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<document id="6529D79F9EDF4C33343ED4688A578C82" ID-DOI="10.1111/j.1096-3642.2012.00863.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5409181" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630350074602" checkinUser="marcus" docAuthor="Criscione, Francesco, Law, Margot Louisa &amp; Köhler, Frank" docDate="2012" docId="03E9878DFFCEFF867000A65D5B48F947" docLanguage="en" docName="zoj863.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 166 (4)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2012.00863.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Exiligada punctata Criscione &amp; Law &amp; Köhler 2012, SP. NOV." docType="treatment" docVersion="4" lastPageNumber="710" masterDocId="FFD0FFF5FFDAFF93710AA04F5F3FFFCD" masterDocTitle="Land snail diversity in the monsoon tropics of Northern Australia: revision of the genus Exiligada Iredale, 1939 (Mollusca: Pulmonata: Camaenidae), with description of 13 new species" masterLastPageNumber="722" masterPageNumber="689" pageNumber="709" updateTime="1699241497941" updateUser="plazi">
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<mods:title id="AE1447F9C654EC1B388BC19D9DED3490">Land snail diversity in the monsoon tropics of Northern Australia: revision of the genus Exiligada Iredale, 1939 (Mollusca: Pulmonata: Camaenidae), with description of 13 new species</mods:title>
</mods:titleInfo>
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<mods:namePart id="B51FC75AB0CF3B1BD432075C35D30079">Criscione, Francesco</mods:namePart>
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<mods:namePart id="BCFF0677EA975615EE75428EDA2441E3">Law, Margot Louisa</mods:namePart>
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<mods:namePart id="32CD9F0F8975AE82833316923A7B1B7A">Köhler, Frank</mods:namePart>
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<mods:title id="B0C9ABC4517F01667939F8E11FE75ECE">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="3A466C697C49E482D4F9FF7D32DF5418">2012</mods:date>
<mods:detail id="2F63CF3F026E7033E78B9FB3D65C9464" type="series">
<mods:title id="776019F86AE66DB8696186896DE25F57">Zool. J. Linn. Soc.</mods:title>
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<mods:number id="75507BC98DC14C862F04F64E963156AA">2012-11-26</mods:number>
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<mods:number id="8F2FF0A368CB07C99EB81EA384411282">166</mods:number>
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<mods:identifier id="04EC0E1ABB3F4CB5C63E1589FD52A79A" type="Zenodo-Dep">5409181</mods:identifier>
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<treatment id="03E9878DFFCEFF867000A65D5B48F947" ID-DOI="http://doi.org/10.5281/zenodo.5480110" ID-Zenodo-Dep="5480110" LSID="urn:lsid:plazi:treatment:03E9878DFFCEFF867000A65D5B48F947" httpUri="http://treatment.plazi.org/id/03E9878DFFCEFF867000A65D5B48F947" lastPageId="21" lastPageNumber="710" pageId="20" pageNumber="709">
<subSubSection id="C35A6510FFCEFF877000A65D5D43F9EA" box="[266,636,1554,1575]" pageId="20" pageNumber="709" type="nomenclature">
<paragraph id="8BFF369BFFCEFF877000A65D5D43F9EA" blockId="20.[266,636,1554,1575]" box="[266,636,1554,1575]" pageId="20" pageNumber="709">
<heading id="D0B781F7FFCEFF877000A65D5D43F9EA" box="[266,636,1554,1575]" centered="true" fontSize="9" level="2" pageId="20" pageNumber="709" reason="4">
<taxonomicName id="4C404D18FFCEFF877000A65D5D32F9EB" ID-CoL="3DL86" authority="Criscione &amp; Law &amp; Köhler, 2012" authorityName="Criscione &amp; Law &amp; Köhler" authorityYear="2012" box="[266,525,1554,1575]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="709" phylum="Mollusca" rank="species" species="punctata" status="sp. nov.">EXILIGADA PUNCTATA</taxonomicName>
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<taxonomicNameLabel id="A20757F2FFCEFF87731DA65A5D43F9EA" box="[535,636,1554,1575]" pageId="20" pageNumber="709" rank="species">SP. NOV.</taxonomicNameLabel>
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<subSubSection id="C35A6510FFCEFF867027A6775B48F947" lastPageId="21" lastPageNumber="710" pageId="20" pageNumber="709" type="description">
<paragraph id="8BFF369BFFCEFF877027A6775D66F983" blockId="20.[301,601,1592,1614]" box="[301,601,1592,1614]" pageId="20" pageNumber="709">
(
<figureCitation id="137B2A1EFFCEFF87703FA6775EAAF980" box="[309,405,1592,1613]" captionStart="Figure 4" captionStartId="8.[144,223,954,973]" captionTargetBox="[144,1425,196,924]" captionTargetId="figure-472@8.[144,1425,196,924]" captionTargetPageId="8" captionText="Figure 4. Photographs of living specimens of Exiligada species (courtesy Vince Kessner, Adelaide River). A, E. negriensis Iredale, 1939. B, E. pallida sp. nov. C, E. nodulicaudata sp. nov. D, E. punctata sp. nov. E, E. qualis Iredale, 1939. F, E. monochroma sp. nov. G, E. unistriata sp. nov. H, E. floraevallis sp. nov. I, E. rivifontis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5409193" httpUri="https://zenodo.org/record/5409193/files/figure.png" pageId="20" pageNumber="709">FIGS 4D</figureCitation>
,
<figureCitation id="137B2A1EFFCEFF8770AFA6775EF6F983" box="[421,457,1592,1614]" captionStart="Figure 6" captionStartId="9.[826,905,1233,1252]" captionTargetBox="[834,1435,197,1200]" captionTargetId="figure-294@9.[830,1439,194,1203]" captionTargetPageId="9" captionText="Figure 6. Shells of Exiligada species. A, E. montejinni sp. nov. (paratype AM C.469982). B, E. nodulicauda sp. nov. (paratype WAM S49160). C, E. pallida sp. nov. (paratype WAM S49215). D, E. punctata sp. nov. (paratype WAM S49221). E, E. qualis Iredale, 1939 (holotype AM C.64866). F, E. rivifontis sp. nov. (WAM S49225). G, E. unistriata sp. nov. (paratype AM C.470460). Scale bar = 2 mm. Note that foot produces from some shells." figureDoi="http://doi.org/10.5281/zenodo.5409195" httpUri="https://zenodo.org/record/5409195/files/figure.png" pageId="20" pageNumber="709">6D</figureCitation>
,
<figureCitation id="137B2A1EFFCEFF8770DDA6775D19F980" box="[471,550,1592,1613]" captionStart="Figure 20" captionStartId="22.[144,223,1220,1239]" captionTargetBox="[303,1262,196,1190]" captionTargetId="figure-272@22.[303,1263,196,1190]" captionTargetPageId="22" captionText="Figure 20. SEM micrographs of Exiligada shells.A-C, E. pallida sp. nov. (paratype WAM S49150). A, Apical view showing protoconch and first whorl. B, Close-up from above of sculpture across entire shell. C, Microsculpture on teleoconch whorl. D-F, E. punctata sp. nov. (paratype WAMS49139). D, Apical view showing protoconch and first whorl. E, Close-up from above of sculpture across entire shell. F, Microsculpture on teleoconch whorl. Scale bars: A-B, D-E = 1 mm, C, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5409227" httpUri="https://zenodo.org/record/5409227/files/figure.png" pageId="20" pageNumber="709">20D–F</figureCitation>
,
<figureCitation id="137B2A1EFFCEFF87733EA6775D6EF983" box="[564,593,1592,1614]" captionStart="Figure 22" captionStartId="23.[164,243,1578,1597]" captionTargetBox="[232,712,880,1547]" captionTargetId="figure-495@23.[232,712,878,1549]" captionTargetPageId="23" captionText="Figure 22. Penial anatomy of E. punctata sp. nov. (holotype WAM S83175). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5409231" httpUri="https://zenodo.org/record/5409231/files/figure.png" pageId="20" pageNumber="709">22</figureCitation>
)
</paragraph>
<paragraph id="8BFF369BFFCEFF87719AA6105DFBF922" blockId="20.[144,759,1631,1775]" pageId="20" pageNumber="709">
<materialsCitation id="3B283CC6FFCEFF87719AA6105DFBF922" accessionNumber="S 83175" collectingDate="2009-09-01" collectionCode="WA, EK, WAM" collectorName="V. Kessner &amp; Kessner, R" country="Australia" latitude="-17.823057" location="Halls Creek District" longLatPrecision="21" longitude="128.24945" municipality="Panton River" pageId="20" pageNumber="709" specimenCode="WA-76" specimenCount="1" typeStatus="holotype">
<emphasis id="B934EA89FFCEFF87719AA6105FC5F9B9" box="[144,250,1631,1652]" italics="true" pageId="20" pageNumber="709">
<typeStatus id="54FB8839FFCEFF87719AA6105FCBF9B9" box="[144,244,1631,1652]" pageId="20" pageNumber="709" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectingCountry id="F357760BFFCEFF87700AA6105EA6F9B9" box="[256,409,1631,1652]" name="Australia" pageId="20" pageNumber="709">AUSTRALIA</collectingCountry>
,
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,
<collectionCode id="ED51AE5EFFCEFF8770D2A6105D3FF9B9" box="[472,512,1631,1652]" pageId="20" pageNumber="709">EK</collectionCode>
,
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, south of
<collectingMunicipality id="6B9BACE1FFCEFF8771FDA6315EADF95E" box="[247,402,1661,1683]" pageId="20" pageNumber="709">Panton River</collectingMunicipality>
, low and open limestone area, numerous gullies, under well-developed spinifex, in talus,
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17°49
<emphasis id="B934EA89FFCEFF877014A6F65E1BF91D" box="[286,292,1721,1744]" italics="true" pageId="20" pageNumber="709">′</emphasis>
23
<emphasis id="B934EA89FFCEFF877035A6F65E74F91D" box="[319,331,1721,1744]" italics="true" pageId="20" pageNumber="709">′′</emphasis>
S
</geoCoordinate>
,
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128°14
<emphasis id="B934EA89FFCEFF8770B7A6F65EFCF91D" box="[445,451,1721,1744]" italics="true" pageId="20" pageNumber="709">′</emphasis>
58
<emphasis id="B934EA89FFCEFF8770D4A6F65ED5F91D" box="[478,490,1721,1744]" italics="true" pageId="20" pageNumber="709">′′</emphasis>
E
</geoCoordinate>
(coll.
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, R. Hokkanen,
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<collectingDate id="EFBAE9B3FFCEFF87701FA6965E41F922" box="[277,382,1753,1775]" pageId="20" pageNumber="709" value="2009-09-01">1.ix.2009</collectingDate>
</date>
;
<specimenCode id="DBE69EE0FFCEFF877081A6965EE7F922" box="[395,472,1753,1775]" collectionCode="WA-" pageId="20" pageNumber="709">WA-76</specimenCode>
/09) (
<collectionCode id="ED51AE5EFFCEFF877305A6965D6FF923" box="[527,592,1753,1774]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="20" pageNumber="709" type="Museum">WAM</collectionCode>
<accessionNumber id="9413AB78FFCEFF877352A6965D87F922" box="[600,696,1753,1775]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S83175" pageId="20" pageNumber="709" type="EnaNcbi">S 83175</accessionNumber>
).
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</paragraph>
<paragraph id="8BFF369BFFCEFF87719AA7535BA2F95D" blockId="20.[144,759,1820,1903]" lastBlockId="20.[806,1422,1413,1680]" pageId="20" pageNumber="709">
<materialsCitation id="3B283CC6FFCEFF87719AA7535D22F8FC" box="[144,541,1820,1841]" collectionCode="WA, EK" country="Australia" location="AUSTRALIA" pageId="20" pageNumber="709" specimenCount="1" typeStatus="paratype">
<emphasis id="B934EA89FFCEFF87719AA7535E38F8FC" box="[144,263,1820,1841]" italics="true" pageId="20" pageNumber="709">
<typeStatus id="54FB8839FFCEFF87719AA7535E3EF8FC" box="[144,257,1820,1841]" pageId="20" pageNumber="709" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectingCountry id="F357760BFFCEFF877019A7535E93F8FC" box="[275,428,1820,1841]" name="Australia" pageId="20" pageNumber="709">AUSTRALIA</collectingCountry>
,
<collectionCode id="ED51AE5EFFCEFF8770B1A7535ED9F8FC" box="[443,486,1820,1841]" pageId="20" pageNumber="709">WA</collectionCode>
,
<collectionCode id="ED51AE5EFFCEFF8770FFA7535D22F8FC" box="[501,541,1820,1841]" pageId="20" pageNumber="709">EK</collectionCode>
</materialsCitation>
,
<materialsCitation id="3B283CC6FFCEFF877326A7535BA6F95D" accessionNumber="S49221, S49220, S49222, S49139" collectingDate="2009-09-01" collectionCode="WAM" collectorName="Nicholson Station H &amp; V. Kessner &amp; Kessner, R" country="Australia" county="Panton River" latitude="-17.779722" location="Halls Creek District" longLatPrecision="21" longitude="128.23305" municipality="Halls Creek District" pageId="20" pageNumber="709" specimenCode="WA-75, WA-77" specimenCount="1" typeStatus="paratype">
same data as for holotype, ten wet (
<collectionCode id="ED51AE5EFFCEFF87707BA7755E8DF882" box="[369,434,1850,1871]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="20" pageNumber="709" type="Museum">WAM</collectionCode>
<accessionNumber id="9413AB78FFCEFF8770B4A7755D2AF89D" box="[446,533,1850,1872]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S49221" pageId="20" pageNumber="709" type="EnaNcbi">S49221</accessionNumber>
);
<location id="8E9F6040FFCEFF877324A7755FFCF8A2" LSID="urn:lsid:plazi:treatment:03E9878DFFCEFF867000A65D5B48F947:8E9F6040FFCEFF877324A7755FFCF8A2" country="Australia" county="Panton River" latitude="-17.779722" longLatPrecision="21" longitude="128.23305" municipality="Halls Creek District" name="Halls Creek District" pageId="20" pageNumber="709">Halls Creek District</location>
,
<quantity id="4CB89B7EFFCEFF8771DAA7165E29F8A2" box="[208,278,1881,1903]" metricMagnitude="4" metricUnit="m" metricValue="5.7" pageId="20" pageNumber="709" unit="km" value="57.0">57 km</quantity>
east of
<collectorName id="26B5534DFFCEFF87707FA7165D5EF8A3" box="[373,609,1881,1903]" pageId="20" pageNumber="709">
<location id="8E9F6040FFCEFF87707FA7165D7DF8A2" LSID="urn:lsid:plazi:treatment:03E9878DFFCEFF867000A65D5B48F947:8E9F6040FFCEFF87707FA7165D7DF8A2" box="[373,578,1881,1903]" country="Australia" county="Panton River" latitude="-17.779722" longLatPrecision="21" longitude="128.23305" municipality="Halls Creek District" name="Nicholson Station" pageId="20" pageNumber="709">Nicholson Station</location>
H
</collectorName>
/S, very open limestone area, well-developed spinifex, talus, under spinifex and slabs,
<geoCoordinate id="EE74505CFFCEFF877504A5EC5BB2FA74" box="[1038,1165,1442,1465]" degrees="17" direction="south" minutes="58" orientation="latitude" pageId="20" pageNumber="709" precision="15" seconds="43" value="-17.978611">
17°58
<emphasis id="B934EA89FFCEFF877544A5ED5B6BFA74" box="[1102,1108,1442,1465]" italics="true" pageId="20" pageNumber="709">′</emphasis>
43
<emphasis id="B934EA89FFCEFF87757AA5ED5B43FA74" box="[1136,1148,1442,1465]" italics="true" pageId="20" pageNumber="709">′′</emphasis>
S
</geoCoordinate>
,
<geoCoordinate id="EE74505CFFCEFF877597A5EB5A14FA74" box="[1181,1323,1442,1465]" degrees="128" direction="east" minutes="21" orientation="longitude" pageId="20" pageNumber="709" precision="15" seconds="34" value="128.35945">
128°21
<emphasis id="B934EA89FFCEFF8775E6A5ED5BCDFA74" box="[1260,1266,1442,1465]" italics="true" pageId="20" pageNumber="709">′</emphasis>
34
<emphasis id="B934EA89FFCEFF877407A5ED5A26FA74" box="[1293,1305,1442,1465]" italics="true" pageId="20" pageNumber="709">′′</emphasis>
E
</geoCoordinate>
(coll.
<collectorName id="26B5534DFFCEFF877472A5EB5CBCFA15" pageId="20" pageNumber="709">V. Kessner</collectorName>
, R. Hokkanen,
<date id="FFFE105BFFCEFF877536A58D5B9AFA15" box="[1084,1189,1474,1496]" pageId="20" pageNumber="709" value="2009-09-01">
<collectingDate id="EFBAE9B3FFCEFF877536A58D5B9AFA15" box="[1084,1189,1474,1496]" pageId="20" pageNumber="709" value="2009-09-01">1.ix.2009</collectingDate>
</date>
;
<specimenCode id="DBE69EE0FFCEFF8775BEA58D5A3CFA15" box="[1204,1283,1474,1496]" collectionCode="WA-" pageId="20" pageNumber="709">WA-75</specimenCode>
/09), 15 wet,
<emphasis id="B934EA89FFCEFF87722CA5AE5C0BFA3B" box="[806,820,1505,1526]" italics="true" pageId="20" pageNumber="709">&gt;</emphasis>
24 dry (
<collectionCode id="ED51AE5EFFCEFF877293A5AE5CE5FA3B" box="[921,986,1505,1526]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="20" pageNumber="709" type="Museum">WAM</collectionCode>
<accessionNumber id="9413AB78FFCEFF8772EFA5AE5B03FA3A" box="[997,1084,1505,1527]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S49220" pageId="20" pageNumber="709" type="EnaNcbi">S49220</accessionNumber>
);
<collectingMunicipality id="6B9BACE1FFCEFF877559A5AE5A7BFA3A" box="[1107,1348,1505,1527]" pageId="20" pageNumber="709">Halls Creek District</collectingMunicipality>
, large rather open limestone area,
<quantity id="4CB89B7EFFCEFF877573A5B05BF9F9D8" box="[1145,1222,1535,1557]" metricMagnitude="3" metricUnit="m" metricValue="4.5" pageId="20" pageNumber="709" unit="km" value="4.5">4.5 km</quantity>
north of
<collectingCounty id="629E4E17FFCEFF877431A64F5C5AF9F9" pageId="20" pageNumber="709">Panton River</collectingCounty>
, numerous gullies, under spinifex, in talus,
<geoCoordinate id="EE74505CFFCEFF87722CA6735C9AF99F" box="[806,933,1595,1618]" degrees="17" direction="south" minutes="46" orientation="latitude" pageId="20" pageNumber="709" precision="15" seconds="47" value="-17.779722">
17°46
<emphasis id="B934EA89FFCEFF87726DA6745C52F99F" box="[871,877,1595,1618]" italics="true" pageId="20" pageNumber="709">′</emphasis>
47
<emphasis id="B934EA89FFCEFF877282A6745CABF99F" box="[904,916,1595,1618]" italics="true" pageId="20" pageNumber="709">′′</emphasis>
S
</geoCoordinate>
,
<geoCoordinate id="EE74505CFFCEFF8772B3A6725B78F99F" box="[953,1095,1595,1618]" degrees="128" direction="east" minutes="13" orientation="longitude" pageId="20" pageNumber="709" precision="15" seconds="59" value="128.23305">
128°13
<emphasis id="B934EA89FFCEFF877502A6745B31F99F" box="[1032,1038,1595,1618]" italics="true" pageId="20" pageNumber="709">′</emphasis>
59
<emphasis id="B934EA89FFCEFF877523A6745B0AF99F" box="[1065,1077,1595,1618]" italics="true" pageId="20" pageNumber="709">′′</emphasis>
E
</geoCoordinate>
(coll.
<collectorName id="26B5534DFFCEFF877590A6725A24F99F" box="[1178,1307,1597,1618]" pageId="20" pageNumber="709">V. Kessner</collectorName>
, R. Hokkanen,
<date id="FFFE105BFFCEFF877287A6145CC9F9BC" box="[909,1014,1627,1649]" pageId="20" pageNumber="709" value="2009-09-01">
<collectingDate id="EFBAE9B3FFCEFF877287A6145CC9F9BC" box="[909,1014,1627,1649]" pageId="20" pageNumber="709" value="2009-09-01">1.ix.2009</collectingDate>
</date>
;
<specimenCode id="DBE69EE0FFCEFF87751EA6145B5BF9BC" box="[1044,1124,1627,1649]" collectionCode="WA-" pageId="20" pageNumber="709">WA-77</specimenCode>
/09), seven wet (
<collectionCode id="ED51AE5EFFCEFF877446A6145AB2F9BD" box="[1356,1421,1627,1648]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="20" pageNumber="709" type="Museum">WAM</collectionCode>
<accessionNumber id="9413AB78FFCEFF87722CA6355C41F95D" box="[806,894,1658,1680]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S49222" pageId="20" pageNumber="709" type="EnaNcbi">S49222</accessionNumber>
), 22 dry (
<collectionCode id="ED51AE5EFFCEFF8772E5A6355B0FF942" box="[1007,1072,1658,1679]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="20" pageNumber="709" type="Museum">WAM</collectionCode>
<accessionNumber id="9413AB78FFCEFF877533A6355BAEF95D" box="[1081,1169,1658,1680]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S49139" pageId="20" pageNumber="709" type="EnaNcbi">S49139</accessionNumber>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BFF369BFFCEFF86722CA68F5B68FE1C" blockId="20.[806,1421,1728,1903]" lastBlockId="21.[826,1441,198,465]" lastPageId="21" lastPageNumber="710" pageId="20" pageNumber="709">
<emphasis id="B934EA89FFCEFF87722CA68F5CE5F918" box="[806,986,1728,1749]" italics="true" pageId="20" pageNumber="709">Other material:</emphasis>
<materialsCitation id="3B283CC6FFCEFF8772ECA68F5BCEF918" box="[998,1265,1728,1749]" collectionCode="WA, EK" country="Australia" location="AUSTRALIA" pageId="20" pageNumber="709" specimenCount="1">
<collectingCountry id="F357760BFFCEFF8772ECA68F5B40F918" box="[998,1151,1728,1749]" name="Australia" pageId="20" pageNumber="709">AUSTRALIA</collectingCountry>
,
<collectionCode id="ED51AE5EFFCEFF877584A68F5B86F918" box="[1166,1209,1728,1749]" pageId="20" pageNumber="709">WA</collectionCode>
,
<collectionCode id="ED51AE5EFFCEFF8775C3A68F5BCEF918" box="[1225,1265,1728,1749]" pageId="20" pageNumber="709">EK</collectionCode>
</materialsCitation>
,
<materialsCitation id="3B283CC6FFCEFF86740AA68F5B6CFE1C" accessionNumber="S49137, S49138, S49144" collectingDate="2009-09-01" collectionCode="WAM" collectorName="Nicholson Station H &amp; V. Kessner &amp; Kessner, R" country="Australia" county="Panton River" lastPageId="21" lastPageNumber="710" latitude="-17.823057" location="Halls Creek District" longLatPrecision="21" longitude="128.24945" municipality="Halls Creek District" pageId="20" pageNumber="709" specimenCode="WA-75, WA-76" specimenCount="1">
<location id="8E9F6040FFCEFF87740AA68F5CBFF939" LSID="urn:lsid:plazi:treatment:03E9878DFFCEFF867000A65D5B48F947:8E9F6040FFCEFF87740AA68F5CBFF939" country="Australia" county="Panton River" latitude="-17.823057" longLatPrecision="21" longitude="128.24945" municipality="Halls Creek District" name="Halls Creek District" pageId="20" pageNumber="709">Halls Creek District</location>
,
<quantity id="4CB89B7EFFCEFF87729BA6915CE8F939" box="[913,983,1758,1780]" metricMagnitude="4" metricUnit="m" metricValue="5.7" pageId="20" pageNumber="709" unit="km" value="57.0">57 km</quantity>
east of
<collectorName id="26B5534DFFCEFF87754AA6915A0CF939" box="[1088,1331,1758,1780]" pageId="20" pageNumber="709">
<location id="8E9F6040FFCEFF87754AA6915A2FF939" LSID="urn:lsid:plazi:treatment:03E9878DFFCEFF867000A65D5B48F947:8E9F6040FFCEFF87754AA6915A2FF939" box="[1088,1296,1758,1780]" country="Australia" county="Panton River" latitude="-17.823057" longLatPrecision="21" longitude="128.24945" municipality="Halls Creek District" name="Nicholson Station" pageId="20" pageNumber="709">Nicholson Station</location>
H
</collectorName>
/S, very open limestone area, well-developed spinifex, talus, under spinifex and slabs,
<geoCoordinate id="EE74505CFFCEFF877566A7545BD4F8FC" box="[1132,1259,1818,1841]" degrees="17" direction="south" minutes="58" orientation="latitude" pageId="20" pageNumber="709" precision="15" seconds="43" value="-17.978611">
17°58
<emphasis id="B934EA89FFCEFF8775A6A7555B8DF8FC" box="[1196,1202,1818,1841]" italics="true" pageId="20" pageNumber="709">′</emphasis>
43
<emphasis id="B934EA89FFCEFF8775C4A7555BE5F8FC" box="[1230,1242,1818,1841]" italics="true" pageId="20" pageNumber="709">′′</emphasis>
S
</geoCoordinate>
,
<geoCoordinate id="EE74505CFFCEFF8775F5A7535AB2F8FC" box="[1279,1421,1818,1841]" degrees="128" direction="east" minutes="21" orientation="longitude" pageId="20" pageNumber="709" precision="15" seconds="34" value="128.35945">
128°21
<emphasis id="B934EA89FFCEFF877444A7555A6BF8FC" box="[1358,1364,1818,1841]" italics="true" pageId="20" pageNumber="709">′</emphasis>
34
<emphasis id="B934EA89FFCEFF877465A7555A44F8FC" box="[1391,1403,1818,1841]" italics="true" pageId="20" pageNumber="709">′′</emphasis>
E
</geoCoordinate>
(coll.
<collectorName id="26B5534DFFCEFF87726CA7745CDDF89D" box="[870,994,1851,1872]" pageId="20" pageNumber="709">V. Kessner</collectorName>
, R. Hokkanen,
<date id="FFFE105BFFCEFF877593A7755A3DF89D" box="[1177,1282,1850,1872]" pageId="20" pageNumber="709" value="2009-09-01">
<collectingDate id="EFBAE9B3FFCEFF877593A7755A3DF89D" box="[1177,1282,1850,1872]" pageId="20" pageNumber="709" value="2009-09-01">1.ix.2009</collectingDate>
</date>
;
<specimenCode id="DBE69EE0FFCEFF87741BA7755A60F89D" box="[1297,1375,1850,1872]" collectionCode="WA-" pageId="20" pageNumber="709">WA-75</specimenCode>
/09), 30 dry (
<collectionCode id="ED51AE5EFFCEFF87728AA7165CFEF8A3" box="[896,961,1881,1902]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="20" pageNumber="709" type="Museum">WAM</collectionCode>
<accessionNumber id="9413AB78FFCEFF8772C0A7165B1EF8A2" box="[970,1057,1881,1903]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S49137" pageId="20" pageNumber="709" type="EnaNcbi">S49137</accessionNumber>
);
<location id="8E9F6040FFCEFF87753CA7165A1CF8A2" LSID="urn:lsid:plazi:treatment:03E9878DFFCEFF867000A65D5B48F947:8E9F6040FFCEFF87753CA7165A1CF8A2" box="[1078,1315,1881,1903]" country="Australia" county="Panton River" latitude="-17.823057" longLatPrecision="21" longitude="128.24945" municipality="Halls Creek District" name="Halls Creek District" pageId="20" pageNumber="709">Halls Creek District</location>
, south of
<location id="8E9F6040FFCFFF867230A0895CEDFF11" LSID="urn:lsid:plazi:treatment:03E9878DFFCEFF867000A65D5B48F947:8E9F6040FFCFFF867230A0895CEDFF11" box="[826,978,198,220]" country="Australia" county="Panton River" latitude="-17.823057" longLatPrecision="21" longitude="128.24945" municipality="Halls Creek District" name="Panton River" pageId="21" pageNumber="710">Panton River</location>
, low and open limestone area, numerous gullies, under well-developed spinifex, in talus,
<geoCoordinate id="EE74505CFFCFFF867230A14C5C86FED5" box="[826,953,258,281]" degrees="17" direction="south" minutes="49" orientation="latitude" pageId="21" pageNumber="710" precision="15" seconds="23" value="-17.823057">
17°49
<emphasis id="B934EA89FFCFFF867271A14D5CBEFED4" box="[891,897,258,281]" italics="true" pageId="21" pageNumber="710">′</emphasis>
23
<emphasis id="B934EA89FFCFFF867296A14D5C97FED4" box="[924,936,258,281]" italics="true" pageId="21" pageNumber="710">′′</emphasis>
S
</geoCoordinate>
,
<geoCoordinate id="EE74505CFFCFFF8672C7A14C5B64FED4" box="[973,1115,258,281]" degrees="128" direction="east" minutes="14" orientation="longitude" pageId="21" pageNumber="710" precision="15" seconds="58" value="128.24945">
128°14
<emphasis id="B934EA89FFCFFF867516A14D5B1DFED4" box="[1052,1058,258,281]" italics="true" pageId="21" pageNumber="710">′</emphasis>
58
<emphasis id="B934EA89FFCFFF867537A14D5B76FED4" box="[1085,1097,258,281]" italics="true" pageId="21" pageNumber="710">′′</emphasis>
E
</geoCoordinate>
(coll.
<collectorName id="26B5534DFFCFFF8675A4A14B5A10FED4" box="[1198,1327,260,281]" pageId="21" pageNumber="710">V. Kessner</collectorName>
, R. Hokkanen,
<date id="FFFE105BFFCFFF867299A16D5CC3FEF5" box="[915,1020,290,312]" pageId="21" pageNumber="710" value="2009-09-01">
<collectingDate id="EFBAE9B3FFCFFF867299A16D5CC3FEF5" box="[915,1020,290,312]" pageId="21" pageNumber="710" value="2009-09-01">1.ix.2009</collectingDate>
</date>
;
<specimenCode id="DBE69EE0FFCFFF867506A16D5B63FEF5" box="[1036,1116,290,312]" collectionCode="WA-" pageId="21" pageNumber="710">WA-76</specimenCode>
/09), 17 dry (
<collectionCode id="ED51AE5EFFCFFF8675F8A16D5A0CFEFA" box="[1266,1331,290,311]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="21" pageNumber="710" type="Museum">WAM</collectionCode>
<accessionNumber id="9413AB78FFCFFF867434A16D5AAAFEF5" box="[1342,1429,290,312]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S49138" pageId="21" pageNumber="710" type="EnaNcbi">S49138</accessionNumber>
);
<collectingMunicipality id="6B9BACE1FFCFFF867230A10F5B17FE9B" box="[826,1064,320,342]" pageId="21" pageNumber="710">Halls Creek District</collectingMunicipality>
, south of
<collectingCounty id="629E4E17FFCFFF867597A10E5A07FE9B" box="[1181,1336,320,342]" pageId="21" pageNumber="710">Panton River</collectingCounty>
, low and open limestone area, numerous gullies, under welldeveloped spinifex, in talus,
<geoCoordinate id="EE74505CFFCFFF86758CA1315A3AFE5E" box="[1158,1285,380,403]" degrees="17" direction="south" minutes="49" orientation="latitude" pageId="21" pageNumber="710" precision="15" seconds="23" value="-17.823057">
17°49
<emphasis id="B934EA89FFCFFF8675CDA1335BF2FE5E" box="[1223,1229,380,403]" italics="true" pageId="21" pageNumber="710">′</emphasis>
23
<emphasis id="B934EA89FFCFFF8675E2A1335BCBFE5E" box="[1256,1268,380,403]" italics="true" pageId="21" pageNumber="710">′′</emphasis>
S
</geoCoordinate>
,
<geoCoordinate id="EE74505CFFCFFF867419A1315A9EFE5E" box="[1299,1441,380,403]" degrees="128" direction="east" minutes="14" orientation="longitude" pageId="21" pageNumber="710" precision="15" seconds="58" value="128.24945">
128°14
<emphasis id="B934EA89FFCFFF867468A1335A57FE5E" box="[1378,1384,380,403]" italics="true" pageId="21" pageNumber="710">′</emphasis>
58
<emphasis id="B934EA89FFCFFF867489A1335AB0FE5E" box="[1411,1423,380,403]" italics="true" pageId="21" pageNumber="710">′′</emphasis>
E
</geoCoordinate>
(coll.
<collectorName id="26B5534DFFCFFF867270A1D25CC9FE7F" box="[890,1014,413,434]" pageId="21" pageNumber="710">V. Kessner</collectorName>
, R. Hokkanen,
<date id="FFFE105BFFCFFF8675A7A1D35A29FE7F" box="[1197,1302,412,434]" pageId="21" pageNumber="710" value="2009-09-01">
<collectingDate id="EFBAE9B3FFCFFF8675A7A1D35A29FE7F" box="[1197,1302,412,434]" pageId="21" pageNumber="710" value="2009-09-01">1.ix.2009</collectingDate>
</date>
;
<specimenCode id="DBE69EE0FFCFFF86742EA1D35A4BFE7F" box="[1316,1396,412,434]" collectionCode="WA-" pageId="21" pageNumber="710">WA-76</specimenCode>
/09), four dry (
<collectionCode id="ED51AE5EFFCFFF8672A3A1F45CD5FE1D" box="[937,1002,443,464]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="21" pageNumber="710" type="Museum">WAM</collectionCode>
S49144)
</materialsCitation>
.
</paragraph>
<paragraph id="8BFF369BFFCFFF867230A2485A73FD96" blockId="21.[826,1441,519,603]" pageId="21" pageNumber="710">
<emphasis id="B934EA89FFCFFF867230A2485C87FDD1" box="[826,952,519,540]" italics="true" pageId="21" pageNumber="710">Etymology:</emphasis>
In reference to the peculiar colour pattern, formed by fine maculations, derived from ‘punctatus’ (Latin = dotted), adjective of feminine gender.
</paragraph>
<paragraph id="8BFF369BFFCFFF867230A2DD5B75FC51" blockId="21.[826,1441,657,1261]" pageId="21" pageNumber="710">
<emphasis id="B934EA89FFCFFF867230A2DD5CFDFD6A" box="[826,962,658,679]" italics="true" pageId="21" pageNumber="710">Description:</emphasis>
Shell (
<figureCitation id="137B2A1EFFCFFF86753BA2DE5BB2FD6A" box="[1073,1165,657,679]" captionStart="Figure 4" captionStartId="8.[144,223,954,973]" captionTargetBox="[144,1425,196,924]" captionTargetId="figure-472@8.[144,1425,196,924]" captionTargetPageId="8" captionText="Figure 4. Photographs of living specimens of Exiligada species (courtesy Vince Kessner, Adelaide River). A, E. negriensis Iredale, 1939. B, E. pallida sp. nov. C, E. nodulicaudata sp. nov. D, E. punctata sp. nov. E, E. qualis Iredale, 1939. F, E. monochroma sp. nov. G, E. unistriata sp. nov. H, E. floraevallis sp. nov. I, E. rivifontis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5409193" httpUri="https://zenodo.org/record/5409193/files/figure.png" pageId="21" pageNumber="710">Figs 4D</figureCitation>
,
<figureCitation id="137B2A1EFFCFFF8675ADA2DE5BF5FD6A" box="[1191,1226,657,679]" captionStart="Figure 6" captionStartId="9.[826,905,1233,1252]" captionTargetBox="[834,1435,197,1200]" captionTargetId="figure-294@9.[830,1439,194,1203]" captionTargetPageId="9" captionText="Figure 6. Shells of Exiligada species. A, E. montejinni sp. nov. (paratype AM C.469982). B, E. nodulicauda sp. nov. (paratype WAM S49160). C, E. pallida sp. nov. (paratype WAM S49215). D, E. punctata sp. nov. (paratype WAM S49221). E, E. qualis Iredale, 1939 (holotype AM C.64866). F, E. rivifontis sp. nov. (WAM S49225). G, E. unistriata sp. nov. (paratype AM C.470460). Scale bar = 2 mm. Note that foot produces from some shells." figureDoi="http://doi.org/10.5281/zenodo.5409195" httpUri="https://zenodo.org/record/5409195/files/figure.png" pageId="21" pageNumber="710">6D</figureCitation>
,
<figureCitation id="137B2A1EFFCFFF8675EFA2DD5A09FD65" box="[1253,1334,658,680]" captionStart="Figure 20" captionStartId="22.[144,223,1220,1239]" captionTargetBox="[303,1262,196,1190]" captionTargetId="figure-272@22.[303,1263,196,1190]" captionTargetPageId="22" captionText="Figure 20. SEM micrographs of Exiligada shells.A-C, E. pallida sp. nov. (paratype WAM S49150). A, Apical view showing protoconch and first whorl. B, Close-up from above of sculpture across entire shell. C, Microsculpture on teleoconch whorl. D-F, E. punctata sp. nov. (paratype WAMS49139). D, Apical view showing protoconch and first whorl. E, Close-up from above of sculpture across entire shell. F, Microsculpture on teleoconch whorl. Scale bars: A-B, D-E = 1 mm, C, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5409227" httpUri="https://zenodo.org/record/5409227/files/figure.png" pageId="21" pageNumber="710">20D–F</figureCitation>
). Large, moderately elevated, moderately thick; micropustulations absent. Shell background colour yellowish horn fading to whitish towards shell base, with several brown-reddish spiral bands, most conspicuous on body whorl, never present on shell base, often interrupted radially; sparse brown maculations in radial clusters present on whole spire and increasing in number on last whorls.
</paragraph>
<caption id="DF3F6613FFCFFF8671AEA2B75EADFC4D" ID-DOI="http://doi.org/10.5281/zenodo.5409221" ID-Zenodo-Dep="5409221" httpUri="https://zenodo.org/record/5409221/files/figure.png" pageId="21" pageNumber="710" startId="21.[164,243,760,779]" targetBox="[169,776,198,729]" targetPageId="21">
<paragraph id="8BFF369BFFCFFF8671AEA2B75EADFC4D" blockId="21.[164,779,760,896]" pageId="21" pageNumber="710">
<emphasis id="B934EA89FFCFFF8671AEA2B75E26FCC6" bold="true" box="[164,281,760,779]" pageId="21" pageNumber="710">Figure 18.</emphasis>
Penial anatomy of
<emphasis id="B934EA89FFCFFF8670F7A2B75C34FCC6" bold="true" box="[509,779,760,779]" pageId="21" pageNumber="710">
<taxonomicName id="4C404D18FFCFFF8670F7A2B75D9DFCC6" authority="Criscione &amp; Law &amp; Köhler, 2012" authorityName="Criscione &amp; Law &amp; Köhler" authorityYear="2012" box="[509,674,760,779]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="710" phylum="Mollusca" rank="species" species="montejinni" status="sp. nov.">
<emphasis id="B934EA89FFCFFF8670F7A2B75D9DFCC6" bold="true" box="[509,674,760,779]" italics="true" pageId="21" pageNumber="710">E. montejinni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20757F2FFCFFF8673A4A2B75C34FCC6" box="[686,779,760,779]" pageId="21" pageNumber="710" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype C.475769). A. External anatomy of penial complex, penial sheath removed. B. Anatomy of inner penial wall and detail of epiphallus–penis junction. Scale bar = 5 mm; a = 1 mm.
</paragraph>
</caption>
<paragraph id="8BFF369BFFCFFF867259A3EA5CECFB20" blockId="21.[826,1441,657,1261]" pageId="21" pageNumber="710">
Genitalia (
<figureCitation id="137B2A1EFFCFFF8672D8A3EA5B1BFC76" box="[978,1060,933,955]" captionStart="Figure 22" captionStartId="23.[164,243,1578,1597]" captionTargetBox="[232,712,880,1547]" captionTargetId="figure-495@23.[232,712,878,1549]" captionTargetPageId="23" captionText="Figure 22. Penial anatomy of E. punctata sp. nov. (holotype WAM S83175). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5409231" httpUri="https://zenodo.org/record/5409231/files/figure.png" pageId="21" pageNumber="710">Fig. 22</figureCitation>
). Epiphallus one third of penis. Vas deferens entering penial sheath from one third up. Penis five times longer than wide, not coiled inside sheath, with inner wall supporting a large apical corrugated pad and six narrow, regularly shaped longitudinal pilasters, five of which interrupted posteriorly and one running along entire wall length. Vagina one third shorter than penis and as long as free oviduct, bursa copulatrix shape variable, without well-differentiated end or with a short to long rounded end.
</paragraph>
<paragraph id="8BFF369BFFCFFF867230A56B5B48F947" blockId="21.[826,1442,1316,1674]" pageId="21" pageNumber="710">
<emphasis id="B934EA89FFCFFF867230A56B5C99FAF4" box="[826,934,1316,1337]" italics="true" pageId="21" pageNumber="710">Remarks:</emphasis>
Shell colour pattern similar to
<taxonomicName id="4C404D18FFCFFF867423A56B5AA5FAF4" authorityName="Criscione &amp; Law &amp; Köhler" authorityYear="2012" box="[1321,1434,1316,1337]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="710" phylum="Mollusca" rank="species" species="pallida">
<emphasis id="B934EA89FFCFFF867423A56B5AA5FAF4" box="[1321,1434,1316,1337]" italics="true" pageId="21" pageNumber="710">E. pallida</emphasis>
</taxonomicName>
,
<taxonomicName id="4C404D18FFCFFF867230A50C5CE7FA9A" authorityName="Criscione &amp; Law &amp; Köhler" authorityYear="2012" box="[826,984,1346,1368]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="710" phylum="Mollusca" rank="species" species="longicauda">
<emphasis id="B934EA89FFCFFF867230A50C5CE7FA9A" box="[826,984,1346,1368]" italics="true" pageId="21" pageNumber="710">E. longicauda</emphasis>
</taxonomicName>
,
<taxonomicName id="4C404D18FFCFFF8672EDA50C5BA9FA9A" authorityName="Criscione &amp; Law &amp; Köhler" authorityYear="2012" box="[999,1174,1346,1368]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="710" phylum="Mollusca" rank="species" species="nodulicauda">
<emphasis id="B934EA89FFCFFF8672EDA50C5BA9FA9A" box="[999,1174,1346,1368]" italics="true" pageId="21" pageNumber="710">E. nodulicauda</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C404D18FFCFFF8675D3A50C5A54FA95" authorityName="Iredale" authorityYear="1939" box="[1241,1387,1347,1368]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="710" phylum="Mollusca" rank="species" species="negriensis">
<emphasis id="B934EA89FFCFFF8675D3A50C5A54FA95" box="[1241,1387,1347,1368]" italics="true" pageId="21" pageNumber="710">E. negriensis</emphasis>
</taxonomicName>
, but easily distinguishable by presence of maculations. Penis length relative to width (pl/pw) and epiphallus length relative to penis (el
<superScript id="7C359BD3FFCFFF86757AA5D15BA9FA65" attach="left" box="[1136,1174,1436,1460]" fontSize="5" pageId="21" pageNumber="710">pl-1</superScript>
) smaller than average. Inner penial wall sculpture similar to
<taxonomicName id="4C404D18FFCFFF867405A5F25A9EFA1F" authorityName="Iredale" authorityYear="1939" box="[1295,1441,1469,1490]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="710" phylum="Mollusca" rank="species" species="negriensis">
<emphasis id="B934EA89FFCFFF867405A5F25A9EFA1F" box="[1295,1441,1469,1490]" italics="true" pageId="21" pageNumber="710">E. negriensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C404D18FFCFFF867267A5935CE1FA3D" authorityName="Criscione &amp; Law &amp; Köhler" authorityYear="2012" box="[877,990,1499,1521]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="710" phylum="Mollusca" rank="species" species="pallida">
<emphasis id="B934EA89FFCFFF867267A5935CE1FA3D" box="[877,990,1499,1521]" italics="true" pageId="21" pageNumber="710">E. pallida</emphasis>
</taxonomicName>
, to be distinguished from these species by narrower posterior pad and longer longitudinal pilasters, always reaching the posterior part of the wall, and by latter by posterior pad being corrugated. Vagina length relative to penis (vl/pl) and relative to free oviduct (vl/ol) average.
</paragraph>
</subSubSection>
</treatment>
</document>
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