɗ, "Kan-ssu [= Jonê, ca
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<quantity id="4CB89B22FFBDFFFCFCDC888D36B0FF61" box="[860,935,152,178]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="9" unit="km" value="50.0">50 km</quantity>
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NW of Minxian], 1885, G. Patanin/
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<date id="FFFE1007FFBDFFFCFAD4888D3083FF61" box="[1364,1428,152,178]" pageId="8" pageNumber="9">10.VI</date>
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" (
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<collectionCode id="ED51AE02FFBDFFFCFF1F88D535F6FF09" box="[159,225,192,218]" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="8" pageNumber="9">ZISP</collectionCode>
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); 1 ɗ, "Kan-ssu, 1885, G. Patain/
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<date id="FFFE1007FFBDFFFCFDEB88D537BDFF09" box="[619,682,192,218]" pageId="8" pageNumber="9">18.VI</date>
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" [= Dangchang, ca
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<quantity id="4CB89B22FFBDFFFCFC1188D536CCFF09" box="[913,987,192,218]" metricMagnitude="4" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="9" unit="km" value="80.0">80 km</quantity>
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SE of Minxian] (
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<collectionCode id="ED51AE02FFBDFFFCFB2888D531FDFF09" box="[1192,1258,192,218]" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="8" pageNumber="9">ZISP</collectionCode>
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||
).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C35A654CFFBDFFFCFF4688F23034FD91" pageId="8" pageNumber="9" type="description">
|
||
<paragraph id="8BFF36C7FFBDFFFCFF4688F23034FD91" blockId="8.[151,1437,151,778]" pageId="8" pageNumber="9">
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<emphasis id="B934EAD5FFBDFFFCFF4688F23440FED2" bold="true" box="[198,343,231,257]" pageId="8" pageNumber="9">Description</emphasis>
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. Very large: body length 8.7–11.0 (usually above 10.0) mm, with or without metallic lustre on dorsum. Head relatively small (WHmax/WPmax =
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<quantity id="4CB89B22FFBDFFFCFCA3890536AFFEF9" box="[803,952,272,298]" metricMagnitude="-2" metricUnit="m" metricValue="1.5621" metricValueMax="1.6002" metricValueMin="1.524" pageId="8" pageNumber="9" unit="in" value="0.615" valueMax="0.63" valueMin="0.6">0.60–0.63 in</quantity>
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||
male and
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<quantity id="4CB89B22FFBDFFFCFBB6890531DCFEF9" box="[1078,1227,272,298]" metricMagnitude="-2" metricUnit="m" metricValue="1.5875" metricValueMax="1.651" metricValueMin="1.524" pageId="8" pageNumber="9" unit="in" value="0.625" valueMax="0.65" valueMin="0.6">0.60–0.65 in</quantity>
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||
female; WHmin/ WPmax =
|
||
<quantity id="4CB89B22FFBDFFFCFE9D892D34A2FE81" box="[285,437,312,338]" metricMagnitude="-2" metricUnit="m" metricValue="1.3716" metricValueMax="1.397" metricValueMin="1.3462" pageId="8" pageNumber="9" unit="in" value="0.54" valueMax="0.55" valueMin="0.53">0.53–0.55 in</quantity>
|
||
male and
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||
<quantity id="4CB89B22FFBDFFFCFDBA892D37C5FE81" box="[570,722,312,338]" metricMagnitude="-2" metricUnit="m" metricValue="1.397" metricValueMax="1.4478" metricValueMin="1.3462" pageId="8" pageNumber="9" unit="in" value="0.55" valueMax="0.57" valueMin="0.53">0.53–0.57 in</quantity>
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female). Pronotum (
|
||
<figureCitation id="137B2A42FFBDFFFCFC52892D310BFE81" box="[978,1052,312,338]" captionStart="FIGURES 7 – 10" captionStartId="4.[151,269,1152,1176]" captionTargetBox="[212,1307,422,1124]" captionTargetId="figure@4.[184,1422,416,1147]" captionTargetPageId="4" captionText="FIGURES 7 – 10. Harapalus viridanus, pronotum. 7 – 8, H. v. angustibasis ssp. n. (7, Qinghai, Yushu Co., paratype; 8, Sichuan, Garzê Co., holotype); 9, H. v. staudingerianus (Minxian, paratype); 10, H. v. viridanus (Russia, Irkutsk env.). Scale = 1.0 mm." httpUri="https://zenodo.org/record/178758/files/figure.png" pageId="8" pageNumber="9">Fig. 9</figureCitation>
|
||
) with comparatively wide base (WPmax/WPmin = 1.03–1.08) and with sides more or less parallel basally, sometimes slightly converging posteriad. Pronotal basal angle right-angled or slightly obtuse, usually with somewhat sharp or slightly blunt apex. Elytral striae deepened and intervals convex, particularly markedly near apex. Dorsal microsculpture visible throughout on head, pronotum and elytra, consisting of distinct, more or less isodiametric meshes. Median lobe of aedeagus (
|
||
<figureCitation id="137B2A42FFBDFFFCFE5E8A153771FDC9" box="[478,614,512,538]" captionStart="FIGURES 11 – 16" captionStartId="7.[151,269,1499,1523]" captionTargetBox="[351,1280,718,1455]" captionTargetId="figure@7.[264,1323,696,1493]" captionTargetPageId="7" captionText="FIGURES 11 – 16. Harpalus viridanus, median lobe of aedeagus (13 – 14, terminal lamella). 11 – 13, H. v. angustibasis ssp. n. (Qinghai, Yushu Co., paratype); 14 – 16, H. v. staudingerianus (Minxian, paratype). 11, 13 – 16, dorsal aspect; 12, 16, lateral aspect. Scales: A = 0.5 mm (Figs 13 – 14), B = 1.0 mm (Figs 11 – 12, 15 – 16)." httpUri="https://zenodo.org/record/178759/files/figure.png" pageId="8" pageNumber="9">Figs 11–12</figureCitation>
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) medium–sized (LB/LML = 3.30–3.54); terminal lamella (
|
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<figureCitation id="137B2A42FFBDFFFCFABA8A153084FDC9" box="[1338,1427,512,538]" captionStart="FIGURES 11 – 16" captionStartId="7.[151,269,1499,1523]" captionTargetBox="[351,1280,718,1455]" captionTargetId="figure@7.[264,1323,696,1493]" captionTargetPageId="7" captionText="FIGURES 11 – 16. Harpalus viridanus, median lobe of aedeagus (13 – 14, terminal lamella). 11 – 13, H. v. angustibasis ssp. n. (Qinghai, Yushu Co., paratype); 14 – 16, H. v. staudingerianus (Minxian, paratype). 11, 13 – 16, dorsal aspect; 12, 16, lateral aspect. Scales: A = 0.5 mm (Figs 13 – 14), B = 1.0 mm (Figs 11 – 12, 15 – 16)." httpUri="https://zenodo.org/record/178759/files/figure.png" pageId="8" pageNumber="9">Fig. 14</figureCitation>
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) comparatively long, with sides more or less notably sinuate, sometimes nearly parallel before apex.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35A654CFFBDFFFCFF468A5A3185FDB9" box="[198,1170,591,618]" pageId="8" pageNumber="9" type="distribution">
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<paragraph id="8BFF36C7FFBDFFFCFF468A5A3185FDB9" blockId="8.[151,1437,151,778]" box="[198,1170,591,618]" pageId="8" pageNumber="9">
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<emphasis id="B934EAD5FFBDFFFCFF468A5A3448FDBA" bold="true" box="[198,351,591,617]" pageId="8" pageNumber="9">Distribution</emphasis>
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. This subspecies seems to have a small range in southernmost Gansu.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35A654CFFBDFFFCFF468A623435FCD9" pageId="8" pageNumber="9" type="discussion">
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<paragraph id="8BFF36C7FFBDFFFCFF468A623435FCD9" blockId="8.[151,1437,151,778]" pageId="8" pageNumber="9">
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<emphasis id="B934EAD5FFBDFFFCFF468A62342FFD42" bold="true" box="[198,312,631,657]" pageId="8" pageNumber="9">Remarks</emphasis>
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. Described from more than
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<specimenCount id="9D46FD4EFFBDFFFCFD0C8A6D363BFD41" box="[652,812,632,658]" pageId="8" pageNumber="9" type="generic">50 specimens</specimenCount>
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captured in Minxian, Gansu,
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<collectingCountry id="F3577657FFBDFFFCFB128A6D31CBFD41" box="[1170,1244,632,658]" name="China" pageId="8" pageNumber="9">China</collectingCountry>
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. In habitus, it is similar to the nominotypical subspecies but distinguished by larger body. This form seems to be distinct although probably is connected with the nominotypical subspecies by the narrow transition zone to the north of Minxian.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |