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<document ID-DOI="10.11646/zootaxa.5264.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7836674" ID-ZooBank="286F4C82-D3A7-455B-8174-354BFBF5D28F" approvalRequired="2" approvalRequired_for_textStreams="2" checkinTime="1681728586229" checkinUser="plazi" docAuthor="Shear, William A. &amp; Marek, Paul E." docDate="2023" docId="CC6F87FEFFF460400AB8FD87FDD0BEFF" docLanguage="en" docName="zootaxa.5264.3.2.pdf" docOrigin="Zootaxa 5264 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5264.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Miniaria richarti Shear &amp; Marek 2023, n. sp." docType="treatment" docVersion="2" lastPageNumber="328" masterDocId="3056FF86FFF160450A2FFFC0FF9CB922" masterDocTitle="The millipede family Striariidae Bollman, 1893. VIII. Three new genera and four new species of minute millipedes from Oregon and Washington, USA (Diplopoda, Chordeumatida, Striarioidea)" masterLastPageNumber="340" masterPageNumber="323" pageNumber="328" updateTime="1681729003861" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>The millipede family Striariidae Bollman, 1893. VIII. Three new genera and four new species of minute millipedes from Oregon and Washington, USA (Diplopoda, Chordeumatida, Striarioidea)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shear, William A.</mods:namePart>
<mods:affiliation>Professor Emeritus, Department of Biology, Hampden-Sydney College, Hampden-Sydney VA 23943 USA, current address: 1950 Price Drive, Farmville VA 23901 USA.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Marek, Paul E.</mods:namePart>
<mods:affiliation>Department of Entomology, Virginia Tech, Blacksburg VA 24061 USA.</mods:affiliation>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-04-14</mods:number>
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<mods:detail type="volume">
<mods:number>5264</mods:number>
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<mods:detail type="issue">
<mods:number>3</mods:number>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5264.3.2</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5264.3.2</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7836674</mods:identifier>
<mods:identifier type="ZooBank">286F4C82-D3A7-455B-8174-354BFBF5D28F</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7836684" ID-Zenodo-Dep="7836684" LSID="urn:lsid:plazi:treatment:CC6F87FEFFF460400AB8FD87FDD0BEFF" httpUri="http://treatment.plazi.org/id/CC6F87FEFFF460400AB8FD87FDD0BEFF" lastPageNumber="328" pageId="5" pageNumber="328">
<subSubSection box="[151,642,582,609]" pageId="5" pageNumber="328" type="nomenclature">
<paragraph blockId="5.[151,642,582,645]" box="[151,642,582,609]" pageId="5" pageNumber="328">
<heading bold="true" box="[151,642,582,609]" fontSize="11" level="1" pageId="5" pageNumber="328" reason="1">
<emphasis bold="true" box="[151,642,582,609]" pageId="5" pageNumber="328">
<taxonomicName authority="Shear &amp; Marek" authorityName="Shear &amp; Marek" authorityYear="2023" box="[151,563,582,609]" class="Diplopoda" family="Striariidae" genus="Miniaria" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="328" phylum="Arthropoda" rank="species" species="richarti" status="n. sp.">
<emphasis bold="true" box="[151,357,582,609]" italics="true" pageId="5" pageNumber="328">Miniaria richarti</emphasis>
Shear &amp; Marek
</taxonomicName>
,
<taxonomicNameLabel box="[575,642,583,609]" pageId="5" pageNumber="328" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection pageId="5" pageNumber="328" type="description">
<paragraph blockId="5.[151,642,582,645]" box="[151,360,618,645]" pageId="5" pageNumber="328">
<figureCitation box="[151,279,618,645]" captionStart-0="FIGURES 914" captionStart-1="FIGURES 1519" captionStart-2="FIGURES 2023" captionStart-3="FIGURES 2430" captionStartId-0="6.[151,264,1848,1872]" captionStartId-1="7.[151,264,1670,1694]" captionStartId-2="8.[151,264,1515,1539]" captionStartId-3="10.[151,264,1820,1844]" captionTargetBox-0="[174,1414,181,1823]" captionTargetBox-1="[162,1425,181,1646]" captionTargetBox-2="[151,1436,184,1491]" captionTargetBox-3="[164,1423,183,1796]" captionTargetId-0="figure-20@6.[174,1414,181,1823]" captionTargetId-1="figure-17@7.[161,1426,181,1646]" captionTargetId-2="figure-20@8.[151,1436,181,1491]" captionTargetId-3="figure-20@10.[164,1423,181,1796]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="10" captionText-0="FIGURES 914. Miniaria n. gen. species. 911. Miniaria ramifera n. gen., n. sp., male holotype. 9. Gonopods, posterior view. 10. Legpair 9 and coxae of legpair 10, anterior view. 11. Tip of gonopod anterior angiocoxite, ventral view. 1214. Miniaria richarti n. gen., n. sp., male from Germany Creek. 12. Head and legpair 1, ventrolateral view. 13. Pleurotergites 11 and 12, lateral view. 14. Pleurotergites 12 and 13, dorsal view. Abbreviations: aac, anterior angiocoxite; cx, gonopod coxa; cx (numeral), coxa of numbered legpair; lab, labrum; L1, legpair 1; m, mandibular stipes; numerals, ring numbers; om, ommatidium." captionText-1="FIGURES 1519. Miniaria richarti n. gen., n. sp., male from Germany Creek. 15. Segmental seta. 16. Telson, lateral view. 17. Tarsus 3, lateral view. 18. Tip of tarsus 4, lateral view. 19. Gonopods, posterior view. Abbreviations: aac, anterior angiocoxite; cx, coxae of gonopods; pac, posterior angiocoxite; s, sternum of gonopods; sp, spinnerets; t, telson; x, comb seta." captionText-2="FIGURES 2023. Miniaria richarti n. gen., n. sp., gonopods of specimen from Lees Camp. 20. Posterior view. 21. Tips of anterior colpocoxites, ventral view. 22. Posterior view showing sheathing of flagellocoxite by posterior angiocoxite. 23. Posterior view showing colpocoxite and origin of flagellocoxite. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cx, gonopod coxa; f, flagellocoxite; pac, posterior angiocoxite." captionText-3="FIGURES 2430. Male striariid millipedes. Figs. 24, 25. Miniaria richarti n. gen., n. sp., male. 24. Gonopods of specimen from Pin Creek, ventral view. 25. Legpair 9 and coxae 10 of specimen from Kennedy Creek, ventral view. Figs. 2630. Tigraria oregonensis n. gen., n. sp. male. 26. Head and first 3 legpairs, ventral view. 27. Collum, lateral view. 28. Pleurotergites 13, 14, dorsal view. 29. Segmental seta. 30. Left leg 2, ventral view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; col, collum; cx, coxa of gonopods; cx (numeral), coxa of numbered legpair; L (numeral), numbered legpair; lab, labrum; m, mandibular stipes; numeral, ring numbers; s, sternite of gonopods; vd, opening of vas deferens." figureDoi-0="http://doi.org/10.5281/zenodo.7836682" figureDoi-1="http://doi.org/10.5281/zenodo.7836686" figureDoi-2="http://doi.org/10.5281/zenodo.7836688" figureDoi-3="http://doi.org/10.5281/zenodo.7836690" httpUri-0="https://zenodo.org/record/7836682/files/figure.png" httpUri-1="https://zenodo.org/record/7836686/files/figure.png" httpUri-2="https://zenodo.org/record/7836688/files/figure.png" httpUri-3="https://zenodo.org/record/7836690/files/figure.png" pageId="5" pageNumber="328">Figs 1225</figureCitation>
,
<figureCitation box="[290,360,618,645]" captionStart-0="FIGURES 3946" captionStart-1="FIGURES 4754" captionStart-2="FIGURES 5561" captionStartId-0="13.[151,262,1817,1841]" captionStartId-1="14.[151,264,1809,1833]" captionStartId-2="15.[151,264,1809,1833]" captionTargetBox-0="[216,1378,182,1788]" captionTargetBox-1="[213,1367,188,1786]" captionTargetBox-2="[192,1400,181,1779]" captionTargetId-0="figure-17@13.[206,1382,181,1792]" captionTargetId-1="figure-20@14.[213,1375,181,1786]" captionTargetId-2="figure-17@15.[186,1400,181,1785]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionText-0="FIGURES 3946. Gonopods and ninth legs of male striariid millipedes. Figs. 3945. Miniaria ramifera n. gen., n. sp. 39. Right gonopod of specimen from Mashel River, mesal view. 40. Same, lateral view. 41. Right gonopod of specimen from Cook- Underwood Road, mesal view. 42. Same, lateral view. 43, Right gonopod of specimen fromAinsworth State Park, mesal view. 44. Right gonopod of specimen from Timber, mesal view. 45. Ninth legpair of specimen from Ainsworth State Park, posterior view. 46. Miniaria richarti n. gen., n. sp., right gonopod of specimen from Hospital Creek, mesal view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cx9, ninth leg coxa; cxp, coxal process; pac, posterior angiocoxite; t9, telopodite of ninth leg." captionText-1="FIGURES 4754. Miniaria richarti n. gen., n. sp., male paratype. 47. Right gonopod of specimen from Hospital Creek, lateral view. 48. Right gonopod of specimen from Centralia, lateral view. 49. Right gonopod of specimen from Toutle, lateral view. 50. Gonopods of specimen from Elochoman River Road, anterior view. 51. Right gonopod of specimen from Kennedy Creek, mesal view. 52. Right gonopod of specimen from Lees Camp, mesal view. 53. Same, Lateral view. 54. Ninth legpair of specimen from Lees Camp, anterior view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cx, coxa; cxp, coxal process; f, flagellum; pac, posterior angiocoxite; t9, telopodite of ninth leg." captionText-2="FIGURES 5561. Gonopods and ninth legs of male striariid millipedes. Figs. 55, 56. Miniaria richarti n. gen., n. sp. 55. Ninth legpair of specimen from Elochoman River Road, anterior view. 56. Left ninth leg of specimen from Hospital Creek, anterior view. Figs. 5759. Tigraria oregonensis n. gen., n. sp., male paratype; 57. Right gonopod, mesal view. 58. Same, lateral view. 59. Right ninth leg, anterior view. Figs. 60, 61. Kingaria prattensis n. gen., n. sp., male holotype. 60. Right gonopod, lateral view. 61. Left ninth leg, anterior view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cx, coxa; cxp, coxal process; f, flagellum; pac, posterior angiocoxite; t9, telopodite of ninth leg." figureDoi-0="http://doi.org/10.5281/zenodo.7836702" figureDoi-1="http://doi.org/10.5281/zenodo.7836704" figureDoi-2="http://doi.org/10.5281/zenodo.7836708" httpUri-0="https://zenodo.org/record/7836702/files/figure.png" httpUri-1="https://zenodo.org/record/7836704/files/figure.png" httpUri-2="https://zenodo.org/record/7836708/files/figure.png" pageId="5" pageNumber="328">4656</figureCitation>
</paragraph>
<paragraph blockId="5.[151,1437,691,2013]" pageId="5" pageNumber="328">
<emphasis box="[151,223,691,716]" italics="true" pageId="5" pageNumber="328">Types:</emphasis>
<materialsCitation collectingDate="2003-11-29" collectionCode="CAS" collectorName="W. Leonard" county="Thurston Co." location="Rainier Road" municipality="Male" pageId="5" pageNumber="328" specimenCount="2" stateProvince="Washington" typeStatus="holotype">
All specimens deposited in
<collectionCode box="[553,611,691,717]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="5" pageNumber="328" type="Museum">CAS</collectionCode>
.
<collectingMunicipality box="[625,683,691,717]" pageId="5" pageNumber="328">Male</collectingMunicipality>
<typeStatus box="[692,790,691,717]" pageId="5" pageNumber="328">holotype</typeStatus>
and female
<typeStatus box="[935,1031,691,717]" pageId="5" pageNumber="328">paratype</typeStatus>
from
<location LSID="urn:lsid:plazi:treatment:CC6F87FEFFF460400AB8FD87FDD0BEFF:41196033FFF460400E7EFD73FB77BBEF" box="[1105,1259,691,717]" county="Thurston Co." municipality="Male" name="Rainier Road" pageId="5" pageNumber="328" stateProvince="Washington">Rainier Road</location>
, 3.0 mi NW of
<location LSID="urn:lsid:plazi:treatment:CC6F87FEFFF460400AB8FD87FDD0BEFF:41196033FFF460400AB8FD17FEA7BBD3" box="[151,315,727,753]" county="Thurston Co." municipality="Male" name="Military Road" pageId="5" pageNumber="328" stateProvince="Washington">Military Road</location>
,
<location LSID="urn:lsid:plazi:treatment:CC6F87FEFFF460400AB8FD87FDD0BEFF:41196033FFF460400B66FD17FD22BBD3" box="[329,702,727,753]" county="Thurston Co." municipality="Male" name="Fort Lewis Military Reservation" pageId="5" pageNumber="328" stateProvince="Washington">Fort Lewis Military Reservation</location>
,
<collectingCounty box="[715,866,727,753]" pageId="5" pageNumber="328">Thurston Co.</collectingCounty>
<collectingRegion box="[875,1011,727,753]" country="United States of America" name="Washington" pageId="5" pageNumber="328">Washington</collectingRegion>
, collected
<date box="[1136,1358,727,753]" pageId="5" pageNumber="328" value="2003-11-29">
<collectingDate box="[1136,1358,727,753]" pageId="5" pageNumber="328" value="2003-11-29">29 November 2003</collectingDate>
</date>
by
<collectorName pageId="5" pageNumber="328">W. Leonard</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="328" type="diagnosis">
<paragraph blockId="5.[151,1437,691,2013]" pageId="5" pageNumber="328">
<emphasis bold="true" box="[199,314,798,824]" pageId="5" pageNumber="328">Diagnosis</emphasis>
. Distinct from
<taxonomicName authorityName="Shear &amp; Marek" authorityYear="2023" box="[501,642,799,825]" class="Diplopoda" family="Striariidae" genus="Miniaria" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="328" phylum="Arthropoda" rank="species" species="ramifera">
<emphasis box="[501,642,799,825]" italics="true" pageId="5" pageNumber="328">M. ramifera</emphasis>
</taxonomicName>
in lacking a posterior subterminal branch on the gonopod anterior angiocoxite, and in having the gonopods closely appressed in the midline. The anterior angiocoxite is sharply elbowed, not evenly curved as in
<emphasis box="[527,670,871,897]" italics="true" pageId="5" pageNumber="328">
<taxonomicName authorityName="Shear &amp; Marek" authorityYear="2023" box="[527,666,871,897]" class="Diplopoda" family="Striariidae" genus="Miniaria" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="328" phylum="Arthropoda" rank="species" species="ramifera">M. ramifera</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="328" type="etymology">
<paragraph blockId="5.[151,1437,691,2013]" pageId="5" pageNumber="328">
<emphasis bold="true" box="[199,328,907,933]" pageId="5" pageNumber="328">Etymology</emphasis>
. The species name honors Dr. Casey Richart, whose collecting has contributed much to our knowledge of the litter fauna of the Pacific Northwest and
<collectingRegion box="[809,926,943,969]" country="United States of America" name="California" pageId="5" pageNumber="328">California</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="328" type="description">
<paragraph blockId="5.[151,1437,691,2013]" box="[199,1127,978,1005]" pageId="5" pageNumber="328">
<emphasis bold="true" box="[199,337,978,1004]" pageId="5" pageNumber="328">Description</emphasis>
. Male from
<collectingCountry box="[478,582,979,1005]" name="Germany" pageId="5" pageNumber="328">Germany</collectingCountry>
Creek. Length 3.0 mm, greatest width
<quantity box="[1020,1120,979,1005]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="328" unit="mm" value="0.3">0.30 mm</quantity>
.
</paragraph>
<paragraph blockId="5.[151,1437,691,2013]" pageId="5" pageNumber="328">
Nonsexual characters and secondary sexual modifications as described for genus. Gonopods (
<figureCitation box="[1253,1382,1015,1041]" captionStart-0="FIGURES 1519" captionStart-1="FIGURES 2023" captionStart-2="FIGURES 2430" captionStartId-0="7.[151,264,1670,1694]" captionStartId-1="8.[151,264,1515,1539]" captionStartId-2="10.[151,264,1820,1844]" captionTargetBox-0="[162,1425,181,1646]" captionTargetBox-1="[151,1436,184,1491]" captionTargetBox-2="[164,1423,183,1796]" captionTargetId-0="figure-17@7.[161,1426,181,1646]" captionTargetId-1="figure-20@8.[151,1436,181,1491]" captionTargetId-2="figure-20@10.[164,1423,181,1796]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="10" captionText-0="FIGURES 1519. Miniaria richarti n. gen., n. sp., male from Germany Creek. 15. Segmental seta. 16. Telson, lateral view. 17. Tarsus 3, lateral view. 18. Tip of tarsus 4, lateral view. 19. Gonopods, posterior view. Abbreviations: aac, anterior angiocoxite; cx, coxae of gonopods; pac, posterior angiocoxite; s, sternum of gonopods; sp, spinnerets; t, telson; x, comb seta." captionText-1="FIGURES 2023. Miniaria richarti n. gen., n. sp., gonopods of specimen from Lees Camp. 20. Posterior view. 21. Tips of anterior colpocoxites, ventral view. 22. Posterior view showing sheathing of flagellocoxite by posterior angiocoxite. 23. Posterior view showing colpocoxite and origin of flagellocoxite. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cx, gonopod coxa; f, flagellocoxite; pac, posterior angiocoxite." captionText-2="FIGURES 2430. Male striariid millipedes. Figs. 24, 25. Miniaria richarti n. gen., n. sp., male. 24. Gonopods of specimen from Pin Creek, ventral view. 25. Legpair 9 and coxae 10 of specimen from Kennedy Creek, ventral view. Figs. 2630. Tigraria oregonensis n. gen., n. sp. male. 26. Head and first 3 legpairs, ventral view. 27. Collum, lateral view. 28. Pleurotergites 13, 14, dorsal view. 29. Segmental seta. 30. Left leg 2, ventral view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; col, collum; cx, coxa of gonopods; cx (numeral), coxa of numbered legpair; L (numeral), numbered legpair; lab, labrum; m, mandibular stipes; numeral, ring numbers; s, sternite of gonopods; vd, opening of vas deferens." figureDoi-0="http://doi.org/10.5281/zenodo.7836686" figureDoi-1="http://doi.org/10.5281/zenodo.7836688" figureDoi-2="http://doi.org/10.5281/zenodo.7836690" httpUri-0="https://zenodo.org/record/7836686/files/figure.png" httpUri-1="https://zenodo.org/record/7836688/files/figure.png" httpUri-2="https://zenodo.org/record/7836690/files/figure.png" pageId="5" pageNumber="328">Figs 1924</figureCitation>
,
<figureCitation captionStart-0="FIGURES 3946" captionStart-1="FIGURES 4754" captionStartId-0="13.[151,262,1817,1841]" captionStartId-1="14.[151,264,1809,1833]" captionTargetBox-0="[216,1378,182,1788]" captionTargetBox-1="[213,1367,188,1786]" captionTargetId-0="figure-17@13.[206,1382,181,1792]" captionTargetId-1="figure-20@14.[213,1375,181,1786]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURES 3946. Gonopods and ninth legs of male striariid millipedes. Figs. 3945. Miniaria ramifera n. gen., n. sp. 39. Right gonopod of specimen from Mashel River, mesal view. 40. Same, lateral view. 41. Right gonopod of specimen from Cook- Underwood Road, mesal view. 42. Same, lateral view. 43, Right gonopod of specimen fromAinsworth State Park, mesal view. 44. Right gonopod of specimen from Timber, mesal view. 45. Ninth legpair of specimen from Ainsworth State Park, posterior view. 46. Miniaria richarti n. gen., n. sp., right gonopod of specimen from Hospital Creek, mesal view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cx9, ninth leg coxa; cxp, coxal process; pac, posterior angiocoxite; t9, telopodite of ninth leg." captionText-1="FIGURES 4754. Miniaria richarti n. gen., n. sp., male paratype. 47. Right gonopod of specimen from Hospital Creek, lateral view. 48. Right gonopod of specimen from Centralia, lateral view. 49. Right gonopod of specimen from Toutle, lateral view. 50. Gonopods of specimen from Elochoman River Road, anterior view. 51. Right gonopod of specimen from Kennedy Creek, mesal view. 52. Right gonopod of specimen from Lees Camp, mesal view. 53. Same, Lateral view. 54. Ninth legpair of specimen from Lees Camp, anterior view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cx, coxa; cxp, coxal process; f, flagellum; pac, posterior angiocoxite; t9, telopodite of ninth leg." figureDoi-0="http://doi.org/10.5281/zenodo.7836702" figureDoi-1="http://doi.org/10.5281/zenodo.7836704" httpUri-0="https://zenodo.org/record/7836702/files/figure.png" httpUri-1="https://zenodo.org/record/7836704/files/figure.png" pageId="5" pageNumber="328">46 53</figureCitation>
) small, anterior angiocoxites (
<emphasis bold="true" box="[527,575,1051,1077]" pageId="5" pageNumber="328">aac,</emphasis>
<figureCitation box="[583,665,1051,1077]" captionStart="FIGURES 1519" captionStartId="7.[151,264,1670,1694]" captionTargetBox="[162,1425,181,1646]" captionTargetId="figure-17@7.[161,1426,181,1646]" captionTargetPageId="7" captionText="FIGURES 1519. Miniaria richarti n. gen., n. sp., male from Germany Creek. 15. Segmental seta. 16. Telson, lateral view. 17. Tarsus 3, lateral view. 18. Tip of tarsus 4, lateral view. 19. Gonopods, posterior view. Abbreviations: aac, anterior angiocoxite; cx, coxae of gonopods; pac, posterior angiocoxite; s, sternum of gonopods; sp, spinnerets; t, telson; x, comb seta." figureDoi="http://doi.org/10.5281/zenodo.7836686" httpUri="https://zenodo.org/record/7836686/files/figure.png" pageId="5" pageNumber="328">Fig. 19</figureCitation>
) distinctly elbowed at near right angle posteriorly, slightly knobbed at tips, distally with prominent fringed margin. Posterior angiocoxites (
<emphasis bold="true" box="[975,1024,1087,1113]" pageId="5" pageNumber="328">pac,</emphasis>
<figureCitation box="[1034,1117,1087,1113]" captionStart="FIGURES 1519" captionStartId="7.[151,264,1670,1694]" captionTargetBox="[162,1425,181,1646]" captionTargetId="figure-17@7.[161,1426,181,1646]" captionTargetPageId="7" captionText="FIGURES 1519. Miniaria richarti n. gen., n. sp., male from Germany Creek. 15. Segmental seta. 16. Telson, lateral view. 17. Tarsus 3, lateral view. 18. Tip of tarsus 4, lateral view. 19. Gonopods, posterior view. Abbreviations: aac, anterior angiocoxite; cx, coxae of gonopods; pac, posterior angiocoxite; s, sternum of gonopods; sp, spinnerets; t, telson; x, comb seta." figureDoi="http://doi.org/10.5281/zenodo.7836686" httpUri="https://zenodo.org/record/7836686/files/figure.png" pageId="5" pageNumber="328">Fig. 19</figureCitation>
) about two-thirds length of anterior ones, fimbriate along posterior margin, divided into medial fimbriate region and longer, thin apical branch, sheathing single curved flagellocoxite or flagellum. Colpocoxite much reduced. Ninth legs (
<figureCitation box="[1215,1305,1158,1185]" captionStart="FIGURES 2430" captionStartId="10.[151,264,1820,1844]" captionTargetBox="[164,1423,183,1796]" captionTargetId="figure-20@10.[164,1423,181,1796]" captionTargetPageId="10" captionText="FIGURES 2430. Male striariid millipedes. Figs. 24, 25. Miniaria richarti n. gen., n. sp., male. 24. Gonopods of specimen from Pin Creek, ventral view. 25. Legpair 9 and coxae 10 of specimen from Kennedy Creek, ventral view. Figs. 2630. Tigraria oregonensis n. gen., n. sp. male. 26. Head and first 3 legpairs, ventral view. 27. Collum, lateral view. 28. Pleurotergites 13, 14, dorsal view. 29. Segmental seta. 30. Left leg 2, ventral view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; col, collum; cx, coxa of gonopods; cx (numeral), coxa of numbered legpair; L (numeral), numbered legpair; lab, labrum; m, mandibular stipes; numeral, ring numbers; s, sternite of gonopods; vd, opening of vas deferens." figureDoi="http://doi.org/10.5281/zenodo.7836690" httpUri="https://zenodo.org/record/7836690/files/figure.png" pageId="5" pageNumber="328">Figs 25</figureCitation>
,
<figureCitation box="[1320,1350,1158,1185]" captionStart="FIGURES 4754" captionStartId="14.[151,264,1809,1833]" captionTargetBox="[213,1367,188,1786]" captionTargetId="figure-20@14.[213,1375,181,1786]" captionTargetPageId="14" captionText="FIGURES 4754. Miniaria richarti n. gen., n. sp., male paratype. 47. Right gonopod of specimen from Hospital Creek, lateral view. 48. Right gonopod of specimen from Centralia, lateral view. 49. Right gonopod of specimen from Toutle, lateral view. 50. Gonopods of specimen from Elochoman River Road, anterior view. 51. Right gonopod of specimen from Kennedy Creek, mesal view. 52. Right gonopod of specimen from Lees Camp, mesal view. 53. Same, Lateral view. 54. Ninth legpair of specimen from Lees Camp, anterior view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cx, coxa; cxp, coxal process; f, flagellum; pac, posterior angiocoxite; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7836704" httpUri="https://zenodo.org/record/7836704/files/figure.png" pageId="5" pageNumber="328">54</figureCitation>
,
<figureCitation box="[1365,1394,1158,1185]" captionStart="FIGURES 5561" captionStartId="15.[151,264,1809,1833]" captionTargetBox="[192,1400,181,1779]" captionTargetId="figure-17@15.[186,1400,181,1785]" captionTargetPageId="15" captionText="FIGURES 5561. Gonopods and ninth legs of male striariid millipedes. Figs. 55, 56. Miniaria richarti n. gen., n. sp. 55. Ninth legpair of specimen from Elochoman River Road, anterior view. 56. Left ninth leg of specimen from Hospital Creek, anterior view. Figs. 5759. Tigraria oregonensis n. gen., n. sp., male paratype; 57. Right gonopod, mesal view. 58. Same, lateral view. 59. Right ninth leg, anterior view. Figs. 60, 61. Kingaria prattensis n. gen., n. sp., male holotype. 60. Right gonopod, lateral view. 61. Left ninth leg, anterior view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cx, coxa; cxp, coxal process; f, flagellum; pac, posterior angiocoxite; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7836708" httpUri="https://zenodo.org/record/7836708/files/figure.png" pageId="5" pageNumber="328">55</figureCitation>
) as described for genus.
</paragraph>
<paragraph blockId="5.[151,1437,691,2013]" box="[199,768,1231,1257]" pageId="5" pageNumber="328">Females similar to males in all nonsexual respects.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="328" type="distribution">
<paragraph blockId="5.[151,1437,691,2013]" pageId="5" pageNumber="328">
<emphasis bold="true" box="[199,345,1266,1292]" pageId="5" pageNumber="328">Distribution</emphasis>
. Oregon: Tillamook Co.:
<quantity box="[644,698,1267,1293]" metricMagnitude="3" metricUnit="m" metricValue="1.609344" pageId="5" pageNumber="328" unit="mi" value="1.0">1 mi</quantity>
W,
<quantity box="[746,821,1266,1293]" metricMagnitude="2" metricUnit="m" metricValue="8.04672" pageId="5" pageNumber="328" unit="mi" value="0.5">0.5 mi</quantity>
S of Lees Camp,
<geoCoordinate box="[1041,1152,1266,1293]" degrees="45.587" direction="north" orientation="latitude" pageId="5" pageNumber="328" precision="55" value="45.587">45.587°N</geoCoordinate>
, -
<geoCoordinate box="[1173,1303,1266,1293]" degrees="123.599" direction="west" orientation="longitude" pageId="5" pageNumber="328" precision="55" value="-123.599">123.599°W</geoCoordinate>
, 700 asl,
<date pageId="5" pageNumber="328" value="1972-11-04">4 November 1972</date>
, E. M. Benedict,
<quantity box="[516,577,1303,1329]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="328" unit="mm" value="2.0">2mm</quantity>
, 2ff.Washington:Cowlitz Co.: SR503,
<quantity box="[1005,1087,1303,1329]" metricMagnitude="4" metricUnit="m" metricValue="1.83465216" pageId="5" pageNumber="328" unit="mi" value="11.4">11.4 mi</quantity>
E of I-5, 410 asl,
<geoCoordinate box="[1279,1432,1302,1329]" degrees="45.968833" direction="north" orientation="latitude" pageId="5" pageNumber="328" precision="1" value="45.968834">45.968833°N</geoCoordinate>
, -
<geoCoordinate box="[159,332,1338,1365]" degrees="122.549817" direction="west" orientation="longitude" pageId="5" pageNumber="328" precision="1" value="-122.54982">122.549817°W</geoCoordinate>
,
<date box="[345,509,1339,1365]" pageId="5" pageNumber="328" value="2004-03-07">7 March 2004</date>
, W. Leonard, m, f; Pin Creek,
<quantity box="[864,938,1338,1365]" metricMagnitude="2" metricUnit="m" metricValue="8.04672" pageId="5" pageNumber="328" unit="mi" value="0.5">0.5 mi</quantity>
E of Carrolls,
<geoCoordinate box="[1112,1223,1339,1365]" degrees="46.267" direction="north" orientation="latitude" pageId="5" pageNumber="328" precision="55" value="46.267">46.267°N</geoCoordinate>
, -
<geoCoordinate box="[1244,1374,1338,1365]" degrees="122.853" direction="west" orientation="longitude" pageId="5" pageNumber="328" precision="55" value="-122.853">122.853°W</geoCoordinate>
, 300 asl.
<date box="[196,358,1375,1401]" pageId="5" pageNumber="328" value="2003-03-07">7 March 2003</date>
. W. Leonard, m;
<collectingCountry box="[550,654,1375,1401]" name="Germany" pageId="5" pageNumber="328">Germany</collectingCountry>
Creek,
<quantity box="[742,814,1374,1401]" metricMagnitude="3" metricUnit="m" metricValue="8.5295232" pageId="5" pageNumber="328" unit="mi" value="5.3">5.3 mi</quantity>
N of SR4,
<geoCoordinate box="[942,1053,1375,1401]" degrees="46.267" direction="north" orientation="latitude" pageId="5" pageNumber="328" precision="55" value="46.267">46.267°N</geoCoordinate>
, -
<geoCoordinate box="[1072,1202,1375,1401]" degrees="123.132" direction="west" orientation="longitude" pageId="5" pageNumber="328" precision="55" value="-123.132">123.132°W</geoCoordinate>
,
<date box="[1214,1432,1375,1401]" pageId="5" pageNumber="328" value="2004-11-11">11 November 2004</date>
, W. Leonard, m. Grays Harbor Co.: Porter Creek Campground,
<geoCoordinate box="[889,1042,1411,1437]" degrees="46.978167" direction="north" orientation="latitude" pageId="5" pageNumber="328" precision="1" value="46.97817">46.978167°N</geoCoordinate>
, -
<geoCoordinate box="[1064,1209,1410,1437]" degrees="123.2565" direction="west" orientation="longitude" pageId="5" pageNumber="328" precision="5" value="-123.2565">123.2565°W</geoCoordinate>
,
<date box="[1224,1391,1410,1437]" pageId="5" pageNumber="328" value="2005-03-03">3 March 2005</date>
, W. Leonard, m. Lewis Co.:604 Roswell Road, Centralia, 280 asl,
<geoCoordinate box="[873,1027,1447,1473]" degrees="46.722317" direction="north" orientation="latitude" pageId="5" pageNumber="328" precision="1" value="46.722317">46.722317°N</geoCoordinate>
, -
<geoCoordinate box="[1047,1192,1447,1473]" degrees="122.9444" direction="west" orientation="longitude" pageId="5" pageNumber="328" precision="5" value="-122.9444">122.9444°W</geoCoordinate>
,
<date box="[1205,1397,1446,1473]" pageId="5" pageNumber="328" value="2004-01-25">25 January 2004</date>
, C. Richart, W. Leonard, m. Mason Co.: Kennedy Creek,
<geoCoordinate box="[775,900,1483,1509]" degrees="47.0876" direction="north" orientation="latitude" pageId="5" pageNumber="328" precision="5" value="47.0876">47.0876°N</geoCoordinate>
, -
<geoCoordinate box="[921,1066,1482,1509]" degrees="123.0957" direction="west" orientation="longitude" pageId="5" pageNumber="328" precision="5" value="-123.0957">123.0957°W</geoCoordinate>
,
<date box="[1080,1259,1483,1509]" pageId="5" pageNumber="328" value="2003-03-30">30 March 2003</date>
, W. Leonard,
<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="328" unit="mm" value="2.0">2 mm</quantity>
, f. Thurston Co.: Hospital Creek, above confluence with Skookumchuck River,
<geoCoordinate box="[1101,1254,1519,1545]" degrees="46.773267" direction="north" orientation="latitude" pageId="5" pageNumber="328" precision="1" value="46.773266">46.773267°N</geoCoordinate>
, -
<geoCoordinate box="[1273,1432,1518,1545]" degrees="122.58555" direction="west" orientation="longitude" pageId="5" pageNumber="328" precision="1" value="-122.58555">122.58555°W</geoCoordinate>
,
<date box="[151,369,1554,1581]" pageId="5" pageNumber="328" value="2003-12-15">15 December 2003</date>
, W. Leonard, K. McAllister,
<quantity box="[696,765,1555,1581]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="328" unit="mm" value="2.0">2 mm</quantity>
; Black Lake and Belmore Road at 66th Avenue, Tumwater,
<geoCoordinate box="[151,262,1591,1617]" degrees="46.989" direction="north" orientation="latitude" pageId="5" pageNumber="328" precision="55" value="46.989">46.989°N</geoCoordinate>
, -
<geoCoordinate box="[283,427,1590,1617]" degrees="122.9665" direction="west" orientation="longitude" pageId="5" pageNumber="328" precision="5" value="-122.9665">122.9665°W</geoCoordinate>
,
<date box="[440,662,1591,1617]" pageId="5" pageNumber="328" value="2004-11-19">19 November 2004</date>
, W. P. Leonard,
<quantity box="[850,919,1591,1617]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="328" unit="mm" value="2.0">2 mm</quantity>
, 5 ff; Summit Lake, 248 asl,
<geoCoordinate box="[1261,1414,1591,1617]" degrees="47.002833" direction="north" orientation="latitude" pageId="5" pageNumber="328" precision="1" value="47.002834">47.002833°N</geoCoordinate>
, -
<geoCoordinate box="[151,319,1627,1653]" degrees="123.130167" direction="north" orientation="latitude" pageId="5" pageNumber="328" precision="1" value="123.130165">123.130167°N</geoCoordinate>
,
<date box="[330,548,1627,1653]" pageId="5" pageNumber="328" value="2004-12-18">18 December 2004</date>
, W. Leonard, m; Olympia, Watershed Park,
<date box="[1047,1249,1627,1653]" pageId="5" pageNumber="328" value="2003-02-11">11 February 2003</date>
, W. Leonard, m, f. Wahakiakum Co.:
<quantity box="[380,465,1663,1689]" metricMagnitude="4" metricUnit="m" metricValue="1.8829324799999998" pageId="5" pageNumber="328" unit="mi" value="11.7">11.7 mi</quantity>
on Elochoman Valley Road from SR4, 450 asl,
<geoCoordinate box="[1002,1156,1663,1689]" degrees="46.316783" direction="north" orientation="latitude" pageId="5" pageNumber="328" precision="1" value="46.316784">46.316783°N</geoCoordinate>
, -
<geoCoordinate box="[1174,1318,1663,1689]" degrees="122.2617" direction="west" orientation="longitude" pageId="5" pageNumber="328" precision="5" value="-122.2617">122.2617°W</geoCoordinate>
,
<date pageId="5" pageNumber="328" value="2004-03-28">28 March 2004</date>
, C. Richart, m.
</paragraph>
<paragraph blockId="5.[151,1437,691,2013]" pageId="5" pageNumber="328">
<emphasis bold="true" box="[199,266,1735,1761]" pageId="5" pageNumber="328">Notes</emphasis>
. As with
<emphasis box="[373,515,1735,1761]" italics="true" pageId="5" pageNumber="328">
<taxonomicName authorityName="Shear &amp; Marek" authorityYear="2023" box="[373,511,1735,1761]" class="Diplopoda" family="Striariidae" genus="Miniaria" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="328" phylum="Arthropoda" rank="species" species="ramifera">M. ramifera</taxonomicName>
,
</emphasis>
variation is evident in the gonopod drawings, but it is not certain if this represents true geographic variation or is simply due to slightly different viewing angles. In the males from Hospital Creek (
<figureCitation box="[159,241,1807,1833]" captionStart="FIGURES 4754" captionStartId="14.[151,264,1809,1833]" captionTargetBox="[213,1367,188,1786]" captionTargetId="figure-20@14.[213,1375,181,1786]" captionTargetPageId="14" captionText="FIGURES 4754. Miniaria richarti n. gen., n. sp., male paratype. 47. Right gonopod of specimen from Hospital Creek, lateral view. 48. Right gonopod of specimen from Centralia, lateral view. 49. Right gonopod of specimen from Toutle, lateral view. 50. Gonopods of specimen from Elochoman River Road, anterior view. 51. Right gonopod of specimen from Kennedy Creek, mesal view. 52. Right gonopod of specimen from Lees Camp, mesal view. 53. Same, Lateral view. 54. Ninth legpair of specimen from Lees Camp, anterior view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cx, coxa; cxp, coxal process; f, flagellum; pac, posterior angiocoxite; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7836704" httpUri="https://zenodo.org/record/7836704/files/figure.png" pageId="5" pageNumber="328">Fig. 47</figureCitation>
) and Centralia, the even curvature of the anterior angiocoxite of
<taxonomicName authorityName="Shear &amp; Marek" authorityYear="2023" box="[967,1102,1807,1833]" class="Diplopoda" family="Striariidae" genus="Miniaria" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="328" phylum="Arthropoda" rank="species" species="ramifera">
<emphasis box="[967,1102,1807,1833]" italics="true" pageId="5" pageNumber="328">M. ramifera</emphasis>
</taxonomicName>
is approached, while in Lees Camp (
<figureCitation box="[233,364,1842,1869]" captionStart="FIGURES 4754" captionStartId="14.[151,264,1809,1833]" captionTargetBox="[213,1367,188,1786]" captionTargetId="figure-20@14.[213,1375,181,1786]" captionTargetPageId="14" captionText="FIGURES 4754. Miniaria richarti n. gen., n. sp., male paratype. 47. Right gonopod of specimen from Hospital Creek, lateral view. 48. Right gonopod of specimen from Centralia, lateral view. 49. Right gonopod of specimen from Toutle, lateral view. 50. Gonopods of specimen from Elochoman River Road, anterior view. 51. Right gonopod of specimen from Kennedy Creek, mesal view. 52. Right gonopod of specimen from Lees Camp, mesal view. 53. Same, Lateral view. 54. Ninth legpair of specimen from Lees Camp, anterior view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cx, coxa; cxp, coxal process; f, flagellum; pac, posterior angiocoxite; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7836704" httpUri="https://zenodo.org/record/7836704/files/figure.png" pageId="5" pageNumber="328">Figs 52, 53</figureCitation>
), Kennedy Creek (
<figureCitation box="[577,658,1842,1869]" captionStart="FIGURES 4754" captionStartId="14.[151,264,1809,1833]" captionTargetBox="[213,1367,188,1786]" captionTargetId="figure-20@14.[213,1375,181,1786]" captionTargetPageId="14" captionText="FIGURES 4754. Miniaria richarti n. gen., n. sp., male paratype. 47. Right gonopod of specimen from Hospital Creek, lateral view. 48. Right gonopod of specimen from Centralia, lateral view. 49. Right gonopod of specimen from Toutle, lateral view. 50. Gonopods of specimen from Elochoman River Road, anterior view. 51. Right gonopod of specimen from Kennedy Creek, mesal view. 52. Right gonopod of specimen from Lees Camp, mesal view. 53. Same, Lateral view. 54. Ninth legpair of specimen from Lees Camp, anterior view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cx, coxa; cxp, coxal process; f, flagellum; pac, posterior angiocoxite; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7836704" httpUri="https://zenodo.org/record/7836704/files/figure.png" pageId="5" pageNumber="328">Fig. 51</figureCitation>
) and Toutle (
<figureCitation box="[809,891,1843,1869]" captionStart="FIGURES 4754" captionStartId="14.[151,264,1809,1833]" captionTargetBox="[213,1367,188,1786]" captionTargetId="figure-20@14.[213,1375,181,1786]" captionTargetPageId="14" captionText="FIGURES 4754. Miniaria richarti n. gen., n. sp., male paratype. 47. Right gonopod of specimen from Hospital Creek, lateral view. 48. Right gonopod of specimen from Centralia, lateral view. 49. Right gonopod of specimen from Toutle, lateral view. 50. Gonopods of specimen from Elochoman River Road, anterior view. 51. Right gonopod of specimen from Kennedy Creek, mesal view. 52. Right gonopod of specimen from Lees Camp, mesal view. 53. Same, Lateral view. 54. Ninth legpair of specimen from Lees Camp, anterior view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cx, coxa; cxp, coxal process; f, flagellum; pac, posterior angiocoxite; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7836704" httpUri="https://zenodo.org/record/7836704/files/figure.png" pageId="5" pageNumber="328">Fig. 49</figureCitation>
) males, the anterior angiocoxite bends at almost a right angle—with the Lees Camp (
<figureCitation box="[567,696,1878,1905]" captionStart="FIGURES 4754" captionStartId="14.[151,264,1809,1833]" captionTargetBox="[213,1367,188,1786]" captionTargetId="figure-20@14.[213,1375,181,1786]" captionTargetPageId="14" captionText="FIGURES 4754. Miniaria richarti n. gen., n. sp., male paratype. 47. Right gonopod of specimen from Hospital Creek, lateral view. 48. Right gonopod of specimen from Centralia, lateral view. 49. Right gonopod of specimen from Toutle, lateral view. 50. Gonopods of specimen from Elochoman River Road, anterior view. 51. Right gonopod of specimen from Kennedy Creek, mesal view. 52. Right gonopod of specimen from Lees Camp, mesal view. 53. Same, Lateral view. 54. Ninth legpair of specimen from Lees Camp, anterior view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cx, coxa; cxp, coxal process; f, flagellum; pac, posterior angiocoxite; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7836704" httpUri="https://zenodo.org/record/7836704/files/figure.png" pageId="5" pageNumber="328">Figs 52, 53</figureCitation>
) and Kennedy Creek (
<figureCitation box="[949,1031,1878,1905]" captionStart="FIGURES 4754" captionStartId="14.[151,264,1809,1833]" captionTargetBox="[213,1367,188,1786]" captionTargetId="figure-20@14.[213,1375,181,1786]" captionTargetPageId="14" captionText="FIGURES 4754. Miniaria richarti n. gen., n. sp., male paratype. 47. Right gonopod of specimen from Hospital Creek, lateral view. 48. Right gonopod of specimen from Centralia, lateral view. 49. Right gonopod of specimen from Toutle, lateral view. 50. Gonopods of specimen from Elochoman River Road, anterior view. 51. Right gonopod of specimen from Kennedy Creek, mesal view. 52. Right gonopod of specimen from Lees Camp, mesal view. 53. Same, Lateral view. 54. Ninth legpair of specimen from Lees Camp, anterior view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cx, coxa; cxp, coxal process; f, flagellum; pac, posterior angiocoxite; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7836704" httpUri="https://zenodo.org/record/7836704/files/figure.png" pageId="5" pageNumber="328">Fig. 51</figureCitation>
) males having a distinct knob at the bend. A common feature appears to be the division of the fimbriate posterior angiocoxite into a thin distal branch and a broader, more proximal portion. Parts of specimens on SEM stubs: Hospital Creek, WS36-10; Kennedy Creek, WS36-3; Lees Camp, WS36-4.
</paragraph>
</subSubSection>
</treatment>
</document>