treatments-xml/data/03/8D/BF/038DBF6E540CCC74FF06F8D9FA50F963.xml
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<document id="F27AA1B1B863A69667D78AE8D505BDCB" ID-DOI="10.11646/zootaxa.5369.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10146170" ID-ZooBank="42563452-CF2B-47A6-B0B0-485AC4F320BD" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1700185584045" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2023" docId="038DBF6E540CCC74FF06F8D9FA50F963" docLanguage="en" docName="zootaxa.5369.2.1.pdf" docOrigin="Zootaxa 5369 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5369.2.1/52237" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Omalium flavotestaceum Shavrin 2023, sp. n." docType="treatment" docVersion="1" lastPageNumber="194" masterDocId="FFB4C7165425CC5FFF91FFD8FFD8FF82" masterDocTitle="A revision of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae) of the Himalayan region" masterLastPageNumber="206" masterPageNumber="151" pageNumber="192" updateTime="1700578467175" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:title id="E053107DAD82208FF29F0151DC12E639">A revision of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae) of the Himalayan region</mods:title>
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<treatment id="038DBF6E540CCC74FF06F8D9FA50F963" LSID="urn:lsid:plazi:treatment:038DBF6E540CCC74FF06F8D9FA50F963" httpUri="http://treatment.plazi.org/id/038DBF6E540CCC74FF06F8D9FA50F963" lastPageId="43" lastPageNumber="194" pageId="41" pageNumber="192">
<subSubSection id="C33E5DF3540CCC76FF06F8D9FD5CF89E" box="[151,644,1793,1820]" pageId="41" pageNumber="192" type="nomenclature">
<paragraph id="8B9B0E78540CCC76FF06F8D9FD5CF89E" blockId="41.[151,644,1793,1856]" box="[151,644,1793,1820]" pageId="41" pageNumber="192">
<heading id="D0D3B914540CCC76FF06F8D9FD5CF89E" bold="true" box="[151,644,1793,1820]" fontSize="11" level="1" pageId="41" pageNumber="192" reason="1">
<emphasis id="B950D26A540CCC76FF06F8D9FD5CF89E" bold="true" box="[151,644,1793,1820]" pageId="41" pageNumber="192">
3.2.5.2.3.
<taxonomicName id="4C2475FB540CCC76FE9CF8D9FDE3F89E" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[269,571,1793,1820]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="192" phylum="Arthropoda" rank="species" species="flavotestaceum" status="sp. nov.">
<emphasis id="B950D26A540CCC76FE9CF8D9FDE3F89E" bold="true" box="[269,571,1793,1820]" italics="true" pageId="41" pageNumber="192">Omalium flavotestaceum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2636F11540CCC76FDD3F8DAFD5CF89E" box="[578,644,1794,1820]" pageId="41" pageNumber="192" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33E5DF3540CCC76FF06F8FEFE17F8C2" box="[151,463,1830,1856]" pageId="41" pageNumber="192" type="description">
<paragraph id="8B9B0E78540CCC76FF06F8FEFE17F8C2" blockId="41.[151,644,1793,1856]" box="[151,463,1830,1856]" pageId="41" pageNumber="192">
(
<figureCitation id="131F12FD540CCC76FF0EF8FEFF2EF8C2" box="[159,246,1830,1856]" captionStart="FIGURE 19" captionStartId="7.[151,250,1891,1915]" captionTargetBox="[151,1435,957,1868]" captionTargetId="figure-299@7.[151,1435,957,1868]" captionTargetPageId="7" captionText="FIGURE 19. Distribution of Omalium in the Himalayan region: O. acutangulum sp. n. (circles), O. amplissimum sp. n. (star), O. beesoni (squares), O. cariosum (diamonds), O. flavotestaceum sp. n. (triangles)." figureDoi="http://doi.org/10.5281/zenodo.10146178" httpUri="https://zenodo.org/record/10146178/files/figure.png" pageId="41" pageNumber="192">Figs 19</figureCitation>
,
<figureCitation id="131F12FD540CCC76FE93F8FEFEF8F8C2" box="[258,288,1830,1856]" captionStart="FIGURES 7072" captionStartId="39.[151,264,1085,1109]" captionTargetBox="[159,1419,187,1052]" captionTargetId="figure-16@39.[151,1436,181,1061]" captionTargetPageId="39" captionText="FIGURES 7072. Habitus of Omalium: 70—O. cariosum (lectotype), 71—O. flavotestaceum (paratype), 72—O. kumari (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10146220" httpUri="https://zenodo.org/record/10146220/files/figure.png" pageId="41" pageNumber="192">71</figureCitation>
,
<figureCitation id="131F12FD540CCC76FEBDF8FEFEADF8C2" box="[300,373,1830,1856]" captionStart="FIGURES 7683" captionStartId="41.[151,264,1651,1675]" captionTargetBox="[154,1432,816,1625]" captionTargetId="figure-275@41.[151,1435,813,1628]" captionTargetPageId="41" captionText="FIGURES 7683. Female accessory sclerite (Figs 7677, 79, 8182) and spermatheca (Figs 78, 80, 83) of Omalium: 76— O.cariosum, 7778—O. flavotestaceum, 7980—O. kumari, 81—O. latocavum, 8283—O. secretum. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146224" httpUri="https://zenodo.org/record/10146224/files/figure.png" pageId="41" pageNumber="192">7778</figureCitation>
,
<figureCitation id="131F12FD540CCC76FE11F8FEFE1FF8C2" box="[384,455,1830,1856]" captionStart="FIGURES 8486" captionStartId="42.[151,264,1700,1724]" captionTargetBox="[154,1432,456,1674]" captionTargetId="figure-135@42.[151,1435,453,1677]" captionTargetPageId="42" captionText="FIGURES 8486. Aedeagus of Omalium flavotestaceum (paratype): 84—aedeagus, parameral view, 85—aedeagus, lateral view, 86—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146226" httpUri="https://zenodo.org/record/10146226/files/figure.png" pageId="41" pageNumber="192">8486</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33E5DF3540CCC75FF06F8B6FBCCFE9F" lastPageId="42" lastPageNumber="193" pageId="41" pageNumber="192" type="materials_examined">
<paragraph id="8B9B0E78540CCC76FF06F8B6FDF0F852" blockId="41.[151,1437,1901,2038]" pageId="41" pageNumber="192">
<emphasis id="B950D26A540CCC76FF06F8B6FE9EF805" bold="true" box="[151,326,1901,1928]" pageId="41" pageNumber="192">Type material.</emphasis>
<materialsCitation id="3B4C0425540CCC76FEDEF8B6FD65F82E" collectorName="W. Bengal" country="India" elevation="3100" location="Darjeeling distr." pageId="41" pageNumber="192" specimenCount="1" typeStatus="holotype">
<typeStatus id="549FB0DA540CCC76FEDEF8B6FE6FF80A" box="[335,439,1902,1928]" pageId="41" pageNumber="192" type="holotype">Holotype</typeStatus>
♁:
<collectingCountry id="F3334EE8540CCC76FE78F8B6FDE4F80A" box="[489,572,1902,1928]" name="India" pageId="41" pageNumber="192">INDIA</collectingCountry>
<collectorName id="26D16BAE540CCC76FDD5F8B6FD62F80A" box="[580,698,1902,1928]" pageId="41" pageNumber="192">W. Bengal</collectorName>
|
<location id="8EFB58A3540CCC76FD40F8B6FC50F80A" LSID="urn:lsid:plazi:treatment:038DBF6E540CCC74FF06F8D9FA50F963:8EFB58A3540CCC76FD40F8B6FC50F80A" box="[721,904,1902,1928]" country="India" name="Darjeeling distr." pageId="41" pageNumber="192">Darjeeling distr.</location>
| Tonglu
<quantity id="4CDCA39D540CCC76FC65F8B6FB9AF80A" box="[1012,1090,1902,1928]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="41" pageNumber="192" unit="m" value="3100.0">
<elevation id="0009E94B540CCC76FC65F8B6FB9AF80A" box="[1012,1090,1902,1928]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="41" pageNumber="192" unit="m" value="3100.0">3100m</elevation>
</quantity>
16.X. | [C.] Besuchet-[I.]Löbl [sifted litter under busges in a pasture] &lt;printed
</materialsCitation>
&gt;,
<typeStatus id="549FB0DA540CCC76FD4DF84AFCADF82E" box="[732,885,1938,1964]" pageId="41" pageNumber="192" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="4C2475FB540CCC76FC1BF84AFC2AF82E" authorityName="Gravenhorst" authorityYear="1802" box="[906,1010,1938,1964]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="192" phylum="Arthropoda" rank="genus">
<emphasis id="B950D26A540CCC76FC1BF84AFC2AF82E" box="[906,1010,1938,1964]" italics="true" pageId="41" pageNumber="192">Omalium</emphasis>
</taxonomicName>
|
<taxonomicName id="4C2475FB540CCC76FB96F84AFB77F82E" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1031,1199,1938,1964]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="192" phylum="Arthropoda" rank="species" species="flavotestaceum" status="sp. nov.">
<emphasis id="B950D26A540CCC76FB96F84AFB77F82E" box="[1031,1199,1938,1964]" italics="true" pageId="41" pageNumber="192">flavotestaceum</emphasis>
</taxonomicName>
<emphasis id="B950D26A540CCC76FB26F84AFB2FF82E" bold="true" box="[1207,1271,1938,1964]" pageId="41" pageNumber="192">
<taxonomicNameLabel id="A2636F11540CCC76FB26F84AFB2FF82E" box="[1207,1271,1938,1964]" pageId="41" pageNumber="192" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.V. des. 2023 &lt;red, printed&gt; (MHNG).
</paragraph>
<paragraph id="8B9B0E78540CCC75FF56F802FBCCFE9F" blockId="41.[151,1437,1901,2038]" lastBlockId="42.[151,1436,151,429]" lastPageId="42" lastPageNumber="193" pageId="41" pageNumber="192">
<materialsCitation id="3B4C0425540CCC76FF56F802FC0EF876" box="[199,982,2010,2038]" collectorName="W. Bengal" country="India" elevation="3100" location="Darjeeling distr." pageId="41" pageNumber="192" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="549FB0DA540CCC76FF56F802FEE0F876" box="[199,312,2010,2036]" pageId="41" pageNumber="192" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D22C5F1540CCC76FED5F802FEAAF876" box="[324,370,2010,2036]" count="1" pageId="41" pageNumber="192" type="female">1 ♀</specimenCount>
: same data as the holotype (MHNG); 1 ♁ (dissected)
</materialsCitation>
,
<materialsCitation id="3B4C0425540CCC75FC73F802FDF5FF57" collectingDate="1998-05-19" collectingDateMax="1998-05-23" collectingDateMin="1998-05-19" collectorName="Fabrizi &amp; Ahrens" country="India" elevation="2800" lastPageId="42" lastPageNumber="193" location="Tonglu" municipality="Darjeeling" pageId="41" pageNumber="192" specimenCount="2" specimenCount-female="2" stateProvince="West Bengal" typeStatus="paratype">
<specimenCount id="9D22C5F1540CCC76FC73F802FBD6F876" box="[994,1038,2010,2036]" count="1" pageId="41" pageNumber="192" type="female">1 ♀</specimenCount>
(dissected):
<collectingCountry id="F3334EE8540CCC76FB35F802FB2FF876" box="[1188,1271,2010,2036]" name="India" pageId="41" pageNumber="192">INDIA</collectingCountry>
,
<collectingRegion id="49E0C09A540CCC76FB6BF802FA56F876" box="[1274,1422,2010,2036]" country="India" name="West Bengal" pageId="41" pageNumber="192">West Bengal</collectingRegion>
|
<collectingMunicipality id="6BFF9402540FCC75FF06FF4FFEC8FF33" box="[151,272,151,177]" pageId="42" pageNumber="193">Darjeeling</collectingMunicipality>
,
<location id="8EFB58A3540FCC75FE8AFF4FFEB2FF33" LSID="urn:lsid:plazi:treatment:038DBF6E540CCC74FF06F8D9FA50F963:8EFB58A3540FCC75FE8AFF4FFEB2FF33" box="[283,362,151,177]" country="India" municipality="Darjeeling" name="Tonglu" pageId="42" pageNumber="193" stateProvince="West Bengal">Tonglu</location>
|
<date id="FF9A28B8540FCC75FEEFFF4FFDF9FF33" box="[382,545,151,177]" pageId="42" pageNumber="193" value="1998-05-19" valueMax="1998-05-23" valueMin="1998-05-19">
<collectingDate id="EFDED150540FCC75FEEFFF4FFDF9FF33" box="[382,545,151,177]" pageId="42" pageNumber="193" value="1998-05-19" valueMax="1998-05-23" valueMin="1998-05-19">19.23.V.1998</collectingDate>
</date>
|
<quantity id="4CDCA39D540FCC75FDA4FF4FFD11FF33" box="[565,713,151,177]" metricMagnitude="3" metricUnit="m" metricValue="2.8" metricValueMax="3.0" metricValueMin="2.6" pageId="42" pageNumber="193" unit="m" value="2800.0" valueMax="3000.0" valueMin="2600.0">
<elevation id="0009E94B540FCC75FDA4FF4FFD11FF33" box="[565,713,151,177]" metricMagnitude="3" metricUnit="m" metricValue="2.8" metricValueMax="3.0" metricValueMin="2.6" pageId="42" pageNumber="193" unit="m" value="2800.0" valueMax="3000.0" valueMin="2600.0">26003000m</elevation>
</quantity>
NN | leg.
<collectorName id="26D16BAE540FCC75FCA0FF4FFC5CFF33" box="[817,900,151,177]" pageId="42" pageNumber="193">Fabrizi</collectorName>
&amp;
<collectorName id="26D16BAE540FCC75FC36FF4FFC20FF33" box="[935,1016,151,177]" pageId="42" pageNumber="193">Ahrens</collectorName>
[under stones along a main ridge] (1 ♁: cSh;
<specimenCount id="9D22C5F1540FCC75FF62FF63FEC6FF57" box="[243,286,187,213]" count="1" pageId="42" pageNumber="193" type="female">1 ♀</specimenCount>
: NME); 1 ♁ (dissected)
</materialsCitation>
,
<materialsCitation id="3B4C0425540FCC75FDA9FF63FF0AFE9F" collectorName="J. Schmidt &amp; All" country="India" elevation="3625" latitude="27.433332" location="Pathibara" longLatPrecision="1237" longitude="87.76667" municipality="Taplejeung" pageId="42" pageNumber="193" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="9D22C5F1540FCC75FDA9FF63FDB9FF57" box="[568,609,187,213]" count="1" pageId="42" pageNumber="193" type="female">1 ♀</specimenCount>
(dissected): E-NEPAL, D:
<collectingMunicipality id="6BFF9402540FCC75FC0BFF63FBCEFF57" box="[922,1046,187,213]" pageId="42" pageNumber="193">Taplejeung</collectingMunicipality>
| E-slope
<location id="8EFB58A3540FCC75FB14FF63FB37FF57" LSID="urn:lsid:plazi:treatment:038DBF6E540CCC74FF06F8D9FA50F963:8EFB58A3540FCC75FB14FF63FB37FF57" box="[1157,1263,187,213]" country="India" latitude="27.433332" longLatPrecision="1237" longitude="87.76667" municipality="Taplejeung" name="Pathibara" pageId="42" pageNumber="193">Pathibara</location>
|
<quantity id="4CDCA39D540FCC75FA90FF62FA40FF57" box="[1281,1432,186,213]" metricMagnitude="3" metricUnit="m" metricValue="3.625" metricValueMax="3.75" metricValueMin="3.5" pageId="42" pageNumber="193" unit="m" value="3625.0" valueMax="3750.0" valueMin="3500.0">
<elevation id="0009E94B540FCC75FA90FF62FA40FF57" box="[1281,1432,186,213]" metricMagnitude="3" metricUnit="m" metricValue="3.625" metricValueMax="3.75" metricValueMin="3.5" pageId="42" pageNumber="193" unit="m" value="3625.0" valueMax="3750.0" valueMin="3500.0">35003750m</elevation>
</quantity>
, 13.V. | 2016, leg.
<collectorName id="26D16BAE540FCC75FEF1FF07FE17FF7B" box="[352,463,223,249]" pageId="42" pageNumber="193">J.Schmidt</collectorName>
|
<geoCoordinate id="EE1068BF540FCC75FE74FF07FD90FF7B" box="[485,584,223,249]" degrees="27" direction="north" minutes="26" orientation="latitude" pageId="42" pageNumber="193" precision="925" value="27.433332">27°26N</geoCoordinate>
,
<geoCoordinate id="EE1068BF540FCC75FDC4FF07FD61FF7B" box="[597,697,223,249]" degrees="87" direction="east" minutes="46" orientation="longitude" pageId="42" pageNumber="193" precision="925" value="87.76667">87°46E</geoCoordinate>
(1 ♁: cSh;
<specimenCount id="9D22C5F1540FCC75FCD1FF07FCB5FF7B" box="[832,877,223,249]" count="1" pageId="42" pageNumber="193" type="female">1 ♀</specimenCount>
: NME).
<collectorName id="26D16BAE540FCC75FC41FF07FC2CFF7B" box="[976,1012,223,249]" pageId="42" pageNumber="193">All</collectorName>
<typeStatus id="549FB0DA540FCC75FC6DFF07FBBFFF7B" box="[1020,1127,223,249]" pageId="42" pageNumber="193" type="paratype">paratypes</typeStatus>
with additional red printed label
</materialsCitation>
:
<typeStatus id="549FB0DA540FCC75FF71FEDBFEA8FE9F" box="[224,368,259,285]" pageId="42" pageNumber="193" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="4C2475FB540FCC75FE15FEDBFE34FE9F" authorityName="Gravenhorst" authorityYear="1802" box="[388,492,259,285]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="193" phylum="Arthropoda" rank="genus">
<emphasis id="B950D26A540FCC75FE15FEDBFE34FE9F" box="[388,492,259,285]" italics="true" pageId="42" pageNumber="193">Omalium</emphasis>
</taxonomicName>
|
<taxonomicName id="4C2475FB540FCC75FD91FEDBFD70FE9F" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[512,680,259,285]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="193" phylum="Arthropoda" rank="species" species="flavotestaceum" status="sp. nov.">
<emphasis id="B950D26A540FCC75FD91FEDBFD70FE9F" box="[512,680,259,285]" italics="true" pageId="42" pageNumber="193">flavotestaceum</emphasis>
</taxonomicName>
<emphasis id="B950D26A540FCC75FD3EFEDBFD37FE9F" bold="true" box="[687,751,259,285]" pageId="42" pageNumber="193">
<taxonomicNameLabel id="A2636F11540FCC75FD3EFEDBFD37FE9F" box="[687,751,259,285]" pageId="42" pageNumber="193" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.V. des. 2023.
</paragraph>
</subSubSection>
<subSubSection id="C33E5DF3540FCC74FF56FEFEFD54FAB3" lastPageId="43" lastPageNumber="194" pageId="42" pageNumber="193" type="description">
<paragraph id="8B9B0E78540FCC75FF56FEFEFE8AFE2F" blockId="42.[151,1436,151,429]" pageId="42" pageNumber="193">
<emphasis id="B950D26A540FCC75FF56FEFEFE81FEC2" bold="true" box="[199,345,294,320]" pageId="42" pageNumber="193">Description.</emphasis>
Measurements (n=6): HW: 0.550.58; HL: 0.420.46; OL: 0.150.16; TL: 0.100.11; AL (
<typeStatus id="549FB0DA540FCC75FF0EFE93FED0FEE7" box="[159,264,331,357]" pageId="42" pageNumber="193" type="holotype">holotype</typeStatus>
): 0.98; PL: 0.410.49; PWmax: 0.650.72; PWmin: 0.550.64; ESL: 0.800.92; EW: 0.850.99; MTbL (
<typeStatus id="549FB0DA540FCC75FF0EFEB7FED0FE0B" box="[159,264,367,393]" pageId="42" pageNumber="193" type="holotype">holotype</typeStatus>
): 0.45; MTrL (
<typeStatus id="549FB0DA540FCC75FE20FEB7FDC2FE0B" box="[433,538,367,393]" pageId="42" pageNumber="193" type="holotype">holotype</typeStatus>
): 0.19 (MTrL 14: 0.08; MTrL 5: 0.11); AW: 0.851.01; AedL: 0.42; BL: 2.49 (
<typeStatus id="549FB0DA540FCC75FF0EFE4AFEDEFE2F" box="[159,262,402,429]" pageId="42" pageNumber="193" type="holotype">holotype</typeStatus>
)3.55.
</paragraph>
<caption id="DF5B5EF0540FCC75FF06F97CFD09F962" ID-DOI="http://doi.org/10.5281/zenodo.10146226" ID-Zenodo-Dep="10146226" httpUri="https://zenodo.org/record/10146226/files/figure.png" pageId="42" pageNumber="193" startId="42.[151,264,1700,1724]" targetBox="[154,1432,456,1674]" targetPageId="42" targetType="figure">
<paragraph id="8B9B0E78540FCC75FF06F97CFD09F962" blockId="42.[151,1436,1700,1760]" pageId="42" pageNumber="193">
<emphasis id="B950D26A540FCC75FF06F97CFE80F93E" bold="true" box="[151,344,1700,1724]" pageId="42" pageNumber="193">FIGURES 8486.</emphasis>
Aedeagus of
<taxonomicName id="4C2475FB540FCC75FE7CF97CFD35F93E" authorityName="Shavrin" authorityYear="2023" box="[493,749,1700,1724]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="193" phylum="Arthropoda" rank="species" species="flavotestaceum">
<emphasis id="B950D26A540FCC75FE7CF97CFD35F93E" box="[493,749,1700,1724]" italics="true" pageId="42" pageNumber="193">Omalium flavotestaceum</emphasis>
</taxonomicName>
(paratype): 84—aedeagus, parameral view, 85—aedeagus, lateral view, 86—median lobe, lateral view. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8B9B0E78540FCC74FF56F8F4FE6DFF7B" blockId="42.[151,1436,1836,2042]" lastBlockId="43.[151,1437,151,1761]" lastPageId="43" lastPageNumber="194" pageId="42" pageNumber="193">
Habitus as in
<figureCitation id="131F12FD540FCC75FEF7F8F4FE63F8C4" box="[358,443,1836,1862]" captionStart="FIGURES 7072" captionStartId="39.[151,264,1085,1109]" captionTargetBox="[159,1419,187,1052]" captionTargetId="figure-16@39.[151,1436,181,1061]" captionTargetPageId="39" captionText="FIGURES 7072. Habitus of Omalium: 70—O. cariosum (lectotype), 71—O. flavotestaceum (paratype), 72—O. kumari (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10146220" httpUri="https://zenodo.org/record/10146220/files/figure.png" pageId="42" pageNumber="193">Fig. 71</figureCitation>
. Head and antennomeres 611 brown; pronotum, elytra and abdomen yellow-brown to reddish-brown (lateral portions of pronotum and paratergites paler; abdomen in some
<typeStatus id="549FB0DA540FCC75FBEFF888FB31F8E8" box="[1150,1257,1872,1898]" pageId="42" pageNumber="193" type="paratype">paratypes</typeStatus>
with yellowish middle parts of abdominal tergites IVVII); mouthparts and antennomeres 15 yellow-brown; legs yellowish, with yellow tarsi. Punctation of head very dense, large and deep, coarser and larger between eyes; nuchal constriction with dense large punctation; punctation of pronotum denser, larger and deeper than that in middle portion of head, sparser in lateral portions, along midline and in mediobasal third; punctation of elytra dense, slightly larger and deeper than that on pronotum, sparser along suture. Anterior portion of clypeus with fine transverse and supra-antennal elevations with diagonal microreticulation; pronotum and elytra without microsculpture; scutellum with fine isodiametric meshes.
</paragraph>
<paragraph id="8B9B0E78540ECC74FF56FEDBFBA2FD2B" blockId="43.[151,1437,151,1761]" pageId="43" pageNumber="194">Head 1.21.3 times as broad as long, with broad clypeus and strongly convex supra-antennal elevations, with very wide and deep anteriomedian depressions. Dorsal surface with distinct elevations in middle and on infraorbital portions: middle area between anteocellar foveae with irregular longitudinal elevations between punctures, portions between anteocellar foveae and eyes with narrow diagonal and moderately long wrinkles, infraorbital portions with three irregular strongly elevated longitudinal wrinkles; portions between posterior margins of eyes and temples with two to three long slightly elevated wrinkles curved posterolaterad. Anteocellar foveae deep and moderately long, slightly convergent latero-apicad, reaching level of middle or anterior third of eyes. Temples about twice shorter than longitudinal length of eyes, without postocular ridges. Distance between ocelli slightly more than twice as long as distance between ocellus and posterior margin of eye. Antenna with antennomeres 610 progressively broadened apicad, with slightly transverse antennomeres 910; antennomere 5 indistinctly broader than 4, 6 slightly longer and broader than 5, 7 slightly broader than 6, 8 distinctly broader than 7, 910 broader than 8, apical antennomere 1.21.3 times as long as 10, from about middle strongly narrowed toward subacute apex.</paragraph>
<paragraph id="8B9B0E78540ECC74FF56FD6BFDB6FCDF" blockId="43.[151,1437,151,1761]" pageId="43" pageNumber="194">Pronotum 1.41.5 times as broad as long, 1.11.2 times as broad as head, from widest apical third more narrowed posteriad than anteriad. Apical angles indistinctly protruded anteriad. Anterior margin widely rounded. Lateral margins widely impressed, deeper in mediolateral portions. Surface of disc with two shallow or deeper and wider longitudinal depressions and oval depression in medioapical portion. Median portion with irregular transverse or diagonal elevations between punctures.</paragraph>
<paragraph id="8B9B0E78540ECC74FF56FCBFFB80FC27" blockId="43.[151,1437,151,1761]" pageId="43" pageNumber="194">Elytra slightly broader than long, about twice as long as pronotum, slightly broadened posteriad. Middle surface of each elytron with strong irregular elevations between punctures, finer along suture.</paragraph>
<paragraph id="8B9B0E78540ECC74FF56FC77FB86FB47" blockId="43.[151,1437,151,1761]" pageId="43" pageNumber="194">
Male. Proportions of preapical antennomeres about as these in females. Posterior margin of abdominal tergite VIII straight or rounded. Posterior margin of abdominal sternite VIII sinuate. Aedeagus with wide basal portion, gradually narrowed toward middle, with long moderately narrow spear-shaped median lobe, with rounded apex; mediolateral portions with elongate sclerotized accessory plates slightly curved and with rounded apices; parameres moderately short, significantly shorter than apex of median lobe, distinctly broadened toward preapical portions and from preapical portions gradually narrowed toward rounded apices, with two moderately long apical and two preapical setae; internal sac narrow and moderately long, without sclerotized structures (
<figureCitation id="131F12FD540ECC74FBE8FB5FFB16FB23" box="[1145,1230,1159,1185]" captionStart="FIGURES 8486" captionStartId="42.[151,264,1700,1724]" captionTargetBox="[154,1432,456,1674]" captionTargetId="figure-135@42.[151,1435,453,1677]" captionTargetPageId="42" captionText="FIGURES 8486. Aedeagus of Omalium flavotestaceum (paratype): 84—aedeagus, parameral view, 85—aedeagus, lateral view, 86—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146226" httpUri="https://zenodo.org/record/10146226/files/figure.png" pageId="43" pageNumber="194">Fig. 84</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="131F12FD540ECC74FEF8FB73FE63FB47" box="[361,443,1194,1221]" captionStart="FIGURES 8486" captionStartId="42.[151,264,1700,1724]" captionTargetBox="[154,1432,456,1674]" captionTargetId="figure-135@42.[151,1435,453,1677]" captionTargetPageId="42" captionText="FIGURES 8486. Aedeagus of Omalium flavotestaceum (paratype): 84—aedeagus, parameral view, 85—aedeagus, lateral view, 86—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146226" httpUri="https://zenodo.org/record/10146226/files/figure.png" pageId="43" pageNumber="194">Fig. 85</figureCitation>
; apical portion moderately wide, rounded apically (
<figureCitation id="131F12FD540ECC74FC6CFB73FB8AFB47" box="[1021,1106,1195,1221]" captionStart="FIGURES 8486" captionStartId="42.[151,264,1700,1724]" captionTargetBox="[154,1432,456,1674]" captionTargetId="figure-135@42.[151,1435,453,1677]" captionTargetPageId="42" captionText="FIGURES 8486. Aedeagus of Omalium flavotestaceum (paratype): 84—aedeagus, parameral view, 85—aedeagus, lateral view, 86—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146226" httpUri="https://zenodo.org/record/10146226/files/figure.png" pageId="43" pageNumber="194">Fig. 86</figureCitation>
).
</paragraph>
<paragraph id="8B9B0E78540ECC74FF56FB17FD54FAB3" blockId="43.[151,1437,151,1761]" pageId="43" pageNumber="194">Female. Posterior margin of abdominal tergite VIII and sternite VIII rounded. Accessory sclertite with very wide basal portion, strongly narrowed toward moderately long and narrow elongate median part, gradually narrrowing toward acute apex. Spermatheca as in Fig. x.</paragraph>
</subSubSection>
<subSubSection id="C33E5DF3540ECC74FF56FAE2FBE5F98B" pageId="43" pageNumber="194" type="diagnosis">
<paragraph id="8B9B0E78540ECC74FF56FAE2FBE5F98B" blockId="43.[151,1437,151,1761]" pageId="43" pageNumber="194">
<emphasis id="B950D26A540ECC74FF56FAE2FE77FAD7" bold="true" box="[199,431,1338,1365]" pageId="43" pageNumber="194">Comparative notes.</emphasis>
Based on the general shape of the temples, distinctly narrowed toward nuchal constriction,
<taxonomicName id="4C2475FB540ECC74FF06FA87FEB8FAFB" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[151,352,1375,1401]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="194" phylum="Arthropoda" rank="species" species="flavotestaceum" status="sp. n.">
<emphasis id="B950D26A540ECC74FF06FA87FF73FAFB" box="[151,171,1375,1401]" italics="true" pageId="43" pageNumber="194">O</emphasis>
.
<emphasis id="B950D26A540ECC74FF29FA87FEB8FAFB" box="[184,352,1375,1401]" italics="true" pageId="43" pageNumber="194">flavotestaceum</emphasis>
</taxonomicName>
<emphasis id="B950D26A540ECC74FEF7FA87FE7DFAFB" bold="true" box="[358,421,1375,1401]" pageId="43" pageNumber="194">
<taxonomicNameLabel id="A2636F11540ECC74FEF7FA87FE7DFAFB" box="[358,421,1375,1401]" pageId="43" pageNumber="194" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="4C2475FB540ECC74FDA3FA87FD63FAFA" box="[562,699,1375,1401]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="194" phylum="Arthropoda" rank="species" species="cariosum">
<emphasis id="B950D26A540ECC74FDA3FA87FD9EFAFB" box="[562,582,1375,1401]" italics="true" pageId="43" pageNumber="194">O</emphasis>
.
<emphasis id="B950D26A540ECC74FDC2FA87FD63FAFA" box="[595,699,1375,1400]" italics="true" pageId="43" pageNumber="194">cariosum</emphasis>
</taxonomicName>
, from which it can be distinguished by the slightly longer temples, the shape of the pronotum widest in apical third, slightly broader elytra, and details of the morphology of the aedeagus (see the key above). Based on the general shape of the pronotum widest in apical third,
<taxonomicName id="4C2475FB540ECC74FB43FA7FFA44FA43" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1234,1436,1447,1473]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="194" phylum="Arthropoda" rank="species" species="flavotestaceum" status="sp. n.">
<emphasis id="B950D26A540ECC74FB43FA7FFB3EFA43" box="[1234,1254,1447,1473]" italics="true" pageId="43" pageNumber="194">O</emphasis>
.
<emphasis id="B950D26A540ECC74FB65FA7FFA44FA43" box="[1268,1436,1447,1473]" italics="true" pageId="43" pageNumber="194">flavotestaceum</emphasis>
</taxonomicName>
<emphasis id="B950D26A540ECC74FF06FA13FF0EFA67" bold="true" box="[151,214,1483,1509]" pageId="43" pageNumber="194">
<taxonomicNameLabel id="A2636F11540ECC74FF06FA13FF0EFA67" box="[151,214,1483,1509]" pageId="43" pageNumber="194" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is somewhat similar to
<taxonomicName id="4C2475FB540ECC74FE4CFA13FD95FA67" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[477,589,1483,1509]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="194" phylum="Arthropoda" rank="species" species="kumari" status="sp. n.">
<emphasis id="B950D26A540ECC74FE4CFA13FE29FA67" box="[477,497,1483,1509]" italics="true" pageId="43" pageNumber="194">O</emphasis>
.
<emphasis id="B950D26A540ECC74FE6FFA13FD95FA67" box="[510,589,1483,1509]" italics="true" pageId="43" pageNumber="194">kumari</emphasis>
</taxonomicName>
<emphasis id="B950D26A540ECC74FDC5FA13FD4BFA67" bold="true" box="[596,659,1483,1509]" pageId="43" pageNumber="194">
<taxonomicNameLabel id="A2636F11540ECC74FDC5FA13FD4BFA67" box="[596,659,1483,1509]" pageId="43" pageNumber="194" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, from which it differs by the paler coloration of the body, shape of the nuchal constriction and different external and internal morphology of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C33E5DF3540ECC74FF56F9CAFD86F9D3" pageId="43" pageNumber="194" type="distribution">
<paragraph id="8B9B0E78540ECC74FF56F9CAFD86F9D3" blockId="43.[151,1437,151,1761]" pageId="43" pageNumber="194">
<emphasis id="B950D26A540ECC74FF56F9CAFEB9F9AE" bold="true" box="[199,353,1554,1580]" pageId="43" pageNumber="194">Distribution.</emphasis>
<taxonomicName id="4C2475FB540ECC74FEF8F9CBFD5AF9AF" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[361,642,1555,1581]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="194" phylum="Arthropoda" rank="species" species="flavotestaceum" status="sp. nov.">
<emphasis id="B950D26A540ECC74FEF8F9CBFD5AF9AF" box="[361,642,1555,1581]" italics="true" pageId="43" pageNumber="194">Omalium flavotestaceum</emphasis>
</taxonomicName>
<emphasis id="B950D26A540ECC74FD1BF9CBFD13F9AF" bold="true" box="[650,715,1555,1581]" pageId="43" pageNumber="194">
<taxonomicNameLabel id="A2636F11540ECC74FD1BF9CBFD13F9AF" box="[650,715,1555,1581]" pageId="43" pageNumber="194" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known from two localities in Pothibara Mt. in eastern
<collectingCountry id="F3334EE8540ECC74FAC8F9CBFA44F9AF" box="[1369,1436,1555,1581]" name="Nepal" pageId="43" pageNumber="194">Nepal</collectingCountry>
and Darjeeling in West Bengal (
<figureCitation id="131F12FD540ECC74FE6CF9EFFD88F9D3" box="[509,592,1591,1617]" captionStart="FIGURE 19" captionStartId="7.[151,250,1891,1915]" captionTargetBox="[151,1435,957,1868]" captionTargetId="figure-299@7.[151,1435,957,1868]" captionTargetPageId="7" captionText="FIGURE 19. Distribution of Omalium in the Himalayan region: O. acutangulum sp. n. (circles), O. amplissimum sp. n. (star), O. beesoni (squares), O. cariosum (diamonds), O. flavotestaceum sp. n. (triangles)." figureDoi="http://doi.org/10.5281/zenodo.10146178" httpUri="https://zenodo.org/record/10146178/files/figure.png" pageId="43" pageNumber="194">Fig. 19</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33E5DF3540ECC74FF66F983FE4DF91B" pageId="43" pageNumber="194" type="etymology">
<paragraph id="8B9B0E78540ECC74FF66F983FE4DF91B" blockId="43.[151,1437,151,1761]" pageId="43" pageNumber="194">
<emphasis id="B950D26A540ECC74FF66F983FEA6F9F7" bold="true" box="[247,382,1627,1653]" pageId="43" pageNumber="194">Etymology.</emphasis>
The specific epither is the Latin
<emphasis id="B950D26A540ECC74FC9FF983FC75F9F7" box="[782,941,1627,1653]" italics="true" pageId="43" pageNumber="194">flavotestaceus</emphasis>
, -
<emphasis id="B950D26A540ECC74FC59F983FC0EF9F6" box="[968,982,1627,1652]" italics="true" pageId="43" pageNumber="194">a</emphasis>
, -
<emphasis id="B950D26A540ECC74FC63F983FBCCF9F6" box="[1010,1044,1627,1652]" italics="true" pageId="43" pageNumber="194">um</emphasis>
(yellowish red). It alludes to the coloration of the body.
</paragraph>
</subSubSection>
<subSubSection id="C33E5DF3540ECC74FF56F97AFA50F963" pageId="43" pageNumber="194" type="biology_ecology">
<paragraph id="8B9B0E78540ECC74FF56F97AFA50F963" blockId="43.[151,1437,151,1761]" pageId="43" pageNumber="194">
<emphasis id="B950D26A540ECC74FF56F97AFE93F93E" bold="true" box="[199,331,1698,1724]" pageId="43" pageNumber="194">Bionomics.</emphasis>
The
<typeStatus id="549FB0DA540ECC74FE16F97BFE6FF93F" box="[391,439,1699,1725]" pageId="43" pageNumber="194">type</typeStatus>
specimen was collected at elevations from
<elevation id="0009E94B540ECC74FC3BF97BFB44F93F" box="[938,1180,1698,1725]" metricMagnitude="3" metricUnit="m" metricValue="3.175" metricValueMax="3.75" metricValueMin="2.6" pageId="43" pageNumber="194" unit="m" value="3175.0" valueMax="3750.0" valueMin="2600.0">
<quantity id="4CDCA39D540ECC74FC3BF97BFB87F93F" box="[938,1119,1698,1725]" metricMagnitude="3" metricUnit="m" metricValue="3.175" metricValueMax="3.75" metricValueMin="2.6" pageId="43" pageNumber="194" unit="m" value="3175.0" valueMax="3750.0" valueMin="2600.0">2600 to 3750 m</quantity>
a.s.l.
</elevation>
<location id="8EFB58A3540ECC74FB35F97BFAC5F93F" LSID="urn:lsid:plazi:treatment:038DBF6E540CCC74FF06F8D9FA50F963:8EFB58A3540ECC74FB35F97BFAC5F93F" box="[1188,1309,1699,1725]" name="Specimens" pageId="43" pageNumber="194" stateProvince="West Bengal">Specimens</location>
from
<collectingRegion id="49E0C09A540ECC74FAF4F97BFF3EF963" country="India" name="West Bengal" pageId="43" pageNumber="194">West Bengal</collectingRegion>
were taken by sifting litter under bushes in a pasture or were collected under stones along the main ridge.
</paragraph>
</subSubSection>
</treatment>
</document>