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<document id="D7787365CBEA02F05B305A44F50A25E2" ID-DOI="10.5281/zenodo.186485" ID-GBIF-Dataset="2eec0d6a-510b-44e7-a8e3-602dc5a88838" ID-ISSN="1175-5326" ID-Zenodo-Dep="186485" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461052423066" checkinUser="plazi" docAuthor="Joly, Luis J." docDate="2009" docId="E2083F003B0CFFAEFF74FD81FA5D60AE" docLanguage="en" docName="zt02048p064.pdf" docOrigin="Zootaxa 2048" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cyclocephala bicolor" docType="key" docVersion="7" lastPageNumber="49" masterDocId="1E3147783B0DFFACFFE3FB23FFC16441" masterDocTitle="Review of the species in the Cyclocephala bicolor Laporte species group (Coleoptera: Scarabaeidae: Dynastinae)" masterLastPageNumber="64" masterPageNumber="47" pageNumber="48" updateTime="1698579509211" updateUser="plazi">
<mods:mods id="25A6696B1D4B1820D33D7F32C364C751" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C104C3543A4FF8BB11EC5CBC6F7B9B60">
<mods:title id="122583E5ABE822744E1423D04B1FE2EC">Review of the species in the Cyclocephala bicolor Laporte species group (Coleoptera: Scarabaeidae: Dynastinae)</mods:title>
</mods:titleInfo>
<mods:name id="FA4A6ACB06C9C70730818ABD9ECEDB2D" type="personal">
<mods:role id="FDE988BE9A033BD8303C53E3F3A2457C">
<mods:roleTerm id="548A8536732AD90E0386A56968775AB6">Author</mods:roleTerm>
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<mods:namePart id="A6A0C216A9AD50658EE8825B25984377">Joly, Luis J.</mods:namePart>
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<mods:typeOfResource id="ED45F3E4A06A9828DAD3841B53A346DA">text</mods:typeOfResource>
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<mods:title id="0123FCA5D3FD76CE58D93B0CFC5D7F04">Zootaxa</mods:title>
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<mods:part id="3B77379A86FFBDA98A305FBD9BE8ECD9">
<mods:date id="F66E12470A84B77488EB042D53AD044A">2009</mods:date>
<mods:detail id="A4E258D9DC498380F28F24A76E03B770" type="volume">
<mods:number id="9145198AD6AC087F98022450A71F6B5D">2048</mods:number>
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<mods:classification id="B85CE5F29709BC0E18240ABED365E395">journal article</mods:classification>
<mods:identifier id="90D72653BB6DD9074AB823FB58A10754" type="DOI">10.5281/zenodo.186485</mods:identifier>
<mods:identifier id="B6CEBCBB903427116E4A3E2F74E54FAD" type="GBIF-Dataset">2eec0d6a-510b-44e7-a8e3-602dc5a88838</mods:identifier>
<mods:identifier id="0782AD9520FCE0BA4F637BB47D70543B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="706D63DACDD9B7447035F6D70425F892" type="Zenodo-Dep">186485</mods:identifier>
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<treatment id="E2083F003B0CFFAEFF74FD81FA5D60AE" ID-DOI="http://doi.org/10.5281/zenodo.6216456" ID-GBIF-Taxon="119612259" ID-Zenodo-Dep="6216456" LSID="urn:lsid:plazi:treatment:E2083F003B0CFFAEFF74FD81FA5D60AE" httpUri="http://treatment.plazi.org/id/E2083F003B0CFFAEFF74FD81FA5D60AE" lastPageId="2" lastPageNumber="49" pageId="1" pageNumber="48">
<subSubSection id="22BBDD9D3B0CFFADFF74FD81FAFD62FD" box="[151,1340,1698,1724]" pageId="1" pageNumber="48" type="nomenclature">
<paragraph id="6A1E8E163B0CFFADFF74FD81FAFD62FD" blockId="1.[151,1340,1698,1724]" box="[151,1340,1698,1724]" pageId="1" pageNumber="48">
<heading id="3156397A3B0CFFADFF74FD81FAFD62FD" bold="true" box="[151,1340,1698,1724]" fontSize="11" level="1" pageId="1" pageNumber="48" reason="1">
<emphasis id="58D552043B0CFFADFF74FD81FAFD62FD" bold="true" box="[151,1340,1698,1724]" pageId="1" pageNumber="48">
Key for identification of the species of the
<taxonomicName id="ADA1F5953B0CFFADFD47FD81FC6562FD" box="[676,932,1698,1724]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="58D552043B0CFFADFD47FD81FC6562FD" bold="true" box="[676,932,1698,1724]" italics="true" pageId="1" pageNumber="48">Cyclocephala bicolor</emphasis>
</taxonomicName>
group of the genus
<taxonomicName id="ADA1F5953B0CFFADFB7AFD81FAFD62FD" box="[1177,1340,1698,1724]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="58D552043B0CFFADFB7AFD81FAFD62FD" bold="true" box="[1177,1340,1698,1724]" italics="true" pageId="1" pageNumber="48">Cyclocephala</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="22BBDD9D3B0CFFAEFF74FDCFFA5D60AE" lastPageId="2" lastPageNumber="49" pageId="1" pageNumber="48" type="key">
<keyStep id="D15596B33B0CFFADFF74FDCFFA5A6362" pageId="1" pageNumber="48">
<paragraph id="6A1E8E163B0CFFADFF74FDCFFA5A6342" blockId="1.[151,1437,1772,2019]" box="[151,1435,1772,1795]" pageId="1" pageNumber="48">
<keyLead id="D1502D233B0CFFADFF74FDCFFA5A6342" box="[151,1435,1772,1795]" pageId="1" pageNumber="48">1. Anterior tarsi enlarged (males) ..................................................................................................................................... 2</keyLead>
</paragraph>
<paragraph id="6A1E8E163B0CFFADFF74FC2FFA5A6362" blockId="1.[151,1437,1772,2019]" box="[151,1435,1804,1827]" pageId="1" pageNumber="48">
<keyLead id="D1502D233B0CFFADFF74FC2FFA5A6362" box="[151,1435,1804,1827]" pageId="1" pageNumber="48">- Anterior tarsi slender (females) .................................................................................................................................... 6</keyLead>
</paragraph>
</keyStep>
<paragraph id="6A1E8E163B0CFFADFF74FC0FFA5D63C2" blockId="1.[151,1437,1772,2019]" pageId="1" pageNumber="48">
<keyStep id="D15596B33B0CFFADFF74FC0FFA5D63C2" pageId="1" pageNumber="48">
<keyLead id="D1502D233B0CFFADFF74FC0FFA5D63C2" pageId="1" pageNumber="48">
2. Second ventrite with a triangular, flat projection on posterior half; basal tooth of protibiae (
<figureCitation id="F29A92933B0CFFADFB30FC0FFAE76302" box="[1235,1318,1836,1859]" captionStart="FIGURE 24 – 33" captionStartId="7.[151,261,1850,1874]" captionTargetBox="[285,1302,542,1826]" captionTargetId="figure@7.[285,1302,542,1826]" captionTargetPageId="7" captionText="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." httpUri="https://zenodo.org/record/186489/files/figure.png" pageId="1" pageNumber="48">Fig. 29</figureCitation>
) distinctly directed forward, separated from middle tooth by a distance about 2.0 times longer than that separating middle from anterior tooth; clypeus on apical half slightly convex in lateral view (
<figureCitation id="F29A92933B0CFFADFC44FC4FFC3563C2" box="[935,1012,1900,1923]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="1" pageNumber="48">Fig. 12</figureCitation>
) .......................
<taxonomicName id="ADA1F5953B0CFFADFB40FC4FFA5D63C2" authority="Dechambre." authorityName="Dechambre." box="[1187,1436,1900,1923]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="58D552043B0CFFADFB40FC4FFB7463C2" box="[1187,1205,1900,1923]" italics="true" pageId="1" pageNumber="48">C</emphasis>
.
<emphasis id="58D552043B0CFFADFB21FC4FFAD563C2" box="[1218,1300,1900,1923]" italics="true" pageId="1" pageNumber="48">dichroa</emphasis>
Dechambre.
</taxonomicName>
</keyLead>
</keyStep>
</paragraph>
<paragraph id="6A1E8E163B0CFFADFF74FCAFFA5C63A2" blockId="1.[151,1437,1772,2019]" pageId="1" pageNumber="48">
<keyLead id="D1502D233B0CFFADFF74FCAFFA5C63A2" pageId="1" pageNumber="48">
- Second ventrite without triangular projection; basal tooth of protibiae (
<figureCitation id="F29A92933B0CFFADFC54FCAFFB4463E2" box="[951,1157,1932,1955]" captionStart="FIGURE 24 – 33" captionStartId="7.[151,261,1850,1874]" captionTargetBox="[285,1302,542,1826]" captionTargetId="figure@7.[285,1302,542,1826]" captionTargetPageId="7" captionText="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." httpUri="https://zenodo.org/record/186489/files/figure.png" pageId="1" pageNumber="48">Figs. 24, 26, 28, 31</figureCitation>
) with anterior border usually perpendicular to longitudinal axis of tibiae and separated from the middle tooth by a distance 1.5 times or less than that separating middle from anterior tooth; clypeus on apical half more or less flat to slightly concave in lateral
</keyLead>
</paragraph>
<paragraph id="6A1E8E163B0FFFAEFF24FBBBFA5A654E" blockId="2.[151,1437,152,1263]" pageId="2" pageNumber="49">
view (
<figureCitation id="F29A92933B0FFFAEFEEAFBBBFE5764EE" box="[265,406,152,175]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="2" pageNumber="49">Figs. 4, 8, 16</figureCitation>
) .................................................................................................................................................... 3
<keyStep id="D15596B33B0FFFAEFF74FB9BFA5A654E" pageId="2" pageNumber="49">
<keyLead id="D1502D233B0FFFAEFF74FB9BFA5A654E" pageId="2" pageNumber="49">
3. Frontoclypeal suture (
<figureCitation id="F29A92933B0FFFAEFE5BFB9BFDC6648E" box="[440,519,184,207]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="2" pageNumber="49">Fig. 15</figureCitation>
) posteriorly projecting angularly; punctation on scutellum very fine, much more so than on middle of base of pronotum; punctation on apical region of pygidium very fine, distinctly finer than that of basal region, which is moderately fine ...............................................................................................
<taxonomicName id="ADA1F5953B0FFFAEFB0BFBDBFA92654E" box="[1256,1363,248,271]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="anibali" status="sp. nov.">
<emphasis id="58D552043B0FFFAEFB0BFBDBFB3B654E" box="[1256,1274,248,271]" italics="true" pageId="2" pageNumber="49">C</emphasis>
.
<emphasis id="58D552043B0FFFAEFAE4FBDBFA92654E" box="[1287,1363,248,271]" italics="true" pageId="2" pageNumber="49">anibali</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="43E6EF7F3B0FFFAEFA83FBD4FA5A654E" box="[1376,1435,247,271]" pageId="2" pageNumber="49" rank="species">
<emphasis id="58D552043B0FFFAEFA83FBD4FA5A654E" bold="true" box="[1376,1435,247,271]" pageId="2" pageNumber="49">sp. n.</emphasis>
</taxonomicNameLabel>
</keyLead>
</keyStep>
</paragraph>
<paragraph id="6A1E8E163B0FFFAEFF74FA3BFA5A652E" blockId="2.[151,1437,152,1263]" pageId="2" pageNumber="49">
<keyLead id="D1502D233B0FFFAEFF74FA3BFA5A652E" pageId="2" pageNumber="49">
- Frontoclypeal suture (
<figureCitation id="F29A92933B0FFFAEFE50FA3BFDFF656E" box="[435,574,280,303]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="2" pageNumber="49">Figs. 3, 7, 11</figureCitation>
) posteriorly projecting arcuately or almost straight; punctation of scutellum fine, scarcely finer than that on center of pronotal base; punctation on apical region of pygidium fine, scarcely smaller than that of basal region, which is strong to moderately strong .................................................................................. 4
</keyLead>
</paragraph>
<keyStep id="D15596B33B0FFFAEFF74FA5BFA5D666E" pageId="2" pageNumber="49">
<paragraph id="6A1E8E163B0FFFAEFF74FA5BFA5A658E" blockId="2.[151,1437,152,1263]" pageId="2" pageNumber="49">
<keyLead id="D1502D233B0FFFAEFF74FA5BFA5A658E" pageId="2" pageNumber="49">
4. Punctures on base of clypeus not discrete, closely set, mostly confluent, appearing as irregular and fine, more or less transversely striate; frontoclypeal suture posteriorly projecting arcuately (
<figureCitation id="F29A92933B0FFFAEFC31FABBFBD265EE" box="[978,1043,408,431]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="2" pageNumber="49">Fig. 3</figureCitation>
); surface of pygidium smooth (
<date id="1E1FA8D63B0FFFAEFABDFABBFA4E65EE" box="[1374,1423,408,431]" pageId="2" pageNumber="49" value="1950-10">50X</date>
) ...................................................................................................................................................................................... 5
</keyLead>
</paragraph>
<paragraph id="6A1E8E163B0FFFAEFF74FAFBFA5D666E" blockId="2.[151,1437,152,1263]" pageId="2" pageNumber="49">
<keyLead id="D1502D233B0FFFAEFF74FAFBFA5D666E" pageId="2" pageNumber="49">
- Punctures on base of clypeus moderately strong, discrete, somewhat transverse, the distance between most punctures smaller than the diameter of puncture; frontoclypeal suture very slightly arcuately projecting posteriorly (
<figureCitation id="F29A92933B0FFFAEFA92FADBFF17666E" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="2" pageNumber="49">Fig. 7</figureCitation>
); surface of pygidium finely alutaceous (
<date id="1E1FA8D63B0FFFAEFD6FF93BFD7C666E" box="[652,701,536,559]" pageId="2" pageNumber="49" value="1950-10">50X</date>
) ........................................................................
<taxonomicName id="ADA1F5953B0FFFAEFB52F93BFA5D666E" authority="Endrodi" authorityName="Endrodi" box="[1201,1436,536,559]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="bicolorata">
<emphasis id="58D552043B0FFFAEFB52F93BFB02666E" box="[1201,1219,536,559]" italics="true" pageId="2" pageNumber="49">C</emphasis>
.
<emphasis id="58D552043B0FFFAEFB33F93BFAFE666E" box="[1232,1343,536,559]" italics="true" pageId="2" pageNumber="49">bicolorata</emphasis>
Endrödi
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="D15596B33B0FFFAEFF74F91BFA5D676E" pageId="2" pageNumber="49">
<paragraph id="6A1E8E163B0FFFAEFF74F91BFA5D66EE" blockId="2.[151,1437,152,1263]" pageId="2" pageNumber="49">
<keyLead id="D1502D233B0FFFAEFF74F91BFA5D66EE" pageId="2" pageNumber="49">
5. Labium (
<figureCitation id="F29A92933B0FFFAEFECAF91BFEA8660E" box="[297,361,568,591]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="2" pageNumber="49">Fig. 5</figureCitation>
) with strong, dense piliferous punctures on basal two-thirds; metatibiae (
<figureCitation id="F29A92933B0FFFAEFB91F91BFB33660E" box="[1138,1266,568,591]" captionStart="FIGURES 49 – 57" captionStartId="9.[151,278,1631,1655]" captionTargetBox="[368,1218,387,1607]" captionTargetId="figure@9.[368,1219,387,1607]" captionTargetPageId="9" captionText="FIGURES 49 – 57. Figs. 49 – 50. Cyclocephala bicolor: 49) metaleg, male; 50) metaleg, female. Figs. 51 – 52. Cyclocephala bicolorata: 51) metaleg, male; 52) metaleg female. Figs. 53 – 54. Cyclocephala dichroa: 53) metaleg, male; 54) metaleg, female. Figs. 55 – 56. Cyclocephala anibali: 55) metaleg, male (holotype); 56) metaleg, female (allotype). Fig. 57. Cyclocephala sarahae metaleg, male (holotype)." httpUri="https://zenodo.org/record/186491/files/figure.png" pageId="2" pageNumber="49">Figs. 49–50</figureCitation>
) with piliferous punctures almost restricted to those of normal longitudinal series close to posterior border; distances between most punctures on apex of pygidium usually smaller than 2.0 times the diameter of punctures; apex of pygidium (in posterior view) feebly convex-truncate.
<emphasis id="58D552043B0FFFAEFDD5F9BBFB2966EE" box="[566,1256,664,687]" italics="true" pageId="2" pageNumber="49">
....................................................................................................
<taxonomicName id="ADA1F5953B0FFFAEFB35F9BBFB2966EE" authority="Laporte" authorityName="Laporte" box="[1238,1256,664,687]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="bicolor">C</taxonomicName>
</emphasis>
.
<emphasis id="58D552043B0FFFAEFB16F9BBFA8366EE" box="[1269,1346,664,687]" italics="true" pageId="2" pageNumber="49">bicolor</emphasis>
Laporte
</keyLead>
</paragraph>
<paragraph id="6A1E8E163B0FFFAEFF74F99BFA5D676E" blockId="2.[151,1437,152,1263]" pageId="2" pageNumber="49">
<keyLead id="D1502D233B0FFFAEFF74F99BFA5D676E" pageId="2" pageNumber="49">
- Labium with one longitudinal series of long bristles on each side, converging towards base; mesotibia and especially metatibiae with numerous piliferous punctures over all surface, in addition to those of normal longitudinal series; distances between most punctures on apex of pygidium usually 2.0 times or more larger than the diameter of punctures; apex of pygidium (in posterior view) rounded.....................................................................
<taxonomicName id="ADA1F5953B0FFFAEFB27F83BFA5D676E" authority="Ratcliffe" authorityName="Ratcliffe" box="[1220,1436,792,815]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="sarahae">
<emphasis id="58D552043B0FFFAEFB27F83BFB17676E" box="[1220,1238,792,815]" italics="true" pageId="2" pageNumber="49">C</emphasis>
.
<emphasis id="58D552043B0FFFAEFB00F83BFAF8676E" box="[1251,1337,792,815]" italics="true" pageId="2" pageNumber="49">sarahae</emphasis>
Ratcliffe
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="D15596B33B0FFFAEFF74F81BFA5A67AE" pageId="2" pageNumber="49">
<paragraph id="6A1E8E163B0FFFAEFF74F81BFA5D67CE" blockId="2.[151,1437,152,1263]" pageId="2" pageNumber="49">
<keyLead id="D1502D233B0FFFAEFF74F81BFA5D67CE" pageId="2" pageNumber="49">
6. Basal tooth of protibiae distinctly anteriorly projected, separated from the middle tooth by a distance about 2.0 times larger than that separating the middle from the anterior (
<figureCitation id="F29A92933B0FFFAEFCFDF87BFCAC672E" box="[798,877,856,879]" captionStart="FIGURE 24 – 33" captionStartId="7.[151,261,1850,1874]" captionTargetBox="[285,1302,542,1826]" captionTargetId="figure@7.[285,1302,542,1826]" captionTargetPageId="7" captionText="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." httpUri="https://zenodo.org/record/186489/files/figure.png" pageId="2" pageNumber="49">Fig. 30</figureCitation>
); clypeus convex in lateral view (
<figureCitation id="F29A92933B0FFFAEFB37F87BFAE0672E" box="[1236,1313,856,879]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="2" pageNumber="49">Fig. 12</figureCitation>
) ................
<emphasis id="58D552043B0FFFAEFF2DF85BFB7A67CE" box="[206,1211,888,911]" italics="true" pageId="2" pageNumber="49">
....................................................................................................................................................
<taxonomicName id="ADA1F5953B0FFFAEFB4AF85BFB7A67CE" authority="Dechambre" authorityName="Dechambre" box="[1193,1211,888,911]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="dichroa">C</taxonomicName>
</emphasis>
.
<emphasis id="58D552043B0FFFAEFB2BF85BFADB67CE" box="[1224,1306,888,911]" italics="true" pageId="2" pageNumber="49">dichroa</emphasis>
Dechambre
</keyLead>
</paragraph>
<paragraph id="6A1E8E163B0FFFAEFF74F8BBFA5A67AE" blockId="2.[151,1437,152,1263]" pageId="2" pageNumber="49">
<keyLead id="D1502D233B0FFFAEFF74F8BBFA5A67AE" pageId="2" pageNumber="49">
- Basal tooth of protibiae more diagonal and/or its anterior border more or less perpendicular to longitudinal axis of tibiae, separated from the middle tooth by a distance 1.5 or less than that separating the middle from the anterior (
<figureCitation id="F29A92933B0FFFAEFF2CF8FBFE6067AE" box="[207,417,984,1007]" captionStart="FIGURE 24 – 33" captionStartId="7.[151,261,1850,1874]" captionTargetBox="[285,1302,542,1826]" captionTargetId="figure@7.[285,1302,542,1826]" captionTargetPageId="7" captionText="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." httpUri="https://zenodo.org/record/186489/files/figure.png" pageId="2" pageNumber="49">Figs. 25, 27, 29, 32</figureCitation>
); clypeus more or less flat in lateral view (
<figureCitation id="F29A92933B0FFFAEFC84F8FBFC3567AE" box="[871,1012,984,1007]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="2" pageNumber="49">Figs. 4, 8, 16</figureCitation>
) .......................................................... 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="D15596B33B0FFFAEFF74F8DBFA5A600E" pageId="2" pageNumber="49">
<paragraph id="6A1E8E163B0FFFAEFF74F8DBFA5A604E" blockId="2.[151,1437,152,1263]" box="[151,1435,1015,1039]" pageId="2" pageNumber="49">
<keyLead id="D1502D233B0FFFAEFF74F8DBFA5A604E" box="[151,1435,1015,1039]" pageId="2" pageNumber="49">
7. Punctures on apical region of pygidium distinctly smaller than those of basal region. ......................
<taxonomicName id="ADA1F5953B0FFFAEFB0DF8DBFA98604E" box="[1262,1369,1016,1039]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="anibali" status="sp. nov.">
<emphasis id="58D552043B0FFFAEFB0DF8DBFAC1604E" box="[1262,1280,1016,1039]" italics="true" pageId="2" pageNumber="49">C</emphasis>
.
<emphasis id="58D552043B0FFFAEFAEDF8DBFA98604E" box="[1294,1369,1016,1039]" italics="true" pageId="2" pageNumber="49">anibali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43E6EF7F3B0FFFAEFA83F8D4FA5A604E" box="[1376,1435,1015,1039]" pageId="2" pageNumber="49" rank="species">
<emphasis id="58D552043B0FFFAEFA83F8D4FA5A604E" bold="true" box="[1376,1435,1015,1039]" pageId="2" pageNumber="49">sp. n.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="6A1E8E163B0FFFAEFF74FF3BFA5A600E" blockId="2.[151,1437,152,1263]" pageId="2" pageNumber="49">
<keyLead id="D1502D233B0FFFAEFF74FF3BFA5A600E" pageId="2" pageNumber="49">- Punctures on apical region of pygidium scarcely smaller than those of basal region, which are strong to moderately strong ........................................................................................................................................................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="D15596B33B0FFFAEFF74FF7BFA5D60AE" pageId="2" pageNumber="49">
<paragraph id="6A1E8E163B0FFFAEFF74FF7BFA5D60CE" blockId="2.[151,1437,152,1263]" pageId="2" pageNumber="49">
<keyLead id="D1502D233B0FFFAEFF74FF7BFA5D60CE" pageId="2" pageNumber="49">
8. Pygidium feebly convex in lateral view, surface smooth, punctures more or less strong almost to apex, on apical region somewhat smaller, usually dense, sometimes scattered .......................................................
<taxonomicName id="ADA1F5953B0FFFAEFB35FF5BFA5D60CE" authority="Laporte" authorityName="Laporte" box="[1238,1436,1144,1167]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="58D552043B0FFFAEFB35FF5BFB2960CE" box="[1238,1256,1144,1167]" italics="true" pageId="2" pageNumber="49">C</emphasis>
.
<emphasis id="58D552043B0FFFAEFB16FF5BFA8360CE" box="[1269,1346,1144,1167]" italics="true" pageId="2" pageNumber="49">bicolor</emphasis>
Laporte
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="6A1E8E163B0FFFAEFF74FFBBFA5D60AE" blockId="2.[151,1437,152,1263]" pageId="2" pageNumber="49">
<keyLead id="D1502D233B0FFFAEFF74FFBBFA5D60AE" pageId="2" pageNumber="49">
- Pygidium strongly convex in lateral view, surface finely alutaceous (
<date id="1E1FA8D63B0FFFAEFC54FFBBFC2860EE" box="[951,1001,1176,1199]" pageId="2" pageNumber="49" value="1950-10">50X</date>
), punctures more or less strong at base, smaller on apex, most distances between punctures 2.0 or more times larger than diameter of puncture. .................... ....................................................................................................................................................
<taxonomicName id="ADA1F5953B0FFFAEFB52FFFBFA5D60AE" authority="Endrodi" authorityName="Endrodi" box="[1201,1436,1240,1263]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="bicolorata">
<emphasis id="58D552043B0FFFAEFB52FFFBFB0260AE" box="[1201,1219,1240,1263]" italics="true" pageId="2" pageNumber="49">C</emphasis>
.
<emphasis id="58D552043B0FFFAEFB33FFFBFAFE60AE" box="[1232,1343,1240,1263]" italics="true" pageId="2" pageNumber="49">bicolorata</emphasis>
Endrödi
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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