treatments-xml/data/D8/8E/E9/D88EE9CDA2485D8B8CACE550AEE4BE79.xml

<document ID-DOI="http://dx.doi.org/10.3897/contrib.entomol.74.e114543" ID-Pensoft-Pub="2511-6428-1-13" ID-Pensoft-UUID="F4C1C6A2DFD95D3BAF5AF1CE09226837" ID-ZooBank="589077DFF8BB45ABB30D7E0DE1C307FE" ModsDocID="2511-6428-74-1-13" checkinTime="1706792588976" checkinUser="pensoft" docAuthor="Irmler, Ulrich" docDate="2024" docId="D88EE9CDA2485D8B8CACE550AEE4BE79" docLanguage="en" docName="ContribEntomol 74(1): 13-34" docOrigin="Contributions to Entomology 74 (1)" docPubDate="2024-01-31" docSource="http://dx.doi.org/10.3897/contrib.entomol.74.e114543" docTitle="Somoleptus luteicornis Irmler 2024, sp. nov." docType="treatment" docUuid="9F985E1F-6421-4BD2-AF90-4412F6ED7724" docUuidSource="ZooBank" docVersion="1" id="F4C1C6A2DFD95D3BAF5AF1CE09226837" lastPageNumber="13" masterDocId="F4C1C6A2DFD95D3BAF5AF1CE09226837" masterDocTitle="Additions to Neotropical species of the genera Lithocharodes Sharp, 1881 and Somoleptus Sharp, 1881 (Coleoptera, Staphylinidae)" masterLastPageNumber="34" masterPageNumber="13" pageNumber="13" updateTime="1706792588976" updateUser="pensoft">
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<mods:title id="D367FBE461E9AFEC7D7A994CB88B36CD">Additions to Neotropical species of the genera Lithocharodes Sharp, 1881 and Somoleptus Sharp, 1881 (Coleoptera, Staphylinidae)</mods:title>
</mods:titleInfo>
<mods:name id="F02795AEC4A7CAE39785DFCE492B4BC4" type="personal">
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<mods:namePart id="8632BBB4928C7F1CA27493350387B438">Irmler, Ulrich</mods:namePart>
<mods:affiliation id="E23BE10E07C135598AD867310CD802AE">Institute for Ecosystem Research, University of Kiel, Olshausenstrasse 40, 24098 Kiel, Germany</mods:affiliation>
<mods:nameIdentifier id="BF55A9CD9783529F00E6FA407E78DDCD" type="email">uirmler@ecology.uni-kiel.de</mods:nameIdentifier>
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<mods:titleInfo id="A2C00E4E7150F3AACEF673DBB31BE0E8">
<mods:title id="4341D2FA601BF23F85D29BFF8E497D76">Contributions to Entomology</mods:title>
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<mods:part id="EA9B91289F827D7679F50801F0EF36B5">
<mods:date id="879F4866A8E5D28750C05878D3A1A0FA">2024</mods:date>
<mods:detail id="9D19D012DD20D67ADE7833AB19456D31" type="pubDate">
<mods:number id="71E500E65D6E8F330C1FA63BFDED6722">2024-01-31</mods:number>
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<mods:number id="1C428273DFF4AD793DADEBAC392358C7">74</mods:number>
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<mods:url id="C0D3F3CDDF1FC53C5834AEC51978ADC9">http://dx.doi.org/10.3897/contrib.entomol.74.e114543</mods:url>
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<mods:classification id="8E3BA79B41CEE7280037BB750EDBBD35">journal article</mods:classification>
<mods:identifier id="2C1E5BE934F0F946AA7687718DF9834E" type="DOI">http://dx.doi.org/10.3897/contrib.entomol.74.e114543</mods:identifier>
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<treatment id="D88EE9CDA2485D8B8CACE550AEE4BE79" LSID="urn:lsid:zoobank.org:act:9F985E1F-6421-4BD2-AF90-4412F6ED7724" httpUri="http://treatment.plazi.org/id/D88EE9CDA2485D8B8CACE550AEE4BE79" lastPageNumber="13" pageId="0" pageNumber="13">
<subSubSection id="0481B5B444AED35622078A3BB5EA03FB" pageId="0" pageNumber="13" type="nomenclature">
<paragraph id="4ECE725A5A88A43AEC2E20248FBB684A" pageId="0" pageNumber="13">
<taxonomicName id="A9EFB70F42DA53E2D56761D7EDDBE1CF" LSID="https://zoobank.org/9F985E1F-6421-4BD2-AF90-4412F6ED7724" authority="Irmler, 2024" authorityName="Irmler" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Somoleptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Somoleptus luteicornis" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="luteicornis" status="sp. nov.">Somoleptus luteicornis</taxonomicName>
<taxonomicNameLabel id="AD20CDE6C8AE963AE8D8B84A28D9723D" pageId="0" pageNumber="13">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="FA8ACEDE2FCF368059A705B56AD85316" pageId="0" pageNumber="13" type="description">
<paragraph id="99EC9BEFF66B5FD927C5E12B2D08AF0F" pageId="0" pageNumber="13">
<figureCitation id="28FD1CD36C95679D02522F2EBA79E533" captionStart="Figures 13–16" captionStartId="F4" captionText="Figures 13 - 16. 13. Somoleptus densiceps; 14. S. guianensis; 15. S. luteicornis; 16. S. paramocola. a. Aedeagus; b. Paramere; c. Sternite VII; d. Tergite VII. Scale bars: 0.5 mm (a, c, d); 0.1 mm (b)." figureDoi="10.3897/contrib.entomol.74.e114543.figures13-16" httpUri="https://binary.pensoft.net/fig/975718" pageId="0" pageNumber="13">Figs 15a-d</figureCitation>
<figureCitation id="DD7B4C7420C16B4D156617FAB66B8B68" captionStart="Figure 21" captionStartId="F8" captionText="Figure 21. A. Fore-body of Somoleptus densiceps; B. S. guianensis; C. S. luteicornis; D. S. paramocola; E. S. puntarenae; F. S. umicola. Scale bars: 0.5 mm." figureDoi="10.3897/contrib.entomol.74.e114543.figure21" httpUri="https://binary.pensoft.net/fig/975722" pageId="0" pageNumber="13">, 21C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="3895A2DCEB7BAE3C0720661668D744DC" pageId="0" pageNumber="13" type="materials_examined">
<paragraph id="34D93BD952B2DC0BD4ECAB8368A81F83" pageId="0" pageNumber="13">Material examined.</paragraph>
<paragraph id="BB4A8A12523620E941E76F702EA7023A" pageId="0" pageNumber="13">
<materialsCitation id="548AEBBB89ED04CA0E35ADFB0B10DEBE" collectingDate="1998-06-21" collectorName="Berlese, R. Anderson" country="COSTA RICA" elevation="1720" latitude="8.973888" location="Puntarenas" longLatPrecision="21" longitude="-82.83445" specimenCount="1" stateProvince="Puntarenas" typeStatus="Holotype">
<emphasis id="427D7851CDD360BC631C3A381EDAEB21" bold="true" italics="true" pageId="0" pageNumber="13">
<typeStatus id="CFDCE09CCD35999C9E355E38714FCF7F" type="holotype">Holotype</typeStatus>
</emphasis>
: 
<collectingCountry id="35C681474F3DBDF0FB662A45CB5B8C20" name="Costa Rica">COSTA RICA</collectingCountry>
male; 
<collectingRegion id="2188D2D35CA1C940762BF9324E2D6070" country="Costa Rica" name="Puntarenas">Puntarenas</collectingRegion>
; Estac. Biol. Las Alturas; 
<quantity id="9ED15C9A634494617A269E661F1BA27F" metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="km" value="2.0">2 km</quantity>
NE; 
<geoCoordinate id="3F7D7C12512525AA975AFECE533C02D8" degrees="8" direction="north" minutes="58" orientation="latitude" precision="15" seconds="26" value="8.973888">8°58'26&quot;N</geoCoordinate>
, 
<geoCoordinate id="617C3C701346615FBAD6E69FFC98330B" degrees="82" direction="west" minutes="50" orientation="longitude" precision="15" seconds="4" value="-82.83445">82°50'4&quot;W</geoCoordinate>
; 
<quantity id="5F04AE8FC281B980A2C536E32BEEA28D" metricMagnitude="3" metricUnit="m" metricValue="1.72" unit="m" value="1720.0">
<elevation id="63FE0B426B11104D84292F6A52F61374" metricMagnitude="3" metricUnit="m" metricValue="1.72" unit="m" value="1720.0">1720 m</elevation>
</quantity>
elevation; 
<collectorName id="02D776E5C8F42971B8CE505ABE2C3D46">Berlese</collectorName>
; leaf litter; 
<collectingDate id="BE1E742D5D9506828CDDB1BBCB44BD72" value="1998-06-21">21 Jun 1998</collectingDate>
; 
<collectorName id="6BFFDB3852ADB5BEC6BF75738FAD08F9">R. Anderson</collectorName>
leg.; KNHM CR1A98 106.
</materialsCitation>
</paragraph>
<paragraph id="F39476F60A655CD47C1C7B17B159017E" pageId="0" pageNumber="13">
<materialsCitation id="AE4332CACA1AC5109042052C48B7630E" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" country="COSTA RICA" location="COSTA RICA" specimenCount="1" specimenCount-male="1" typeStatus="Paratypes">
<emphasis id="92088BF7639BB07DF5F0463A564AFE0C" bold="true" italics="true" pageId="0" pageNumber="13">
<typeStatus id="0089354E488B7561A2A7FFD368888E69" type="paratype">Paratypes</typeStatus>
</emphasis>
: 
<collectingCountry id="DA36BE8D7C4FCD66F5077D6B24DD7924" name="Costa Rica">COSTA RICA</collectingCountry>
<specimenCount id="2D063BDFA9D33B8FFCF0B535C84333D7" count="1" type="male">1 male</specimenCount>
</materialsCitation>
; 
<specimenCount id="E83456C7007857B1250CCD26F8068909" count="1" type="female">1 female</specimenCount>
; 
<materialsCitation id="5351D0952F451C02F8FE2B17A41DA807" collectingDate="1997-06-25" collectorName="Berlese, R. Anderson" country="Costa Rica" elevation="1800" latitude="9.466666" location="Hwy." longLatPrecision="21" longitude="-83.70555" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
<collectingRegion id="346C2A33A616187ECE2D8D44B3D1D72A" country="Costa Rica" name="San Jose">
San 
<normalizedToken id="CD71FD16D7C97317173AD4C8EF2D7DC3" originalValue="José">Jose</normalizedToken>
</collectingRegion>
; km 117 Pan-Am 
<location id="9626251F7E962363BD73935F54C7AAFA" LSID="urn:lsid:plazi:treatment:D88EE9CDA2485D8B8CACE550AEE4BE79:9626251F7E962363BD73935F54C7AAFA" country="Costa Rica" latitude="9.466666" longLatPrecision="21" longitude="-83.70555" name="Hwy." stateProvince="San Jose">Hwy.</location>
; 
<locationDeviation id="5847D8768105FF217090F9FFE3760E71" location="San Isidro">
<quantity id="BB90823E920BF291956A3813A2010D0A" metricMagnitude="4" metricUnit="m" metricValue="1.9" unit="km" value="19.0">19 km</quantity>
N San Isidro
</locationDeviation>
; 
<geoCoordinate id="0E494BCC3F7E5740AC89E792218D95D9" degrees="9" direction="north" minutes="28" orientation="latitude" precision="15" seconds="0" value="9.466666">9°28'0&quot;N</geoCoordinate>
, 
<geoCoordinate id="C96BC18D05FB116231DA3756CC6566A9" degrees="83" direction="west" minutes="42" orientation="longitude" precision="15" seconds="20" value="-83.70555">83°42'20&quot;W</geoCoordinate>
; 
<quantity id="B8FC135272652FD6ED7D82B9B6828912" metricMagnitude="3" metricUnit="m" metricValue="1.8" unit="m" value="1800.0">
<elevation id="6028C587CFD8AE98ED46E21C1108E09B" metricMagnitude="3" metricUnit="m" metricValue="1.8" unit="m" value="1800.0">1800 m</elevation>
</quantity>
elevation; 
<collectorName id="081ADC4F07324ED4CF34951EE7B004DE">Berlese</collectorName>
; forest litter; 
<collectingDate id="FC92D6A4303E5681B1D2CC97C39923BC" value="1997-06-25">25 Jun 1997</collectingDate>
; 
<collectorName id="60C7BEC016F1C252EAB75380FFD54125">R. Anderson</collectorName>
leg.: KNHM CR1A97 035A
</materialsCitation>
; 
<specimenCount id="42961EC52EF30178A4C4E4207118D9EF" count="2" type="female">2 females</specimenCount>
; 
<materialsCitation id="DB763B1FFA2B68C44B35B3EDA2506A4A" collectingDate="1997-06-14" collectorName="Berlese, R. Anderson" country="Costa Rica" elevation="960" latitude="10.224999" location="San Ramon" longLatPrecision="21" longitude="-84.59167" municipality="San Ramon" specimenCount="1" stateProvince="Alajuela" typeStatus="paratype">
<collectingRegion id="F3CA43970255E3409D8AF0273A690FAE" country="Costa Rica" name="Alajuela">Alajuela</collectingRegion>
; E.B. 
<collectingMunicipality id="E808453FF4D6B6727A4741600A27A340">San Ramon</collectingMunicipality>
; R.B. 
<location id="480054401A960BF5992161E641632D40" LSID="urn:lsid:plazi:treatment:D88EE9CDA2485D8B8CACE550AEE4BE79:480054401A960BF5992161E641632D40" country="Costa Rica" latitude="10.224999" longLatPrecision="21" longitude="-84.59167" municipality="San Ramon" name="San Ramon" stateProvince="Alajuela">San Ramon</location>
; 
<quantity id="A9638BB3803A9BAFE843D3408FE79BEE" metricMagnitude="4" metricUnit="m" metricValue="2.7" unit="km" value="27.0">27 km</quantity>
N and 
<locationDeviation id="49A587D3DF674EF21DE666BA11AEB7BB" location="San Ramon">
<quantity id="B5017B5E59D71695691C840E747B3ADF" metricMagnitude="3" metricUnit="m" metricValue="8.0" unit="km" value="8.0">8 km</quantity>
W San Ramon
</locationDeviation>
; 
<geoCoordinate id="8463498FA69610FA57DDB5DB21C61BF0" degrees="10" direction="north" minutes="13" orientation="latitude" precision="15" seconds="30" value="10.224999">10°13'30&quot;N</geoCoordinate>
, 
<geoCoordinate id="970BA1042DA4C13E1E8ABAA107B45113" degrees="84" direction="west" minutes="35" orientation="longitude" precision="15" seconds="30" value="-84.59167">84°35'30&quot;W</geoCoordinate>
; 
<quantity id="B2639F1A8175CC0069E241C90A55D1F8" metricMagnitude="2" metricUnit="m" metricValue="9.6" unit="m" value="960.0">
<elevation id="B6E23F0B3A41CDC65DECB25EFE762146" metricMagnitude="2" metricUnit="m" metricValue="9.6" unit="m" value="960.0">960 m</elevation>
</quantity>
elevation, 
<collectorName id="F84D0938188DAEAB99BE69B592AF6BF8">Berlese</collectorName>
; forest litter, 
<collectingDate id="A6010EBEC0343303C9549381F4D551DF" value="1997-06-14">14 Jun 1997</collectingDate>
, leg. 
<collectorName id="AD5794B5FFBAF80F907EBE45CE4A8F1E">R. Anderson</collectorName>
(CR1A97 014A) (KNHM)
</materialsCitation>
; 
<specimenCount id="9F98EABB5F162BD7777FA0E5903C5AF7" count="1" type="female">1 female</specimenCount>
; 
<materialsCitation id="4C0DF8A2448C2AA8A41B134C62F3CA4C" collectingDate="2000-06-13" collectingDateMax="2000-06-15" collectingDateMin="2000-06-13" collectingMethod="flight intercept trap" collectorName="J. Ashe, R. Brooks, Z. Falin" country="Costa Rica" elevation="610" latitude="10.989445" location="Patilla Biological Station" longLatPrecision="21" longitude="-85.42583" specimenCount="1" stateProvince="Guanacaste" typeStatus="paratype">
<collectingRegion id="5C06D0A304CE1CE0529D858FC9530A3D" country="Costa Rica" name="Guanacaste">Guanacaste</collectingRegion>
; 
<location id="7A0A0F6DC55C1FC1AFF8D1B894A2AEF5" LSID="urn:lsid:plazi:treatment:D88EE9CDA2485D8B8CACE550AEE4BE79:7A0A0F6DC55C1FC1AFF8D1B894A2AEF5" country="Costa Rica" latitude="10.989445" longLatPrecision="21" longitude="-85.42583" name="Patilla Biological Station" stateProvince="Guanacaste">Patilla Biological Station</location>
; 
<geoCoordinate id="F3540FFD3C40EA62ECAD03974B7AC443" degrees="10" direction="north" minutes="59" orientation="latitude" precision="15" seconds="22" value="10.989445">10°59'22&quot;N</geoCoordinate>
, 
<geoCoordinate id="8BA172A4DA057E56B5F73039D3B172C1" degrees="85" direction="west" minutes="25" orientation="longitude" precision="15" seconds="33" value="-85.42583">85°25'33&quot;W</geoCoordinate>
; 
<quantity id="17DD297C70BC0E493B6244C6748C66D2" metricMagnitude="2" metricUnit="m" metricValue="6.1" unit="m" value="610.0">
<elevation id="8C33B18DAF28679EB80FCA01BCA6183C" metricMagnitude="2" metricUnit="m" metricValue="6.1" unit="m" value="610.0">610 m</elevation>
</quantity>
elevation; 
<collectingMethod id="BEC01F19285AAA1FBA85B66A2F810DA1">flight intercept trap</collectingMethod>
; 
<collectingDate id="A54A14DB03D11B1C78C05B16FA24C2B7" value="2000-06-13" valueMax="2000-06-15" valueMin="2000-06-13">13-15 Jun 2000</collectingDate>
; 
<collectorName id="341F834B7A6DD461969EF7E0B0810873">J. Ashe</collectorName>
, 
<collectorName id="EF2200CE488C3A8F00A671816696D8D5">R. Brooks</collectorName>
, 
<collectorName id="D2E8EB45705AA6177501B380A124D112">Z. Falin</collectorName>
leg.; KNHM CR1ABF00 135
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3338A00D0E523DD09C758B650F9D58A6" pageId="0" pageNumber="13" type="diagnosis">
<paragraph id="DC7454D6F651F05D58C760FB98ADD509" pageId="0" pageNumber="13">Diagnosis.</paragraph>
<paragraph id="E67449EA7EBA327DAC82DFF588E858FB" pageId="0" pageNumber="13">
The species belongs to the 
<taxonomicName id="7562684BF4C51614772D9EB59E0952B9" baseAuthorityName="LeConte" baseAuthorityYear="1863" class="Insecta" family="Staphylinidae" genus="Somoleptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Somoleptus longicollis" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="longicollis">Somoleptus longicollis</taxonomicName>
-subgroup due to the structure of the aedeagus and the sternite VII of male with long central spine. According to the shape of the head and the proportion of the elytra, 
<taxonomicName id="549CA4E1C2B36D124A708D6510A4CE78" lsidName="S. luteicornis" pageId="0" pageNumber="13" rank="species" species="luteicornis">
<emphasis id="6FF44C5B346231855D1051323BC78847" italics="true" pageId="0" pageNumber="13">S. luteicornis</emphasis>
</taxonomicName>
resembles 
<taxonomicName id="51BDED4378925A058D48F5BA6F3AEBF8" lsidName="S. punctulatus" pageId="0" pageNumber="13" rank="species" species="punctulatus">
<emphasis id="5D4D40E9F93E3097193D5ECF790FD40B" italics="true" pageId="0" pageNumber="13">S. punctulatus</emphasis>
</taxonomicName>
Sharp, 1885, but the colouration is different. 
<taxonomicName id="8B7E6CA97D0C66D6365573F7EFEC5A23" authorityName="Irmler" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Somoleptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Somoleptus luteicornis" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="luteicornis">
<emphasis id="BC84339DE0DB39C945E50A0683F217B1" italics="true" pageId="0" pageNumber="13">Somoleptus luteicornis</emphasis>
</taxonomicName>
is totally black, whereas the pronotum of 
<taxonomicName id="308F01B1F25C1A44794C216E0AB74148" lsidName="S. punctulatus" pageId="0" pageNumber="13" rank="species" species="punctulatus">
<emphasis id="AA8AD048F8DB16FF4BE3B21AD4B7AD5E" italics="true" pageId="0" pageNumber="13">S. punctulatus</emphasis>
</taxonomicName>
is lighter reddish-brown. It differs also by the yellow antennomeres IV to XI contrasting with the dark basal three antennomeres. In 
<taxonomicName id="ED5B45F54EE61A090ED43BBDC9201DE1" lsidName="S. punctulatus" pageId="0" pageNumber="13" rank="species" species="punctulatus">
<emphasis id="12C5B8D8E94E40DE8DB536522C2B2178" italics="true" pageId="0" pageNumber="13">S. punctulatus</emphasis>
</taxonomicName>
, the antennomeres are totally yellow. Moreover, the apical cones at the orifice of the aedeagus are longer in 
<taxonomicName id="62C96AA8A6C6AA0A1569F109D6EAC861" lsidName="S. luteicornis" pageId="0" pageNumber="13" rank="species" species="luteicornis">
<emphasis id="578A10B625FC8AABD6219CF648117332" italics="true" pageId="0" pageNumber="13">S. luteicornis</emphasis>
</taxonomicName>
with C:A ratio of 0.3, whereas the ratio is 0.15 in 
<taxonomicName id="25FD6A302661087D0B42D134D3C629FC" lsidName="S. punctulatus" pageId="0" pageNumber="13" rank="species" species="punctulatus">
<emphasis id="63A8BD1F30BE0B40BFD3BF45B89A6219" italics="true" pageId="0" pageNumber="13">S. punctulatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="A4E2DA7B6CCBC2ED96017A7256D04897" pageId="0" pageNumber="13" type="description">
<paragraph id="C91C0EBE3DFA2713612F641B66061E72" pageId="0" pageNumber="13">Description.</paragraph>
<paragraph id="830D882235367C6428F1A0EE5CD4B5B0" pageId="0" pageNumber="13">Length: 3.89 mm; Colouration: completely black including legs and base of antennae; antennomeres IV to XI contrasting yellow from the three basal antennomeres.</paragraph>
<paragraph id="3ABCD4262860E364E6BDBF79A2EA8B04" pageId="0" pageNumber="13">Head: 0.67 mm long, 0.49 mm wide; eyes slightly prominent; moderately large; PS:E ratio 3.5; postocular sides nearly parallel; posterior angles sub-rectangular; setiferous punctation deep and dense; on average, interstices between punctures half as wide as diameter of punctures; small spot at posterior head more sparsely punctate; surface without microsculpture; polished; antennae with first antennomere elongate; slightly longer than half-length of head; second and third antennomere longer than wide; conical; combined half as long as first antennomere; following antennomeres wider than long; anteriad increasing in width. Pronotum: 0.75 mm long, 0.44 mm wide; widest at anterior third; conically narrowed to neck; posteriad, sides slightly convergent to sub-rectangular posterior angles; posterior margin convex; setiferous punctation deep and dense; on average, interstices between punctures 0.8 times as wide as diameter of punctures; wide mid-line impunctate; surface without microsculpture; polished. Elytra: 0.78 mm long, 0.66 mm wide; humeral angles obtuse; sides nearly parallel; posterior angles sub-rectangular; posterior margin convex; retreated to suture; setiferous punctation deep and dense; irregular; close to suture nearly coriaceous; laterally, on average, interstices between punctures as wide as diameter of punctures; meso- and meta-tibia with 3 ctenidia each. Abdomen with setiferous punctation finer, but denser than on elytra; sternite VII of male with long triangularly prominent posterior margin; posterior margin of tergite VII of male convex. Aedeagus long oval; dorsal plate covering nearly total central lobe; endophallus with one torsion; central part of endophallus straight covered by distinct lobes; cones at apical orifice long; C:A ratio 0.38; paramere bilobed; outer transparent lobe reniform; inner elongate lobe slender with several setae at outer edge.</paragraph>
</subSubSection>
<subSubSection id="6F76E6BC030D7E672E0DE59CDC0E0383" pageId="0" pageNumber="13" type="etymology">
<paragraph id="6D0C4DEAFE434126267B64CD8E5002D7" pageId="0" pageNumber="13">Etymology.</paragraph>
<paragraph id="28D87A0A822FD6D7E2C8423A8097DD48" pageId="0" pageNumber="13">
The species name is a combination of the Latin words 
<emphasis id="2B7A689FB3A4114C0803C463DE7294B1" italics="true" pageId="0" pageNumber="13">luteus</emphasis>
meaning 
<normalizedToken id="EE5171DBAB4D5D61E55070BC37D2E8A8" originalValue="“yellow”">&quot;yellow&quot;</normalizedToken>
and 
<emphasis id="4A9E20B6577276B13A270CCA1A6C66A8" italics="true" pageId="0" pageNumber="13">cornu</emphasis>
meaning the antennae of insects and refers to the yellow antennomeres IV to XI.
</paragraph>
</subSubSection>
<subSubSection id="5C38794A25ECCEDE4DD121C7BC4E7160" pageId="0" pageNumber="13" type="geography">
<paragraph id="FB0F9334E1340460D0F991B0C4FA9C14" pageId="0" pageNumber="13">Geography.</paragraph>
<paragraph id="EA8C4987810648A816C705CE15E2FF06" pageId="0" pageNumber="13">Costa Rica.</paragraph>
</subSubSection>
<subSubSection id="A60F9C778F3D2E0694ECB0E35638BE52" pageId="0" pageNumber="13" type="ecology">
<paragraph id="2E4FC410603B555E0B1A822E245FAA2A" pageId="0" pageNumber="13">Ecology.</paragraph>
<paragraph id="3589297B2AE160BC9F886B34FE3DFE42" pageId="0" pageNumber="13">Floor of montane and cloud forest of 500 to 1500 m elevation in June.</paragraph>
</subSubSection>
</treatment>
</document>