386 lines
49 KiB
XML
386 lines
49 KiB
XML
<document id="1C7038B15D94A02DD3A914079D549811" ID-DOI="10.11646/zootaxa.5369.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10146170" ID-ZooBank="42563452-CF2B-47A6-B0B0-485AC4F320BD" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1700185584045" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2023" docId="038DBF6E5422CC55FF06FD0EFAE4FEC3" docLanguage="en" docName="zootaxa.5369.2.1.pdf" docOrigin="Zootaxa 5369 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5369.2.1/52237" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Omalium amicorum Shavrin 2023, sp. n." docType="treatment" docVersion="1" lastPageNumber="161" masterDocId="FFB4C7165425CC5FFF91FFD8FFD8FF82" masterDocTitle="A revision of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae) of the Himalayan region" masterLastPageNumber="206" masterPageNumber="151" pageNumber="158" updateTime="1700578467175" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:title id="519DC04265A075D10E69021F8042AA36">A revision of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae) of the Himalayan region</mods:title>
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<mods:namePart id="9B3C907CFB3EAF171350F1E05D807127">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="5AA2E31EA7AE4ED20BAF381E286093FF">Zootaxa</mods:title>
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<mods:date id="17E4EBC9CA87B23947BAFCB4B58E4A5E">2023</mods:date>
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<mods:number id="27C802B5A4AA24BAE4C13D6DF162CF92">2023-11-09</mods:number>
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<mods:number id="28BD750C296B5B164ACBE05E311B08F2">5369</mods:number>
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<mods:classification id="CAF6F6FE64CBA1BF06BC912B06D01AE7">journal article</mods:classification>
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<mods:identifier id="DF51D9FF020C3D1824B854CAE278CA5F" type="DOI">10.11646/zootaxa.5369.2.1</mods:identifier>
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<mods:identifier id="AA89AE3FAC26D6BA2383BBB6929F354E" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="038DBF6E5422CC55FF06FD0EFAE4FEC3" LSID="urn:lsid:plazi:treatment:038DBF6E5422CC55FF06FD0EFAE4FEC3" httpUri="http://treatment.plazi.org/id/038DBF6E5422CC55FF06FD0EFAE4FEC3" lastPageId="10" lastPageNumber="161" pageId="7" pageNumber="158">
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<subSubSection id="C33E5DF35422CC58FF06FD0EFD93FD73" box="[151,587,726,753]" pageId="7" pageNumber="158" type="nomenclature">
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<paragraph id="8B9B0E785422CC58FF06FD0EFD93FD73" blockId="7.[151,587,726,789]" box="[151,587,726,753]" pageId="7" pageNumber="158">
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<heading id="D0D3B9145422CC58FF06FD0EFD93FD73" bold="true" box="[151,587,726,753]" fontSize="11" level="1" pageId="7" pageNumber="158" reason="1">
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<emphasis id="B950D26A5422CC58FF06FD0EFD93FD73" bold="true" box="[151,587,726,753]" pageId="7" pageNumber="158">
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3.2.1.2.2.
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<taxonomicName id="4C2475FB5422CC58FE9CFD0EFDDAFD73" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[269,514,726,753]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="amicorum" status="sp. nov.">
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<emphasis id="B950D26A5422CC58FE9CFD0EFDDAFD73" bold="true" box="[269,514,726,753]" italics="true" pageId="7" pageNumber="158">Omalium amicorum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2636F115422CC58FD98FD0FFD93FD73" box="[521,587,727,753]" pageId="7" pageNumber="158" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33E5DF35422CC58FF06FD23FE51FC97" box="[151,393,763,789]" pageId="7" pageNumber="158" type="description">
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<paragraph id="8B9B0E785422CC58FF06FD23FE51FC97" blockId="7.[151,587,726,789]" box="[151,393,763,789]" pageId="7" pageNumber="158">
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(
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<figureCitation id="131F12FD5422CC58FF0EFD23FEDCFC97" box="[159,260,763,789]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1260,1284]" captionTargetBox="[168,1420,194,1223]" captionTargetId="figure-22@4.[155,1432,181,1236]" captionTargetPageId="4" captionText="FIGURES 1–4. Habitus (Figs 1, 3) and dorsolateral view of head and pronotum (Figs 2, 4) of Omalium: 1–2—O. acutangulum (paratype), 3–4—O. amicorum (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10146172" httpUri="https://zenodo.org/record/10146172/files/figure.png" pageId="7" pageNumber="158">Figs 3–4</figureCitation>
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,
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<figureCitation id="131F12FD5422CC58FE81FD23FEF6FC97" box="[272,302,763,789]" captionStart="FIGURES 8–18" captionStartId="6.[151,264,1896,1920]" captionTargetBox="[154,1433,564,1870]" captionTargetId="figure-184@6.[151,1436,561,1873]" captionTargetPageId="6" captionText="FIGURES 8–18. Female accessory sclerite (Figs 8, 10–16, 18) and spermatheca (Figs 9, 17) of Omalium: 8–9—O. acutangulum, 10—O. amicorum, 11—O. amplissimum, 12—O. bilobum, 13—O. alticola, 14—O.beesoni, 15—O. altivagans, 16–17—O. marginatum, 18—O. scapulare. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146176" httpUri="https://zenodo.org/record/10146176/files/figure.png" pageId="7" pageNumber="158">10</figureCitation>
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,
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<figureCitation id="131F12FD5422CC58FEABFD23FE59FC97" box="[314,385,763,789]" captionStart-0="FIGURE 19" captionStart-1="FIGURES 20–22" captionStartId-0="7.[151,250,1891,1915]" captionStartId-1="8.[151,264,1882,1906]" captionTargetBox-0="[151,1435,957,1868]" captionTargetBox-1="[151,1433,600,1855]" captionTargetId-0="figure-299@7.[151,1435,957,1868]" captionTargetId-1="figure-219@8.[151,1436,597,1858]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURE 19. Distribution of Omalium in the Himalayan region: O. acutangulum sp. n. (circles), O. amplissimum sp. n. (star), O. beesoni (squares), O. cariosum (diamonds), O. flavotestaceum sp. n. (triangles)." captionText-1="FIGURES 20–22. Aedeagus of Omalium amicorum (holotype): 20—aedeagus, parameral view, 21—aedeagus, lateral view, 22—median lobe, lateral view). Scale bar: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10146178" figureDoi-1="http://doi.org/10.5281/zenodo.10146180" httpUri-0="https://zenodo.org/record/10146178/files/figure.png" httpUri-1="https://zenodo.org/record/10146180/files/figure.png" pageId="7" pageNumber="158">19–22</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C33E5DF35422CC57FF06FC9BFA59FE2F" lastPageId="8" lastPageNumber="159" pageId="7" pageNumber="158" type="materials_examined">
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<paragraph id="8B9B0E785422CC58FF06FC9BFDC9FC27" blockId="7.[151,1437,834,933]" pageId="7" pageNumber="158">
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<emphasis id="B950D26A5422CC58FF06FC9BFE9DFCDE" bold="true" box="[151,325,834,861]" pageId="7" pageNumber="158">Type material.</emphasis>
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<materialsCitation id="3B4C04255422CC58FEDDFC9BFD24FC03" collectingDate="2005-05-15" country="Nepal" elevation="2150" latitude="28.366667" location="Manaslu Mts." longLatPrecision="1232" longitude="84.48333" municipality="Ngadi" pageId="7" pageNumber="158" specimenCount="1" typeStatus="holotype">
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<typeStatus id="549FB0DA5422CC58FEDDFC9BFE6CFCDF" box="[332,436,835,861]" pageId="7" pageNumber="158" type="holotype">Holotype</typeStatus>
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♁ (dissected): ‘
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<collectingCountry id="F3334EE85422CC58FDF2FC9BFD67FCDF" box="[611,703,835,861]" name="Nepal" pageId="7" pageNumber="158">NEPAL</collectingCountry>
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<location id="8EFB58A35422CC58FD57FC9BFCB9FCDF" LSID="urn:lsid:plazi:treatment:038DBF6E5422CC55FF06FD0EFAE4FEC3:8EFB58A35422CC58FD57FC9BFCB9FCDF" box="[710,865,835,861]" country="Nepal" latitude="28.366667" longLatPrecision="1232" longitude="84.48333" municipality="Ngadi" name="Manaslu Mts." pageId="7" pageNumber="158">Manaslu Mts.</location>
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|
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<geoCoordinate id="EE1068BF5422CC58FCE5FC9BFC0DFCDF" box="[884,981,835,861]" degrees="28" direction="north" minutes="22" orientation="latitude" pageId="7" pageNumber="158" precision="925" value="28.366667">28°22’N</geoCoordinate>
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<geoCoordinate id="EE1068BF5422CC58FC4AFC9BFBE1FCDF" box="[987,1081,835,861]" degrees="84" direction="east" minutes="29" orientation="longitude" pageId="7" pageNumber="158" precision="925" value="84.48333">84°29’E</geoCoordinate>
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| E slope of
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<collectingMunicipality id="6BFF94025422CC58FB54FC9BFAD1FCDF" box="[1221,1289,835,861]" pageId="7" pageNumber="158">Ngadi</collectingMunicipality>
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| Khola Vall. |
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<quantity id="4CDCA39D5422CC58FF35FCBFFE99FC03" box="[164,321,871,897]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="7" pageNumber="158" unit="m" value="2150.0" valueMax="2300.0" valueMin="2000.0">
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<elevation id="0009E94B5422CC58FF35FCBFFE99FC03" box="[164,321,871,897]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="7" pageNumber="158" unit="m" value="2150.0" valueMax="2300.0" valueMin="2000.0">2000–2300 m</elevation>
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</quantity>
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||
, leg. | [J.] Schmidt
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||
<date id="FF9A28B85422CC58FD89FCBEFD56FC03" box="[536,654,870,897]" pageId="7" pageNumber="158" value="2005-05-15">
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||
<collectingDate id="EFDED1505422CC58FD89FCBEFD56FC03" box="[536,654,870,897]" pageId="7" pageNumber="158" value="2005-05-15">15.V.2005</collectingDate>
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||
</date>
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’ <printed
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||
</materialsCitation>
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||
>, ‘
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||
<typeStatus id="549FB0DA5422CC58FC8BFCBFFC6BFC03" box="[794,947,871,897]" pageId="7" pageNumber="158" type="holotype">HOLOTYPE</typeStatus>
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|
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||
<taxonomicName id="4C2475FB5422CC58FC56FCBFFBF7FC03" authorityName="Gravenhorst" authorityYear="1802" box="[967,1071,871,897]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B950D26A5422CC58FC56FCBFFBF7FC03" box="[967,1071,871,897]" italics="true" pageId="7" pageNumber="158">Omalium</emphasis>
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</taxonomicName>
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|
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<taxonomicName id="4C2475FB5422CC58FBD3FCBFFB6CFC02" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1090,1204,871,896]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="amicorum" status="sp. nov.">
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||
<emphasis id="B950D26A5422CC58FBD3FCBFFB6CFC02" box="[1090,1204,871,896]" italics="true" pageId="7" pageNumber="158">amicorum</emphasis>
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</taxonomicName>
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<emphasis id="B950D26A5422CC58FB2AFCBFFB22FC03" bold="true" box="[1211,1274,871,897]" pageId="7" pageNumber="158">
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||
<taxonomicNameLabel id="A2636F115422CC58FB2AFCBFFB22FC03" box="[1211,1274,871,897]" pageId="7" pageNumber="158" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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| Shavrin A.V. des. 2023’ <red, printed> (NME).
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</paragraph>
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<caption id="DF5B5EF05422CC58FF06F8BBFC3BF81D" ID-DOI="http://doi.org/10.5281/zenodo.10146178" ID-Zenodo-Dep="10146178" httpUri="https://zenodo.org/record/10146178/files/figure.png" pageId="7" pageNumber="158" startId="7.[151,250,1891,1915]" targetBox="[151,1435,957,1868]" targetPageId="7" targetType="figure">
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<paragraph id="8B9B0E785422CC58FF06F8BBFC3BF81D" blockId="7.[151,1436,1891,1951]" pageId="7" pageNumber="158">
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||
<emphasis id="B950D26A5422CC58FF06F8BBFEF8F8F9" bold="true" box="[151,288,1891,1915]" pageId="7" pageNumber="158">FIGURE 19.</emphasis>
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Distribution of
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<taxonomicName id="4C2475FB5422CC58FE52F8BBFDF9F8F9" authorityName="Gravenhorst" authorityYear="1802" box="[451,545,1891,1915]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
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<emphasis id="B950D26A5422CC58FE52F8BBFDF9F8F9" box="[451,545,1891,1915]" italics="true" pageId="7" pageNumber="158">Omalium</emphasis>
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||
</taxonomicName>
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in the Himalayan region:
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<taxonomicName id="4C2475FB5422CC58FCBDF8BCFC17F8F9" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[812,975,1891,1915]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="acutangulum" status="sp. n.">
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||
<emphasis id="B950D26A5422CC58FCBDF8BCFC17F8F9" box="[812,975,1891,1915]" italics="true" pageId="7" pageNumber="158">O. acutangulum</emphasis>
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</taxonomicName>
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||
<emphasis id="B950D26A5422CC58FC47F8BBFBD7F8F9" bold="true" box="[982,1039,1891,1915]" pageId="7" pageNumber="158">
|
||
<taxonomicNameLabel id="A2636F115422CC58FC47F8BBFBD7F8F9" box="[982,1039,1891,1915]" pageId="7" pageNumber="158" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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||
(circles),
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||
<taxonomicName id="4C2475FB5422CC58FBE4F8BCFAC2F8F9" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1141,1306,1891,1915]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="amplissimum" status="sp. n.">
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||
<emphasis id="B950D26A5422CC58FBE4F8BCFAC2F8F9" box="[1141,1306,1891,1915]" italics="true" pageId="7" pageNumber="158">O. amplissimum</emphasis>
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||
</taxonomicName>
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||
<emphasis id="B950D26A5422CC58FAB0F8BBFA82F8F9" bold="true" box="[1313,1370,1891,1915]" pageId="7" pageNumber="158">
|
||
<taxonomicNameLabel id="A2636F115422CC58FAB0F8BBFA82F8F9" box="[1313,1370,1891,1915]" pageId="7" pageNumber="158" rank="species">sp. n.</taxonomicNameLabel>
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||
</emphasis>
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||
(star),
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||
<taxonomicName id="4C2475FB5422CC58FF06F850FEDBF81D" authorityName="Cameron" authorityYear="1924" box="[151,259,1927,1951]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="beesoni">
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||
<emphasis id="B950D26A5422CC58FF06F850FEDBF81D" box="[151,259,1927,1951]" italics="true" pageId="7" pageNumber="158">O. beesoni</emphasis>
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||
</taxonomicName>
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||
(squares),
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||
<taxonomicName id="4C2475FB5422CC58FEE2F850FE28F81D" authorityName="Cameron" authorityYear="1924" box="[371,496,1928,1951]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="cariosum">
|
||
<emphasis id="B950D26A5422CC58FEE2F850FE28F81D" box="[371,496,1928,1951]" italics="true" pageId="7" pageNumber="158">O. cariosum</emphasis>
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||
</taxonomicName>
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||
(diamonds),
|
||
<taxonomicName id="4C2475FB5422CC58FDE6F850FCF6F81D" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[631,814,1927,1951]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="flavotestaceum" status="sp. n.">
|
||
<emphasis id="B950D26A5422CC58FDE6F850FCF6F81D" box="[631,814,1927,1951]" italics="true" pageId="7" pageNumber="158">O. flavotestaceum</emphasis>
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||
</taxonomicName>
|
||
<emphasis id="B950D26A5422CC58FCA5F85FFCB5F81D" bold="true" box="[820,877,1927,1951]" pageId="7" pageNumber="158">
|
||
<taxonomicNameLabel id="A2636F115422CC58FCA5F85FFCB5F81D" box="[820,877,1927,1951]" pageId="7" pageNumber="158" rank="species">sp. n.</taxonomicNameLabel>
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||
</emphasis>
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||
(triangles).
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||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9B0E78542DCC57FF56FF4FFA59FE2F" blockId="8.[151,1437,151,573]" pageId="8" pageNumber="159">
|
||
<typeStatus id="549FB0DA542DCC57FF56FF4FFEE0FF33" box="[199,312,151,177]" pageId="8" pageNumber="159" type="paratype">Paratypes</typeStatus>
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||
: 2 ♁♁ (
|
||
<specimenCount id="9D22C5F1542DCC57FE02FF4FFDF7FF33" box="[403,559,151,177]" count="1" pageId="8" pageNumber="159" type="generic">one specimen</specimenCount>
|
||
dissected),
|
||
<materialsCitation id="3B4C0425542DCC57FD27FF4FFBA2FF7B" collectingDate="2009-05-06" collectorName="N Banduk & J. Schmidt'" country="Nepal" elevation="2100" latitude="28.46861" location="Himal" longLatPrecision="20" longitude="83.58611" pageId="8" pageNumber="159" specimenCount="2" specimenCount-female="2" stateProvince="Dhawalagiri" typeStatus="paratype">
|
||
<specimenCount id="9D22C5F1542DCC57FD27FF4FFD3BFF33" box="[694,739,151,177]" count="1" pageId="8" pageNumber="159" type="female">1 ♀</specimenCount>
|
||
: ‘
|
||
<collectingCountry id="F3334EE8542DCC57FD65FF4FFC8CFF33" box="[756,852,151,177]" name="Nepal" pageId="8" pageNumber="159">NEPAL</collectingCountry>
|
||
,
|
||
<collectingRegion id="49E0C09A542DCC57FCF1FF4FFC00FF33" box="[864,984,151,177]" country="Nepal" name="Dhawalagiri" pageId="8" pageNumber="159">Dhaulagiri</collectingRegion>
|
||
<location id="8EFB58A3542DCC57FC71FF4FFBFEFF33" LSID="urn:lsid:plazi:treatment:038DBF6E5422CC55FF06FD0EFAE4FEC3:8EFB58A3542DCC57FC71FF4FFBFEFF33" box="[992,1062,151,177]" country="Nepal" latitude="28.46861" longLatPrecision="20" longitude="83.58611" name="Himal" pageId="8" pageNumber="159" stateProvince="Dhawalagiri">Himal</location>
|
||
| s-slope,
|
||
<collectorName id="26D16BAE542DCC57FB08FF4FFAD4FF33" box="[1177,1292,151,177]" pageId="8" pageNumber="159">N Banduk</collectorName>
|
||
vill. |
|
||
<quantity id="4CDCA39D542DCC57FAC7FF4FFF30FF57" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.3" metricValueMin="1.9" pageId="8" pageNumber="159" unit="m" value="2100.0" valueMax="2300.0" valueMin="1900.0">
|
||
<elevation id="0009E94B542DCC57FAC7FF4FFF30FF57" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.3" metricValueMin="1.9" pageId="8" pageNumber="159" unit="m" value="2100.0" valueMax="2300.0" valueMin="1900.0">1900– 2300m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate id="EE1068BF542DCC57FF60FF63FE5AFF57" box="[241,386,187,213]" degrees="28" direction="north" minutes="27" orientation="latitude" pageId="8" pageNumber="159" precision="15" seconds="22" value="28.456112">28°27’22’’N</geoCoordinate>
|
||
, |
|
||
<geoCoordinate id="EE1068BF542DCC57FE07FF62FDF8FF57" box="[406,544,186,213]" degrees="83" direction="east" minutes="35" orientation="longitude" pageId="8" pageNumber="159" precision="15" seconds="13" value="83.586945">83°35’13’’E</geoCoordinate>
|
||
to 28°28’07’’N | 83°35’10’’E,
|
||
<date id="FF9A28B8542DCC57FCE7FF63FC31FF57" box="[886,1001,187,213]" pageId="8" pageNumber="159" value="2009-05-06">
|
||
<collectingDate id="EFDED150542DCC57FCE7FF63FC31FF57" box="[886,1001,187,213]" pageId="8" pageNumber="159" value="2009-05-06">06.V.2009</collectingDate>
|
||
</date>
|
||
| leg.
|
||
<collectorName id="26D16BAE542DCC57FBB9FF63FB7CFF57" box="[1064,1188,187,213]" pageId="8" pageNumber="159">J. Schmidt’</collectorName>
|
||
<printed>, ‘collection | NATURKUNDE | MUSEUM ERFURT’ <yellow, printed> (1 ♁: cSh; 1 ♁,
|
||
<specimenCount id="9D22C5F1542DCC57FC64FF07FBFAFF7B" box="[1013,1058,223,249]" count="1" pageId="8" pageNumber="159" type="female">1 ♀</specimenCount>
|
||
: NME)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B4C0425542DCC57FB14FF07FE8AFEC3" collectingDate="2009-05-12" collectorName="J. Schmidt'" country="Nepal" elevation="2600" latitude="28.53861" location="Pathlekharka" longLatPrecision="20" longitude="83.49027" municipality="Mts." pageId="8" pageNumber="159" specimenCount="4" specimenCount-female="4" stateProvince="Dhawalagiri" typeStatus="paratype">
|
||
<specimenCount id="9D22C5F1542DCC57FB14FF07FB1FFF7B" box="[1157,1223,223,249]" count="2" pageId="8" pageNumber="159" type="female">2 ♀♀</specimenCount>
|
||
: ‘
|
||
<collectingCountry id="F3334EE8542DCC57FB49FF07FAE0FF7B" box="[1240,1336,223,249]" name="Nepal" pageId="8" pageNumber="159">NEPAL</collectingCountry>
|
||
, S slope,
|
||
<collectingRegion id="49E0C09A542DCC57FF06FEDBFED7FE9F" box="[151,271,259,285]" country="Nepal" name="Dhawalagiri" pageId="8" pageNumber="159">Dhaulagiri</collectingRegion>
|
||
|
|
||
<collectingMunicipality id="6BFF9402542DCC57FEB3FEDBFE8AFE9F" box="[290,338,259,285]" pageId="8" pageNumber="159">Mts.</collectingMunicipality>
|
||
above
|
||
<location id="8EFB58A3542DCC57FE0FFEDBFDE8FE9F" LSID="urn:lsid:plazi:treatment:038DBF6E5422CC55FF06FD0EFAE4FEC3:8EFB58A3542DCC57FE0FFEDBFDE8FE9F" box="[414,560,259,285]" country="Nepal" latitude="28.53861" longLatPrecision="20" longitude="83.49027" municipality="Mts." name="Pathlekharka" pageId="8" pageNumber="159" stateProvince="Dhawalagiri">Pathlekharka</location>
|
||
|
|
||
<quantity id="4CDCA39D542DCC57FDD3FEDAFD07FE9F" box="[578,735,258,285]" metricMagnitude="3" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="8" pageNumber="159" unit="m" value="2600.0" valueMax="2700.0" valueMin="2500.0">
|
||
<elevation id="0009E94B542DCC57FDD3FEDAFD07FE9F" box="[578,735,258,285]" metricMagnitude="3" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="8" pageNumber="159" unit="m" value="2600.0" valueMax="2700.0" valueMin="2500.0">2500–2700 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FF9A28B8542DCC57FD7BFEDBFC85FE9F" box="[746,861,259,285]" pageId="8" pageNumber="159" value="2009-05-12">
|
||
<collectingDate id="EFDED150542DCC57FD7BFEDBFC85FE9F" box="[746,861,259,285]" pageId="8" pageNumber="159" value="2009-05-12">12.V.2009</collectingDate>
|
||
</date>
|
||
|
|
||
<geoCoordinate id="EE1068BF542DCC57FCE1FEDBFC27FE9F" box="[880,1023,259,285]" degrees="28" direction="north" minutes="32" orientation="latitude" pageId="8" pageNumber="159" precision="15" seconds="19" value="28.53861">28°32’19’’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1068BF542DCC57FB9BFEDAFB4FFE9F" box="[1034,1175,258,285]" degrees="83" direction="east" minutes="29" orientation="longitude" pageId="8" pageNumber="159" precision="15" seconds="25" value="83.49027">83°29’25’’E</geoCoordinate>
|
||
, | leg.
|
||
<collectorName id="26D16BAE542DCC57FB4FFEDBFA83FE9F" box="[1246,1371,259,285]" pageId="8" pageNumber="159">J. Schmidt’</collectorName>
|
||
(
|
||
<specimenCount id="9D22C5F1542DCC57FAFCFEDBFA4EFE9F" box="[1389,1430,259,285]" count="1" pageId="8" pageNumber="159" type="female">1 ♀</specimenCount>
|
||
: cSh;
|
||
<specimenCount id="9D22C5F1542DCC57FF5EFEFFFF23FEC3" box="[207,251,295,321]" count="1" pageId="8" pageNumber="159" type="female">1 ♀</specimenCount>
|
||
: NME)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B4C0425542DCC57FECCFEFFFB8DFEE7" collectorName="E. Mangsingma" country="Nepal" elevation="1900" location="Smetana" municipality="Lobl" pageId="8" pageNumber="159" specimenCount="1" specimenCount-female="1" stateProvince="Kosi" typeStatus="paratype">
|
||
<specimenCount id="9D22C5F1542DCC57FECCFEFFFE5EFEC3" box="[349,390,295,321]" count="1" pageId="8" pageNumber="159" type="female">1 ♀</specimenCount>
|
||
(left antennomeres 3–11 and left hind leg missing): ‘E.
|
||
<collectingCountry id="F3334EE8542DCC57FC62FEFFFB97FEC3" box="[1011,1103,295,321]" name="Nepal" pageId="8" pageNumber="159">NEPAL</collectingCountry>
|
||
:
|
||
<collectingRegion id="49E0C09A542DCC57FBC8FEFFFB42FEC3" box="[1113,1178,295,321]" country="Nepal" name="Kosi" pageId="8" pageNumber="159">KOSI</collectingRegion>
|
||
|
|
||
<quantity id="4CDCA39D542DCC57FB3CFEFFFB3DFEC3" box="[1197,1253,295,321]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="159" unit="km" value="2.0">2 km</quantity>
|
||
<collectorName id="26D16BAE542DCC57FB7AFEFFFA44FEC3" box="[1259,1436,295,321]" pageId="8" pageNumber="159">E. Mangsingma</collectorName>
|
||
|
|
||
<quantity id="4CDCA39D542DCC57FF32FE93FF21FEE7" box="[163,249,331,357]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="8" pageNumber="159" unit="m" value="1900.0">
|
||
<elevation id="0009E94B542DCC57FF32FE93FF21FEE7" box="[163,249,331,357]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="8" pageNumber="159" unit="m" value="1900.0">1900 m</elevation>
|
||
</quantity>
|
||
, 19.IV.[19]84 | [I.]
|
||
<collectingMunicipality id="6BFF9402542DCC57FE5CFE93FDDAFEE7" box="[461,514,331,357]" pageId="8" pageNumber="159">Löbl</collectingMunicipality>
|
||
- [A.]
|
||
<location id="8EFB58A3542DCC57FDDAFE93FD73FEE7" LSID="urn:lsid:plazi:treatment:038DBF6E5422CC55FF06FD0EFAE4FEC3:8EFB58A3542DCC57FDDAFE93FD73FEE7" box="[587,683,331,357]" country="Nepal" municipality="Lobl" name="Smetana" pageId="8" pageNumber="159" stateProvince="Kosi">Smetana</location>
|
||
[sifted moss and leaf litter]’ (MHNG)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B4C0425542DCC57FBCEFE93FAACFE0B" country="Nepal" county="Val. Induwa Kola" elevation="2050" location="Smetana" municipality="Lobl" pageId="8" pageNumber="159" specimenCount="1" specimenCount-female="1" stateProvince="Kosi" typeStatus="paratype">
|
||
<specimenCount id="9D22C5F1542DCC57FBCEFE93FB50FEE7" box="[1119,1160,331,357]" count="1" pageId="8" pageNumber="159" type="female">1 ♀</specimenCount>
|
||
(dissected): ‘E.
|
||
<collectingCountry id="F3334EE8542DCC57FAACFE93FA41FEE7" box="[1341,1433,331,357]" name="Nepal" pageId="8" pageNumber="159">NEPAL</collectingCountry>
|
||
:
|
||
<collectingRegion id="49E0C09A542DCC57FF06FEB7FF00FE0B" box="[151,216,367,393]" country="Nepal" name="Kosi" pageId="8" pageNumber="159">KOSI</collectingRegion>
|
||
|
|
||
<collectingCounty id="62FA76F4542DCC57FF7BFEB7FE74FE0B" box="[234,428,367,393]" pageId="8" pageNumber="159">Val. Induwa Kola</collectingCounty>
|
||
|
|
||
<quantity id="4CDCA39D542DCC57FE2EFEB6FDCDFE0B" box="[447,533,366,393]" metricMagnitude="3" metricUnit="m" metricValue="2.05" pageId="8" pageNumber="159" unit="m" value="2050.0">
|
||
<elevation id="0009E94B542DCC57FE2EFEB6FDCDFE0B" box="[447,533,366,393]" metricMagnitude="3" metricUnit="m" metricValue="2.05" pageId="8" pageNumber="159" unit="m" value="2050.0">2050 m</elevation>
|
||
</quantity>
|
||
, 17.IV.[19]84 | [I.]
|
||
<collectingMunicipality id="6BFF9402542DCC57FD78FEB7FCC6FE0B" box="[745,798,367,393]" pageId="8" pageNumber="159">Löbl</collectingMunicipality>
|
||
- [A.]
|
||
<location id="8EFB58A3542DCC57FCF9FEB7FC10FE0B" LSID="urn:lsid:plazi:treatment:038DBF6E5422CC55FF06FD0EFAE4FEC3:8EFB58A3542DCC57FCF9FEB7FC10FE0B" box="[872,968,367,393]" country="Nepal" county="Val. Induwa Kola" municipality="Lobl" name="Smetana" pageId="8" pageNumber="159" stateProvince="Kosi">Smetana</location>
|
||
[bank of a river]’ <printed> (MHNG).
|
||
</materialsCitation>
|
||
<location id="8EFB58A3542DCC57FAE9FEB7FA44FE0B" LSID="urn:lsid:plazi:treatment:038DBF6E5422CC55FF06FD0EFAE4FEC3:8EFB58A3542DCC57FAE9FEB7FA44FE0B" box="[1400,1436,367,393]" country="Nepal" county="Val. Induwa Kola" municipality="Lobl" name="All" pageId="8" pageNumber="159" stateProvince="Kosi">All</location>
|
||
<typeStatus id="549FB0DA542DCC57FF06FE4BFEDAFE2F" box="[151,258,403,429]" pageId="8" pageNumber="159" type="paratype">paratypes</typeStatus>
|
||
with additional red printed label: ‘
|
||
<typeStatus id="549FB0DA542DCC57FD12FE4BFCCBFE2F" box="[643,787,403,429]" pageId="8" pageNumber="159" type="paratype">PARATYPE</typeStatus>
|
||
|
|
||
<taxonomicName id="4C2475FB542DCC57FCB6FE4BFC57FE2F" authorityName="Gravenhorst" authorityYear="1802" box="[807,911,403,429]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B950D26A542DCC57FCB6FE4BFC57FE2F" box="[807,911,403,429]" italics="true" pageId="8" pageNumber="159">Omalium</emphasis>
|
||
</taxonomicName>
|
||
|
|
||
<taxonomicName id="4C2475FB542DCC57FC32FE4BFBCDFE2E" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[931,1045,403,428]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="amicorum" status="sp. nov.">
|
||
<emphasis id="B950D26A542DCC57FC32FE4BFBCDFE2E" box="[931,1045,403,428]" italics="true" pageId="8" pageNumber="159">amicorum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B950D26A542DCC57FB8DFE4BFB83FE2F" bold="true" box="[1052,1115,403,429]" pageId="8" pageNumber="159">
|
||
<taxonomicNameLabel id="A2636F11542DCC57FB8DFE4BFB83FE2F" box="[1052,1115,403,429]" pageId="8" pageNumber="159" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
| Shavrin A.V. des. 2023’.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33E5DF3542DCC56FF56FE6EFC9EF8C2" lastPageId="9" lastPageNumber="160" pageId="8" pageNumber="159" type="description">
|
||
<paragraph id="8B9B0E78542DCC57FF56FE6EFE12FDBF" blockId="8.[151,1437,151,573]" pageId="8" pageNumber="159">
|
||
<emphasis id="B950D26A542DCC57FF56FE6EFE81FE52" bold="true" box="[199,345,438,464]" pageId="8" pageNumber="159">Description.</emphasis>
|
||
Measurements (n=7): HW: 0.59–0.62; HL: 0.42–0.47; OL: 0.17–0.20; TL: 0.07–0.09; AL (
|
||
<typeStatus id="549FB0DA542DCC57FF0EFE03FED0FE77" box="[159,264,475,501]" pageId="8" pageNumber="159" type="holotype">holotype</typeStatus>
|
||
): 1.37; PL: 0.51–0.57; PWmax: 0.74–0.75; PWmin: 0.63–0.70; ESL: 0.91–1.07; EW: 1.11–1.17; MTbL (
|
||
<typeStatus id="549FB0DA542DCC57FF0EFE27FED0FD9B" box="[159,264,511,537]" pageId="8" pageNumber="159" type="holotype">holotype</typeStatus>
|
||
): 0.60; MTrL (
|
||
<typeStatus id="549FB0DA542DCC57FE21FE27FDC1FD9B" box="[432,537,511,537]" pageId="8" pageNumber="159" type="holotype">holotype</typeStatus>
|
||
): 0.26 (MTrL 1–4: 0.11; MTrL 5: 0.15); AW: 0.91–1.10; AedL: 0.72–0.80; BL: 3.15–3.50 (
|
||
<typeStatus id="549FB0DA542DCC57FE87FDFBFEA6FDBF" box="[278,382,547,573]" pageId="8" pageNumber="159" type="holotype">holotype</typeStatus>
|
||
: 3.25).
|
||
</paragraph>
|
||
<caption id="DF5B5EF0542DCC57FF06F882FD47F814" ID-DOI="http://doi.org/10.5281/zenodo.10146180" ID-Zenodo-Dep="10146180" httpUri="https://zenodo.org/record/10146180/files/figure.png" pageId="8" pageNumber="159" startId="8.[151,264,1882,1906]" targetBox="[151,1433,600,1855]" targetPageId="8" targetType="figure">
|
||
<paragraph id="8B9B0E78542DCC57FF06F882FD47F814" blockId="8.[151,1436,1882,1942]" pageId="8" pageNumber="159">
|
||
<emphasis id="B950D26A542DCC57FF06F882FE8EF8F0" bold="true" box="[151,342,1882,1906]" pageId="8" pageNumber="159">FIGURES 20–22.</emphasis>
|
||
Aedeagus of
|
||
<taxonomicName id="4C2475FB542DCC57FE79F882FD6DF8F0" authorityName="Shavrin" authorityYear="2023" box="[488,693,1882,1906]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="amicorum">
|
||
<emphasis id="B950D26A542DCC57FE79F882FD6DF8F0" box="[488,693,1882,1906]" italics="true" pageId="8" pageNumber="159">Omalium amicorum</emphasis>
|
||
</taxonomicName>
|
||
(holotype): 20—aedeagus, parameral view, 21—aedeagus, lateral view, 22—median lobe, lateral view). Scale bar: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9B0E78542CCC56FF56FF4FFB16FE70" blockId="9.[151,1437,151,2036]" pageId="9" pageNumber="160">
|
||
Habitus as in
|
||
<figureCitation id="131F12FD542CCC56FEF1FF4FFE7BFF33" box="[352,419,151,177]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1260,1284]" captionTargetBox="[168,1420,194,1223]" captionTargetId="figure-22@4.[155,1432,181,1236]" captionTargetPageId="4" captionText="FIGURES 1–4. Habitus (Figs 1, 3) and dorsolateral view of head and pronotum (Figs 2, 4) of Omalium: 1–2—O. acutangulum (paratype), 3–4—O. amicorum (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10146172" httpUri="https://zenodo.org/record/10146172/files/figure.png" pageId="9" pageNumber="160">Fig. 3</figureCitation>
|
||
; head and pronotum dorsolaterally as in
|
||
<figureCitation id="131F12FD542CCC56FCFBFF4FFC76FF33" box="[874,942,151,177]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1260,1284]" captionTargetBox="[168,1420,194,1223]" captionTargetId="figure-22@4.[155,1432,181,1236]" captionTargetPageId="4" captionText="FIGURES 1–4. Habitus (Figs 1, 3) and dorsolateral view of head and pronotum (Figs 2, 4) of Omalium: 1–2—O. acutangulum (paratype), 3–4—O. amicorum (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10146172" httpUri="https://zenodo.org/record/10146172/files/figure.png" pageId="9" pageNumber="160">Fig. 4</figureCitation>
|
||
. Body reddish-brown, with distinctly darker head, sometimes with paler lateral portions of pronotum and abdomen; antennomeres 6–11 brown; mouthparts, antennomeres 1–5 and legs yellow-brown or slightly darker. Punctation of head moderately large and dense, finer and denser on apical portion of clypeus, markedly denser in middle between anteocellar foveae and slightly coarser on infraorbital portions (
|
||
<typeStatus id="549FB0DA542CCC56FE3AFEFEFDD7FEC2" box="[427,527,294,320]" pageId="9" pageNumber="160" type="holotype">holotype</typeStatus>
|
||
with sparser median punctation); nuchal constriction with moderately dense and large punctation, sometimes finer in middle; punctation of pronotum dense, slightly larger and deeper than that in middle part of head, sparser and finer in medioapical and mediobasal portions; punctation of elytra dense, about as that on pronotum, denser around scutellum, finer and sparser along suture. Anterior portion of clypeus with very fine and sometimes indistinct transverse microreticulation; nuchal constriction with dense isodiametric meshes; pronotum and elytra without microsculpture; scutellum with fine isodiametric microreticulation.
|
||
</paragraph>
|
||
<paragraph id="8B9B0E78542CCC56FF56FE24FA57FC65" blockId="9.[151,1437,151,2036]" pageId="9" pageNumber="160">
|
||
Head 1.3–1.4 times as broad as long, with broad clypeus, distinctly convex supra-antennal elevations and strongly elevated middle portion between eyes and infraorbital portions, with very deep and wide anteriomedian depressions, sometimes significantly stretching posteriad and reaching apical part of anteocellar foveae; portions between infraorbital parts and middle deeply depressed. Surface without or with shallow elevations between anteocellar foveae, with slightly elevated narrow diagonal wrinkles on basal portions of clypeus, infraorbital portions without or with irregular longitudinal elevations between punctures. Anteocellar foveae very deep and wide, diagonally stretching apicad toward level of anterior third of eyes. Temples more than twice shorter than longitudinal length of eyes, strongly narrowed posteriad toward distinctly protruded basad subacute angles. Apical half of nuchal constriction significantly depressed, broader in middle; medioapical part of nuchal constriction strongly elevated, rounded, distinctly protruded antreriad (
|
||
<figureCitation id="131F12FD542CCC56FDC2FCE6FD40FCDA" box="[595,664,830,856]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1260,1284]" captionTargetBox="[168,1420,194,1223]" captionTargetId="figure-22@4.[155,1432,181,1236]" captionTargetPageId="4" captionText="FIGURES 1–4. Habitus (Figs 1, 3) and dorsolateral view of head and pronotum (Figs 2, 4) of Omalium: 1–2—O. acutangulum (paratype), 3–4—O. amicorum (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10146172" httpUri="https://zenodo.org/record/10146172/files/figure.png" pageId="9" pageNumber="160">Fig. 4</figureCitation>
|
||
). Ocelli very large, located close to basal impression, sometimes with concave strong depression between basal margins; distance between ocelli 1.6 times to twice as long as distance between ocellus and posterior margin of eye. Antenna with antennomeres 9–10 slightly transverse; antennomere 4 small, slightly longer than broad, shorter than 3, 5 slightly longer than 4, 6 slightly longer and indistinctly broader than 5, 7–8 slightly broader than 6, 8–9 distinctly broader than 7, apical antennomere 1.2–1.3 times as long as 10.
|
||
</paragraph>
|
||
<paragraph id="8B9B0E78542CCC56FF56FC29FE31FA87" blockId="9.[151,1437,151,2036]" pageId="9" pageNumber="160">
|
||
Pronotum distinctly convex, 1.3–1.4 times as broad as long, 1.2–1.5 times as broad as head, from widest apical third slightly narrowed anteriad and distinctly narrowed posteriad, with widely concave laterobasal margins, with apical angles indistinctly protruded anteriad. Anterior margin rounded, sometimes significantly protruded apicad in some specimens (
|
||
<figureCitation id="131F12FD542CCC56FECDFB84FE47FBF4" box="[348,415,1116,1142]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1260,1284]" captionTargetBox="[168,1420,194,1223]" captionTargetId="figure-22@4.[155,1432,181,1236]" captionTargetPageId="4" captionText="FIGURES 1–4. Habitus (Figs 1, 3) and dorsolateral view of head and pronotum (Figs 2, 4) of Omalium: 1–2—O. acutangulum (paratype), 3–4—O. amicorum (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10146172" httpUri="https://zenodo.org/record/10146172/files/figure.png" pageId="9" pageNumber="160">Fig. 4</figureCitation>
|
||
), about as long as straight or slightly rounded basal margin. Surface of disc with two strong, wide and deep longitudinal depression, significantly broadened posteriad, and with very deep oval depression in medioapical portion; lateral portions widely impressed, distinctly deeper laterobasally; surface between all pronotal depressions strongly elevated. Middle surface of disc with irregular and sometimes indistinct diagonal and transverse elevations between punctures.
|
||
</paragraph>
|
||
<paragraph id="8B9B0E78542CCC56FF56FAD6FF28FAF2" blockId="9.[151,1437,151,2036]" pageId="9" pageNumber="160">Elytra slightly broader than long, 1.7–1.8 times as long as pronotum, distinctly broadened posteriad. Middle surface of each elytron with irregular diagonal and transverse elevations between punctures, coarser in parascutellar portion.</paragraph>
|
||
<paragraph id="8B9B0E78542CCC56FF56FAA1FDFFFA11" blockId="9.[151,1437,151,2036]" box="[199,551,1401,1428]" pageId="9" pageNumber="160">Abdomen narrower than elytra.</paragraph>
|
||
<paragraph id="8B9B0E78542CCC56FF56FA45FD4CF957" blockId="9.[151,1437,151,2036]" pageId="9" pageNumber="160">
|
||
Male. Antennomeres 8–10 slightly transverse. Posterior margin of abdominal tergite VIII straight or slightly rounded. Posterior margin of abdominal sternite VIII slightly concave. Aedeagus with wide basal portion, gradually narrowed toward middle, with long and moderately narrow median lobe, from widest middle gradually narrowed apically toward small rounded apex; mediolateral portions with wide accessory plates widely broadened apically; parameres moderately wide, distinctly shorter than apex of median lobe, from widest middle distinctly narrowed apically toward small rounded apices, with three long apical and one preapical setae; internal sac wide and moderately long, without sclerotized structures in basal portion (
|
||
<figureCitation id="131F12FD542CCC56FD61F9ACFC9DF90C" box="[752,837,1652,1678]" captionStart="FIGURES 20–22" captionStartId="8.[151,264,1882,1906]" captionTargetBox="[151,1433,600,1855]" captionTargetId="figure-219@8.[151,1436,597,1858]" captionTargetPageId="8" captionText="FIGURES 20–22. Aedeagus of Omalium amicorum (holotype): 20—aedeagus, parameral view, 21—aedeagus, lateral view, 22—median lobe, lateral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146180" httpUri="https://zenodo.org/record/10146180/files/figure.png" pageId="9" pageNumber="160">Fig. 20</figureCitation>
|
||
). Lateral aspect of the aedeagus as in
|
||
<figureCitation id="131F12FD542CCC56FB69F9ACFA94F90C" box="[1272,1356,1652,1678]" captionStart="FIGURES 20–22" captionStartId="8.[151,264,1882,1906]" captionTargetBox="[151,1433,600,1855]" captionTargetId="figure-219@8.[151,1436,597,1858]" captionTargetPageId="8" captionText="FIGURES 20–22. Aedeagus of Omalium amicorum (holotype): 20—aedeagus, parameral view, 21—aedeagus, lateral view, 22—median lobe, lateral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146180" httpUri="https://zenodo.org/record/10146180/files/figure.png" pageId="9" pageNumber="160">Fig. 21</figureCitation>
|
||
; apical portion of median lobe moderately narrow, broadened in preapical portion, with curved acute apex and two obtuse teeth on inner side of preapical part (
|
||
<figureCitation id="131F12FD542CCC56FDA2F963FD5FF957" box="[563,647,1723,1749]" captionStart="FIGURES 20–22" captionStartId="8.[151,264,1882,1906]" captionTargetBox="[151,1433,600,1855]" captionTargetId="figure-219@8.[151,1436,597,1858]" captionTargetPageId="8" captionText="FIGURES 20–22. Aedeagus of Omalium amicorum (holotype): 20—aedeagus, parameral view, 21—aedeagus, lateral view, 22—median lobe, lateral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146180" httpUri="https://zenodo.org/record/10146180/files/figure.png" pageId="9" pageNumber="160">Fig. 22</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9B0E78542CCC56FF56F907FC9EF8C2" blockId="9.[151,1437,151,2036]" pageId="9" pageNumber="160">
|
||
Female. Antennomeres 8–10 less transverse. Posterior margin of abdominal tergite VIII straight. Posterior margin of abdominal sternite VIII rounded.Accessory sclerite short, with very wide basal portion, strongly narrowed toward subacute apex (
|
||
<figureCitation id="131F12FD542CCC56FE08F8FEFE34F8C2" box="[409,492,1830,1856]" captionStart="FIGURES 8–18" captionStartId="6.[151,264,1896,1920]" captionTargetBox="[154,1433,564,1870]" captionTargetId="figure-184@6.[151,1436,561,1873]" captionTargetPageId="6" captionText="FIGURES 8–18. Female accessory sclerite (Figs 8, 10–16, 18) and spermatheca (Figs 9, 17) of Omalium: 8–9—O. acutangulum, 10—O. amicorum, 11—O. amplissimum, 12—O. bilobum, 13—O. alticola, 14—O.beesoni, 15—O. altivagans, 16–17—O. marginatum, 18—O. scapulare. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146176" httpUri="https://zenodo.org/record/10146176/files/figure.png" pageId="9" pageNumber="160">Fig. 10</figureCitation>
|
||
). Spermatheca not recognized.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33E5DF3542CCC56FF56F892FEC2F876" pageId="9" pageNumber="160" type="diagnosis">
|
||
<paragraph id="8B9B0E78542CCC56FF56F892FEC2F876" blockId="9.[151,1437,151,2036]" pageId="9" pageNumber="160">
|
||
<emphasis id="B950D26A542CCC56FF56F892FE6DF8E6" bold="true" box="[199,437,1866,1892]" pageId="9" pageNumber="160">Comparative notes.</emphasis>
|
||
<taxonomicName id="4C2475FB542CCC56FE50F892FD7FF8E6" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[449,679,1866,1892]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="amicorum" status="sp. nov.">
|
||
<emphasis id="B950D26A542CCC56FE50F892FD7FF8E6" box="[449,679,1866,1892]" italics="true" pageId="9" pageNumber="160">Omalium amicorum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B950D26A542CCC56FD23F892FD2EF8E6" bold="true" box="[690,758,1866,1892]" pageId="9" pageNumber="160">
|
||
<taxonomicNameLabel id="A2636F11542CCC56FD23F892FD2EF8E6" box="[690,758,1866,1892]" pageId="9" pageNumber="160" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from
|
||
<taxonomicName id="4C2475FB542CCC56FBAFF893FB2FF8E6" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1086,1271,1866,1893]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="acutangulum" status="sp. n.">
|
||
<emphasis id="B950D26A542CCC56FBAFF893FB8AF8E7" box="[1086,1106,1867,1893]" italics="true" pageId="9" pageNumber="160">O</emphasis>
|
||
.
|
||
<emphasis id="B950D26A542CCC56FBF4F892FB2FF8E6" box="[1125,1271,1866,1892]" italics="true" pageId="9" pageNumber="160">acutangulum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B950D26A542CCC56FA92F892FA9FF8E6" bold="true" box="[1283,1351,1866,1892]" pageId="9" pageNumber="160">
|
||
<taxonomicNameLabel id="A2636F11542CCC56FA92F892FA9FF8E6" box="[1283,1351,1866,1892]" pageId="9" pageNumber="160" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the temples strongly narrowed posteriad, finer sculpture of middle and infraorbital portions of head, by the presense of an unusual deep impression in medioapical part and strongly elevated mediobasal part of nuchal constriction, narrower preapical part of the pronotum, and details of external and internal morphology of the aedeagus (see the key above).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33E5DF3542FCC55FF56FF4EFA49FF33" box="[199,1425,150,177]" pageId="10" pageNumber="161" type="distribution">
|
||
<paragraph id="8B9B0E78542FCC55FF56FF4EFA49FF33" blockId="10.[151,1436,150,321]" box="[199,1425,150,177]" pageId="10" pageNumber="161">
|
||
<emphasis id="B950D26A542FCC55FF56FF4EFE86FF33" bold="true" box="[199,350,150,177]" pageId="10" pageNumber="161">Distribution.</emphasis>
|
||
<taxonomicName id="4C2475FB542FCC55FEF5FF4FFD98FF32" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[356,576,151,177]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="amicorum" status="sp. nov.">
|
||
<emphasis id="B950D26A542FCC55FEF5FF4FFD98FF32" box="[356,576,151,177]" italics="true" pageId="10" pageNumber="161">Omalium amicorum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B950D26A542FCC55FDD6FF4FFD5DFF33" bold="true" box="[583,645,151,177]" pageId="10" pageNumber="161">
|
||
<taxonomicNameLabel id="A2636F11542FCC55FDD6FF4FFD5DFF33" box="[583,645,151,177]" pageId="10" pageNumber="161" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is known from several localities in central and eastern
|
||
<collectingCountry id="F3334EE8542FCC55FB73FF4FFAFCFF33" box="[1250,1316,151,177]" name="Nepal" pageId="10" pageNumber="161">Nepal</collectingCountry>
|
||
(
|
||
<figureCitation id="131F12FD542FCC55FAA2FF4FFA5CFF33" box="[1331,1412,151,177]" captionStart="FIGURE 19" captionStartId="7.[151,250,1891,1915]" captionTargetBox="[151,1435,957,1868]" captionTargetId="figure-299@7.[151,1435,957,1868]" captionTargetPageId="7" captionText="FIGURE 19. Distribution of Omalium in the Himalayan region: O. acutangulum sp. n. (circles), O. amplissimum sp. n. (star), O. beesoni (squares), O. cariosum (diamonds), O. flavotestaceum sp. n. (triangles)." figureDoi="http://doi.org/10.5281/zenodo.10146178" httpUri="https://zenodo.org/record/10146178/files/figure.png" pageId="10" pageNumber="161">Fig. 19</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33E5DF3542FCC55FF56FF63FDA4FF7B" pageId="10" pageNumber="161" type="etymology">
|
||
<paragraph id="8B9B0E78542FCC55FF56FF63FDA4FF7B" blockId="10.[151,1436,150,321]" pageId="10" pageNumber="161">
|
||
<emphasis id="B950D26A542FCC55FF56FF63FE96FF57" bold="true" box="[199,334,187,213]" pageId="10" pageNumber="161">Etymology.</emphasis>
|
||
The specific epithet is the Latin
|
||
<taxonomicName id="4C2475FB542FCC55FD27FF63FCF0FF56" authorityName="Shavrin" authorityYear="2023" box="[694,808,187,212]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="amicorum">
|
||
<emphasis id="B950D26A542FCC55FD27FF63FCF0FF56" box="[694,808,187,212]" italics="true" pageId="10" pageNumber="161">amicorum</emphasis>
|
||
</taxonomicName>
|
||
, -
|
||
<emphasis id="B950D26A542FCC55FCAFFF63FC94FF56" box="[830,844,187,212]" italics="true" pageId="10" pageNumber="161">a</emphasis>
|
||
, -
|
||
<emphasis id="B950D26A542FCC55FCF3FF63FCA3FF56" box="[866,891,187,212]" italics="true" pageId="10" pageNumber="161">us</emphasis>
|
||
(friends). The species is dedicated to A. Smetana and I. Löbl, collectors of several
|
||
<typeStatus id="549FB0DA542FCC55FD98FF07FDA0FF7B" box="[521,632,223,249]" pageId="10" pageNumber="161" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33E5DF3542FCC55FF56FEDAFAE4FEC3" pageId="10" pageNumber="161" type="biology_ecology">
|
||
<paragraph id="8B9B0E78542FCC55FF56FEDAFAE4FEC3" blockId="10.[151,1436,150,321]" pageId="10" pageNumber="161">
|
||
<emphasis id="B950D26A542FCC55FF56FEDAFE93FE9E" bold="true" box="[199,331,258,284]" pageId="10" pageNumber="161">Bionomics.</emphasis>
|
||
Specimens were collected at elevations from
|
||
<quantity id="4CDCA39D542FCC55FCD2FEDBFC37FE9F" box="[835,1007,259,285]" metricMagnitude="3" metricUnit="m" metricValue="2.3" metricValueMax="2.7" metricValueMin="1.9" pageId="10" pageNumber="161" unit="m" value="2300.0" valueMax="2700.0" valueMin="1900.0">1900 to 2700 m</quantity>
|
||
a.s.l. The specimen from Mangsingma was collected by sifting moss and leaf litter. The specimen from Induwa Kola was collected near the river.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |