143 lines
17 KiB
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143 lines
17 KiB
XML
<document id="8705A931AE62DB85D39151CEA24A0442" ID-DOI="10.1206/3695.2" ID-ISSN="0003-0082" ID-Zenodo-Dep="5359065" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630226598931" checkinUser="carolina" docAuthor="Ojanguren-Affilastro, Andrés A., Ochoa, José A., Mattoni, Camilo I. & Prendini, Lorenzo" docDate="2010" docId="033187C9FFD55106FDDDFEC6C785FB86" docLanguage="en" docName="N3695.pdf" docOrigin="American Museum Novitates 2010 (3695)" docSource="http://www.bioone.org/doi/abs/10.1206/3695.2" docStyle="DocumentStyle:26C3580A17D4BBE58BD8D8AC6AE09AC4.6:AmMusNovit.2010-.journal_article" docStyleId="26C3580A17D4BBE58BD8D8AC6AE09AC4" docStyleName="AmMusNovit.2010-.journal_article" docStyleVersion="6" docTitle="Urophonius granulatus Pocock 1898" docType="treatment" docVersion="4" lastPageNumber="5" masterDocId="FF08FFB1FFD15102FFD0FFB1C658FFD0" masterDocTitle="Systematic Revision of the granulatus Group of Urophonius Pocock, 1893 (Scorpiones, Bothriuridae), with Description of a New Species from Central Chile" masterLastPageNumber="44" masterPageNumber="1" pageNumber="5" updateTime="1699192090160" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="7C28C557C1C16700FE686D86F2D43B2A">Systematic Revision of the granulatus Group of Urophonius Pocock, 1893 (Scorpiones, Bothriuridae), with Description of a New Species from Central Chile</mods:title>
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<mods:namePart id="EF1E82BEA0E17B66F55E67D8A818083D">Ojanguren-Affilastro, Andrés A.</mods:namePart>
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<mods:namePart id="35A22B3A01F7D28432AA29C3482C1BAE">Ochoa, José A.</mods:namePart>
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<mods:namePart id="1C2D9416F078AE523455411FBB5D8AE3">Mattoni, Camilo I.</mods:namePart>
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<mods:namePart id="484440455F27930AF7BB8618E1D4D44E">Prendini, Lorenzo</mods:namePart>
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<mods:typeOfResource id="F0433A0788AF098F7FEAEE0C11A6CAC0">text</mods:typeOfResource>
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<mods:title id="C5EF3951F323B66BA4A75147C53DC0CD">American Museum Novitates</mods:title>
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<mods:date id="A4B8436972021E9A49E09A3A6B2057D0">2010</mods:date>
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<mods:number id="9A8E15DAF879A24CC3F43A1A2E2F5205">3695</mods:number>
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<treatment id="033187C9FFD55106FDDDFEC6C785FB86" ID-DOI="http://doi.org/10.5281/zenodo.5454854" ID-Zenodo-Dep="5454854" LSID="urn:lsid:plazi:treatment:033187C9FFD55106FDDDFEC6C785FB86" httpUri="http://treatment.plazi.org/id/033187C9FFD55106FDDDFEC6C785FB86" lastPageNumber="5" pageId="4" pageNumber="5">
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<subSubSection id="C3826554FFD55106FDDDFEC6C54FFE19" pageId="4" pageNumber="5" type="nomenclature">
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<paragraph id="8B2736DFFFD55106FDDDFEC6C53BFE41" blockId="4.[525,867,375,401]" box="[525,867,375,401]" pageId="4" pageNumber="5">
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<heading id="D06F81B3FFD55106FDDDFEC6C53BFE41" box="[525,867,375,401]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="5" reason="5">
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<taxonomicName id="4C984D5CFFD55106FDDDFEC6C54DFE41" ID-CoL="7DY3C" authorityName="Pocock" authorityYear="1898" box="[525,789,375,401]" class="Arachnida" family="Bothriuridae" genus="Urophonius" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="granulatus">
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<emphasis id="B9ECEACDFFD55106FDDDFEC6C54DFE41" box="[525,789,375,401]" italics="true" pageId="4" pageNumber="5">Urophonius granulatus</emphasis>
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</taxonomicName>
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group
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</heading>
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</paragraph>
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<paragraph id="8B2736DFFFD55106FD89FE1EC54FFE19" blockId="4.[601,791,431,457]" box="[601,791,431,457]" pageId="4" pageNumber="5">
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<heading id="D06F81B3FFD55106FD89FE1EC54FFE19" box="[601,791,431,457]" centered="true" fontSize="36" level="3" pageId="4" pageNumber="5" reason="7">
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Figure 1;
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<tableCitation id="C61A0364FFD55106FD19FE1EC54FFE19" box="[713,791,431,457]" captionStart="TABLE 1" captionStartId="1.[96,168,986,1007]" captionText="TABLE 1. Currently recognized species of the scorpion genus Urophonius Pocock, 1893, species groups to which assigned, and countries, provinces (Argentina), states (Brazil), regions (Chile) or departments (Uruguay) in which they have been recorded." pageId="4" pageNumber="5">table 1</tableCitation>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3826554FFD55106FF10FE56C785FB86" pageId="4" pageNumber="5" type="diagnosis">
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<paragraph id="8B2736DFFFD55106FF10FE56C785FB86" blockId="4.[144,1249,487,1110]" pageId="4" pageNumber="5">
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DIAGNOSIS: The four species in the
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<taxonomicName id="4C984D5CFFD55106FDB1FE56C483FDD1" authorityName="Pocock" authorityYear="1898" box="[609,731,487,513]" class="Arachnida" family="Bothriuridae" genus="Urophonius" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="granulatus">
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<emphasis id="B9ECEACDFFD55106FDB1FE56C483FDD1" box="[609,731,487,513]" italics="true" pageId="4" pageNumber="5">granulatus</emphasis>
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</taxonomicName>
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group of
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<taxonomicName id="4C984D5CFFD55106FC9EFE56C58CFDD1" authorityName="Pocock" authorityYear="1893" box="[846,980,487,513]" class="Arachnida" family="Bothriuridae" genus="Urophonius" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="B9ECEACDFFD55106FC9EFE56C58CFDD1" box="[846,980,487,513]" italics="true" pageId="4" pageNumber="5">Urophonius</emphasis>
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</taxonomicName>
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can be separated from other species in the genus by means of the following combination of characters. Metasomal segment I, VSM carinae distinct, usually subparallel throughout length (diverging slightly at anterior margin of segment in some species) (
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<figureCitation id="13A32A5AFFD55106FD63FDE6C50FFDA1" box="[691,855,599,625]" captionStart="FIGURE 3" captionStartId="7.[212,298,1597,1618]" captionTargetBox="[215,1084,224,1567]" captionTargetId="figure-21@7.[212,1086,224,1567]" captionTargetPageId="7" captionText="FIGURE 3. Urophonius granulatus group, sternite VII and metasomal segments I and" figureDoi="http://doi.org/10.5281/zenodo.5359069" httpUri="https://zenodo.org/record/5359069/files/figure.png" pageId="4" pageNumber="5">figs. 3A, C, D</figureCitation>
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,
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<figureCitation id="13A32A5AFFD55106FCB5FDE6C58AFDA1" box="[869,978,599,625]" captionStart="FIGURE 4" captionStartId="8.[177,263,1618,1639]" captionTargetBox="[177,1216,224,1592]" captionTargetId="figure-24@8.[177,1216,224,1593]" captionTargetPageId="8" captionText="FIGURE 4. Urophonius granulatus group, sternite VII and metasomal segments I and II, ventral aspect. A. Urophonius granulatus Pocock, 1898, ♀ (LBRE). B. Urophonius somuncura Acosta, 2001, ♀ (MACN). C. Urophonius tregualemuensis Cekalovic, 1981, ♀ (AMNH). D. Urophonius pizarroi, n. sp., paratype ♀ (MZUC). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5359071" httpUri="https://zenodo.org/record/5359071/files/figure.png" pageId="4" pageNumber="5">4A, C, D</figureCitation>
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), except for
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<emphasis id="B9ECEACDFFD55106FBB7FDE9C692FD46" italics="true" pageId="4" pageNumber="5">
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<taxonomicName id="4C984D5CFFD55106FBB7FDE9C69EFD46" authorityName="Acosta" authorityYear="2001" class="Arachnida" family="Bothriuridae" genus="Urophonius" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="somuncura">U. somuncura</taxonomicName>
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,
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</emphasis>
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in which it is absent or reduced to a few granules (
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<figureCitation id="13A32A5AFFD55106FCECFDCDC5CFFD46" box="[828,919,636,662]" captionStart="FIGURE 3" captionStartId="7.[212,298,1597,1618]" captionTargetBox="[215,1084,224,1567]" captionTargetId="figure-21@7.[212,1086,224,1567]" captionTargetPageId="7" captionText="FIGURE 3. Urophonius granulatus group, sternite VII and metasomal segments I and" figureDoi="http://doi.org/10.5281/zenodo.5359069" httpUri="https://zenodo.org/record/5359069/files/figure.png" pageId="4" pageNumber="5">figs. 3B</figureCitation>
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,
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<figureCitation id="13A32A5AFFD55106FC75FDCDC591FD46" box="[933,969,636,662]" captionStart="FIGURE 4" captionStartId="8.[177,263,1618,1639]" captionTargetBox="[177,1216,224,1592]" captionTargetId="figure-24@8.[177,1216,224,1593]" captionTargetPageId="8" captionText="FIGURE 4. Urophonius granulatus group, sternite VII and metasomal segments I and II, ventral aspect. A. Urophonius granulatus Pocock, 1898, ♀ (LBRE). B. Urophonius somuncura Acosta, 2001, ♀ (MACN). C. Urophonius tregualemuensis Cekalovic, 1981, ♀ (AMNH). D. Urophonius pizarroi, n. sp., paratype ♀ (MZUC). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5359071" httpUri="https://zenodo.org/record/5359071/files/figure.png" pageId="4" pageNumber="5">4B</figureCitation>
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). Metasomal segment I with two or three pairs of ventral macrosetae; II and III each with three pairs of ventral macrosetae (
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<figureCitation id="13A32A5AFFD55106FF0AFD76C77AFD31" box="[218,290,711,737]" captionStart="FIGURE 3" captionStartId="7.[212,298,1597,1618]" captionTargetBox="[215,1084,224,1567]" captionTargetId="figure-21@7.[212,1086,224,1567]" captionTargetPageId="7" captionText="FIGURE 3. Urophonius granulatus group, sternite VII and metasomal segments I and" figureDoi="http://doi.org/10.5281/zenodo.5359069" httpUri="https://zenodo.org/record/5359069/files/figure.png" pageId="4" pageNumber="5">figs. 3</figureCitation>
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,
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<figureCitation id="13A32A5AFFD55106FEE0FD76C719FD31" box="[304,321,711,737]" captionStart="FIGURE 4" captionStartId="8.[177,263,1618,1639]" captionTargetBox="[177,1216,224,1592]" captionTargetId="figure-24@8.[177,1216,224,1593]" captionTargetPageId="8" captionText="FIGURE 4. Urophonius granulatus group, sternite VII and metasomal segments I and II, ventral aspect. A. Urophonius granulatus Pocock, 1898, ♀ (LBRE). B. Urophonius somuncura Acosta, 2001, ♀ (MACN). C. Urophonius tregualemuensis Cekalovic, 1981, ♀ (AMNH). D. Urophonius pizarroi, n. sp., paratype ♀ (MZUC). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5359071" httpUri="https://zenodo.org/record/5359071/files/figure.png" pageId="4" pageNumber="5">4</figureCitation>
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). Telotarsal spination, leg III: 5–5/5–6; IV: 5–6/6–7. Hemispermatophore, lobe region small to medium sized; capsular concavity very well developed; area between basal lobe and external lobe concave (
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<figureCitation id="13A32A5AFFD55106FE1BFCA0C425FCFC" box="[459,637,785,812]" captionStart="FIGURE 8" captionStartId="13.[136,222,1645,1666]" captionTargetBox="[139,1157,225,1614]" captionTargetId="figure-21@13.[133,1162,221,1620]" captionTargetPageId="13" captionText="FIGURE 8. Urophonius granulatus group, sinistral hemispermatophore, ectal and ental aspects. A, B. Urophonius granulatus Pocock, 1898, ♂ (MACN). C, D. Urophonius somuncura Acosta, 2001, ♂ (MACN). E, F. Urophonius tregualemuensis Cekalovic, 1981, ♂ (MACN). G, H. Urophonius pizarroi, n. sp., paratype ♂ (MZUC). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5359079" httpUri="https://zenodo.org/record/5359079/files/figure.png" pageId="4" pageNumber="5">fig. 8A, C, E, G</figureCitation>
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); internal lobe with small, bilobed apophysis disconnected from distal lamina (
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<figureCitation id="13A32A5AFFD55106FE18FC86C429FC81" box="[456,625,823,849]" captionStart="FIGURE 8" captionStartId="13.[136,222,1645,1666]" captionTargetBox="[139,1157,225,1614]" captionTargetId="figure-21@13.[133,1162,221,1620]" captionTargetPageId="13" captionText="FIGURE 8. Urophonius granulatus group, sinistral hemispermatophore, ectal and ental aspects. A, B. Urophonius granulatus Pocock, 1898, ♂ (MACN). C, D. Urophonius somuncura Acosta, 2001, ♂ (MACN). E, F. Urophonius tregualemuensis Cekalovic, 1981, ♂ (MACN). G, H. Urophonius pizarroi, n. sp., paratype ♂ (MZUC). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5359079" httpUri="https://zenodo.org/record/5359079/files/figure.png" pageId="4" pageNumber="5">fig. 8B, D, F, H</figureCitation>
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). Trichobothrial pattern, neobothriotaxic major
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<typeStatus id="5423887DFFD55106FB77FC86C2B8FC81" box="[1191,1248,823,849]" pageId="4" pageNumber="5">Type</typeStatus>
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C, with one accessory trichobothrium in
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<emphasis id="B9ECEACDFFD55106FDA4FCECC4D0FCA6" box="[628,648,861,886]" italics="true" pageId="4" pageNumber="5">V</emphasis>
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series of chela; femur with 3 trichobothria (
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<emphasis id="B9ECEACDFFD55106FB5FFCEDC297FCA6" box="[1167,1231,860,886]" italics="true" pageId="4" pageNumber="5">d, i, e</emphasis>
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); patella with 19 (3
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<emphasis id="B9ECEACDFFD55106FEB8FC33C727FC4B" box="[360,383,898,923]" italics="true" pageId="4" pageNumber="5">V,</emphasis>
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2
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<emphasis id="B9ECEACDFFD55106FE4DFC30C792FC4B" box="[413,458,897,923]" italics="true" pageId="4" pageNumber="5">d, i,</emphasis>
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3
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<emphasis id="B9ECEACDFFD55106FE38FC33C46AFC4B" box="[488,562,898,923]" italics="true" pageId="4" pageNumber="5">et, est,</emphasis>
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2
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<emphasis id="B9ECEACDFFD55106FD9FFC33C421FC4B" box="[591,633,898,923]" italics="true" pageId="4" pageNumber="5">em,</emphasis>
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2
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<emphasis id="B9ECEACDFFD55106FD47FC30C499FC4B" box="[663,705,897,923]" italics="true" pageId="4" pageNumber="5">esb,</emphasis>
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and 5
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<emphasis id="B9ECEACDFFD55106FCC3FC30C575FC4B" box="[787,813,897,923]" italics="true" pageId="4" pageNumber="5">eb</emphasis>
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); chela with 27 (
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<emphasis id="B9ECEACDFFD55106FC27FC33C279FC4B" box="[1015,1057,898,923]" italics="true" pageId="4" pageNumber="5">Est,</emphasis>
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5
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<emphasis id="B9ECEACDFFD55106FBEFFC33C207FC4B" box="[1087,1119,898,923]" italics="true" pageId="4" pageNumber="5">Et,</emphasis>
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5
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<emphasis id="B9ECEACDFFD55106FBADFC33C292FC4B" box="[1149,1226,897,923]" italics="true" pageId="4" pageNumber="5">V, Esb,</emphasis>
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3
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<emphasis id="B9ECEACDFFD55106FF40FC16C4D5FC11" box="[144,653,935,961]" italics="true" pageId="4" pageNumber="5">Eb, Dt, Db, et, est, esb, eb, dt, dst, dsb, db, ib,</emphasis>
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and
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<emphasis id="B9ECEACDFFD55106FD17FC19C480FC11" box="[711,728,936,961]" italics="true" pageId="4" pageNumber="5">it</emphasis>
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); femoral trichobothrium
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<emphasis id="B9ECEACDFFD55106FBC1FC19C245FC11" box="[1041,1053,936,961]" italics="true" pageId="4" pageNumber="5">e</emphasis>
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situated close to a single dorsal macroseta (M1), variable with respect to macrosetae in different species (figs. 10A, 13A, 16A, 19A). Chelicera, movable finger with two well-developed subdistal teeth. Metasomal segments, ventral surfaces each with paired VL and single VM pigmentation stripes, without paired VSM stripes.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |