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<document ID-DOI="10.11646/zootaxa.5264.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7836674" ID-ZooBank="286F4C82-D3A7-455B-8174-354BFBF5D28F" approvalRequired="2" approvalRequired_for_textStreams="2" checkinTime="1681728586229" checkinUser="plazi" docAuthor="Shear, William A. &amp; Marek, Paul E." docDate="2023" docId="CC6F87FEFFF360470AB8FF57FB88BC0E" docLanguage="en" docName="zootaxa.5264.3.2.pdf" docOrigin="Zootaxa 5264 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5264.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Miniaria Shear &amp; Marek 2023, new genus" docType="treatment" docVersion="2" lastPageNumber="325" masterDocId="3056FF86FFF160450A2FFFC0FF9CB922" masterDocTitle="The millipede family Striariidae Bollman, 1893. VIII. Three new genera and four new species of minute millipedes from Oregon and Washington, USA (Diplopoda, Chordeumatida, Striarioidea)" masterLastPageNumber="340" masterPageNumber="323" pageNumber="325" updateTime="1681729003861" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The millipede family Striariidae Bollman, 1893. VIII. Three new genera and four new species of minute millipedes from Oregon and Washington, USA (Diplopoda, Chordeumatida, Striarioidea)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shear, William A.</mods:namePart>
<mods:affiliation>Professor Emeritus, Department of Biology, Hampden-Sydney College, Hampden-Sydney VA 23943 USA, current address: 1950 Price Drive, Farmville VA 23901 USA.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Marek, Paul E.</mods:namePart>
<mods:affiliation>Department of Entomology, Virginia Tech, Blacksburg VA 24061 USA.</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-04-14</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5264</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>323</mods:start>
<mods:end>340</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5264.3.2</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5264.3.2</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7836674</mods:identifier>
<mods:identifier type="ZooBank">286F4C82-D3A7-455B-8174-354BFBF5D28F</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:CC6F87FEFFF360470AB8FF57FB88BC0E" httpUri="http://treatment.plazi.org/id/CC6F87FEFFF360470AB8FF57FB88BC0E" lastPageNumber="325" pageId="2" pageNumber="325">
<subSubSection box="[151,607,151,177]" pageId="2" pageNumber="325" type="nomenclature">
<paragraph blockId="2.[151,607,151,177]" box="[151,607,151,177]" pageId="2" pageNumber="325">
<heading bold="true" box="[151,607,151,177]" fontSize="11" level="1" pageId="2" pageNumber="325" reason="1">
<emphasis bold="true" box="[151,607,151,177]" pageId="2" pageNumber="325">
<taxonomicName authority="Shear &amp; Marek" authorityName="Shear &amp; Marek" authorityYear="2023" box="[151,464,151,177]" class="Diplopoda" family="Striariidae" genus="Miniaria" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="325" phylum="Arthropoda" rank="genus" status="new genus">
<emphasis bold="true" box="[151,259,151,177]" italics="true" pageId="2" pageNumber="325">Miniaria</emphasis>
Shear &amp; Marek
</taxonomicName>
,
<taxonomicNameLabel box="[477,607,151,177]" pageId="2" pageNumber="325" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,780,219,244]" pageId="2" pageNumber="325" type="materials_examined">
<paragraph blockId="2.[151,780,219,244]" box="[151,780,219,244]" pageId="2" pageNumber="325">
<typeStatus box="[151,201,220,244]" pageId="2" pageNumber="325">Type</typeStatus>
species:
<taxonomicName authority="Shear &amp; Marek" authorityName="Shear &amp; Marek" authorityYear="2023" box="[294,645,220,244]" class="Diplopoda" family="Striariidae" genus="Miniaria" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="325" phylum="Arthropoda" rank="species" species="ramifera" status="new species">
<emphasis box="[294,480,220,244]" italics="true" pageId="2" pageNumber="325">Miniaria ramifera</emphasis>
Shear &amp; Marek
</taxonomicName>
,
<emphasis bold="true" box="[655,780,219,244]" pageId="2" pageNumber="325">
<taxonomicNameLabel box="[655,780,219,244]" pageId="2" pageNumber="325" rank="species">new species</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="325" type="diagnosis">
<paragraph blockId="2.[151,1437,289,1324]" pageId="2" pageNumber="325">
<emphasis bold="true" box="[151,274,289,315]" pageId="2" pageNumber="325">Diagnosis.</emphasis>
A genus of
<taxonomicName authorityName="Bollman" authorityYear="1893" box="[414,531,290,316]" class="Diplopoda" family="Striariidae" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="324" phylum="Arthropoda" rank="subFamily" subFamily="Striariinae">Striariinae</taxonomicName>
consisting of two species, distinct from other small striariids in the form of the male labrum, which is convex distally and with sharp lateral corners, but lacking curved processes (
<emphasis bold="true" box="[1304,1341,326,352]" pageId="2" pageNumber="325">lab</emphasis>
,
<figureCitation box="[1358,1429,326,352]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1908,1932]" captionTargetBox="[188,1400,525,1884]" captionTargetId="figure-137@3.[188,1400,525,1884]" captionTargetPageId="3" captionText="FIGURES 1–4. Miniaria ramifera n. gen., n. sp., male holotype. 1. Head and legpairs 1 and 2, anterioventral view. 2. Left leg 1, posterior view. 3. Coxae of legpair 2, ventral view. 4. Legpairs 3 and 4, ventral view. Abbreviations: cf, coxal flask of coxa 3; cx3, body of coxa 3; lab, labrum; L (numeral), numbered legpair; m, mandibular stipes; om, ommatidium; pf (numeral), prefemur of numbered legpair; vd, opening of vas deferens." figureDoi="http://doi.org/10.5281/zenodo.7836676" httpUri="https://zenodo.org/record/7836676/files/figure.png" pageId="2" pageNumber="325">Figs 1</figureCitation>
,
<figureCitation box="[151,182,362,388]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1848,1872]" captionTargetBox="[174,1414,181,1823]" captionTargetId="figure-20@6.[174,1414,181,1823]" captionTargetPageId="6" captionText="FIGURES 9–14. Miniaria n. gen. species. 9–11. Miniaria ramifera n. gen., n. sp., male holotype. 9. Gonopods, posterior view. 10. Legpair 9 and coxae of legpair 10, anterior view. 11. Tip of gonopod anterior angiocoxite, ventral view. 12–14. Miniaria richarti n. gen., n. sp., male from Germany Creek. 12. Head and legpair 1, ventrolateral view. 13. Pleurotergites 11 and 12, lateral view. 14. Pleurotergites 12 and 13, dorsal view. Abbreviations: aac, anterior angiocoxite; cx, gonopod coxa; cx (numeral), coxa of numbered legpair; lab, labrum; L1, legpair 1; m, mandibular stipes; numerals, ring numbers; om, ommatidium." figureDoi="http://doi.org/10.5281/zenodo.7836682" httpUri="https://zenodo.org/record/7836682/files/figure.png" pageId="2" pageNumber="325">12</figureCitation>
). Additional distinguishing characters of the males: first legpair (
<figureCitation box="[931,1001,362,388]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1908,1932]" captionTargetBox="[188,1400,525,1884]" captionTargetId="figure-137@3.[188,1400,525,1884]" captionTargetPageId="3" captionText="FIGURES 1–4. Miniaria ramifera n. gen., n. sp., male holotype. 1. Head and legpairs 1 and 2, anterioventral view. 2. Left leg 1, posterior view. 3. Coxae of legpair 2, ventral view. 4. Legpairs 3 and 4, ventral view. Abbreviations: cf, coxal flask of coxa 3; cx3, body of coxa 3; lab, labrum; L (numeral), numbered legpair; m, mandibular stipes; om, ommatidium; pf (numeral), prefemur of numbered legpair; vd, opening of vas deferens." figureDoi="http://doi.org/10.5281/zenodo.7836676" httpUri="https://zenodo.org/record/7836676/files/figure.png" pageId="2" pageNumber="325">Fig. 2</figureCitation>
) lacks needle setae; flasks of coxae 3 very short, with subterminal processes; fourth legpair largest but with short, wide podomeres (
<emphasis bold="true" box="[1203,1243,398,424]" pageId="2" pageNumber="325">L4,</emphasis>
<figureCitation box="[1249,1317,398,424]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1908,1932]" captionTargetBox="[188,1400,525,1884]" captionTargetId="figure-137@3.[188,1400,525,1884]" captionTargetPageId="3" captionText="FIGURES 1–4. Miniaria ramifera n. gen., n. sp., male holotype. 1. Head and legpairs 1 and 2, anterioventral view. 2. Left leg 1, posterior view. 3. Coxae of legpair 2, ventral view. 4. Legpairs 3 and 4, ventral view. Abbreviations: cf, coxal flask of coxa 3; cx3, body of coxa 3; lab, labrum; L (numeral), numbered legpair; m, mandibular stipes; om, ommatidium; pf (numeral), prefemur of numbered legpair; vd, opening of vas deferens." figureDoi="http://doi.org/10.5281/zenodo.7836676" httpUri="https://zenodo.org/record/7836676/files/figure.png" pageId="2" pageNumber="325">Fig. 4</figureCitation>
); gonopod colpocoxites strongly reduced (
<figureCitation box="[508,582,434,460]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1848,1872]" captionTargetBox="[174,1414,181,1823]" captionTargetId="figure-20@6.[174,1414,181,1823]" captionTargetPageId="6" captionText="FIGURES 9–14. Miniaria n. gen. species. 9–11. Miniaria ramifera n. gen., n. sp., male holotype. 9. Gonopods, posterior view. 10. Legpair 9 and coxae of legpair 10, anterior view. 11. Tip of gonopod anterior angiocoxite, ventral view. 12–14. Miniaria richarti n. gen., n. sp., male from Germany Creek. 12. Head and legpair 1, ventrolateral view. 13. Pleurotergites 11 and 12, lateral view. 14. Pleurotergites 12 and 13, dorsal view. Abbreviations: aac, anterior angiocoxite; cx, gonopod coxa; cx (numeral), coxa of numbered legpair; lab, labrum; L1, legpair 1; m, mandibular stipes; numerals, ring numbers; om, ommatidium." figureDoi="http://doi.org/10.5281/zenodo.7836682" httpUri="https://zenodo.org/record/7836682/files/figure.png" pageId="2" pageNumber="325">Figs 9</figureCitation>
,
<figureCitation box="[596,627,434,460]" captionStart="FIGURES 15–19" captionStartId="7.[151,264,1670,1694]" captionTargetBox="[162,1425,181,1646]" captionTargetId="figure-17@7.[161,1426,181,1646]" captionTargetPageId="7" captionText="FIGURES 15–19. Miniaria richarti n. gen., n. sp., male from Germany Creek. 15. Segmental seta. 16. Telson, lateral view. 17. Tarsus 3, lateral view. 18. Tip of tarsus 4, lateral view. 19. Gonopods, posterior view. Abbreviations: aac, anterior angiocoxite; cx, coxae of gonopods; pac, posterior angiocoxite; s, sternum of gonopods; sp, spinnerets; t, telson; x, comb seta." figureDoi="http://doi.org/10.5281/zenodo.7836686" httpUri="https://zenodo.org/record/7836686/files/figure.png" pageId="2" pageNumber="325">19</figureCitation>
); ninth legpair with coxal pores and curved coxal processes (
<emphasis bold="true" box="[1330,1380,434,460]" pageId="2" pageNumber="325">cxp,</emphasis>
<figureCitation captionStart="FIGURES 9–14" captionStartId="6.[151,264,1848,1872]" captionTargetBox="[174,1414,181,1823]" captionTargetId="figure-20@6.[174,1414,181,1823]" captionTargetPageId="6" captionText="FIGURES 9–14. Miniaria n. gen. species. 9–11. Miniaria ramifera n. gen., n. sp., male holotype. 9. Gonopods, posterior view. 10. Legpair 9 and coxae of legpair 10, anterior view. 11. Tip of gonopod anterior angiocoxite, ventral view. 12–14. Miniaria richarti n. gen., n. sp., male from Germany Creek. 12. Head and legpair 1, ventrolateral view. 13. Pleurotergites 11 and 12, lateral view. 14. Pleurotergites 12 and 13, dorsal view. Abbreviations: aac, anterior angiocoxite; cx, gonopod coxa; cx (numeral), coxa of numbered legpair; lab, labrum; L1, legpair 1; m, mandibular stipes; numerals, ring numbers; om, ommatidium." figureDoi="http://doi.org/10.5281/zenodo.7836682" httpUri="https://zenodo.org/record/7836682/files/figure.png" pageId="2" pageNumber="325">Figs 10</figureCitation>
,
<figureCitation box="[193,225,470,497]" captionStart="FIGURES 24–30" captionStartId="10.[151,264,1820,1844]" captionTargetBox="[164,1423,183,1796]" captionTargetId="figure-20@10.[164,1423,181,1796]" captionTargetPageId="10" captionText="FIGURES 24–30. Male striariid millipedes. Figs. 24, 25. Miniaria richarti n. gen., n. sp., male. 24. Gonopods of specimen from Pin Creek, ventral view. 25. Legpair 9 and coxae 10 of specimen from Kennedy Creek, ventral view. Figs. 26–30. Tigraria oregonensis n. gen., n. sp. male. 26. Head and first 3 legpairs, ventral view. 27. Collum, lateral view. 28. Pleurotergites 13, 14, dorsal view. 29. Segmental seta. 30. Left leg 2, ventral view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; col, collum; cx, coxa of gonopods; cx (numeral), coxa of numbered legpair; L (numeral), numbered legpair; lab, labrum; m, mandibular stipes; numeral, ring numbers; s, sternite of gonopods; vd, opening of vas deferens." figureDoi="http://doi.org/10.5281/zenodo.7836690" httpUri="https://zenodo.org/record/7836690/files/figure.png" pageId="2" pageNumber="325">25</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection box="[199,1238,506,532]" pageId="2" pageNumber="325" type="etymology">
<paragraph blockId="2.[151,1437,289,1324]" box="[199,1238,506,532]" pageId="2" pageNumber="325">
<emphasis bold="true" box="[199,334,506,532]" pageId="2" pageNumber="325">Etymology.</emphasis>
The genus name is feminine in gender and is an arbitrary combination of letters.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="325" type="description">
<paragraph blockId="2.[151,1437,289,1324]" pageId="2" pageNumber="325">
<emphasis bold="true" box="[199,337,541,567]" pageId="2" pageNumber="325">Description</emphasis>
. Small species of
<taxonomicName authorityName="Bollman" authorityYear="1893" box="[547,664,542,568]" class="Diplopoda" family="Striariidae" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="324" phylum="Arthropoda" rank="subFamily" subFamily="Striariinae">Striariinae</taxonomicName>
with 28 postcephalic rings. Length
<quantity box="[1073,1169,542,568]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="325" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
, width about
<quantity metricMagnitude="-4" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="2" pageNumber="325" unit="mm" value="0.275" valueMax="0.3" valueMin="0.25">0.25–0.30 mm</quantity>
. Color white, with single black ommatidium on each side of head. Antennae relatively short, robust, clavate. Head, except for labrum, covered with fine setae and small tubercles (
<figureCitation box="[954,1021,614,640]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1908,1932]" captionTargetBox="[188,1400,525,1884]" captionTargetId="figure-137@3.[188,1400,525,1884]" captionTargetPageId="3" captionText="FIGURES 1–4. Miniaria ramifera n. gen., n. sp., male holotype. 1. Head and legpairs 1 and 2, anterioventral view. 2. Left leg 1, posterior view. 3. Coxae of legpair 2, ventral view. 4. Legpairs 3 and 4, ventral view. Abbreviations: cf, coxal flask of coxa 3; cx3, body of coxa 3; lab, labrum; L (numeral), numbered legpair; m, mandibular stipes; om, ommatidium; pf (numeral), prefemur of numbered legpair; vd, opening of vas deferens." figureDoi="http://doi.org/10.5281/zenodo.7836676" httpUri="https://zenodo.org/record/7836676/files/figure.png" pageId="2" pageNumber="325">Fig. 1</figureCitation>
). Collum ornamented by closely set small tubercles, coalescing into vague crests in posterior fifth of collum length. Metazonites with twelve subequal crests (
<figureCitation box="[233,370,686,712]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1848,1872]" captionTargetBox="[174,1414,181,1823]" captionTargetId="figure-20@6.[174,1414,181,1823]" captionTargetPageId="6" captionText="FIGURES 9–14. Miniaria n. gen. species. 9–11. Miniaria ramifera n. gen., n. sp., male holotype. 9. Gonopods, posterior view. 10. Legpair 9 and coxae of legpair 10, anterior view. 11. Tip of gonopod anterior angiocoxite, ventral view. 12–14. Miniaria richarti n. gen., n. sp., male from Germany Creek. 12. Head and legpair 1, ventrolateral view. 13. Pleurotergites 11 and 12, lateral view. 14. Pleurotergites 12 and 13, dorsal view. Abbreviations: aac, anterior angiocoxite; cx, gonopod coxa; cx (numeral), coxa of numbered legpair; lab, labrum; L1, legpair 1; m, mandibular stipes; numerals, ring numbers; om, ommatidium." figureDoi="http://doi.org/10.5281/zenodo.7836682" httpUri="https://zenodo.org/record/7836682/files/figure.png" pageId="2" pageNumber="325">Figs 13, 14</figureCitation>
); metazonital setae long, prominent, with longitudinal grooves and feathered tips (
<figureCitation box="[1332,1423,686,713]" captionStart="FIGURES 15–19" captionStartId="7.[151,264,1670,1694]" captionTargetBox="[162,1425,181,1646]" captionTargetId="figure-17@7.[161,1426,181,1646]" captionTargetPageId="7" captionText="FIGURES 15–19. Miniaria richarti n. gen., n. sp., male from Germany Creek. 15. Segmental seta. 16. Telson, lateral view. 17. Tarsus 3, lateral view. 18. Tip of tarsus 4, lateral view. 19. Gonopods, posterior view. Abbreviations: aac, anterior angiocoxite; cx, coxae of gonopods; pac, posterior angiocoxite; s, sternum of gonopods; sp, spinnerets; t, telson; x, comb seta." figureDoi="http://doi.org/10.5281/zenodo.7836686" httpUri="https://zenodo.org/record/7836686/files/figure.png" pageId="2" pageNumber="325">Figs 15</figureCitation>
). Telson with three lobes only shallowly separated, not prominent; spinnerets directed posteriad (
<emphasis bold="true" box="[1235,1269,722,748]" pageId="2" pageNumber="325">sp,</emphasis>
<figureCitation box="[1277,1360,722,748]" captionStart="FIGURES 15–19" captionStartId="7.[151,264,1670,1694]" captionTargetBox="[162,1425,181,1646]" captionTargetId="figure-17@7.[161,1426,181,1646]" captionTargetPageId="7" captionText="FIGURES 15–19. Miniaria richarti n. gen., n. sp., male from Germany Creek. 15. Segmental seta. 16. Telson, lateral view. 17. Tarsus 3, lateral view. 18. Tip of tarsus 4, lateral view. 19. Gonopods, posterior view. Abbreviations: aac, anterior angiocoxite; cx, coxae of gonopods; pac, posterior angiocoxite; s, sternum of gonopods; sp, spinnerets; t, telson; x, comb seta." figureDoi="http://doi.org/10.5281/zenodo.7836686" httpUri="https://zenodo.org/record/7836686/files/figure.png" pageId="2" pageNumber="325">Fig. 16</figureCitation>
). Legs with specialized flattened setae with median ridges and long, filamentous extensions (
<figureCitation box="[1113,1181,758,784]" captionStart="FIGURES 5–8" captionStartId="4.[151,264,1492,1516]" captionTargetBox="[155,1434,181,1467]" captionTargetId="figure-20@4.[153,1434,181,1468]" captionTargetPageId="4" captionText="FIGURES 5–8. Miniaria ramifera n. gen., n. sp., male holotype. 5. Legpairs 2 and 3, ventral view. 6. Tarsus of left leg 3, lateral view. 7. Comb seta of tarsus 3. 8. Legpairs 4–6, ventral view. Abbreviations: cf, coxal flask of leg 3; L (numeral), numbered legpair;pf(numeral),prefemurofnumberedleg;w,accessory clawoftarsus3,x,combsetae;y,recessedporeplate;z,sensory seta." figureDoi="http://doi.org/10.5281/zenodo.7836680" httpUri="https://zenodo.org/record/7836680/files/figure.png" pageId="2" pageNumber="325">Fig. 6</figureCitation>
).
</paragraph>
<paragraph blockId="2.[151,1437,289,1324]" pageId="2" pageNumber="325">
The following secondary sexual characters occur in males. Head frontally flattened with slight swellings below antennal sockets. Labrum flat and smooth, slightly concave, distal corners distinctly angular, not rounded (
<emphasis bold="true" box="[1392,1437,830,856]" pageId="2" pageNumber="325">lab,</emphasis>
<figureCitation box="[151,216,866,892]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1908,1932]" captionTargetBox="[188,1400,525,1884]" captionTargetId="figure-137@3.[188,1400,525,1884]" captionTargetPageId="3" captionText="FIGURES 1–4. Miniaria ramifera n. gen., n. sp., male holotype. 1. Head and legpairs 1 and 2, anterioventral view. 2. Left leg 1, posterior view. 3. Coxae of legpair 2, ventral view. 4. Legpairs 3 and 4, ventral view. Abbreviations: cf, coxal flask of coxa 3; cx3, body of coxa 3; lab, labrum; L (numeral), numbered legpair; m, mandibular stipes; om, ommatidium; pf (numeral), prefemur of numbered legpair; vd, opening of vas deferens." figureDoi="http://doi.org/10.5281/zenodo.7836676" httpUri="https://zenodo.org/record/7836676/files/figure.png" pageId="2" pageNumber="325">Fig. 1</figureCitation>
). Mandibular stipes with serrate anteriodistal margin, prominent distal tooth (
<emphasis bold="true" box="[1090,1120,866,892]" pageId="2" pageNumber="325">m,</emphasis>
<figureCitation box="[1127,1184,866,892]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1908,1932]" captionTargetBox="[188,1400,525,1884]" captionTargetId="figure-137@3.[188,1400,525,1884]" captionTargetPageId="3" captionText="FIGURES 1–4. Miniaria ramifera n. gen., n. sp., male holotype. 1. Head and legpairs 1 and 2, anterioventral view. 2. Left leg 1, posterior view. 3. Coxae of legpair 2, ventral view. 4. Legpairs 3 and 4, ventral view. Abbreviations: cf, coxal flask of coxa 3; cx3, body of coxa 3; lab, labrum; L (numeral), numbered legpair; m, mandibular stipes; om, ommatidium; pf (numeral), prefemur of numbered legpair; vd, opening of vas deferens." figureDoi="http://doi.org/10.5281/zenodo.7836676" httpUri="https://zenodo.org/record/7836676/files/figure.png" pageId="2" pageNumber="325">Fig 1</figureCitation>
). First legpair (
<figureCitation box="[1360,1428,866,892]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1908,1932]" captionTargetBox="[188,1400,525,1884]" captionTargetId="figure-137@3.[188,1400,525,1884]" captionTargetPageId="3" captionText="FIGURES 1–4. Miniaria ramifera n. gen., n. sp., male holotype. 1. Head and legpairs 1 and 2, anterioventral view. 2. Left leg 1, posterior view. 3. Coxae of legpair 2, ventral view. 4. Legpairs 3 and 4, ventral view. Abbreviations: cf, coxal flask of coxa 3; cx3, body of coxa 3; lab, labrum; L (numeral), numbered legpair; m, mandibular stipes; om, ommatidium; pf (numeral), prefemur of numbered legpair; vd, opening of vas deferens." figureDoi="http://doi.org/10.5281/zenodo.7836676" httpUri="https://zenodo.org/record/7836676/files/figure.png" pageId="2" pageNumber="325">Fig. 2</figureCitation>
) enlarged, femora curved, lacking needle setae, distal podomeres with few long, flattened setae ventrally. Second legpair larger than first; openings of
<emphasis box="[575,734,938,964]" italics="true" pageId="2" pageNumber="325">vas deferentia</emphasis>
separate, subtended by long, flattened, ribbon-like setae (
<emphasis bold="true" box="[1400,1437,938,964]" pageId="2" pageNumber="325">vd,</emphasis>
<figureCitation box="[151,224,974,1000]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1908,1932]" captionTargetBox="[188,1400,525,1884]" captionTargetId="figure-137@3.[188,1400,525,1884]" captionTargetPageId="3" captionText="FIGURES 1–4. Miniaria ramifera n. gen., n. sp., male holotype. 1. Head and legpairs 1 and 2, anterioventral view. 2. Left leg 1, posterior view. 3. Coxae of legpair 2, ventral view. 4. Legpairs 3 and 4, ventral view. Abbreviations: cf, coxal flask of coxa 3; cx3, body of coxa 3; lab, labrum; L (numeral), numbered legpair; m, mandibular stipes; om, ommatidium; pf (numeral), prefemur of numbered legpair; vd, opening of vas deferens." figureDoi="http://doi.org/10.5281/zenodo.7836676" httpUri="https://zenodo.org/record/7836676/files/figure.png" pageId="2" pageNumber="325">Figs 3</figureCitation>
,
<figureCitation box="[240,256,974,1001]" captionStart="FIGURES 5–8" captionStartId="4.[151,264,1492,1516]" captionTargetBox="[155,1434,181,1467]" captionTargetId="figure-20@4.[153,1434,181,1468]" captionTargetPageId="4" captionText="FIGURES 5–8. Miniaria ramifera n. gen., n. sp., male holotype. 5. Legpairs 2 and 3, ventral view. 6. Tarsus of left leg 3, lateral view. 7. Comb seta of tarsus 3. 8. Legpairs 4–6, ventral view. Abbreviations: cf, coxal flask of leg 3; L (numeral), numbered legpair;pf(numeral),prefemurofnumberedleg;w,accessory clawoftarsus3,x,combsetae;y,recessedporeplate;z,sensory seta." figureDoi="http://doi.org/10.5281/zenodo.7836680" httpUri="https://zenodo.org/record/7836680/files/figure.png" pageId="2" pageNumber="325">5</figureCitation>
). Third legpair robust, coxal flasks short, with subterminal process (
<figureCitation box="[1049,1120,974,1001]" captionStart="FIGURES 5–8" captionStartId="4.[151,264,1492,1516]" captionTargetBox="[155,1434,181,1467]" captionTargetId="figure-20@4.[153,1434,181,1468]" captionTargetPageId="4" captionText="FIGURES 5–8. Miniaria ramifera n. gen., n. sp., male holotype. 5. Legpairs 2 and 3, ventral view. 6. Tarsus of left leg 3, lateral view. 7. Comb seta of tarsus 3. 8. Legpairs 4–6, ventral view. Abbreviations: cf, coxal flask of leg 3; L (numeral), numbered legpair;pf(numeral),prefemurofnumberedleg;w,accessory clawoftarsus3,x,combsetae;y,recessedporeplate;z,sensory seta." figureDoi="http://doi.org/10.5281/zenodo.7836680" httpUri="https://zenodo.org/record/7836680/files/figure.png" pageId="2" pageNumber="325">Fig. 5</figureCitation>
); tarsi with special sensory array including comb setae and recessed pore plate (
<figureCitation box="[732,800,1010,1036]" captionStart="FIGURES 5–8" captionStartId="4.[151,264,1492,1516]" captionTargetBox="[155,1434,181,1467]" captionTargetId="figure-20@4.[153,1434,181,1468]" captionTargetPageId="4" captionText="FIGURES 5–8. Miniaria ramifera n. gen., n. sp., male holotype. 5. Legpairs 2 and 3, ventral view. 6. Tarsus of left leg 3, lateral view. 7. Comb seta of tarsus 3. 8. Legpairs 4–6, ventral view. Abbreviations: cf, coxal flask of leg 3; L (numeral), numbered legpair;pf(numeral),prefemurofnumberedleg;w,accessory clawoftarsus3,x,combsetae;y,recessedporeplate;z,sensory seta." figureDoi="http://doi.org/10.5281/zenodo.7836680" httpUri="https://zenodo.org/record/7836680/files/figure.png" pageId="2" pageNumber="325">Fig. 6</figureCitation>
; see Discussion section below for further details). Fourth legpair the largest; prefemora and femora nearly as wide as long (
<emphasis bold="true" box="[901,940,1046,1072]" pageId="2" pageNumber="325">L4,</emphasis>
<figureCitation box="[948,1021,1046,1072]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1908,1932]" captionTargetBox="[188,1400,525,1884]" captionTargetId="figure-137@3.[188,1400,525,1884]" captionTargetPageId="3" captionText="FIGURES 1–4. Miniaria ramifera n. gen., n. sp., male holotype. 1. Head and legpairs 1 and 2, anterioventral view. 2. Left leg 1, posterior view. 3. Coxae of legpair 2, ventral view. 4. Legpairs 3 and 4, ventral view. Abbreviations: cf, coxal flask of coxa 3; cx3, body of coxa 3; lab, labrum; L (numeral), numbered legpair; m, mandibular stipes; om, ommatidium; pf (numeral), prefemur of numbered legpair; vd, opening of vas deferens." figureDoi="http://doi.org/10.5281/zenodo.7836676" httpUri="https://zenodo.org/record/7836676/files/figure.png" pageId="2" pageNumber="325">Figs 4</figureCitation>
,
<figureCitation box="[1034,1050,1046,1072]" captionStart="FIGURES 5–8" captionStartId="4.[151,264,1492,1516]" captionTargetBox="[155,1434,181,1467]" captionTargetId="figure-20@4.[153,1434,181,1468]" captionTargetPageId="4" captionText="FIGURES 5–8. Miniaria ramifera n. gen., n. sp., male holotype. 5. Legpairs 2 and 3, ventral view. 6. Tarsus of left leg 3, lateral view. 7. Comb seta of tarsus 3. 8. Legpairs 4–6, ventral view. Abbreviations: cf, coxal flask of leg 3; L (numeral), numbered legpair;pf(numeral),prefemurofnumberedleg;w,accessory clawoftarsus3,x,combsetae;y,recessedporeplate;z,sensory seta." figureDoi="http://doi.org/10.5281/zenodo.7836680" httpUri="https://zenodo.org/record/7836680/files/figure.png" pageId="2" pageNumber="325">8</figureCitation>
). Fifth, sixth and seventh legpairs enlarged; seventh coxae unmodified. Tenth coxae with glands, not modified or enlarged (
<emphasis bold="true" box="[1149,1211,1082,1108]" pageId="2" pageNumber="325">cx10,</emphasis>
<figureCitation box="[1218,1300,1082,1108]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1848,1872]" captionTargetBox="[174,1414,181,1823]" captionTargetId="figure-20@6.[174,1414,181,1823]" captionTargetPageId="6" captionText="FIGURES 9–14. Miniaria n. gen. species. 9–11. Miniaria ramifera n. gen., n. sp., male holotype. 9. Gonopods, posterior view. 10. Legpair 9 and coxae of legpair 10, anterior view. 11. Tip of gonopod anterior angiocoxite, ventral view. 12–14. Miniaria richarti n. gen., n. sp., male from Germany Creek. 12. Head and legpair 1, ventrolateral view. 13. Pleurotergites 11 and 12, lateral view. 14. Pleurotergites 12 and 13, dorsal view. Abbreviations: aac, anterior angiocoxite; cx, gonopod coxa; cx (numeral), coxa of numbered legpair; lab, labrum; L1, legpair 1; m, mandibular stipes; numerals, ring numbers; om, ommatidium." figureDoi="http://doi.org/10.5281/zenodo.7836682" httpUri="https://zenodo.org/record/7836682/files/figure.png" pageId="2" pageNumber="325">Fig. 10</figureCitation>
).
</paragraph>
<paragraph blockId="2.[151,1437,289,1324]" pageId="2" pageNumber="325">
Gonopods (
<figureCitation box="[327,436,1118,1144]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1848,1872]" captionTargetBox="[174,1414,181,1823]" captionTargetId="figure-20@6.[174,1414,181,1823]" captionTargetPageId="6" captionText="FIGURES 9–14. Miniaria n. gen. species. 9–11. Miniaria ramifera n. gen., n. sp., male holotype. 9. Gonopods, posterior view. 10. Legpair 9 and coxae of legpair 10, anterior view. 11. Tip of gonopod anterior angiocoxite, ventral view. 12–14. Miniaria richarti n. gen., n. sp., male from Germany Creek. 12. Head and legpair 1, ventrolateral view. 13. Pleurotergites 11 and 12, lateral view. 14. Pleurotergites 12 and 13, dorsal view. Abbreviations: aac, anterior angiocoxite; cx, gonopod coxa; cx (numeral), coxa of numbered legpair; lab, labrum; L1, legpair 1; m, mandibular stipes; numerals, ring numbers; om, ommatidium." figureDoi="http://doi.org/10.5281/zenodo.7836682" httpUri="https://zenodo.org/record/7836682/files/figure.png" pageId="2" pageNumber="325">Figs 9, 11</figureCitation>
,
<figureCitation box="[446,518,1118,1144]" captionStart-0="FIGURES 15–19" captionStart-1="FIGURES 20–23" captionStart-2="FIGURES 24–30" captionStartId-0="7.[151,264,1670,1694]" captionStartId-1="8.[151,264,1515,1539]" captionStartId-2="10.[151,264,1820,1844]" captionTargetBox-0="[162,1425,181,1646]" captionTargetBox-1="[151,1436,184,1491]" captionTargetBox-2="[164,1423,183,1796]" captionTargetId-0="figure-17@7.[161,1426,181,1646]" captionTargetId-1="figure-20@8.[151,1436,181,1491]" captionTargetId-2="figure-20@10.[164,1423,181,1796]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="10" captionText-0="FIGURES 15–19. Miniaria richarti n. gen., n. sp., male from Germany Creek. 15. Segmental seta. 16. Telson, lateral view. 17. Tarsus 3, lateral view. 18. Tip of tarsus 4, lateral view. 19. Gonopods, posterior view. Abbreviations: aac, anterior angiocoxite; cx, coxae of gonopods; pac, posterior angiocoxite; s, sternum of gonopods; sp, spinnerets; t, telson; x, comb seta." captionText-1="FIGURES 20–23. Miniaria richarti n. gen., n. sp., gonopods of specimen from Lee’s Camp. 20. Posterior view. 21. Tips of anterior colpocoxites, ventral view. 22. Posterior view showing sheathing of flagellocoxite by posterior angiocoxite. 23. Posterior view showing colpocoxite and origin of flagellocoxite. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cx, gonopod coxa; f, flagellocoxite; pac, posterior angiocoxite." captionText-2="FIGURES 24–30. Male striariid millipedes. Figs. 24, 25. Miniaria richarti n. gen., n. sp., male. 24. Gonopods of specimen from Pin Creek, ventral view. 25. Legpair 9 and coxae 10 of specimen from Kennedy Creek, ventral view. Figs. 26–30. Tigraria oregonensis n. gen., n. sp. male. 26. Head and first 3 legpairs, ventral view. 27. Collum, lateral view. 28. Pleurotergites 13, 14, dorsal view. 29. Segmental seta. 30. Left leg 2, ventral view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; col, collum; cx, coxa of gonopods; cx (numeral), coxa of numbered legpair; L (numeral), numbered legpair; lab, labrum; m, mandibular stipes; numeral, ring numbers; s, sternite of gonopods; vd, opening of vas deferens." figureDoi-0="http://doi.org/10.5281/zenodo.7836686" figureDoi-1="http://doi.org/10.5281/zenodo.7836688" figureDoi-2="http://doi.org/10.5281/zenodo.7836690" httpUri-0="https://zenodo.org/record/7836686/files/figure.png" httpUri-1="https://zenodo.org/record/7836688/files/figure.png" httpUri-2="https://zenodo.org/record/7836690/files/figure.png" pageId="2" pageNumber="325">19–24</figureCitation>
,
<figureCitation box="[527,683,1118,1145]" captionStart-0="FIGURES 39–46" captionStart-1="FIGURES 47–54" captionStartId-0="13.[151,262,1817,1841]" captionStartId-1="14.[151,264,1809,1833]" captionTargetBox-0="[216,1378,182,1788]" captionTargetBox-1="[213,1367,188,1786]" captionTargetId-0="figure-17@13.[206,1382,181,1792]" captionTargetId-1="figure-20@14.[213,1375,181,1786]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURES 39–46. Gonopods and ninth legs of male striariid millipedes. Figs. 39–45. Miniaria ramifera n. gen., n. sp. 39. Right gonopod of specimen from Mashel River, mesal view. 40. Same, lateral view. 41. Right gonopod of specimen from Cook- Underwood Road, mesal view. 42. Same, lateral view. 43, Right gonopod of specimen fromAinsworth State Park, mesal view. 44. Right gonopod of specimen from Timber, mesal view. 45. Ninth legpair of specimen from Ainsworth State Park, posterior view. 46. Miniaria richarti n. gen., n. sp., right gonopod of specimen from Hospital Creek, mesal view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cx9, ninth leg coxa; cxp, coxal process; pac, posterior angiocoxite; t9, telopodite of ninth leg." captionText-1="FIGURES 47–54. Miniaria richarti n. gen., n. sp., male paratype. 47. Right gonopod of specimen from Hospital Creek, lateral view. 48. Right gonopod of specimen from Centralia, lateral view. 49. Right gonopod of specimen from Toutle, lateral view. 50. Gonopods of specimen from Elochoman River Road, anterior view. 51. Right gonopod of specimen from Kennedy Creek, mesal view. 52. Right gonopod of specimen from Lee’s Camp, mesal view. 53. Same, Lateral view. 54. Ninth legpair of specimen from Lee’s Camp, anterior view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cx, coxa; cxp, coxal process; f, flagellum; pac, posterior angiocoxite; t9, telopodite of ninth leg." figureDoi-0="http://doi.org/10.5281/zenodo.7836702" figureDoi-1="http://doi.org/10.5281/zenodo.7836704" httpUri-0="https://zenodo.org/record/7836702/files/figure.png" httpUri-1="https://zenodo.org/record/7836704/files/figure.png" pageId="2" pageNumber="325">39–44, 46–53</figureCitation>
), separate from prominent, transverse sternum, coxae large, with two or three setae. Anterior angiocoxites curved, with fimbriate posterior margin distally. Posterior angiocoxites smaller than anterior ones, fimbriate, sheathing single flagellum or flagellocoxite. Colpocoxites membranous-fimbriate, much reduced. Ninth legs (
<figureCitation box="[469,560,1226,1252]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1848,1872]" captionTargetBox="[174,1414,181,1823]" captionTargetId="figure-20@6.[174,1414,181,1823]" captionTargetPageId="6" captionText="FIGURES 9–14. Miniaria n. gen. species. 9–11. Miniaria ramifera n. gen., n. sp., male holotype. 9. Gonopods, posterior view. 10. Legpair 9 and coxae of legpair 10, anterior view. 11. Tip of gonopod anterior angiocoxite, ventral view. 12–14. Miniaria richarti n. gen., n. sp., male from Germany Creek. 12. Head and legpair 1, ventrolateral view. 13. Pleurotergites 11 and 12, lateral view. 14. Pleurotergites 12 and 13, dorsal view. Abbreviations: aac, anterior angiocoxite; cx, gonopod coxa; cx (numeral), coxa of numbered legpair; lab, labrum; L1, legpair 1; m, mandibular stipes; numerals, ring numbers; om, ommatidium." figureDoi="http://doi.org/10.5281/zenodo.7836682" httpUri="https://zenodo.org/record/7836682/files/figure.png" pageId="2" pageNumber="325">Figs 10</figureCitation>
,
<figureCitation box="[576,606,1226,1253]" captionStart="FIGURES 24–30" captionStartId="10.[151,264,1820,1844]" captionTargetBox="[164,1423,183,1796]" captionTargetId="figure-20@10.[164,1423,181,1796]" captionTargetPageId="10" captionText="FIGURES 24–30. Male striariid millipedes. Figs. 24, 25. Miniaria richarti n. gen., n. sp., male. 24. Gonopods of specimen from Pin Creek, ventral view. 25. Legpair 9 and coxae 10 of specimen from Kennedy Creek, ventral view. Figs. 26–30. Tigraria oregonensis n. gen., n. sp. male. 26. Head and first 3 legpairs, ventral view. 27. Collum, lateral view. 28. Pleurotergites 13, 14, dorsal view. 29. Segmental seta. 30. Left leg 2, ventral view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; col, collum; cx, coxa of gonopods; cx (numeral), coxa of numbered legpair; L (numeral), numbered legpair; lab, labrum; m, mandibular stipes; numeral, ring numbers; s, sternite of gonopods; vd, opening of vas deferens." figureDoi="http://doi.org/10.5281/zenodo.7836690" httpUri="https://zenodo.org/record/7836690/files/figure.png" pageId="2" pageNumber="325">25</figureCitation>
,
<figureCitation box="[622,653,1226,1253]" captionStart="FIGURES 39–46" captionStartId="13.[151,262,1817,1841]" captionTargetBox="[216,1378,182,1788]" captionTargetId="figure-17@13.[206,1382,181,1792]" captionTargetPageId="13" captionText="FIGURES 39–46. Gonopods and ninth legs of male striariid millipedes. Figs. 39–45. Miniaria ramifera n. gen., n. sp. 39. Right gonopod of specimen from Mashel River, mesal view. 40. Same, lateral view. 41. Right gonopod of specimen from Cook- Underwood Road, mesal view. 42. Same, lateral view. 43, Right gonopod of specimen fromAinsworth State Park, mesal view. 44. Right gonopod of specimen from Timber, mesal view. 45. Ninth legpair of specimen from Ainsworth State Park, posterior view. 46. Miniaria richarti n. gen., n. sp., right gonopod of specimen from Hospital Creek, mesal view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cx9, ninth leg coxa; cxp, coxal process; pac, posterior angiocoxite; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7836702" httpUri="https://zenodo.org/record/7836702/files/figure.png" pageId="2" pageNumber="325">45</figureCitation>
), variably free or partially fused to sternum; coxae and telopodites fused, coxae with prominent anterior lobe bearing gland pore, also with strongly curved single process; telopodites strongly flattened, setose, with ornament of pointed tubercles (as on other legs).
</paragraph>
</subSubSection>
</treatment>
</document>
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