168 lines
20 KiB
XML
168 lines
20 KiB
XML
<document id="D967F286753DA6710C8F2DAE34A80A49" ID-DOI="10.11646/zootaxa.5369.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10146170" ID-ZooBank="42563452-CF2B-47A6-B0B0-485AC4F320BD" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1700185584045" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2023" docId="038DBF6E542FCC55FF06FE4AFA44F83C" docLanguage="en" docName="zootaxa.5369.2.1.pdf" docOrigin="Zootaxa 5369 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5369.2.1/52237" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Omalium Gravenhorst 1802" docType="treatment" docVersion="1" lastPageNumber="161" masterDocId="FFB4C7165425CC5FFF91FFD8FFD8FF82" masterDocTitle="A revision of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae) of the Himalayan region" masterLastPageNumber="206" masterPageNumber="151" pageNumber="161" updateTime="1700578467175" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:title id="0EFBBD3DA4EC0DC5DF4F4CBF122EF5FF">A revision of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae) of the Himalayan region</mods:title>
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<mods:namePart id="B8BB3208233C197B0E1BA1F2CFF1E4D6">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="4D699665A77919702C8B9EE8A497D84E">Zootaxa</mods:title>
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<mods:part id="2AD071FA2EF25D23C4040F527C9CFA69">
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<mods:date id="9E504D7955DF1FD94982F48DFED90370">2023</mods:date>
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<mods:number id="809867A3FAF278C00EEEB43C531E8BE2">2023-11-09</mods:number>
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<mods:number id="0FF3EB91F3404ED21172F5A140085D69">5369</mods:number>
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<mods:url id="2FB7FBF16B95F03B4366BB377FAB607A">https://www.mapress.com/zt/article/download/zootaxa.5369.2.1/52237</mods:url>
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<mods:classification id="4AE14782C201DC6C98FA3E73A180F9E2">journal article</mods:classification>
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<mods:identifier id="F911E399BBF86CE53BE3538DE7AC6C6B" type="DOI">10.11646/zootaxa.5369.2.1</mods:identifier>
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<mods:identifier id="DD6368E525D1B962E0B9D6EF60795891" type="ISSN">1175-5326</mods:identifier>
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<treatment id="038DBF6E542FCC55FF06FE4AFA44F83C" LSID="urn:lsid:plazi:treatment:038DBF6E542FCC55FF06FE4AFA44F83C" httpUri="http://treatment.plazi.org/id/038DBF6E542FCC55FF06FE4AFA44F83C" lastPageNumber="161" pageId="10" pageNumber="161">
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<subSubSection id="C33E5DF3542FCC55FF06FE4AFE07FE2F" box="[151,479,402,429]" pageId="10" pageNumber="161" type="nomenclature">
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<paragraph id="8B9B0E78542FCC55FF06FE4AFE07FE2F" blockId="10.[151,479,402,429]" box="[151,479,402,429]" pageId="10" pageNumber="161">
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<heading id="D0D3B914542FCC55FF06FE4AFE07FE2F" bold="true" box="[151,479,402,429]" fontSize="11" level="1" pageId="10" pageNumber="161" reason="1">
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<emphasis id="B950D26A542FCC55FF06FE4AFE07FE2F" bold="true" box="[151,479,402,429]" pageId="10" pageNumber="161">
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3.2.2.
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<taxonomicName id="4C2475FB542FCC55FF4EFE4AFE55FE2E" ID-CoL="8VRX7" authority="Gravenhorst, 1802" authorityName="Gravenhorst" authorityYear="1802" box="[223,397,402,428]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="genus">Amplissimum</taxonomicName>
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group
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33E5DF3542FCC55FF06FE0CFAD2FD90" pageId="10" pageNumber="161" type="diagnosis">
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<paragraph id="8B9B0E78542FCC55FF06FE0CFAD2FD90" blockId="10.[151,1436,468,674]" pageId="10" pageNumber="161">
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<emphasis id="B950D26A542FCC55FF06FE0CFECAFE6C" bold="true" box="[151,274,468,494]" pageId="10" pageNumber="161">Diagnosis.</emphasis>
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Aedeagus very wide and short, with elongate median lobe and somewhat truncate (
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<figureCitation id="131F12FD542FCC55FB5BFE0CFAC5FE6C" box="[1226,1309,468,494]" captionStart="FIGURES 27–29" captionStartId="13.[151,264,1505,1529]" captionTargetBox="[154,1433,456,1479]" captionTargetId="figure-158@13.[151,1436,453,1482]" captionTargetPageId="13" captionText="FIGURES 27–29. Aedeagus of Omalium amplissimum (paratype): 27—aedeagus, parameral view, 28—aedeagus, lateral view, 29—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146186" httpUri="https://zenodo.org/record/10146186/files/figure.png" pageId="10" pageNumber="161">Fig. 27</figureCitation>
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) or incised (
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<figureCitation id="131F12FD542FCC55FF0EFE20FF29FD90" box="[159,241,504,530]" captionStart="FIGURES 30–32" captionStartId="14.[151,264,1598,1622]" captionTargetBox="[154,1433,672,1570]" captionTargetId="figure-236@14.[151,1436,669,1575]" captionTargetPageId="14" captionText="FIGURES 30–32. Aedeagus of Omalium bilobum (holotype): 30—aedeagus, parameral view, 31—aedeagus, lateral view, 32—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146188" httpUri="https://zenodo.org/record/10146188/files/figure.png" pageId="10" pageNumber="161">Fig. 30</figureCitation>
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) apex; parameres wide, slighty or significantly (
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<figureCitation id="131F12FD542FCC55FC9EFE20FCB9FD90" box="[783,865,504,530]" captionStart="FIGURES 27–29" captionStartId="13.[151,264,1505,1529]" captionTargetBox="[154,1433,456,1479]" captionTargetId="figure-158@13.[151,1436,453,1482]" captionTargetPageId="13" captionText="FIGURES 27–29. Aedeagus of Omalium amplissimum (paratype): 27—aedeagus, parameral view, 28—aedeagus, lateral view, 29—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146186" httpUri="https://zenodo.org/record/10146186/files/figure.png" pageId="10" pageNumber="161">Fig. 27</figureCitation>
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) longer than apex of the median lobe.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33E5DF3542FCC55FF56FDC4FCB3FDB4" box="[199,875,540,567]" pageId="10" pageNumber="161" type="taxon_list">
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<paragraph id="8B9B0E78542FCC55FF56FDC4FCB3FDB4" blockId="10.[151,1436,468,674]" box="[199,875,540,567]" pageId="10" pageNumber="161">
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<emphasis id="B950D26A542FCC55FF56FDC4FE4DFDB4" bold="true" box="[199,405,540,566]" pageId="10" pageNumber="161">Species included:</emphasis>
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<taxonomicName id="4C2475FB542FCC55FE0DFDC4FD8BFDB4" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[412,595,540,567]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="amplissimum" status="sp. n.">
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<emphasis id="B950D26A542FCC55FE0DFDC4FE68FDB4" box="[412,432,540,566]" italics="true" pageId="10" pageNumber="161">O</emphasis>
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.
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<emphasis id="B950D26A542FCC55FE2EFDC4FD8BFDB4" box="[447,595,540,566]" italics="true" pageId="10" pageNumber="161">amplissimum</emphasis>
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</taxonomicName>
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<emphasis id="B950D26A542FCC55FDC8FDC4FD41FDB4" bold="true" box="[601,665,540,566]" pageId="10" pageNumber="161">
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<taxonomicNameLabel id="A2636F11542FCC55FDC8FDC4FD41FDB4" box="[601,665,540,566]" pageId="10" pageNumber="161" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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,
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<taxonomicName id="4C2475FB542FCC55FD37FDC4FCFDFDB4" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[678,805,540,567]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="bilobum" status="sp. n.">
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<emphasis id="B950D26A542FCC55FD37FDC4FD62FDB4" box="[678,698,540,566]" italics="true" pageId="10" pageNumber="161">O</emphasis>
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.
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<emphasis id="B950D26A542FCC55FD58FDC4FCFDFDB4" box="[713,805,540,566]" italics="true" pageId="10" pageNumber="161">bilobum</emphasis>
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</taxonomicName>
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<emphasis id="B950D26A542FCC55FCBAFDC4FCB3FDB4" bold="true" box="[811,875,540,566]" pageId="10" pageNumber="161">
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<taxonomicNameLabel id="A2636F11542FCC55FCBAFDC4FCB3FDB4" box="[811,875,540,566]" pageId="10" pageNumber="161" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33E5DF3542FCC55FF56FD98FD77FD20" pageId="10" pageNumber="161" type="discussion">
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<paragraph id="8B9B0E78542FCC55FF56FD98FD77FD20" blockId="10.[151,1436,468,674]" pageId="10" pageNumber="161">
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<emphasis id="B950D26A542FCC55FF56FD98FEE3FDD8" bold="true" box="[199,315,576,602]" pageId="10" pageNumber="161">Remarks.</emphasis>
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Regarding the general shape and ground sculpture of the body, species of the Amplissimum group are similar to the Himalayan species of the
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<taxonomicName id="4C2475FB542FCC55FD11FDBCFCCCFDFC" authorityName="Gravenhorst" authorityYear="1806" box="[640,788,612,638]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="oxyacanthae">
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<emphasis id="B950D26A542FCC55FD11FDBCFCCCFDFC" box="[640,788,612,638]" italics="true" pageId="10" pageNumber="161">Oxyacanthae</emphasis>
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</taxonomicName>
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and the Rivulare group, but differs by shapes of the wide aedeagus and apical portion of the median lobe.
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</paragraph>
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</subSubSection>
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||
<caption id="DF5B5EF0542FCC55FF06F9EBFCE9F9C9" ID-DOI="http://doi.org/10.5281/zenodo.10146182" ID-Zenodo-Dep="10146182" box="[151,817,1587,1612]" httpUri="https://zenodo.org/record/10146182/files/figure.png" pageId="10" pageNumber="161" startId="10.[151,250,1587,1611]" targetBox="[151,1436,698,1564]" targetPageId="10" targetType="figure">
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<paragraph id="8B9B0E78542FCC55FF06F9EBFCE9F9C9" blockId="10.[151,817,1587,1612]" box="[151,817,1587,1612]" pageId="10" pageNumber="161">
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<emphasis id="B950D26A542FCC55FF06F9EBFEF8F9C9" bold="true" box="[151,288,1587,1611]" pageId="10" pageNumber="161">FIGURE 23.</emphasis>
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Distribution of
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<taxonomicName id="4C2475FB542FCC55FE53F9EBFD56F9C9" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[450,654,1587,1611]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="amicorum" status="sp. nov.">
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<emphasis id="B950D26A542FCC55FE53F9EBFD56F9C9" box="[450,654,1587,1611]" italics="true" pageId="10" pageNumber="161">Omalium amicorum</emphasis>
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</taxonomicName>
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<emphasis id="B950D26A542FCC55FD05F9ECFD15F9CE" bold="true" box="[660,717,1588,1612]" pageId="10" pageNumber="161">
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<taxonomicNameLabel id="A2636F11542FCC55FD05F9ECFD15F9CE" box="[660,717,1588,1612]" pageId="10" pageNumber="161" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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in Nepal.
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</paragraph>
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</caption>
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<subSubSection id="C33E5DF3542FCC55FF06F945FD96F93A" box="[151,590,1693,1720]" pageId="10" pageNumber="161" type="multiple">
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<paragraph id="8B9B0E78542FCC55FF06F945FD96F93A" blockId="10.[151,590,1693,1720]" box="[151,590,1693,1720]" pageId="10" pageNumber="161">
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<heading id="D0D3B914542FCC55FF06F945FD96F93A" bold="true" box="[151,590,1693,1720]" fontSize="11" level="1" pageId="10" pageNumber="161" reason="1">
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<emphasis id="B950D26A542FCC55FF06F945FD96F93A" bold="true" box="[151,590,1693,1720]" pageId="10" pageNumber="161">3.2.2.1. Key to Amplissimum group</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33E5DF3542FCC55FF06F904FA44F83C" pageId="10" pageNumber="161" type="key">
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<keyStep id="30D016DD542FCC55FF06F904FA44F83C" pageId="10" pageNumber="161">
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<key id="FFB55949542FCC55FF06F904FA44F83C" pageId="10" pageNumber="161">
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<paragraph id="8B9B0E78542FCC55FF06F904FA44F8CB" blockId="10.[151,1436,1756,1983]" pageId="10" pageNumber="161">
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<keyLead id="30D5AD4D542FCC55FF06F904FA44F8CB" pageId="10" pageNumber="161">
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1 Pronotum widest in apical third, with apical angles not protruded anteriad. Punctation and microsculpture of head and pronotum distinctly coarse. Elytra about as long as broad. Median lobe with truncate apex; parameres very wide, significantly longer than apex of median lobe (
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<figureCitation id="131F12FD542FCC55FE23F8CFFE22F8AE" box="[434,506,1815,1837]" captionStart="FIGURES 27–29" captionStartId="13.[151,264,1505,1529]" captionTargetBox="[154,1433,456,1479]" captionTargetId="figure-158@13.[151,1436,453,1482]" captionTargetPageId="13" captionText="FIGURES 27–29. Aedeagus of Omalium amplissimum (paratype): 27—aedeagus, parameral view, 28—aedeagus, lateral view, 29—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146186" httpUri="https://zenodo.org/record/10146186/files/figure.png" pageId="10" pageNumber="161">Fig. 27</figureCitation>
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). Female accessory sclerite as in
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<figureCitation id="131F12FD542FCC55FCA8F8CFFCA6F8AE" box="[825,894,1815,1837]" captionStart="FIGURES 8–18" captionStartId="6.[151,264,1896,1920]" captionTargetBox="[154,1433,564,1870]" captionTargetId="figure-184@6.[151,1436,561,1873]" captionTargetPageId="6" captionText="FIGURES 8–18. Female accessory sclerite (Figs 8, 10–16, 18) and spermatheca (Figs 9, 17) of Omalium: 8–9—O. acutangulum, 10—O. amicorum, 11—O. amplissimum, 12—O. bilobum, 13—O. alticola, 14—O.beesoni, 15—O. altivagans, 16–17—O. marginatum, 18—O. scapulare. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146176" httpUri="https://zenodo.org/record/10146176/files/figure.png" pageId="10" pageNumber="161">Fig. 11</figureCitation>
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. Habitus as in
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<figureCitation id="131F12FD542FCC55FB9AF8CFFB8AF8AE" box="[1035,1106,1815,1837]" captionStart="FIGURES 24–26" captionStartId="11.[151,264,1490,1514]" captionTargetBox="[164,1420,463,1452]" captionTargetId="figure-64@11.[151,1435,453,1467]" captionTargetPageId="11" captionText="FIGURES 24–26. Habitus of Omalium: 24—O. amplissimum (paratype), 25—O. bilobum (paratype), 26—O. altivagans (Kashmir). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10146184" httpUri="https://zenodo.org/record/10146184/files/figure.png" pageId="10" pageNumber="161">Fig. 24</figureCitation>
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||
. Body length:
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||
<quantity id="4CDCA39D542FCC55FB4FF8CEFAB1F8AE" box="[1246,1385,1814,1836]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" metricValueMax="3.5" metricValueMin="2.7" pageId="10" pageNumber="161" unit="mm" value="3.1" valueMax="3.5" valueMin="2.7">2.70–3.50 mm</quantity>
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......................................................................................
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<taxonomicName id="4C2475FB542FCC55FB55F8ECFAB9F8CB" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1220,1377,1844,1865]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="amplissimum" status="sp. n.">
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||
<emphasis id="B950D26A542FCC55FB55F8ECFB0DF8CB" box="[1220,1237,1844,1865]" italics="true" pageId="10" pageNumber="161">O</emphasis>
|
||
.
|
||
<emphasis id="B950D26A542FCC55FB73F8ECFAB9F8CB" box="[1250,1377,1844,1865]" italics="true" pageId="10" pageNumber="161">amplissimum</emphasis>
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||
</taxonomicName>
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||
<emphasis id="B950D26A542FCC55FAF7F8ECFA44F8CB" bold="true" box="[1382,1436,1844,1865]" pageId="10" pageNumber="161">
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||
<taxonomicNameLabel id="A2636F11542FCC55FAF7F8ECFA44F8CB" box="[1382,1436,1844,1865]" pageId="10" pageNumber="161" rank="species">sp. n.</taxonomicNameLabel>
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||
</emphasis>
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||
</keyLead>
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||
</paragraph>
|
||
<paragraph id="8B9B0E78542FCC55FF06F88AFA44F83C" blockId="10.[151,1436,1756,1983]" pageId="10" pageNumber="161">
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||
<keyLead id="30D5AD4D542FCC55FF06F88AFA44F83C" pageId="10" pageNumber="161">
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||
- Pronotum widest slightly above middle, with anterior angles slightly protruded anteriad. Punctation and microsculpture of head and pronotum moderately fine. Elytra slightly broader than long. Median lobe with wide apical excision; parameres moderately wide, slightly longer than apex of median lobe (
|
||
<figureCitation id="131F12FD542FCC55FD25F854FD24F823" box="[692,764,1932,1954]" captionStart="FIGURES 30–32" captionStartId="14.[151,264,1598,1622]" captionTargetBox="[154,1433,672,1570]" captionTargetId="figure-236@14.[151,1436,669,1575]" captionTargetPageId="14" captionText="FIGURES 30–32. Aedeagus of Omalium bilobum (holotype): 30—aedeagus, parameral view, 31—aedeagus, lateral view, 32—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146188" httpUri="https://zenodo.org/record/10146188/files/figure.png" pageId="10" pageNumber="161">Fig. 30</figureCitation>
|
||
). Female accessory sclerite as in
|
||
<figureCitation id="131F12FD542FCC55FBD1F854FB5FF823" box="[1088,1159,1932,1954]" captionStart="FIGURES 8–18" captionStartId="6.[151,264,1896,1920]" captionTargetBox="[154,1433,564,1870]" captionTargetId="figure-184@6.[151,1436,561,1873]" captionTargetPageId="6" captionText="FIGURES 8–18. Female accessory sclerite (Figs 8, 10–16, 18) and spermatheca (Figs 9, 17) of Omalium: 8–9—O. acutangulum, 10—O. amicorum, 11—O. amplissimum, 12—O. bilobum, 13—O. alticola, 14—O.beesoni, 15—O. altivagans, 16–17—O. marginatum, 18—O. scapulare. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146176" httpUri="https://zenodo.org/record/10146176/files/figure.png" pageId="10" pageNumber="161">Fig. 12</figureCitation>
|
||
. Habitus as in
|
||
<figureCitation id="131F12FD542FCC55FA86F854FA86F820" box="[1303,1374,1932,1954]" captionStart="FIGURES 24–26" captionStartId="11.[151,264,1490,1514]" captionTargetBox="[164,1420,463,1452]" captionTargetId="figure-64@11.[151,1435,453,1467]" captionTargetPageId="11" captionText="FIGURES 24–26. Habitus of Omalium: 24—O. amplissimum (paratype), 25—O. bilobum (paratype), 26—O. altivagans (Kashmir). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10146184" httpUri="https://zenodo.org/record/10146184/files/figure.png" pageId="10" pageNumber="161">Fig. 25</figureCitation>
|
||
. Body length:
|
||
<quantity id="4CDCA39D542FCC55FEBAF871FE6EF83D" box="[299,438,1961,1983]" metricMagnitude="-3" metricUnit="m" metricValue="2.885" metricValueMax="2.97" metricValueMin="2.8" pageId="10" pageNumber="161" unit="mm" value="2.885" valueMax="2.97" valueMin="2.8">2.80–2.97 mm</quantity>
|
||
.....................................................................
|
||
<taxonomicName id="4C2475FB542FCC55FB65F872FAB9F83C" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1268,1377,1961,1983]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="bilobum" status="sp. n.">
|
||
<emphasis id="B950D26A542FCC55FB65F872FADDF83D" box="[1268,1285,1962,1983]" italics="true" pageId="10" pageNumber="161">O</emphasis>
|
||
.
|
||
<emphasis id="B950D26A542FCC55FA83F871FAB9F83C" box="[1298,1377,1961,1982]" italics="true" pageId="10" pageNumber="161">bilobum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B950D26A542FCC55FAF7F871FA44F83C" bold="true" box="[1382,1436,1961,1982]" pageId="10" pageNumber="161">
|
||
<taxonomicNameLabel id="A2636F11542FCC55FAF7F871FA44F83C" box="[1382,1436,1961,1982]" pageId="10" pageNumber="161" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</key>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |