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<document id="A71E28590514CBDE93E9599459D29242" ID-DOI="10.1111/j.1096-3642.2011.00731.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5442095" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630452512153" checkinUser="valdenar" docAuthor="Li, Liang &amp; Zhou, Hong-Zhang" docDate="2011" docId="E0275714D95C0E3EFCEDEABCFE99AADE" docLanguage="en" docName="j.1096-3642.2011.00731.x.pdf" docOrigin="Zoological Journal of the Linnean Society 163 (3)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2011.00731.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Pseudohesperus sparsipunctatus Li &amp; Zhou 2011, SP. NOV." docType="treatment" docVersion="3" lastPageNumber="714" masterDocId="1C1E2F6CD97D0E1DFFA5EC4DFFF6AE0C" masterDocTitle="Revision and phylogenetic assessment of the rove beetle genus Pseudohesperus Hayashi, with broad reference to the subtribe Philonthina (Coleoptera: Staphylinidae: Staphylinini)" masterLastPageNumber="722" masterPageNumber="679" pageNumber="712" updateTime="1699275715089" updateUser="plazi">
<mods:mods id="B986898EE0427EE00ED9C9B0A198C241" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="2E9354580CCC6F1675427CFEB281D3D2">Revision and phylogenetic assessment of the rove beetle genus Pseudohesperus Hayashi, with broad reference to the subtribe Philonthina (Coleoptera: Staphylinidae: Staphylinini)</mods:title>
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<mods:name id="D18B57EFE2E78159AD7D8C496D93D8D8" type="personal">
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<mods:namePart id="863EB7A286AF1998EFF26E46A084799B">Li, Liang</mods:namePart>
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<mods:namePart id="FB7E9764D4D080E42FFFC3BF50D76FC6">Zhou, Hong-Zhang</mods:namePart>
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<mods:typeOfResource id="12838153B4A2A1E80913859E703B613D">text</mods:typeOfResource>
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<mods:title id="BABCD706E55A30C602CC5F5CBF1484CE">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="D135E56CC4723E53E38948B4911A6A12">
<mods:date id="2A8FA89E8F661373F3605282F646E604">2011</mods:date>
<mods:detail id="E99978AE69E7CC83B74195096BFE2FA8" type="pubDate">
<mods:number id="FCA8B5080E7B7D7F72B75F8D3E72B6A4">2011-10-25</mods:number>
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<mods:url id="37FE724189C0891D64A76C7B3903C490">http://dx.doi.org/10.1111/j.1096-3642.2011.00731.x</mods:url>
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<mods:classification id="2D7D30461A075B06BD0E9B3BEDF25E5E">journal article</mods:classification>
<mods:identifier id="AF32A1B2555E24A2EB75509D13646B17" type="DOI">10.1111/j.1096-3642.2011.00731.x</mods:identifier>
<mods:identifier id="AF56BA0BB2034B2649E898C296167F11" type="ISSN">0024-4082</mods:identifier>
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</mods:mods>
<treatment id="E0275714D95C0E3EFCEDEABCFE99AADE" LSID="urn:lsid:plazi:treatment:E0275714D95C0E3EFCEDEABCFE99AADE" httpUri="http://treatment.plazi.org/id/E0275714D95C0E3EFCEDEABCFE99AADE" lastPageId="35" lastPageNumber="714" pageId="33" pageNumber="712">
<subSubSection id="2094B589D95C0E3CFCEDEABCFB1BA92B" pageId="33" pageNumber="712" type="nomenclature">
<paragraph id="6831E602D95C0E3CFCEDEABCFB1BA92B" blockId="33.[840,1427,1777,1831]" pageId="33" pageNumber="712">
<heading id="3379516ED95C0E3CFCEDEABCFB1BA92B" centered="true" fontSize="9" level="2" pageId="33" pageNumber="712" reason="4">
<taxonomicName id="AF8E9D81D95C0E3CFCEDEABCFA65A90A" authority="Li &amp; Zhou" authorityName="Li &amp; Zhou" authorityYear="2011" box="[840,1427,1777,1798]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="714" phylum="Arthropoda" rank="species" species="sparsipunctatus" status="sp. nov.">
<emphasis id="5AFA3A10D95C0E3CFCEDEABCFAFEA90A" bold="true" box="[840,1288,1777,1798]" italics="true" pageId="33" pageNumber="712">PSEUDOHESPERUS SPARSIPUNCTATUS</emphasis>
LI &amp; ZHOU
</taxonomicName>
<emphasis id="5AFA3A10D95C0E3CFC4AEB59FBA2A92A" bold="true" box="[1007,1108,1809,1830]" pageId="33" pageNumber="712">
<taxonomicNameLabel id="41C9876BD95C0E3CFC4AEB59FBA2A92A" box="[1007,1108,1809,1830]" pageId="33" pageNumber="712" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="F0B5FA87D95C0E3CFBC3EB5CFB13A92B" box="[1126,1253,1809,1831]" captionStart="Figure 28" captionStartId="34.[806,885,741,760]" captionTargetBox="[808,1421,198,707]" captionTargetId="figure-419@34.[806,1422,197,711]" captionTargetPageId="34" captionText="Figure 28. Pseudohesperus sparsipunctatus sp. nov. A, aedeagus, dorsal (parameral side) view; B, median lobe of aedeagus, lateral view; C, D, underside of paramere; E, tergite X of female; F, sternite VIII of male; G, stylus of tergite IX of male; H, sternite IX of male; I, tergite X of male. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5442149" httpUri="https://zenodo.org/record/5442149/files/figure.png" pageId="33" pageNumber="712">FIG. 28AI</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2094B589D95C0E3EFC9FEB75FE99AADE" lastPageId="35" lastPageNumber="714" pageId="33" pageNumber="712" type="materials_examined">
<paragraph id="6831E602D95C0E3CFC9FEB75FA57A960" blockId="33.[826,1441,1848,1900]" pageId="33" pageNumber="712">
<emphasis id="5AFA3A10D95C0E3CFC9FEB75FC16A941" box="[826,992,1848,1869]" italics="true" pageId="33" pageNumber="712">Type material:</emphasis>
<materialsCitation id="D8E6EC5FD95C0E3CFC4EEB75FA57A960" collectingDate="2008-08-09" country="China" elevation="400" latitude="30.2" location="Shunxi" longLatPrecision="1222" longitude="119.566666" municipality="Linan" pageId="33" pageNumber="712" specimenCount="1" specimenCount-male="1" stateProvince="Zhejiang" typeStatus="holotype">
<typeStatus id="B73558A0D95C0E3CFC4EEB75FBA3A942" box="[1003,1109,1848,1870]" pageId="33" pageNumber="712" type="holotype">Holotype</typeStatus>
:
<specimenCount id="7E882D8BD95C0E3CFBC7EB77FB8DA940" box="[1122,1147,1850,1868]" pageId="33" pageNumber="712" type="male"></specimenCount>
,
<collectingCountry id="1099A692D95C0E3CFB2EEB75FB22A942" box="[1163,1236,1848,1870]" name="China" pageId="33" pageNumber="712">China</collectingCountry>
:
<collectingRegion id="AA4A28E0D95C0E3CFB47EB75FABFA942" box="[1250,1353,1848,1870]" country="China" name="Zhejiang" pageId="33" pageNumber="712">Zhejiang</collectingRegion>
:
<collectingMunicipality id="88557C78D95C0E3CFAF3EB75FA6BA942" box="[1366,1437,1848,1870]" pageId="33" pageNumber="712">Linan</collectingMunicipality>
:
<location id="6D51B0D9D95C0E3CFC9FEB1BFC7AA960" LSID="urn:lsid:plazi:treatment:E0275714D95C0E3EFCEDEABCFE99AADE:6D51B0D9D95C0E3CFC9FEB1BFC7AA960" box="[826,908,1878,1900]" country="China" latitude="30.2" longLatPrecision="1222" longitude="119.566666" municipality="Linan" name="Shunxi" pageId="33" pageNumber="712" stateProvince="Zhejiang">Shunxi</location>
(
<geoCoordinate id="0DBA80C5D95C0E3CFC06EB1BFBF6A960" box="[931,1024,1877,1900]" degrees="30" direction="north" minutes="12" orientation="latitude" pageId="33" pageNumber="712" precision="925" value="30.2">30°12N</geoCoordinate>
,
<geoCoordinate id="0DBA80C5D95C0E3CFBB1EB1BFB8DA960" box="[1044,1147,1877,1900]" degrees="119" direction="east" minutes="34" orientation="longitude" pageId="33" pageNumber="712" precision="925" value="119.566666">119°34E</geoCoordinate>
),
<elevation id="E3A30131D95C0E3CFB32EB1BFAE6A960" box="[1175,1296,1878,1900]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="33" pageNumber="712" unit="m" value="400.0">
<quantity id="AF764BE7D95C0E3CFB32EB1BFB2BA960" box="[1175,1245,1878,1900]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="33" pageNumber="712" unit="m" value="400.0">400 m</quantity>
asl
</elevation>
,
<date id="1C30C0C2D95C0E3CFA81EB1BFA6AA960" box="[1316,1436,1878,1900]" pageId="33" pageNumber="712" value="2008-08-09">
<collectingDate id="0C74392AD95C0E3CFA81EB1BFA6AA960" box="[1316,1436,1878,1900]" pageId="33" pageNumber="712" value="2008-08-09">9.viii.2008</collectingDate>
</date>
,
</materialsCitation>
</paragraph>
<caption id="3CF1B68AD95F0E3FFC83EEA8FBB3AD87" ID-DOI="http://doi.org/10.5281/zenodo.5442149" ID-Zenodo-Dep="5442149" httpUri="https://zenodo.org/record/5442149/files/figure.png" pageId="34" pageNumber="713" startId="34.[806,885,741,760]" targetBox="[808,1421,198,707]" targetPageId="34">
<paragraph id="6831E602D95F0E3FFC83EEA8FBB3AD87" blockId="34.[806,1421,741,907]" pageId="34" pageNumber="713">
<emphasis id="5AFA3A10D95F0E3FFC83EEA8FA7BACF4" bold="true" box="[806,1421,741,760]" pageId="34" pageNumber="713">
Figure 28.
<taxonomicName id="AF8E9D81D95F0E3FFC0DEEA8FADCACF4" authority="Li &amp; Zhou, 2011" authorityName="Li &amp; Zhou" authorityYear="2011" box="[936,1322,741,760]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="713" phylum="Arthropoda" rank="species" species="sparsipunctatus" status="sp. nov.">
<emphasis id="5AFA3A10D95F0E3FFC0DEEA8FADCACF4" bold="true" box="[936,1322,741,760]" italics="true" pageId="34" pageNumber="713">Pseudohesperus sparsipunctatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="41C9876BD95F0E3FFA96EEA8FA7BACF4" box="[1331,1421,741,760]" pageId="34" pageNumber="713" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, aedeagus, dorsal (parameral side) view; B, median lobe of aedeagus, lateral view; C, D, underside of paramere; E, tergite X of female; F, sternite VIII of male; G, stylus of tergite IX of male; H, sternite IX of male; I, tergite X of male. Scale bars = 0.5 mm.
</paragraph>
</caption>
<caption id="3CF1B68AD95F0E3FFF35EF9FFEDDAABE" ID-DOI="http://doi.org/10.5281/zenodo.5442151" ID-Zenodo-Dep="5442151" httpUri="https://zenodo.org/record/5442151/files/figure.png" pageId="34" pageNumber="713" startId="34.[144,223,978,997]" targetBox="[146,758,197,945]" targetPageId="34">
<paragraph id="6831E602D95F0E3FFF35EF9FFEDDAABE" blockId="34.[144,759,978,1202]" pageId="34" pageNumber="713">
<emphasis id="5AFA3A10D95F0E3FFF35EF9FFEF3ADE9" bold="true" box="[144,261,978,997]" pageId="34" pageNumber="713">Figure 27.</emphasis>
<taxonomicName id="AF8E9D81D95F0E3FFEB4EF9FFD37ADE9" authority="(Sharp)" authorityName="" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[273,705,978,997]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="713" phylum="Arthropoda" rank="species" species="rutiliventris">
<emphasis id="5AFA3A10D95F0E3FFEB4EF9FFDA4ADE9" box="[273,594,978,997]" italics="true" pageId="34" pageNumber="713">Pseudohesperus rutiliventris</emphasis>
(Sharp)
</taxonomicName>
. A, aedeagus, dorsal (parameral side) view; B, median lobe of aedeagus, lateral view; C, D, underside of paramere; E, sternite IX of male; F, sternite VIII of male; G, stylus of tergite IX of male; H, gonocoxites of genital segment of female; I, tergite X of female; J, tergite VIII of male; K, sternite VIII of female; L, tergite X of male. Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="6831E602D95F0E3FFF35E8A1FDB3AB2D" blockId="34.[144,759,1260,1313]" pageId="34" pageNumber="713">
<materialsCitation id="D8E6EC5FD95F0E3FFF35E8A1FDB7AB2D" collectingDate="2008-08-09" country="China" elevation="400" latitude="30.2" location="Shunxi" longLatPrecision="1222" longitude="119.566666" municipality="Linan" pageId="34" pageNumber="713" specimenCount="10" specimenCount-female="6" specimenCount-male="4" stateProvince="Zhejiang" typeStatus="paratype">
Zhou Yulingzi collected (IZ-CAS);
<typeStatus id="B73558A0D95F0E3FFDB7E8A1FD7FAB0D" box="[530,649,1260,1281]" pageId="34" pageNumber="713" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="7E882D8BD95F0E3FFD31E8A1FD2CAB0D" box="[660,730,1260,1281]" pageId="34" pageNumber="713" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="7E882D8BD95F0E3FFD4CE8A1FF4FAB13" pageId="34" pageNumber="713" type="female">6 ♀♀</specimenCount>
, same data as holotype (IZ-CAS)
</materialsCitation>
.
</paragraph>
<paragraph id="6831E602D95F0E3FFF35E91CFE51A831" blockId="34.[144,759,1361,1903]" pageId="34" pageNumber="713">
<emphasis id="5AFA3A10D95F0E3FFF35E91CFEEEAB6A" box="[144,280,1361,1382]" italics="true" pageId="34" pageNumber="713">Description:</emphasis>
Head and pronotum black, shining. Antennae black-brown, with antennomeres IIII to various extent reddish-brown, antennomere XI or antennomeres XXI sometimes reddish-brown. Elytra black with posterior margin markedly reddish-brown. Scutellum black. Abdomen black with strongly iridescent lustre. Mandibles, maxillary and labial palpi, and legs reddish-brown.
</paragraph>
<paragraph id="6831E602D95F0E3FFF0DEA08FBF8AA5F" blockId="34.[144,759,1361,1903]" lastBlockId="34.[806,1422,963,1903]" pageId="34" pageNumber="713">
Body
<quantity id="AF764BE7D95F0E3FFF53EA0BFE60A857" box="[246,406,1606,1627]" metricMagnitude="-3" metricUnit="m" metricValue="9.39" metricValueMax="9.8" metricValueMin="8.98" pageId="34" pageNumber="713" unit="mm" value="9.39" valueMax="9.8" valueMin="8.98">8.989.80 mm</quantity>
long (
<collectionCode id="0E9F7EC7D95F0E3FFE53EA0BFDD9A857" box="[502,559,1606,1627]" pageId="34" pageNumber="713">HPL</collectionCode>
:
<quantity id="AF764BE7D95F0E3FFDE2EA08FD1DA857" box="[583,747,1605,1627]" metricMagnitude="-3" metricUnit="m" metricValue="2.815" metricValueMax="2.94" metricValueMin="2.69" pageId="34" pageNumber="713" unit="mm" value="2.815" valueMax="2.94" valueMin="2.69">2.692.94 mm</quantity>
). Head of rounded quadrangular shape,
<quantity id="AF764BE7D95F0E3FFDF3EA29FD01A875" box="[598,759,1636,1657]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.22" metricValueMin="0.98" pageId="34" pageNumber="713" unit="mm" value="1.1" valueMax="1.22" valueMin="0.98">0.981.22 mm</quantity>
long,
<quantity id="AF764BE7D95F0E3FFF77EACFFE84A894" box="[210,370,1666,1688]" metricMagnitude="-3" metricUnit="m" metricValue="1.43" metricValueMax="1.55" metricValueMin="1.31" pageId="34" pageNumber="713" unit="mm" value="1.43" valueMax="1.55" valueMin="1.31">1.311.55 mm</quantity>
wide, distinctly wider than long (
<collectionCode id="0E9F7EC7D95F0E3FFF32EAECFF33A8BA" box="[151,197,1697,1718]" pageId="34" pageNumber="713">HW</collectionCode>
:
<collectionCode id="0E9F7EC7D95F0E3FFF7CEAEFFF08A8BB" box="[217,254,1698,1719]" pageId="34" pageNumber="713">HL</collectionCode>
= 1.231.33). Tempora
<quantity id="AF764BE7D95F0E3FFDB7EAECFD45A8BB" box="[530,691,1697,1719]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" metricValueMax="3.3" metricValueMin="2.5" pageId="34" pageNumber="713" unit="mm" value="0.29" valueMax="0.33" valueMin="0.25">0.250.33 mm</quantity>
long, almost evenly rounded, sparsely and coarsely punctate; eyes very large, moderately prominent,
<quantity id="AF764BE7D95F0E3FFD1CEA92FF04A91E" metricMagnitude="-4" metricUnit="m" metricValue="5.3" metricValueMax="5.699999999999999" metricValueMin="4.9" pageId="34" pageNumber="713" unit="mm" value="0.53" valueMax="0.57" valueMin="0.49">0.49 0.57 mm</quantity>
long, 1.752.33 times as long as tempora. Lateral portions of head with scattered, large setiferous punctuation; vertex largely impunctate. Entire head without microsculpture. Antennae long, antennomere I very long, thickened towards apex, antennomere III much longer than antennomere II, following antennomeres gradually decreasing in length, but all longer than wide, antennomere XI obliquely truncated.
</paragraph>
<paragraph id="6831E602D95F0E3FFC9AE811FADFAB07" blockId="34.[806,1422,963,1903]" pageId="34" pageNumber="713">
Pronotum slightly narrowed anteriad,
<quantity id="AF764BE7D95F0E3FFAEAE811FC7FAA9C" metricMagnitude="-3" metricUnit="m" metricValue="1.5499999999999998" metricValueMax="1.63" metricValueMin="1.47" pageId="34" pageNumber="713" unit="mm" value="1.5499999999999998" valueMax="1.63" valueMin="1.47">1.47 1.63 mm</quantity>
long,
<quantity id="AF764BE7D95F0E3FFC41E836FB73AA9C" box="[996,1157,1147,1168]" metricMagnitude="-3" metricUnit="m" metricValue="1.5899999999999999" metricValueMax="1.71" metricValueMin="1.47" pageId="34" pageNumber="713" unit="mm" value="1.5899999999999999" valueMax="1.71" valueMin="1.47">1.471.71 mm</quantity>
wide, slightly wider than head (
<collectionCode id="0E9F7EC7D95F0E3FFC08E8D4FC21AAA2" box="[941,983,1177,1198]" country="Thailand" name="Paleontological Collections" pageId="34" pageNumber="713">PW</collectionCode>
:
<collectionCode id="0E9F7EC7D95F0E3FFC4EE8D4FBEEAAA2" box="[1003,1048,1177,1198]" pageId="34" pageNumber="713">HW</collectionCode>
= 1.101.19); each side of broad impunctate midline with 1215 scatter punctures, sometimes eight to ten punctures forming an irregular, dorsal row, disc without microsculpture.
</paragraph>
<paragraph id="6831E602D95F0E3FFC9AE95EFC56ABCE" blockId="34.[806,1422,963,1903]" pageId="34" pageNumber="713">
Elytra
<quantity id="AF764BE7D95F0E3FFC39E959FBCBAB25" box="[924,1085,1300,1321]" metricMagnitude="-3" metricUnit="m" metricValue="1.92" metricValueMax="2.04" metricValueMin="1.8" pageId="34" pageNumber="713" unit="mm" value="1.92" valueMax="2.04" valueMin="1.8">1.802.04 mm</quantity>
long,
<quantity id="AF764BE7D95F0E3FFB39E959FACBAB25" box="[1180,1341,1300,1321]" metricMagnitude="-3" metricUnit="m" metricValue="2.205" metricValueMax="2.37" metricValueMin="2.04" pageId="34" pageNumber="713" unit="mm" value="2.205" valueMax="2.37" valueMin="2.04">2.042.37 mm</quantity>
wide, along sides 1.161.25 times as long as pronotum, densely and finely punctate, punctures separated by one to two times their diameter. Scutellum large, triangular, densely and finely punctate, and pubescent.
</paragraph>
<paragraph id="6831E602D95F0E3FFC9AE986FB7BA8D9" blockId="34.[806,1422,963,1903]" pageId="34" pageNumber="713">
Abdomen a little narrowed behind, widest
<quantity id="AF764BE7D95F0E3FFAEAE986FC7DABF3" metricMagnitude="-3" metricUnit="m" metricValue="1.88" metricValueMax="1.96" metricValueMin="1.8" pageId="34" pageNumber="713" unit="mm" value="1.88" valueMax="1.96" valueMin="1.8">1.80 1.96 mm</quantity>
, densely and finely punctate, punctures separated by two to three times their diameter at base on tergites IIIV and punctures separated by less than one puncture diameter at base on tergites VIVII, punctation gradually becoming sparser toward apex of each tergite; tergites IIIV with two basal carinae, elevated area between basal carinae densely and coarsely punctate.
</paragraph>
<paragraph id="6831E602D95F0E3EFC9AEA93FD79AF5A" blockId="34.[806,1422,963,1903]" lastBlockId="35.[164,779,198,740]" lastPageId="35" lastPageNumber="714" pageId="34" pageNumber="713">
Male: Tergite VIII with arcuate apex. Sternite VIII with moderately wide, obtusely triangular medioapical emargination, partly filled by semimembranous extension (
<figureCitation id="F0B5FA87D95F0E3FFC03EB77FBFEA95C" box="[934,1032,1850,1872]" captionStart="Figure 28" captionStartId="34.[806,885,741,760]" captionTargetBox="[808,1421,198,707]" captionTargetId="figure-419@34.[806,1422,197,711]" captionTargetPageId="34" captionText="Figure 28. Pseudohesperus sparsipunctatus sp. nov. A, aedeagus, dorsal (parameral side) view; B, median lobe of aedeagus, lateral view; C, D, underside of paramere; E, tergite X of female; F, sternite VIII of male; G, stylus of tergite IX of male; H, sternite IX of male; I, tergite X of male. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5442149" httpUri="https://zenodo.org/record/5442149/files/figure.png" pageId="34" pageNumber="713">Fig. 28F</figureCitation>
). Genital segment with stylus of tergite IX simple, densely setose apically (
<figureCitation id="F0B5FA87D95F0E3FFAB9EB14FA77A963" box="[1308,1409,1881,1903]" captionStart="Figure 28" captionStartId="34.[806,885,741,760]" captionTargetBox="[808,1421,198,707]" captionTargetId="figure-419@34.[806,1422,197,711]" captionTargetPageId="34" captionText="Figure 28. Pseudohesperus sparsipunctatus sp. nov. A, aedeagus, dorsal (parameral side) view; B, median lobe of aedeagus, lateral view; C, D, underside of paramere; E, tergite X of female; F, sternite VIII of male; G, stylus of tergite IX of male; H, sternite IX of male; I, tergite X of male. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5442149" httpUri="https://zenodo.org/record/5442149/files/figure.png" pageId="34" pageNumber="713">Fig. 28G</figureCitation>
). Sternite IX with asymmetrical basal portion, apex deeply emarginate, each lobe prolong, forming modified rod-like extensions (
<figureCitation id="F0B5FA87D95E0E3EFE5DED4EFDA9AF15" box="[504,607,259,281]" captionStart="Figure 28" captionStartId="34.[806,885,741,760]" captionTargetBox="[808,1421,198,707]" captionTargetId="figure-419@34.[806,1422,197,711]" captionTargetPageId="34" captionText="Figure 28. Pseudohesperus sparsipunctatus sp. nov. A, aedeagus, dorsal (parameral side) view; B, median lobe of aedeagus, lateral view; C, D, underside of paramere; E, tergite X of female; F, sternite VIII of male; G, stylus of tergite IX of male; H, sternite IX of male; I, tergite X of male. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5442149" httpUri="https://zenodo.org/record/5442149/files/figure.png" pageId="35" pageNumber="714">Fig. 28H</figureCitation>
). Tergite X triangular, inconspicuously emarginate at apex, with variable number of apical setae (
<figureCitation id="F0B5FA87D95E0E3EFD82ED0DFD74AF5A" box="[551,642,320,342]" captionStart="Figure 28" captionStartId="34.[806,885,741,760]" captionTargetBox="[808,1421,198,707]" captionTargetId="figure-419@34.[806,1422,197,711]" captionTargetPageId="34" captionText="Figure 28. Pseudohesperus sparsipunctatus sp. nov. A, aedeagus, dorsal (parameral side) view; B, median lobe of aedeagus, lateral view; C, D, underside of paramere; E, tergite X of female; F, sternite VIII of male; G, stylus of tergite IX of male; H, sternite IX of male; I, tergite X of male. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5442149" httpUri="https://zenodo.org/record/5442149/files/figure.png" pageId="35" pageNumber="714">Fig. 28I</figureCitation>
).
</paragraph>
<paragraph id="6831E602D95E0E3EFF19ED12FEC9AC46" blockId="35.[164,779,198,740]" pageId="35" pageNumber="714">
Aedeagus with median lobe distinctly exceeding paramere, narrowed into obtusely rounded apex (
<figureCitation id="F0B5FA87D95E0E3EFF09EDD1FEE6AFBE" box="[172,272,412,434]" captionStart="Figure 28" captionStartId="34.[806,885,741,760]" captionTargetBox="[808,1421,198,707]" captionTargetId="figure-419@34.[806,1422,197,711]" captionTargetPageId="34" captionText="Figure 28. Pseudohesperus sparsipunctatus sp. nov. A, aedeagus, dorsal (parameral side) view; B, median lobe of aedeagus, lateral view; C, D, underside of paramere; E, tergite X of female; F, sternite VIII of male; G, stylus of tergite IX of male; H, sternite IX of male; I, tergite X of male. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5442149" httpUri="https://zenodo.org/record/5442149/files/figure.png" pageId="35" pageNumber="714">Fig. 28A</figureCitation>
); in lateral view, apical portion of median lobe slightly dilated (
<figureCitation id="F0B5FA87D95E0E3EFE3EEDF6FE08AFDD" box="[411,510,443,465]" captionStart="Figure 28" captionStartId="34.[806,885,741,760]" captionTargetBox="[808,1421,198,707]" captionTargetId="figure-419@34.[806,1422,197,711]" captionTargetPageId="34" captionText="Figure 28. Pseudohesperus sparsipunctatus sp. nov. A, aedeagus, dorsal (parameral side) view; B, median lobe of aedeagus, lateral view; C, D, underside of paramere; E, tergite X of female; F, sternite VIII of male; G, stylus of tergite IX of male; H, sternite IX of male; I, tergite X of male. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5442149" httpUri="https://zenodo.org/record/5442149/files/figure.png" pageId="35" pageNumber="714">Fig. 28B</figureCitation>
); paramere widened at subapical portion, narrowed into obtusely pointed apex, face adjacent to median lobe with peg setae irregularly arranged along apical portion of paramere (
<figureCitation id="F0B5FA87D95E0E3EFF09EE78FEC5AC46" box="[172,307,565,587]" captionStart="Figure 28" captionStartId="34.[806,885,741,760]" captionTargetBox="[808,1421,198,707]" captionTargetId="figure-419@34.[806,1422,197,711]" captionTargetPageId="34" captionText="Figure 28. Pseudohesperus sparsipunctatus sp. nov. A, aedeagus, dorsal (parameral side) view; B, median lobe of aedeagus, lateral view; C, D, underside of paramere; E, tergite X of female; F, sternite VIII of male; G, stylus of tergite IX of male; H, sternite IX of male; I, tergite X of male. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5442149" httpUri="https://zenodo.org/record/5442149/files/figure.png" pageId="35" pageNumber="714">Fig. 28C, D</figureCitation>
).
</paragraph>
<paragraph id="6831E602D95E0E3EFF19EE19FE03ACE8" blockId="35.[164,779,198,740]" pageId="35" pageNumber="714">
Female: Sternite VIII with arcuate apex, lacking medioapical emargination. Genital segment with second gonocoxites moderately long, each with minute stylus bearing two long apical setae. Tergite X subtruncated at apex (
<figureCitation id="F0B5FA87D95E0E3EFE20EE83FE1FACE8" box="[389,489,718,740]" captionStart="Figure 28" captionStartId="34.[806,885,741,760]" captionTargetBox="[808,1421,198,707]" captionTargetId="figure-419@34.[806,1422,197,711]" captionTargetPageId="34" captionText="Figure 28. Pseudohesperus sparsipunctatus sp. nov. A, aedeagus, dorsal (parameral side) view; B, median lobe of aedeagus, lateral view; C, D, underside of paramere; E, tergite X of female; F, sternite VIII of male; G, stylus of tergite IX of male; H, sternite IX of male; I, tergite X of male. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5442149" httpUri="https://zenodo.org/record/5442149/files/figure.png" pageId="35" pageNumber="714">Fig. 28E</figureCitation>
).
</paragraph>
<paragraph id="6831E602D95E0E3EFF01EF40FE98AD72" blockId="35.[164,779,781,895]" pageId="35" pageNumber="714">
<emphasis id="5AFA3A10D95E0E3EFF01EF40FED4AD2E" box="[164,290,781,802]" italics="true" pageId="35" pageNumber="714">Etymology:</emphasis>
The specific epithet is derived from a combination of the Latin words
<emphasis id="5AFA3A10D95E0E3EFD8DEF61FD9BAD4D" box="[552,621,812,833]" italics="true" pageId="35" pageNumber="714">spars-</emphasis>
(sparse) and
<emphasis id="5AFA3A10D95E0E3EFF01EF06FF1AAD6C" box="[164,236,843,864]" italics="true" pageId="35" pageNumber="714">punct-</emphasis>
(puncture), and refers to the sparse punctation on the pronotum.
</paragraph>
<caption id="3CF1B68AD95E0E3EFC9FEF2FFB1EAA04" ID-DOI="http://doi.org/10.5281/zenodo.5442153" ID-Zenodo-Dep="5442153" httpUri="https://zenodo.org/record/5442153/files/figure.png" pageId="35" pageNumber="714" startId="35.[826,905,866,885]" targetBox="[827,1441,199,835]" targetPageId="35">
<paragraph id="6831E602D95E0E3EFC9FEF2FFB1EAA04" blockId="35.[826,1441,866,1032]" pageId="35" pageNumber="714">
<emphasis id="5AFA3A10D95E0E3EFC9FEF2FFA8FAD79" bold="true" box="[826,1401,866,885]" pageId="35" pageNumber="714">
Figure 29.
<taxonomicName id="AF8E9D81D95E0E3EFC19EF2FFAF3AD79" authority="Li &amp; Zhou, 2011" authorityName="Li &amp; Zhou" authorityYear="2011" box="[956,1285,866,885]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="714" phylum="Arthropoda" rank="species" species="tripartitus" status="sp. nov.">
<emphasis id="5AFA3A10D95E0E3EFC19EF2FFAF3AD79" bold="true" box="[956,1285,866,885]" italics="true" pageId="35" pageNumber="714">Pseudohesperus tripartitus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="41C9876BD95E0E3EFAB2EF2FFA8FAD79" box="[1303,1401,866,885]" pageId="35" pageNumber="714" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, aedeagus, dorsal (parameral side) view; B, median lobe of aedeagus, lateral view; C, underside of paramere; D, sternite IX of male; E, sternite VIII of male; F, tergite X of male; G, tergite X of female; H, gonocoxites of genital segment of female. Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="6831E602D95E0E3EFF01EFEAFEE9ADD0" blockId="35.[164,778,935,988]" pageId="35" pageNumber="714">
<emphasis id="5AFA3A10D95E0E3EFF01EFEAFEC3ADB0" box="[164,309,935,956]" italics="true" pageId="35" pageNumber="714">Distribution:</emphasis>
At present only known from
<collectingCountry id="1099A692D95E0E3EFD60EFEAFCFCADB1" box="[709,778,935,957]" name="China" pageId="35" pageNumber="714">China</collectingCountry>
(
<collectingRegion id="AA4A28E0D95E0E3EFF0EEF8BFEE5ADD0" box="[171,275,966,988]" country="China" name="Zhejiang" pageId="35" pageNumber="714">Zhejiang</collectingRegion>
).
</paragraph>
<paragraph id="6831E602D95E0E3EFF01E848FE99AADE" blockId="35.[164,778,1029,1234]" pageId="35" pageNumber="714">
<emphasis id="5AFA3A10D95E0E3EFF01E848FEF9AA16" box="[164,271,1029,1050]" italics="true" pageId="35" pageNumber="714">Remarks:</emphasis>
<taxonomicName id="AF8E9D81D95E0E3EFEBDE848FCFCAA17" authority="Li &amp; Zhou" authorityName="Li &amp; Zhou" authorityYear="2011" box="[280,778,1029,1051]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="714" phylum="Arthropoda" rank="species" species="sparsipunctatus" status="sp. nov.">
<emphasis id="5AFA3A10D95E0E3EFEBDE848FD7DAA16" box="[280,651,1029,1050]" italics="true" pageId="35" pageNumber="714">Pseudohesperus sparsipunctatus</emphasis>
Li &amp; Zhou
</taxonomicName>
<taxonomicNameLabel id="41C9876BD95E0E3EFF01E869FF0BAA35" box="[164,253,1060,1081]" pageId="35" pageNumber="714" rank="species">sp. nov.</taxonomicNameLabel>
is closely related to
<taxonomicName id="AF8E9D81D95E0E3EFE5DE869FF10AA54" authority="(Kraatz, 1859)" baseAuthorityName="Kraatz" baseAuthorityYear="1859" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="714" phylum="Arthropoda" rank="species" species="eustilbus">
<emphasis id="5AFA3A10D95E0E3EFE5DE869FDFBAA35" box="[504,525,1060,1081]" italics="true" pageId="35" pageNumber="714">P.</emphasis>
cf.
<emphasis id="5AFA3A10D95E0E3EFD99E86EFD54AA34" box="[572,674,1059,1080]" italics="true" pageId="35" pageNumber="714">eustilbus</emphasis>
(
<bibRefCitation id="0C1F9BF3D95E0E3EFD10E869FF28AA54" author="Kraatz G" pageId="35" pageNumber="714" pagination="1 - 196" refId="ref20995" refString="Kraatz G. 1859. Die Staphylinen-Fauna von Ostindien, insbesondere der Insel Ceylan. Archiv fur Naturgeschichte 25: 1 - 196." type="journal article" year="1859">Kraatz, 1859</bibRefCitation>
)
</taxonomicName>
.
<taxonomicName id="AF8E9D81D95E0E3EFF5FE80FFCFCAA54" authority="Li &amp; Zhou" authorityName="Li &amp; Zhou" authorityYear="2011" box="[250,778,1090,1112]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="714" phylum="Arthropoda" rank="species" species="sparsipunctatus" status="sp. nov.">
<emphasis id="5AFA3A10D95E0E3EFF5FE80FFD82AA54" box="[250,628,1090,1112]" italics="true" pageId="35" pageNumber="714">Pseudohesperus sparsipunctatus</emphasis>
Li &amp; Zhou
</taxonomicName>
<taxonomicNameLabel id="41C9876BD95E0E3EFF01E82CFEFEAA7A" box="[164,264,1121,1142]" pageId="35" pageNumber="714" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="AF8E9D81D95E0E3EFE6DE82CFDAEAA7A" baseAuthorityName="Kraatz" baseAuthorityYear="1859" box="[456,600,1121,1142]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="714" phylum="Arthropoda" rank="species" species="eustilbus">
<emphasis id="5AFA3A10D95E0E3EFE6DE82CFDAEAA7A" box="[456,600,1121,1142]" italics="true" pageId="35" pageNumber="714">P. eustilbus</emphasis>
</taxonomicName>
by distinctly sparser punctures on pronotum, much wider paramere, and larger number of peg setae on underside of paramere.
</paragraph>
</subSubSection>
</treatment>
</document>