treatments-xml/data/E0/27/57/E0275714D95E0E39FF76E944FD00AB41.xml

<document id="6009B6EBA010007D6A710FDDD55CEDC9" ID-DOI="10.1111/j.1096-3642.2011.00731.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5442095" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630452512153" checkinUser="valdenar" docAuthor="Li, Liang &amp; Zhou, Hong-Zhang" docDate="2011" docId="E0275714D95E0E39FF76E944FD00AB41" docLanguage="en" docName="j.1096-3642.2011.00731.x.pdf" docOrigin="Zoological Journal of the Linnean Society 163 (3)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2011.00731.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Pseudohesperus tripartitus Li &amp; Zhou 2011, SP. NOV." docType="treatment" docVersion="4" lastPageNumber="715" masterDocId="1C1E2F6CD97D0E1DFFA5EC4DFFF6AE0C" masterDocTitle="Revision and phylogenetic assessment of the rove beetle genus Pseudohesperus Hayashi, with broad reference to the subtribe Philonthina (Coleoptera: Staphylinidae: Staphylinini)" masterLastPageNumber="722" masterPageNumber="679" pageNumber="714" updateTime="1699275715089" updateUser="plazi">
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<mods:title id="DC25A47D9DA833F1138555665E77B862">Revision and phylogenetic assessment of the rove beetle genus Pseudohesperus Hayashi, with broad reference to the subtribe Philonthina (Coleoptera: Staphylinidae: Staphylinini)</mods:title>
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<mods:namePart id="3119A5ADE2E0BB4A775288FE24A693A9">Li, Liang</mods:namePart>
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<mods:namePart id="3CE3158F8F465BE0FDCC0C4FAA09598F">Zhou, Hong-Zhang</mods:namePart>
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<mods:date id="587710CA93AEA649B1E4F54B75AE96E2">2011</mods:date>
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<mods:number id="13A35E7DA808371DB1835A534BF0F8E6">2011-10-25</mods:number>
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<treatment id="E0275714D95E0E39FF76E944FD00AB41" ID-DOI="http://doi.org/10.5281/zenodo.5492141" ID-Zenodo-Dep="5492141" LSID="urn:lsid:plazi:treatment:E0275714D95E0E39FF76E944FD00AB41" httpUri="http://treatment.plazi.org/id/E0275714D95E0E39FF76E944FD00AB41" lastPageId="36" lastPageNumber="715" pageId="35" pageNumber="714">
<subSubSection id="2094B589D95E0E3EFF76E944FDAAAB33" pageId="35" pageNumber="714" type="nomenclature">
<paragraph id="6831E602D95E0E3EFF76E944FDAAAB33" blockId="35.[211,731,1289,1343]" pageId="35" pageNumber="714">
<heading id="3379516ED95E0E3EFF76E944FDAAAB33" centered="true" fontSize="9" level="2" pageId="35" pageNumber="714" reason="4">
<taxonomicName id="AF8E9D81D95E0E3EFF76E944FD2DAB11" authority="Li &amp; Zhou" authorityName="Li &amp; Zhou" authorityYear="2011" box="[211,731,1289,1310]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="715" phylum="Arthropoda" rank="species" species="tripartitus" status="sp. nov.">
<emphasis id="5AFA3A10D95E0E3EFF76E944FDA6AB11" bold="true" box="[211,592,1289,1310]" italics="true" pageId="35" pageNumber="714">PSEUDOHESPERUS TRIPARTITUS</emphasis>
LI &amp; ZHOU
</taxonomicName>
<emphasis id="5AFA3A10D95E0E3EFEF6E961FE4EAB32" bold="true" box="[339,440,1321,1342]" pageId="35" pageNumber="714">
<taxonomicNameLabel id="41C9876BD95E0E3EFEF6E961FE4EAB32" box="[339,440,1321,1342]" pageId="35" pageNumber="714" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="F0B5FA87D95E0E3EFE6FE964FDA2AB33" box="[458,596,1321,1343]" captionStart="Figure 29" captionStartId="35.[826,905,866,885]" captionTargetBox="[827,1441,199,835]" captionTargetId="figure-520@35.[826,1442,197,836]" captionTargetPageId="35" captionText="Figure 29. Pseudohesperus tripartitus sp. nov. A, aedeagus, dorsal (parameral side) view; B, median lobe of aedeagus, lateral view; C, underside of paramere; D, sternite IX of male; E, sternite VIII of male; F, tergite X of male; G, tergite X of female; H, gonocoxites of genital segment of female. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5442153" httpUri="https://zenodo.org/record/5442153/files/figure.png" pageId="35" pageNumber="714">FIG. 29A–H</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2094B589D95E0E39FF01E91DFD00AB41" lastPageId="36" lastPageNumber="715" pageId="35" pageNumber="714" type="materials_examined">
<paragraph id="6831E602D95E0E3EFF01E91DFE22ABEC" blockId="35.[164,779,1359,1504]" pageId="35" pageNumber="714">
<emphasis id="5AFA3A10D95E0E3EFF01E91DFEBEAB68" box="[164,328,1359,1381]" italics="true" pageId="35" pageNumber="714">Type material:</emphasis>
<materialsCitation id="D8E6EC5FD95E0E3EFEF4E902FE22ABEC" collectingDate="2004-06-06" collectingDateMax="2004-07-09" collectingDateMin="2004-06-06" country="China" elevation="2220" latitude="31.466667" longLatPrecision="1216" longitude="103.6" pageId="35" pageNumber="714" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Sichuan" typeStatus="holotype">
<typeStatus id="B73558A0D95E0E3EFEF4E902FE4DAB69" box="[337,443,1359,1381]" pageId="35" pageNumber="714" type="holotype">Holotype</typeStatus>
: 
<specimenCount id="7E882D8BD95E0E3EFE62E91FFE16AB68" box="[455,480,1362,1380]" pageId="35" pageNumber="714" type="male">♂</specimenCount>
, 
<collectingCountry id="1099A692D95E0E3EFE4AE902FDCEAB69" box="[495,568,1359,1381]" name="China" pageId="35" pageNumber="714">China</collectingCountry>
: 
<collectingRegion id="AA4A28E0D95E0E3EFDE1E902FD52AB69" box="[580,676,1359,1381]" country="China" name="Sichuan" pageId="35" pageNumber="714">Sichuan</collectingRegion>
: 
<taxonomicName id="AF8E9D81D95E0E3EFD15E902FEF7AB88" authority=": Wulitun" authorityName="Wulitun" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="714" phylum="Arthropoda" rank="species" species="wolong">Wolong: Wulitun</taxonomicName>
(
<geoCoordinate id="0DBA80C5D95E0E3EFEB4E923FE9BAB8F" box="[273,365,1388,1411]" degrees="31" direction="north" minutes="28" orientation="latitude" pageId="35" pageNumber="714" precision="925" value="31.466667">31°28′N</geoCoordinate>
, 
<geoCoordinate id="0DBA80C5D95E0E3EFEDFE923FE14AB8F" box="[378,482,1388,1412]" degrees="103" direction="east" minutes="36" orientation="longitude" pageId="35" pageNumber="714" precision="925" value="103.6">103°36′E</geoCoordinate>
), 
<elevation id="E3A30131D95E0E3EFE52E923FD83AB88" box="[503,629,1390,1412]" metricMagnitude="3" metricUnit="m" metricValue="2.2199999999999998" pageId="35" pageNumber="714" unit="m" value="2220.0">
<quantity id="AF764BE7D95E0E3EFE52E923FDBCAB8F" box="[503,586,1390,1411]" metricMagnitude="3" metricUnit="m" metricValue="2.2199999999999998" pageId="35" pageNumber="714" unit="m" value="2220.0">2220 m</quantity>
asl
</elevation>
, 
<date id="1C30C0C2D95E0E3EFD24E923FCF0AB88" box="[641,774,1390,1412]" pageId="35" pageNumber="714" value="2004-06-06" valueMax="2004-06-09" valueMin="2004-06-06">
<collectingDate id="0C74392AD95E0E3EFD24E923FCF0AB88" box="[641,774,1390,1412]" pageId="35" pageNumber="714" value="2004-06-06" valueMax="2004-06-09" valueMin="2004-06-06">6–9.vi.2004</collectingDate>
</date>
, Yu Xiaodong collected (IZ-CAS); 
<typeStatus id="B73558A0D95E0E3EFD94E9C0FD53ABAE" box="[561,677,1421,1442]" pageId="35" pageNumber="714" type="paratype">paratypes</typeStatus>
: 
<specimenCount id="7E882D8BD95E0E3EFD12E9C1FCF2ABAD" box="[695,772,1420,1441]" pageId="35" pageNumber="714" type="male">2 ♂♂</specimenCount>
, 
<specimenCount id="7E882D8BD95E0E3EFF01E9E6FF22ABCC" box="[164,212,1451,1472]" pageId="35" pageNumber="714" type="female">1 ♀</specimenCount>
, same data as holotype, but 
<elevation id="E3A30131D95E0E3EFD9BE9E6FCF0ABCD" box="[574,774,1451,1473]" metricMagnitude="3" metricUnit="m" metricValue="2.315" metricValueMax="2.41" metricValueMin="2.2199999999999998" pageId="35" pageNumber="714" unit="m" value="2315.0" valueMax="2410.0" valueMin="2220.0">
<quantity id="AF764BE7D95E0E3EFD9BE9E6FD20ABCC" box="[574,726,1451,1472]" metricMagnitude="3" metricUnit="m" metricValue="2.315" metricValueMax="2.41" metricValueMin="2.2199999999999998" pageId="35" pageNumber="714" unit="m" value="2315.0" valueMax="2410.0" valueMin="2220.0">2220–2410 m</quantity>
asl
</elevation>
, 
<date id="1C30C0C2D95E0E3EFF01E984FEA8ABD3" box="[164,350,1481,1503]" pageId="35" pageNumber="714" value="2004-06-21" valueMax="2004-07-09" valueMin="2004-06-21">
<collectingDate id="0C74392AD95E0E3EFF01E984FEA8ABD3" box="[164,350,1481,1503]" pageId="35" pageNumber="714" value="2004-06-21" valueMax="2004-07-09" valueMin="2004-06-21">21.vi.-9.vii.2004</collectingDate>
</date>
(IZ-CAS).
</materialsCitation>
</paragraph>
<paragraph id="6831E602D95E0E3EFF01EA44FDF0A8BB" blockId="35.[164,779,1544,1903]" pageId="35" pageNumber="714">
<emphasis id="5AFA3A10D95E0E3EFF01EA44FEDAA812" box="[164,300,1545,1566]" italics="true" pageId="35" pageNumber="714">Description:</emphasis>
Head and pronotum black, shining. Antennae dark brown, with antennomere XI or antennomeres X–XI reddish-brown. Elytra reddish-brown. Scutellum dark brown. Abdomen black, with strongly iridescent lustre. Mandibles, maxillary and labial palpi, and legs reddish-brown.
</paragraph>
<paragraph id="6831E602D95E0E3EFF19EA8DFADBAB6B" blockId="35.[164,779,1544,1903]" lastBlockId="35.[826,1441,1085,1903]" pageId="35" pageNumber="714">
Body 9.55–10.0 mm long (
<collectionCode id="0E9F7EC7D95E0E3EFDAEEA8DFDB2A8D9" box="[523,580,1728,1749]" pageId="35" pageNumber="714">HPL</collectionCode>
: 
<quantity id="AF764BE7D95E0E3EFDFEEA8DFD09A8D9" box="[603,767,1728,1750]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="2.94" metricValueMin="2.86" pageId="35" pageNumber="714" unit="mm" value="2.9" valueMax="2.94" valueMin="2.86">2.86–2.94 mm</quantity>
). Head of rounded quadrangular shape, 
<quantity id="AF764BE7D95E0E3EFDCFEA92FCFDA8F8" box="[618,779,1759,1780]" metricMagnitude="-3" metricUnit="m" metricValue="1.305" metricValueMax="1.39" metricValueMin="1.22" pageId="35" pageNumber="714" unit="mm" value="1.305" valueMax="1.39" valueMin="1.22">1.22–1.39 mm</quantity>
long, 
<quantity id="AF764BE7D95E0E3EFF46EAB0FE75A91E" box="[227,387,1789,1810]" metricMagnitude="-3" metricUnit="m" metricValue="1.225" metricValueMax="1.31" metricValueMin="1.14" pageId="35" pageNumber="714" unit="mm" value="1.225" valueMax="1.31" valueMin="1.14">1.14–1.31 mm</quantity>
wide, slightly narrower than long (
<collectionCode id="0E9F7EC7D95E0E3EFF0EEB51FF2FA93D" box="[171,217,1820,1841]" pageId="35" pageNumber="714">HW</collectionCode>
: 
<collectionCode id="0E9F7EC7D95E0E3EFF48EB51FEE4A93D" box="[237,274,1820,1841]" pageId="35" pageNumber="714">HL</collectionCode>
= 0.93–1.00). Tempora 
<quantity id="AF764BE7D95E0E3EFD82EB51FD31A93D" box="[551,711,1820,1841]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="4.9" metricValueMin="4.1" pageId="35" pageNumber="714" unit="mm" value="0.44999999999999996" valueMax="0.49" valueMin="0.41">0.41–0.49 mm</quantity>
long, almost evenly rounded, sparsely and coarsely punctate; eyes small, slightly prominent, 
<quantity id="AF764BE7D95E0E3EFDCFEB14FCFDA962" box="[618,779,1881,1902]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" metricValueMax="4.1" metricValueMin="3.3" pageId="35" pageNumber="714" unit="mm" value="0.37" valueMax="0.41" valueMin="0.33">0.33–0.41 mm</quantity>
long, 0.80–0.83 times as long as tempora. Dorsal surface of head with moderately numerous, large setiferous punctures, becoming sparser toward vertex; vertex largely impunctate. Entire head with distinct and profound microsculpture of transverse waves. Antennae long, antennomere I very long, thickened towards apex, antennomere III much longer than antennomere II, following antennomeres gradually decreasing in length, but all longer than wide, antennomere XI obliquely truncated.
</paragraph>
<paragraph id="6831E602D95E0E3EFCF6E93DFC22A831" blockId="35.[826,1441,1085,1903]" pageId="35" pageNumber="714">
Pronotum slightly narrowed anteriad, 
<quantity id="AF764BE7D95E0E3EFAC6E922FC6BABAF" metricMagnitude="-3" metricUnit="m" metricValue="1.51" metricValueMax="1.55" metricValueMin="1.47" pageId="35" pageNumber="714" unit="mm" value="1.51" valueMax="1.55" valueMin="1.47">1.47– 1.55 mm</quantity>
long, 
<quantity id="AF764BE7D95E0E3EFC5DE9C3FB6FABAF" box="[1016,1177,1422,1443]" metricMagnitude="-3" metricUnit="m" metricValue="1.39" metricValueMax="1.47" metricValueMin="1.31" pageId="35" pageNumber="714" unit="mm" value="1.39" valueMax="1.47" valueMin="1.31">1.31–1.47 mm</quantity>
wide, slightly wider than head (
<collectionCode id="0E9F7EC7D95E0E3EFC1BE9E0FC1EABCE" box="[958,1000,1453,1474]" country="Thailand" name="Paleontological Collections" pageId="35" pageNumber="714">PW</collectionCode>
: 
<collectionCode id="0E9F7EC7D95E0E3EFC59E9E0FBDFABCE" box="[1020,1065,1453,1474]" pageId="35" pageNumber="714">HW</collectionCode>
= 1.06–1.20), sparsely and finely punctate, punctures separated by three to five times their diameter, narrowly impunctate along midline, disc with distinct and profound microsculpture of oblique lines.
</paragraph>
<paragraph id="6831E602D95E0E3EFCF6EA08FC42A8F8" blockId="35.[826,1441,1085,1903]" pageId="35" pageNumber="714">
Elytra 
<quantity id="AF764BE7D95E0E3EFC15EA0BFBA6A857" box="[944,1104,1606,1627]" metricMagnitude="-3" metricUnit="m" metricValue="1.92" metricValueMax="2.04" metricValueMin="1.8" pageId="35" pageNumber="714" unit="mm" value="1.92" valueMax="2.04" valueMin="1.8">1.80–2.04 mm</quantity>
long, 
<quantity id="AF764BE7D95E0E3EFB15EA08FAA7A857" box="[1200,1361,1605,1627]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.04" metricValueMin="1.96" pageId="35" pageNumber="714" unit="mm" value="2.0" valueMax="2.04" valueMin="1.96">1.96–2.04 mm</quantity>
wide, along sides 1.16–1.33 times as long as pronotum, densely and finely punctate, punctures separated by one to two times their diameter. Scutellum large, triangular, densely and finely punctate, and pubescent.
</paragraph>
<paragraph id="6831E602D95E0E39FCF6EAB0FD8AAEF6" blockId="35.[826,1441,1085,1903]" lastBlockId="36.[144,759,198,954]" lastPageId="36" lastPageNumber="715" pageId="35" pageNumber="714">
Abdomen a little narrowed posteriad, widest 
<quantity id="AF764BE7D95E0E3EFAC6EAB0FC69A93D" metricMagnitude="-3" metricUnit="m" metricValue="1.755" metricValueMax="1.88" metricValueMin="1.63" pageId="35" pageNumber="714" unit="mm" value="1.755" valueMax="1.88" valueMin="1.63">1.63– 1.88 mm</quantity>
, densely and finely punctate, punctures separated by two to three times their diameter at base, gradually becoming sparser toward apex; tergites III–V with two basal carinae, elevated area between basal carinae almost impunctate.
</paragraph>
<paragraph id="6831E602D9590E39FF0DED4EFD8FAFE3" blockId="36.[144,759,198,954]" pageId="36" pageNumber="715">
Male: Sternite VIII with moderately wide, obtusely triangular medioapical emargination, partly filled by semimembranous extension (
<figureCitation id="F0B5FA87D9590E39FD85ED0DFD75AF5A" box="[544,643,320,342]" captionStart="Figure 29" captionStartId="35.[826,905,866,885]" captionTargetBox="[827,1441,199,835]" captionTargetId="figure-520@35.[826,1442,197,836]" captionTargetPageId="35" captionText="Figure 29. Pseudohesperus tripartitus sp. nov. A, aedeagus, dorsal (parameral side) view; B, median lobe of aedeagus, lateral view; C, underside of paramere; D, sternite IX of male; E, sternite VIII of male; F, tergite X of male; G, tergite X of female; H, gonocoxites of genital segment of female. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5442153" httpUri="https://zenodo.org/record/5442153/files/figure.png" pageId="36" pageNumber="715">Fig. 29E</figureCitation>
). Genital segment with stylus of tergite IX simple, densely setose apically. Sternite IX with asymmetrical basal portion, apex deeply emarginate, without modified rod-like extensions (
<figureCitation id="F0B5FA87D9590E39FE21EDF6FE1FAFDD" box="[388,489,443,465]" captionStart="Figure 29" captionStartId="35.[826,905,866,885]" captionTargetBox="[827,1441,199,835]" captionTargetId="figure-520@35.[826,1442,197,836]" captionTargetPageId="35" captionText="Figure 29. Pseudohesperus tripartitus sp. nov. A, aedeagus, dorsal (parameral side) view; B, median lobe of aedeagus, lateral view; C, underside of paramere; D, sternite IX of male; E, sternite VIII of male; F, tergite X of male; G, tergite X of female; H, gonocoxites of genital segment of female. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5442153" httpUri="https://zenodo.org/record/5442153/files/figure.png" pageId="36" pageNumber="715">Fig. 29D</figureCitation>
). Tergite X triangular with medioapical emargination (
<figureCitation id="F0B5FA87D9590E39FDAFED94FD9AAFE3" box="[522,620,473,495]" captionStart="Figure 29" captionStartId="35.[826,905,866,885]" captionTargetBox="[827,1441,199,835]" captionTargetId="figure-520@35.[826,1442,197,836]" captionTargetPageId="35" captionText="Figure 29. Pseudohesperus tripartitus sp. nov. A, aedeagus, dorsal (parameral side) view; B, median lobe of aedeagus, lateral view; C, underside of paramere; D, sternite IX of male; E, sternite VIII of male; F, tergite X of male; G, tergite X of female; H, gonocoxites of genital segment of female. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5442153" httpUri="https://zenodo.org/record/5442153/files/figure.png" pageId="36" pageNumber="715">Fig. 29F</figureCitation>
).
</paragraph>
<paragraph id="6831E602D9590E39FF0DEDB5FEFEAD0F" blockId="36.[144,759,198,954]" pageId="36" pageNumber="715">
Aedeagus similar to that of 
<taxonomicName id="AF8E9D81D9590E39FDBDEDB4FF19AC20" authority="Li &amp; Zhou" authorityName="Li &amp; Zhou" authorityYear="2011" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="715" phylum="Arthropoda" rank="species" species="pedatiformis" status="sp. nov.">
<emphasis id="5AFA3A10D9590E39FDBDEDB4FD3BAC01" box="[536,717,504,526]" italics="true" pageId="36" pageNumber="715">P. pedatiformis</emphasis>
Li &amp; Zhou
</taxonomicName>
<taxonomicNameLabel id="41C9876BD9590E39FEA4EE5AFE95AC20" box="[257,355,535,556]" pageId="36" pageNumber="715" rank="species">sp. nov.</taxonomicNameLabel>
; median lobe slightly exceeding paramere, narrowed into subacute apex (
<figureCitation id="F0B5FA87D9590E39FD22EE78FD1DAC47" box="[647,747,565,587]" captionStart="Figure 29" captionStartId="35.[826,905,866,885]" captionTargetBox="[827,1441,199,835]" captionTargetId="figure-520@35.[826,1442,197,836]" captionTargetPageId="35" captionText="Figure 29. Pseudohesperus tripartitus sp. nov. A, aedeagus, dorsal (parameral side) view; B, median lobe of aedeagus, lateral view; C, underside of paramere; D, sternite IX of male; E, sternite VIII of male; F, tergite X of male; G, tergite X of female; H, gonocoxites of genital segment of female. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5442153" httpUri="https://zenodo.org/record/5442153/files/figure.png" pageId="36" pageNumber="715">Fig. 29A</figureCitation>
); in lateral view, apical portion slightly bent dorsad (parameral side) (
<figureCitation id="F0B5FA87D9590E39FECFEE3FFE3AAC84" box="[362,460,626,648]" captionStart="Figure 29" captionStartId="35.[826,905,866,885]" captionTargetBox="[827,1441,199,835]" captionTargetId="figure-520@35.[826,1442,197,836]" captionTargetPageId="35" captionText="Figure 29. Pseudohesperus tripartitus sp. nov. A, aedeagus, dorsal (parameral side) view; B, median lobe of aedeagus, lateral view; C, underside of paramere; D, sternite IX of male; E, sternite VIII of male; F, tergite X of male; G, tergite X of female; H, gonocoxites of genital segment of female. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5442153" httpUri="https://zenodo.org/record/5442153/files/figure.png" pageId="36" pageNumber="715">Fig. 29B</figureCitation>
); paramere trilobed apically, middle lobe distinctly longer and wider than lateral lobes, face adjacent to median lobe with peg setae arranged along apical portion of paramere (
<figureCitation id="F0B5FA87D9590E39FF3DEEA0FF0DAD0F" box="[152,251,749,771]" captionStart="Figure 29" captionStartId="35.[826,905,866,885]" captionTargetBox="[827,1441,199,835]" captionTargetId="figure-520@35.[826,1442,197,836]" captionTargetPageId="35" captionText="Figure 29. Pseudohesperus tripartitus sp. nov. A, aedeagus, dorsal (parameral side) view; B, median lobe of aedeagus, lateral view; C, underside of paramere; D, sternite IX of male; E, sternite VIII of male; F, tergite X of male; G, tergite X of female; H, gonocoxites of genital segment of female. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5442153" httpUri="https://zenodo.org/record/5442153/files/figure.png" pageId="36" pageNumber="715">Fig. 29C</figureCitation>
).
</paragraph>
<paragraph id="6831E602D9590E39FF0DEF46FE20ADB6" blockId="36.[144,759,198,954]" pageId="36" pageNumber="715">
Female: Sternite VIII with arcuate apex, lacking medioapical emargination. Genital segment with second gonocoxites moderately long, each with minute stylus bearing two long apical setae (
<figureCitation id="F0B5FA87D9590E39FD21EF2AFD1DAD71" box="[644,747,871,893]" captionStart="Figure 29" captionStartId="35.[826,905,866,885]" captionTargetBox="[827,1441,199,835]" captionTargetId="figure-520@35.[826,1442,197,836]" captionTargetPageId="35" captionText="Figure 29. Pseudohesperus tripartitus sp. nov. A, aedeagus, dorsal (parameral side) view; B, median lobe of aedeagus, lateral view; C, underside of paramere; D, sternite IX of male; E, sternite VIII of male; F, tergite X of male; G, tergite X of female; H, gonocoxites of genital segment of female. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5442153" httpUri="https://zenodo.org/record/5442153/files/figure.png" pageId="36" pageNumber="715">Fig. 29H</figureCitation>
). Tergite X triangular, subtruncated at apex, with two long apical setae (
<figureCitation id="F0B5FA87D9590E39FEC0EFE9FE3CADB6" box="[357,458,932,954]" captionStart="Figure 29" captionStartId="35.[826,905,866,885]" captionTargetBox="[827,1441,199,835]" captionTargetId="figure-520@35.[826,1442,197,836]" captionTargetPageId="35" captionText="Figure 29. Pseudohesperus tripartitus sp. nov. A, aedeagus, dorsal (parameral side) view; B, median lobe of aedeagus, lateral view; C, underside of paramere; D, sternite IX of male; E, sternite VIII of male; F, tergite X of male; G, tergite X of female; H, gonocoxites of genital segment of female. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5442153" httpUri="https://zenodo.org/record/5442153/files/figure.png" pageId="36" pageNumber="715">Fig. 29G</figureCitation>
).
</paragraph>
<caption id="3CF1B68AD9590E39FC83EF02FC34ADD4" ID-DOI="http://doi.org/10.5281/zenodo.5442155" ID-Zenodo-Dep="5442155" httpUri="https://zenodo.org/record/5442155/files/figure.png" pageId="36" pageNumber="715" startId="36.[806,885,847,866]" targetBox="[808,1417,198,816]" targetPageId="36">
<paragraph id="6831E602D9590E39FC83EF02FC34ADD4" blockId="36.[806,1421,847,984]" pageId="36" pageNumber="715">
<emphasis id="5AFA3A10D9590E39FC83EF02FB1CAD6F" bold="true" box="[806,1258,847,867]" pageId="36" pageNumber="715">
Figure 30. 
<taxonomicName id="AF8E9D81D9590E39FC0DEF02FB82AD6E" authority="Li &amp; Zhou, 2011" authorityName="Li &amp; Zhou" authorityYear="2011" box="[936,1140,847,866]" class="Insecta" family="Staphylinidae" genus="Bisnius" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="715" phylum="Arthropoda" rank="species" species="lubricus" status="sp. nov.">
<emphasis id="5AFA3A10D9590E39FC0DEF02FB82AD6E" bold="true" box="[936,1140,847,866]" italics="true" pageId="36" pageNumber="715">Bisnius lubricus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="41C9876BD9590E39FB22EF1DFB1CAD6F" box="[1159,1258,848,867]" pageId="36" pageNumber="715" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, aedeagus, dorsal (parameral side) view; B, median lobe of aedeagus, lateral view; C, underside of paramere; D, sternite IX of male; E, sternite VIII of male; F, tergite X of male. Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="6831E602D9590E39FF35EFAEFDFAAA3A" blockId="36.[144,758,995,1078]" pageId="36" pageNumber="715">
<emphasis id="5AFA3A10D9590E39FF35EFAEFEFBADF4" box="[144,269,995,1016]" italics="true" pageId="36" pageNumber="715">Etymology:</emphasis>
The specific epithet is derived from the Latin word 
<emphasis id="5AFA3A10D9590E39FEB8E84FFE74AA1B" box="[285,386,1026,1047]" italics="true" pageId="36" pageNumber="715">tripartit-</emphasis>
(tripart), and refers to the trilobed paramere of the aedeagus.
</paragraph>
<paragraph id="6831E602D9590E39FF35E812FE46AA98" blockId="36.[144,758,1119,1172]" pageId="36" pageNumber="715">
<emphasis id="5AFA3A10D9590E39FF35E812FED7AA78" box="[144,289,1119,1140]" italics="true" pageId="36" pageNumber="715">Distribution:</emphasis>
At present known only from the 
<typeStatus id="B73558A0D9590E39FD60E812FD00AA78" box="[709,758,1119,1140]" pageId="36" pageNumber="715">type</typeStatus>
locality 
<collectingCountry id="1099A692D9590E39FF49E833FEC7AA98" box="[236,305,1150,1172]" name="China" pageId="36" pageNumber="715">China</collectingCountry>
(
<collectingRegion id="AA4A28E0D9590E39FEE6E833FE52AA98" box="[323,420,1150,1172]" country="China" name="Sichuan" pageId="36" pageNumber="715">Sichuan</collectingRegion>
).
</paragraph>
<paragraph id="6831E602D9590E39FF35E8F1FD00AB41" blockId="36.[144,759,1212,1357]" pageId="36" pageNumber="715">
<emphasis id="5AFA3A10D9590E39FF35E8F1FF0DAADD" box="[144,251,1212,1233]" italics="true" pageId="36" pageNumber="715">Remarks:</emphasis>
<taxonomicName id="AF8E9D81D9590E39FEADE8F1FD3FAADE" authority="Li &amp; Zhou" authorityName="Li &amp; Zhou" authorityYear="2011" box="[264,713,1212,1234]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="715" phylum="Arthropoda" rank="species" species="tripartitus" status="sp. nov.">
<emphasis id="5AFA3A10D9590E39FEADE8F1FDC8AADE" box="[264,574,1212,1234]" italics="true" pageId="36" pageNumber="715">Pseudohesperus tripartitus</emphasis>
Li &amp; Zhou
</taxonomicName>
<taxonomicNameLabel id="41C9876BD9590E39FD73E8F0FF4BAAFC" pageId="36" pageNumber="715" rank="species">sp. nov.</taxonomicNameLabel>
is closely related to 
<taxonomicName id="AF8E9D81D9590E39FE02E891FD38AAFD" authority="Li &amp; Zhou" authorityName="Li &amp; Zhou" authorityYear="2011" box="[423,718,1243,1265]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="715" phylum="Arthropoda" rank="species" species="pedatiformis" status="sp. nov.">
<emphasis id="5AFA3A10D9590E39FE02E891FDA5AAFC" box="[423,595,1243,1265]" italics="true" pageId="36" pageNumber="715">P. pedatiformis</emphasis>
Li &amp; Zhou
</taxonomicName>
<taxonomicNameLabel id="41C9876BD9590E39FD73E896FF4BAB03" pageId="36" pageNumber="715" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicName id="AF8E9D81D9590E39FF6FE8B7FD78AB1C" authority="Li &amp; Zhou" authorityName="Li &amp; Zhou" authorityYear="2011" box="[202,654,1274,1296]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="715" phylum="Arthropoda" rank="species" species="tripartitus" status="sp. nov.">
<emphasis id="5AFA3A10D9590E39FF6FE8B7FDF6AB03" box="[202,512,1274,1295]" italics="true" pageId="36" pageNumber="715">Pseudohesperus tripartitus</emphasis>
Li &amp; Zhou
</taxonomicName>
<taxonomicNameLabel id="41C9876BD9590E39FD39E8B7FD01AB03" box="[668,759,1274,1295]" pageId="36" pageNumber="715" rank="species">sp. nov.</taxonomicNameLabel>
differs from the latter species by distinctly smaller eyes and completely different paramere of aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>