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<document id="418EEECB74EE76D7D80F1FFFF3F919B2" ID-DOI="10.11646/zootaxa.5428.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10847364" ID-ZooBank="E438D8DA-B142-4A0C-BA25-F6B6EC45C197" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1711009717928" checkinUser="plazi" docAuthor="Almeida-Silva, Lina M. &amp; Brescovit, Antonio D." docDate="2024" docId="B86D8799FF92FFF58CDDF946FC9F7229" docLanguage="en" docName="zootaxa.5428.2.1.pdf" docOrigin="Zootaxa 5428 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5428.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Goeldia goytacazes Almeida-Silva &amp; Brescovit 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="170" masterDocId="4454FFE1FF83FFE68C4AFFDCFFC37030" masterDocTitle="Unraveling the mysteries of Goeldia Keyserling, 1891: revision, description of seven new species and first record from USA (Araneae: Titanoecidae)" masterLastPageNumber="193" masterPageNumber="151" pageNumber="168" updateTime="1711010311489" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="703DF62FC93C848D1175583029821116">Unraveling the mysteries of Goeldia Keyserling, 1891: revision, description of seven new species and first record from USA (Araneae: Titanoecidae)</mods:title>
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<mods:namePart id="711A4B7FD7B349CA815A57C38B353DCD">Almeida-Silva, Lina M.</mods:namePart>
<mods:affiliation id="EB73C880B7AFE75C86BBF1EEA19C099A">Laboratório de Genômica Evolutiva, Universidade Estadual de Campinas, Instituto de Biologia, Departamento de Genética, Evolução, Microbiologia e Imunologia, Cidade Universitária “ Zeferino Vaz ” Rua Monteiro Lobato, 255, 13083 - 970 - Campinas, SP, Brazil.</mods:affiliation>
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<mods:namePart id="0835DFBFE520509A5183717C81B01E1D">Brescovit, Antonio D.</mods:namePart>
<mods:affiliation id="2FEE4BD18DC09B6FA16ABFEB66466926">Laboratório de Genômica Evolutiva, Universidade Estadual de Campinas, Instituto de Biologia, Departamento de Genética, Evolução, Microbiologia e Imunologia, Cidade Universitária “ Zeferino Vaz ” Rua Monteiro Lobato, 255, 13083 - 970 - Campinas, SP, Brazil. &amp; Laboratório de Coleções Zoológicas-LCZ, Instituto Butantan, Avenida Vital Brazil, 1500, 05503 - 900, São Paulo, SP, Brazil &amp; Laboratório de Genômica Evolutiva, Universidade Estadual de Campinas, Instituto de Biologia, Departamento de Genética, Evolução, Microbiologia e Imunologia, Cidade Universitária “ Zeferino Vaz ” Rua Monteiro Lobato, 255, 13083 - 970 - Campinas, SP, Brazil.</mods:affiliation>
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<mods:title id="295AE069CCF3979EE98D0D652B4DAFD9">Zootaxa</mods:title>
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<mods:date id="5F0FEA6B7BD4DC2405D4AE912625E22A">2024</mods:date>
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<mods:number id="F5163E89FE7577EC0D63A3D181D6831C">2024-03-21</mods:number>
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<mods:number id="8E8FDD934426FCD3DA3E79F9E5FF4D65">5428</mods:number>
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<mods:number id="D0BD02DA02278FBD2CCE9B46789E06D4">2</mods:number>
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<mods:identifier id="2805699A174301440F96245824AA19EE" type="ISSN">1175-5326</mods:identifier>
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<treatment id="B86D8799FF92FFF58CDDF946FC9F7229" ID-DOI="http://doi.org/10.5281/zenodo.10847406" ID-Zenodo-Dep="10847406" LSID="urn:lsid:plazi:treatment:B86D8799FF92FFF58CDDF946FC9F7229" httpUri="http://treatment.plazi.org/id/B86D8799FF92FFF58CDDF946FC9F7229" lastPageId="19" lastPageNumber="170" pageId="17" pageNumber="168">
<subSubSection id="78DE6504FF92FFF78CDDF946FE207685" box="[151,483,1690,1717]" pageId="17" pageNumber="168" type="nomenclature">
<paragraph id="307B368FFF92FFF78CDDF946FE207685" blockId="17.[151,483,1690,1753]" box="[151,483,1690,1717]" pageId="17" pageNumber="168">
<heading id="6B3381E3FF92FFF78CDDF946FE207685" bold="true" box="[151,483,1690,1717]" fontSize="11" level="1" pageId="17" pageNumber="168" reason="1">
<emphasis id="02B0EA9DFF92FFF78CDDF946FE207685" bold="true" box="[151,483,1690,1717]" pageId="17" pageNumber="168">
<taxonomicName id="F7C44D0CFF92FFF78CDDF946FEBD7685" authority="Almeida-Silva &amp; Brescovit, 2024" authorityName="Almeida-Silva &amp; Brescovit" authorityYear="2024" box="[151,382,1690,1717]" class="Arachnida" family="Titanoecidae" genus="Goeldia" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="168" phylum="Arthropoda" rank="species" species="goytacazes" status="sp. nov.">
<emphasis id="02B0EA9DFF92FFF78CDDF946FEBD7685" bold="true" box="[151,382,1690,1717]" italics="true" pageId="17" pageNumber="168">Goeldia goytacazes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="198357E6FF92FFF78DCFF947FE207685" box="[389,483,1691,1717]" pageId="17" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="78DE6504FF92FFF78CDDF963FEEB76E9" box="[151,296,1727,1753]" pageId="17" pageNumber="168" type="description">
<paragraph id="307B368FFF92FFF78CDDF963FEEB76E9" blockId="17.[151,483,1690,1753]" box="[151,296,1727,1753]" pageId="17" pageNumber="168">
(
<figureCitation id="A8FF2A0AFF92FFF78CD5F963FF3476E9" box="[159,247,1727,1753]" captionStart="FIGURE 12" captionStartId="18.[151,250,1937,1961]" captionTargetBox="[158,1436,786,1910]" captionTargetId="figure-284@18.[151,1437,777,1913]" captionTargetPageId="18" captionText="FIGURE 12. A–F. Goeldia goytacazes sp. nov. A–D: male, left palp; E–F: female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847404" httpUri="https://zenodo.org/record/10847404/files/figure.png" pageId="17" pageNumber="168">Figs 12</figureCitation>
;
<figureCitation id="A8FF2A0AFF92FFF78D49F963FEE376E9" box="[259,288,1727,1753]" captionStart="FIGURE 27" captionStartId="40.[151,250,1068,1092]" captionTargetBox="[232,1356,186,1038]" captionTargetId="figure-21@40.[225,1363,181,1044]" captionTargetPageId="40" captionText="FIGURE 27. Distribution of Goeldia spp. in South America: Goeldia luteipes (square—purple); G. zyngierae (star—red) and G. gauderio (triangle—yellow); G. goytacazes (pentagon—orange); G. patellaris (circle—green). Type species are marked as black versions of each symbol when there are records to multiple localities." figureDoi="http://doi.org/10.5281/zenodo.10847453" httpUri="https://zenodo.org/record/10847453/files/figure.png" pageId="17" pageNumber="168">27</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="78DE6504FF92FFF78CDDF8DBFDBC77BD" pageId="17" pageNumber="168" type="materials_examined">
<paragraph id="307B368FFF92FFF78CDDF8DBFDEA7759" blockId="17.[151,1436,1799,2005]" pageId="17" pageNumber="168">
<emphasis id="02B0EA9DFF92FFF78CDDF8DBFE857711" bold="true" box="[151,326,1799,1825]" pageId="17" pageNumber="168">Type material.</emphasis>
<materialsCitation id="80AC3CD2FF92FFF78D05F8DBFC3C7775" collectingDate="2003-09-24" collectorName="de Goytacazes &amp; C. C. L. Teixeira" country="Brazil" latitude="-21.75389" location="Campos de Goytacazes" longLatPrecision="21" longitude="-41.323887" pageId="17" pageNumber="168" specimenCount="2" stateProvince="Rio de Janeiro" typeStatus="holotype">
<typeStatus id="EF7F882DFF92FFF78D05F8DBFE747711" box="[335,439,1799,1825]" pageId="17" pageNumber="168" type="holotype">Holotype</typeStatus>
male,
<typeStatus id="EF7F882DFF92FFF78E4FF8DBFDA67711" box="[517,613,1799,1825]" pageId="17" pageNumber="168" type="paratype">paratype</typeStatus>
female from
<location id="351B6054FF92FFF78F48F8DBFBC97711" LSID="urn:lsid:plazi:treatment:B86D8799FF92FFF58CDDF946FC9F7229:351B6054FF92FFF78F48F8DBFBC97711" box="[770,1034,1799,1825]" country="Brazil" latitude="-21.75389" longLatPrecision="21" longitude="-41.323887" name="Campos de Goytacazes" pageId="17" pageNumber="168" stateProvince="Rio de Janeiro">
Campos
<collectorName id="9D315359FF92FFF78F2CF8DBFBC97711" box="[870,1034,1799,1825]" pageId="17" pageNumber="168">de Goytacazes</collectorName>
</location>
(
<geoCoordinate id="55F05048FF92FFF78851F8DBFB637712" box="[1051,1184,1799,1826]" degrees="21" direction="south" minutes="45" orientation="latitude" pageId="17" pageNumber="168" precision="15" seconds="14" value="-21.75389">21°45’14”S</geoCoordinate>
<geoCoordinate id="55F05048FF92FFF788E3F8DBFAF87711" box="[1193,1339,1799,1825]" degrees="41" direction="west" minutes="19" orientation="longitude" pageId="17" pageNumber="168" precision="15" seconds="26" value="-41.323887">41°19’26”W</geoCoordinate>
),
<collectingRegion id="F200F86DFF92FFF7891BF8DBFF287775" country="Brazil" name="Rio de Janeiro" pageId="17" pageNumber="168">Rio de Janeiro</collectingRegion>
,
<collectingCountry id="48D3761FFF92FFF78CB3F8F7FE837775" box="[249,320,1835,1861]" name="Brazil" pageId="17" pageNumber="168">Brazil</collectingCountry>
,
<date id="447A104FFF92FFF78D04F8F7FE137775" box="[334,464,1835,1861]" pageId="17" pageNumber="168" value="2003-09-24">
<collectingDate id="543EE9A7FF92FFF78D04F8F7FE137775" box="[334,464,1835,1861]" pageId="17" pageNumber="168" value="2003-09-24">24.IX.2003</collectingDate>
</date>
,
<collectorName id="9D315359FF92FFF78D94F8F7FD4E7775" box="[478,653,1835,1861]" pageId="17" pageNumber="168">C.C.L. Teixeira</collectorName>
coll., deposited at MCN 39868
</materialsCitation>
;
<typeStatus id="EF7F882DFF92FFF78847F8F7FBAC7775" box="[1037,1135,1835,1861]" pageId="17" pageNumber="168" type="paratype">Paratype</typeStatus>
female with same data as holotype, deposited at MCN 43434.
</paragraph>
<paragraph id="307B368FFF92FFF78C8DF8AFFDBC77BD" blockId="17.[151,1436,1799,2005]" box="[199,639,1907,1933]" pageId="17" pageNumber="168">
<emphasis id="02B0EA9DFF92FFF78C8DF8AFFDF777BD" bold="true" box="[199,564,1907,1933]" pageId="17" pageNumber="168">Additional material examined.</emphasis>
None.
</paragraph>
</subSubSection>
<subSubSection id="78DE6504FF92FFF78C8DF84BFDE577E5" pageId="17" pageNumber="168" type="etymology">
<paragraph id="307B368FFF92FFF78C8DF84BFDE577E5" blockId="17.[151,1436,1799,2005]" pageId="17" pageNumber="168">
<emphasis id="02B0EA9DFF92FFF78C8DF84BFE8B7781" bold="true" box="[199,328,1943,1969]" pageId="17" pageNumber="168">Etymology</emphasis>
. The specific name is a noun in apposition in honor to the indigenous people of the Goytacazes tribe, who lived in the
<typeStatus id="EF7F882DFF92FFF78DDFF867FE0677E5" box="[405,453,1979,2005]" pageId="17" pageNumber="168">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="78DE6504FF91FFF48C8DFF4AFE2C7171" pageId="18" pageNumber="169" type="diagnosis">
<paragraph id="307B368FFF91FFF48C8DFF4AFE2C7171" blockId="18.[151,1437,150,753]" pageId="18" pageNumber="169">
<emphasis id="02B0EA9DFF91FFF48C8DFF4AFEF97080" bold="true" box="[199,314,150,176]" pageId="18" pageNumber="169">Diagnosis</emphasis>
.
<taxonomicName id="F7C44D0CFF91FFF48D03FF4BFDE57080" authorityName="Almeida-Silva &amp; Brescovit" authorityYear="2024" box="[329,550,151,177]" class="Arachnida" family="Titanoecidae" genus="Goeldia" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="goytacazes">
<emphasis id="02B0EA9DFF91FFF48D03FF4BFDE57080" box="[329,550,151,177]" italics="true" pageId="18" pageNumber="169">Goeldia goytacazes</emphasis>
</taxonomicName>
resembles
<taxonomicName id="F7C44D0CFF91FFF48EEFFF4BFCC97080" authorityName="Almeida-Silva &amp; Brescovit" authorityYear="2024" box="[677,778,151,177]" class="Arachnida" family="Titanoecidae" genus="Goeldia" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="mirim">
<emphasis id="02B0EA9DFF91FFF48EEFFF4BFCC97080" box="[677,778,151,177]" italics="true" pageId="18" pageNumber="169">G. mirim</emphasis>
</taxonomicName>
by the general shape of the male palp and epigynum (
<figureCitation id="A8FF2A0AFF91FFF48925FF4BFF7670E5" captionStart="FIGURE 16" captionStartId="27.[151,250,1917,1941]" captionTargetBox="[156,1434,822,1888]" captionTargetId="figure-283@27.[151,1436,813,1894]" captionTargetPageId="27" captionText="FIGURE 16. A–F. Goeldia mirim sp. nov. A–D: male, left palp; E–F: female, epigynum: A. Ventral view; B. Tegular process, frontal view. C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847416" httpUri="https://zenodo.org/record/10847416/files/figure.png" pageId="18" pageNumber="169">Fig. 16</figureCitation>
), but differs by the patellar apophysis pointing posteriorly (
<figureCitation id="A8FF2A0AFF91FFF48F0CFF67FC6870E5" box="[838,939,187,213]" captionStart="FIGURE 12" captionStartId="18.[151,250,1937,1961]" captionTargetBox="[158,1436,786,1910]" captionTargetId="figure-284@18.[151,1437,777,1913]" captionTargetPageId="18" captionText="FIGURE 12. A–F. Goeldia goytacazes sp. nov. A–D: male, left palp; E–F: female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847404" httpUri="https://zenodo.org/record/10847404/files/figure.png" pageId="18" pageNumber="169">Fig. 12C</figureCitation>
); by the enlarged base of the MLT (
<figureCitation id="A8FF2A0AFF91FFF4897BFF67FA5770E5" box="[1329,1428,187,213]" captionStart="FIGURE 12" captionStartId="18.[151,250,1937,1961]" captionTargetBox="[158,1436,786,1910]" captionTargetId="figure-284@18.[151,1437,777,1913]" captionTargetPageId="18" captionText="FIGURE 12. A–F. Goeldia goytacazes sp. nov. A–D: male, left palp; E–F: female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847404" httpUri="https://zenodo.org/record/10847404/files/figure.png" pageId="18" pageNumber="169">Fig. 12C</figureCitation>
) and tegular process lanceolate (
<figureCitation id="A8FF2A0AFF91FFF48DB9FF03FD9470C9" box="[499,599,223,249]" captionStart="FIGURE 12" captionStartId="18.[151,250,1937,1961]" captionTargetBox="[158,1436,786,1910]" captionTargetId="figure-284@18.[151,1437,777,1913]" captionTargetPageId="18" captionText="FIGURE 12. A–F. Goeldia goytacazes sp. nov. A–D: male, left palp; E–F: female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847404" httpUri="https://zenodo.org/record/10847404/files/figure.png" pageId="18" pageNumber="169">Fig. 12A</figureCitation>
) in the male palp. The female epigynum is also similar but in
<taxonomicName id="F7C44D0CFF91FFF4894AFF03FA5F70C8" authorityName="Almeida-Silva &amp; Brescovit" authorityYear="2024" box="[1280,1436,223,249]" class="Arachnida" family="Titanoecidae" genus="Goeldia" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="goytacazes">
<emphasis id="02B0EA9DFF91FFF4894AFF03FA5F70C8" box="[1280,1436,223,249]" italics="true" pageId="18" pageNumber="169">G. goytacazes</emphasis>
</taxonomicName>
it covers almost completely the copulatory duct wall (
<figureCitation id="A8FF2A0AFF91FFF48EB9FEDFFC95712D" box="[755,854,259,285]" captionStart="FIGURE 12" captionStartId="18.[151,250,1937,1961]" captionTargetBox="[158,1436,786,1910]" captionTargetId="figure-284@18.[151,1437,777,1913]" captionTargetPageId="18" captionText="FIGURE 12. A–F. Goeldia goytacazes sp. nov. A–D: male, left palp; E–F: female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847404" httpUri="https://zenodo.org/record/10847404/files/figure.png" pageId="18" pageNumber="169">Fig. 12E</figureCitation>
) and the vertical spermathecae is much shorter than the copulatory duct (
<figureCitation id="A8FF2A0AFF91FFF48D35FEFBFE217171" box="[383,482,295,321]" captionStart="FIGURE 12" captionStartId="18.[151,250,1937,1961]" captionTargetBox="[158,1436,786,1910]" captionTargetId="figure-284@18.[151,1437,777,1913]" captionTargetPageId="18" captionText="FIGURE 12. A–F. Goeldia goytacazes sp. nov. A–D: male, left palp; E–F: female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847404" httpUri="https://zenodo.org/record/10847404/files/figure.png" pageId="18" pageNumber="169">Fig. 12F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="78DE6504FF91FFF58C8DFE96FB9271C5" lastPageId="19" lastPageNumber="170" pageId="18" pageNumber="169" type="description">
<paragraph id="307B368FFF91FFF48C8DFE96FF1B72C1" blockId="18.[151,1437,150,753]" pageId="18" pageNumber="169">
<emphasis id="02B0EA9DFF91FFF48C8DFE96FE9A7155" bold="true" box="[199,345,330,357]" pageId="18" pageNumber="169">Description.</emphasis>
Male (
<typeStatus id="EF7F882DFF91FFF48DE7FE97FDDF7155" box="[429,540,331,357]" pageId="18" pageNumber="169" type="holotype">Holotype</typeStatus>
). Coloration: carapace dark brown. Sternum and labium black, endites brown. Leg coxae and femora black, other leg segments yellowish brown except distal half of patella and tibia III and IV black and metatarsus IV grayish. Opisthosoma grayish brown. Total length 3.15. Carapace: lenght 1.59; width 1.16; height 0.75. Opisthosoma: length 1.65; width 1.38. Clypeus: height 0.13. Eye diameters and interdistances: AME 0.04; ALE 0.08; PME 0.06; PLE 0.06; AME–AME 0.08; AME–ALE 0.07; PME–PME 0.1; PME–PLE 0.15; AME–PME 0.08; ALE–PLE 0.07. Chelicerae paturon: length 0.65. Leg measurements: I: femur 1.23/ patella 0.50/ tibia 1.05/ metatarsus 0.98/ tarsus 0.62/ total 4.37; II: 1.13/ 0.48/ 0.83/1.03/ 0.48/ 3.93; III: 1.05/ 0.4/ 0.93/ 0.75/ 0.34/ 3.46; IV: 1.25/ 1.20/ 0.5/ 1.06/ 0.44/ 4.45. Leg formula 4123. Leg spination: femur I pap; tibia I v1r-2(1r-
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)- 2(1r-
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)-0, p0-1-0, r0-1-0-1-0; III–IV: p1ap, r1ap; metatarsus I v1r-1r-
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, p1ap, r1ap; II v1r-0-1r-
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, p1ap, r1ap; IV v1r-1p-1map, p0-1-1ap, r1ap. Patellar apophysis pointing dorsally and posteriorly (
<figureCitation id="A8FF2A0AFF91FFF48824FD53FB3A7299" box="[1134,1273,655,681]" captionStart="FIGURE 12" captionStartId="18.[151,250,1937,1961]" captionTargetBox="[158,1436,786,1910]" captionTargetId="figure-284@18.[151,1437,777,1913]" captionTargetPageId="18" captionText="FIGURE 12. A–F. Goeldia goytacazes sp. nov. A–D: male, left palp; E–F: female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847404" httpUri="https://zenodo.org/record/10847404/files/figure.png" pageId="18" pageNumber="169">Fig. 12C–D</figureCitation>
). RLT with an extra basal fold (
<figureCitation id="A8FF2A0AFF91FFF48D13FD6FFE0172FD" box="[345,450,691,717]" captionStart="FIGURE 12" captionStartId="18.[151,250,1937,1961]" captionTargetBox="[158,1436,786,1910]" captionTargetId="figure-284@18.[151,1437,777,1913]" captionTargetPageId="18" captionText="FIGURE 12. A–F. Goeldia goytacazes sp. nov. A–D: male, left palp; E–F: female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847404" httpUri="https://zenodo.org/record/10847404/files/figure.png" pageId="18" pageNumber="169">Fig. 12C</figureCitation>
). Tegular process as in illustration (
<figureCitation id="A8FF2A0AFF91FFF48F11FD6FFC0672FD" box="[859,965,691,717]" captionStart="FIGURE 12" captionStartId="18.[151,250,1937,1961]" captionTargetBox="[158,1436,786,1910]" captionTargetId="figure-284@18.[151,1437,777,1913]" captionTargetPageId="18" captionText="FIGURE 12. A–F. Goeldia goytacazes sp. nov. A–D: male, left palp; E–F: female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847404" httpUri="https://zenodo.org/record/10847404/files/figure.png" pageId="18" pageNumber="169">Fig. 12A</figureCitation>
).
<emphasis id="02B0EA9DFF91FFF48F90FD6FFBB172FD" box="[986,1138,691,717]" italics="true" pageId="18" pageNumber="169">Pars pendula</emphasis>
thickening subdistal (
<figureCitation id="A8FF2A0AFF91FFF48925FD6FFF0F72C1" captionStart="FIGURE 12" captionStartId="18.[151,250,1937,1961]" captionTargetBox="[158,1436,786,1910]" captionTargetId="figure-284@18.[151,1437,777,1913]" captionTargetPageId="18" captionText="FIGURE 12. A–F. Goeldia goytacazes sp. nov. A–D: male, left palp; E–F: female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847404" httpUri="https://zenodo.org/record/10847404/files/figure.png" pageId="18" pageNumber="169">Fig. 12D</figureCitation>
).
</paragraph>
<caption id="64BB6607FF91FFF48CDDF84DFA5577FD" ID-DOI="http://doi.org/10.5281/zenodo.10847404" ID-Zenodo-Dep="10847404" httpUri="https://zenodo.org/record/10847404/files/figure.png" pageId="18" pageNumber="169" startId="18.[151,250,1937,1961]" targetBox="[158,1436,786,1910]" targetPageId="18" targetType="figure">
<paragraph id="307B368FFF91FFF48CDDF84DFA5577FD" blockId="18.[151,1437,1937,1997]" pageId="18" pageNumber="169">
<emphasis id="02B0EA9DFF91FFF48CDDF84DFEE37799" bold="true" box="[151,288,1937,1961]" pageId="18" pageNumber="169">FIGURE 12.</emphasis>
A–F.
<taxonomicName id="F7C44D0CFF91FFF48D17F84DFDE67799" authority="Almeida-Silva &amp; Brescovit, 2024" authorityName="Almeida-Silva &amp; Brescovit" authorityYear="2024" box="[349,549,1937,1961]" class="Arachnida" family="Titanoecidae" genus="Goeldia" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="goytacazes" status="sp. nov.">
<emphasis id="02B0EA9DFF91FFF48D17F84DFDE67799" box="[349,549,1937,1961]" italics="true" pageId="18" pageNumber="169">Goeldia goytacazes</emphasis>
</taxonomicName>
<emphasis id="02B0EA9DFF91FFF48E66F84DFDBE7799" bold="true" box="[556,637,1937,1961]" pageId="18" pageNumber="169">
<taxonomicNameLabel id="198357E6FF91FFF48E66F84DFDBE7799" box="[556,637,1937,1961]" pageId="18" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A–D: male, left palp; E–F: female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view.
</paragraph>
</caption>
<paragraph id="307B368FFF90FFF58C8DFF4BFC1671E1" blockId="19.[151,1437,151,537]" pageId="19" pageNumber="170">
Female (
<typeStatus id="EF7F882DFF90FFF58D61FF4BFE577081" box="[299,404,151,177]" pageId="19" pageNumber="170" type="paratype">Paratype</typeStatus>
): Coloration as in male except, femora black, tarsus I and II reddish brown and other leg articles yellowish brown. Total length 3.84. Carapace: length 1.80; width 1.24; height 0.75. Opisthosoma: lentgth 1.24; width 1.48. Clypeus: height 1.40. Eye diameters and interdistances: AME 0.05; ALE 0.09; PME 0.06; PLE 0.08; AME–AME 0.07; AME–ALE 0.11; PME–PME 0.10; PME–PLE 0.19; AME–PME 0.07; ALE–PLE 0.05. Chelicerae paturon: length 0.75. Leg measurements: I: femur 1.20/ patella 0.48/ tibia 1.04/ metatarsus 0.88/ tarsus 0.56/ total 4.16; II: 1.10/ 0.54/ 0.75/ 0.92/ 0.54/ 3.85; III: 1.00/ 0.52/ 0.72/ 0.74/ 0.44/ 3.42; IV: 1.48/ 1.10/ 0.48/ 0.90/ 0.42/ 4.38. Leg formula 1243. Leg spination: metatarsus II–IV v1map. Copulatory ducts almost completely covered by the median field (
<figureCitation id="A8FF2A0AFF90FFF58DAEFE4FFD88719D" box="[484,587,403,429]" captionStart="FIGURE 12" captionStartId="18.[151,250,1937,1961]" captionTargetBox="[158,1436,786,1910]" captionTargetId="figure-284@18.[151,1437,777,1913]" captionTargetPageId="18" captionText="FIGURE 12. A–F. Goeldia goytacazes sp. nov. A–D: male, left palp; E–F: female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847404" httpUri="https://zenodo.org/record/10847404/files/figure.png" pageId="19" pageNumber="170">Fig. 12E</figureCitation>
). Posterior edge of the epigynum round (
<figureCitation id="A8FF2A0AFF90FFF58857FE4FFB40719D" box="[1053,1155,403,429]" captionStart="FIGURE 12" captionStartId="18.[151,250,1937,1961]" captionTargetBox="[158,1436,786,1910]" captionTargetId="figure-284@18.[151,1437,777,1913]" captionTargetPageId="18" captionText="FIGURE 12. A–F. Goeldia goytacazes sp. nov. A–D: male, left palp; E–F: female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847404" httpUri="https://zenodo.org/record/10847404/files/figure.png" pageId="19" pageNumber="170">Fig. 12E</figureCitation>
). Apex of the copulatory ducts bent (
<figureCitation id="A8FF2A0AFF90FFF58D53FE6BFEBE71E1" box="[281,381,439,465]" captionStart="FIGURE 12" captionStartId="18.[151,250,1937,1961]" captionTargetBox="[158,1436,786,1910]" captionTargetId="figure-284@18.[151,1437,777,1913]" captionTargetPageId="18" captionText="FIGURE 12. A–F. Goeldia goytacazes sp. nov. A–D: male, left palp; E–F: female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847404" httpUri="https://zenodo.org/record/10847404/files/figure.png" pageId="19" pageNumber="170">Fig. 12F</figureCitation>
). Horizontal spermathecae slightly curved (
<figureCitation id="A8FF2A0AFF90FFF58F2FFE6BFC0B71E1" box="[869,968,439,465]" captionStart="FIGURE 12" captionStartId="18.[151,250,1937,1961]" captionTargetBox="[158,1436,786,1910]" captionTargetId="figure-284@18.[151,1437,777,1913]" captionTargetPageId="18" captionText="FIGURE 12. A–F. Goeldia goytacazes sp. nov. A–D: male, left palp; E–F: female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847404" httpUri="https://zenodo.org/record/10847404/files/figure.png" pageId="19" pageNumber="170">Fig. 12F</figureCitation>
).
</paragraph>
<paragraph id="307B368FFF90FFF58C8DFE06FB9271C5" blockId="19.[151,1437,151,537]" box="[199,1105,474,501]" pageId="19" pageNumber="170">
<emphasis id="02B0EA9DFF90FFF58C8DFE06FEFB71C4" bold="true" box="[199,312,474,500]" pageId="19" pageNumber="170">Variation</emphasis>
. Length (
<specimenCount id="26C2FD06FF90FFF58DEDFE07FE0871C5" box="[423,459,475,501]" count="2" pageId="19" pageNumber="170" type="female">2♀</specimenCount>
): total 3.24–3.84; carapace 1.56–1.80; femur I 1.03–1.20.
</paragraph>
</subSubSection>
<subSubSection id="78DE6504FF90FFF58C8DFE22FC9F7229" box="[199,860,510,537]" pageId="19" pageNumber="170" type="distribution">
<paragraph id="307B368FFF90FFF58C8DFE22FC9F7229" blockId="19.[151,1437,151,537]" box="[199,860,510,537]" pageId="19" pageNumber="170">
<emphasis id="02B0EA9DFF90FFF58C8DFE22FE9A7228" bold="true" box="[199,345,510,536]" pageId="19" pageNumber="170">Distribution</emphasis>
. Only known from the
<typeStatus id="EF7F882DFF90FFF58E2AFE23FD537229" box="[608,656,511,537]" pageId="19" pageNumber="170">type</typeStatus>
locality (
<figureCitation id="A8FF2A0AFF90FFF58EB1FE23FC8C7229" box="[763,847,511,537]" captionStart="FIGURE 27" captionStartId="40.[151,250,1068,1092]" captionTargetBox="[232,1356,186,1038]" captionTargetId="figure-21@40.[225,1363,181,1044]" captionTargetPageId="40" captionText="FIGURE 27. Distribution of Goeldia spp. in South America: Goeldia luteipes (square—purple); G. zyngierae (star—red) and G. gauderio (triangle—yellow); G. goytacazes (pentagon—orange); G. patellaris (circle—green). Type species are marked as black versions of each symbol when there are records to multiple localities." figureDoi="http://doi.org/10.5281/zenodo.10847453" httpUri="https://zenodo.org/record/10847453/files/figure.png" pageId="19" pageNumber="170">Fig. 27</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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