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<mods:title id="6B94754FCEA491BA1A0E8317DFE8B2F3">Taxonomy of Diamesa steinboecki group (Diptera: Chironomidae: Diamesinae) with description and DNA barcoding of new species. I. Subgroups steinboecki and longipes</mods:title>
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<mods:namePart id="B3CECC6C4D08DC379FDD76CC10FE5A5B">Palatov, Dmitry M.</mods:namePart>
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<taxonomicName id="3DBE4D769452FFEDF5C96078FD036906" ID-CoL="359GS" authority="Kownacki et Kownacka" authorityName="Kownacki et Kownacka (Moubayed &amp; Laville" authorityYear="1983" box="[151,701,1632,1659]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="501" phylum="Arthropoda" rank="species" species="sakartvella">
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<emphasis id="C8CAEAE79452FFEDF5C96078FE326906" bold="true" box="[151,396,1632,1659]" italics="true" pageId="18" pageNumber="501">Diamesa sakartvella</emphasis>
Kownacki 
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Kownacka
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(
<figureCitation id="62852A709452FFEDF5C1609CFE9669E2" box="[159,296,1668,1695]" captionStart="FIGURES 37–44" captionStartId="14.[151,264,1880,1904]" captionTargetBox="[151,1435,349,1857]" captionTargetId="figure-94@14.[151,1435,345,1857]" captionTargetPageId="14" captionText="FIGURES 37–44. Males of Diamesa zagrosica sp. nov. (37, 41), D. sakartvella Kownacki et Kownacka (38, 42) and D. moubayedi sp. nov. (39–40, 43–44). 37, head; 38, 41, hypopygium in dorsal view; 40, hypopygium in dorsal view without tergite IX; 42, wing; 43, tergite IX; 44, anal point in lateral view." figureDoi="http://doi.org/10.5281/zenodo.6457273" httpUri="https://zenodo.org/record/6457273/files/figure.png" pageId="18" pageNumber="501">Figs. 38, 42</figureCitation>
, 
<figureCitation id="62852A709452FFEDF46D609CFE7169E2" box="[307,463,1668,1695]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="18" pageNumber="501">45, 50–51, 57</figureCitation>
, 
<figureCitation id="62852A709452FFEDF485609DFE4669E2" box="[475,504,1669,1695]" captionStart="FIGURES 68–74" captionStartId="23.[151,264,1617,1641]" captionTargetBox="[151,1435,181,1592]" captionTargetId="figure-18@23.[151,1435,181,1593]" captionTargetPageId="23" captionText="FIGURES 68–74. Type localities of Diamesa dragani sp. nov. (68), D. kownackii sp. nov. (69–70), D. maisaraensis sp. nov. (71), D. marinskiyi sp. nov. (72), D. steinboecki Goetghebuer (73) and D. sakartvella Kownacki et Kownacka (74). 68, unnamed stream of Bolshoi On River basin, alt. 2063 m a. sea l., Sayan Mountains (photo by S.V. Dragan); 69, unnamed stream of Somnitelnaya River basin, Wrangel Island (photo by E.A. Makarchenko); 70, Somnitelnaya River, upper stream, alt. 205–210 m a. sea l., Wrangel Island (photo by O.A. Khruleva); 71, Maisara River near Maisara Pass, alt. 4168 m a. sea l., Pamir Mountains (photo by D.M. Palatov); 72, Zienzan River, Tien Shan Mountains, alt. 1923 m a. sea l., Xinjiang Uygur Autonomous Region, China (photo by D.M. Palatov); 73, Balm stream, Massif of Aiguilles–Rouges, French Alps (photo by Pierre Clévenot); 74, Skazdon River near the Tsey (Tseyskoe gorge), alt. 1943 m a. sea l., North Caucasus (photo by D.M. Palatov)." figureDoi="http://doi.org/10.5281/zenodo.6457281" httpUri="https://zenodo.org/record/6457281/files/figure.png" pageId="18" pageNumber="501">74</figureCitation>
)
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<treatmentCitation id="7B1F10E49452FFEDF5C960DCFD7569A1" author="Kownacki A. &amp; Kownacka, M." box="[151,715,1732,1756]" page="21" pageId="18" pageNumber="501" year="1973">
<taxonomicName id="3DBE4D769452FFEDF5C960DCFD7569A1" ID-CoL="359GS" authority="Kownacki et Kownacka, 1973: 21" authorityName="Kownacki et Kownacka (Moubayed &amp; Laville" authorityPageNumber="21" authorityYear="1983" box="[151,715,1732,1756]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="501" phylum="Arthropoda" rank="species" species="sakartvella">
<emphasis id="C8CAEAE79452FFEDF5C960DCFED769A1" box="[151,361,1732,1756]" italics="true" pageId="18" pageNumber="501">Diamesa sakartvella</emphasis>
<bibRefCitation id="9E2F4B049452FFEDF43060DCFD7569A1" author="Kownacki A. &amp; Kownacka, M." box="[366,715,1732,1756]" pageId="18" pageNumber="501" pagination="27 - 37" refId="ref18733" refString="Kownacki A., Kownacka, M. (1973) Chironomidae (Diptera) from the Caucasus, Diamesa Waltl group steinboecki. Bulletin de l'Academie Polonaise des Sciences, Cl. II, Serie des Sciences Biologiques, 21, 27 - 37." type="journal article" year="1973">Kownacki et Kownacka, 1973: 21</bibRefCitation>
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<bibRefCitation id="9E2F4B049452FFEDF78B60DCFC5569A1" author="Langton, P. &amp; Visser, H." box="[725,1003,1732,1756]" pageId="18" pageNumber="501" refId="ref18902" refString="Langton, P. &amp; Visser, H. (2003) Chironomidae exuviae. A key to pupal exuviae of the West Palaearctic Region. Vols. 1 &amp; 2. Freshwater Biological Association. Scientific Publication 64. Expert Center for Taxonomic Information, Amsterdam, 239 pp., CD ROM." type="book" year="2003">Langton &amp; Visser 2003: 43</bibRefCitation>
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; 
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<bibRefCitation id="9E2F4B049452FFEDF6AA60DCFAA169A1" author="Ashe, P. &amp; O'Connor, J. P." box="[1012,1311,1732,1756]" pageId="18" pageNumber="501" refId="ref18121" refString="Ashe, P. &amp; O'Connor, J. P. (2009) A World Catalogue of Chironomidae (Diptera). Part 1. Buchonomyiinae, Chilenomyiinae, Podonominae, Aphroteniinae, Tanypodinae, Usambaromyiinae, Diamesinae, Prodiamesinae and Telmatogetoninae. Irish Biogeographical Society &amp; National Museum of Ireland, Dublin, 445 pp." type="book" year="2009">Ashe &amp; O’Connor 2009: 284</bibRefCitation>
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.
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<emphasis id="C8CAEAE79452FFEDF5C9611CFE366862" bold="true" box="[151,392,1796,1823]" pageId="18" pageNumber="501">Material examened.</emphasis>
<materialsCitation id="4AD63CA89452FFEDF4CD611DFDE1681B" ID-GBIF-Occurrence="3751071312" collectingDate="2019-07-27" collectorName="D. Palatov." country="Russia" county="Greater Caucasus" elevation="1943" latitude="42.78417" location="Alagirsky District" longLatPrecision="1" longitude="43.903667" municipality="Republic" pageId="18" pageNumber="501" specimenCount="11" specimenCount-adult="11" stateProvince="North Ossetia">
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males, 
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: 
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, 
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of 
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, 
<location id="FF61602E9452FFEDF070611CFF50683F" LSID="urn:lsid:plazi:treatment:721787E39452FFECF5C96078FCAF68A0:FF61602E9452FFEDF070611CFF50683F" country="Russia" county="Greater Caucasus" latitude="42.78417" longLatPrecision="1" longitude="43.903667" municipality="Republic" name="Alagirsky District" pageId="18" pageNumber="501" stateProvince="North Ossetia">Alagirsky District</location>
, 
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near the 
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(Tseyskoe gorge) ski resort, altitude 
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<quantity id="3D469B109452FFEDF6896131FB92683E" box="[983,1068,1833,1859]" metricMagnitude="3" metricUnit="m" metricValue="1.943" pageId="18" pageNumber="501" unit="m" value="1943.0">1943 m</quantity>
above sea level
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, 
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, 
<geoCoordinate id="9F8A50329452FFEDF0D56131FEAF681B" degrees="43.903667" direction="east" orientation="longitude" pageId="18" pageNumber="501" precision="1" value="43.903667">E 43.903667</geoCoordinate>
, 
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<collectingDate id="9E44E9DD9452FFEDF4426155FE16681A" box="[284,424,1869,1895]" pageId="18" pageNumber="501" value="2019-07-27">27.VII.2019</collectingDate>
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, leg. 
<collectorName id="574B53239452FFEDF4BD6155FDE1681B" box="[483,607,1868,1895]" pageId="18" pageNumber="501">D. Palatov.</collectorName>
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<emphasis id="C8CAEAE79452FFEDF5996168FEEF68F7" bold="true" box="[199,337,1904,1930]" pageId="18" pageNumber="501">Description</emphasis>
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<emphasis id="C8CAEAE79452FFEDF599618CFEF868D2" bold="true" box="[199,326,1940,1967]" italics="true" pageId="18" pageNumber="501">Adult male</emphasis>
(n = 5, except when otherwise stated). Total length 
<quantity id="3D469B109452FFEDF6D5618DFC4168D2" box="[907,1023,1941,1967]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="3.1" metricValueMin="2.9" pageId="18" pageNumber="501" unit="m" value="3.0" valueMax="3.1" valueMin="2.9">2.9–3.1 m</quantity>
. Total length/wing length 1.04–1.19.
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<paragraph id="FA0136F59452FFEDF59961A0FE3C688B" blockId="18.[151,1436,1796,2038]" pageId="18" pageNumber="501">Coloration. Brown to dark brown. Head, thorax and abdomen dark brown. Legs brown. Wings brownish gray, with brownish veins.</paragraph>
<paragraph id="FA0136F59453FFECF599668FFC166EF4" blockId="19.[151,1437,151,609]" pageId="19" pageNumber="502">
Head. Eyes hairy, reniform. Temporal setae including 7–8 frontals, 7–8 orbitals, 7–9 verticals and 3–4 postorbitals. Frontal tubercles 12 μm height and 56–60 μm width. Clypeus with 9 setae. Antenna with 8 flagellomeres and reduced plume of setae; number and length of these setae on 1–7 flagellomeres respectively: 1–2 (52 μm), 2 (60–72 μm), 2 (60–76 μm), 1–2 (76 μm), 0, 0; terminal flagellomere with 3–4 setae, 60–80 μm long in basal part and with 2 subapical setae, 28–32 μm long. Length of 1–8 flagellomeres (μm): 80, 40, 40, 32, 32, 28, 32, 164; AR 0.58. Palpomere length (μm): 40, 76, 128, 92, 112. Palpomere 
<quantity id="3D469B109453FFECF6196753FCCC6E18" box="[839,882,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="19" pageNumber="502" unit="in" value="3.0">3 in</quantity>
distal part with sensilla capitata with diameter 28 μm. Head width/palpal length 0.91. Antennal length/palpal length 1.0.
</paragraph>
<paragraph id="FA0136F59453FFECF599678BFC4B6EAC" blockId="19.[151,1437,151,609]" pageId="19" pageNumber="502">Thorax. Antepronotum with 11–13 ventrolateral setae, 36–48 μm long. Dorsocentrals 11–18, 92–120 μm long (2 rows in front); prealars 3–7, 92–104 μm long. Scutellum with 10–12 setae.</paragraph>
<paragraph id="FA0136F59453FFECF59967C3FACC6D64" blockId="19.[151,1437,151,609]" pageId="19" pageNumber="502">
Wing (
<figureCitation id="62852A709453FFECF44A67C3FED66E88" box="[276,360,475,501]" captionStart="FIGURES 37–44" captionStartId="14.[151,264,1880,1904]" captionTargetBox="[151,1435,349,1857]" captionTargetId="figure-94@14.[151,1435,345,1857]" captionTargetPageId="14" captionText="FIGURES 37–44. Males of Diamesa zagrosica sp. nov. (37, 41), D. sakartvella Kownacki et Kownacka (38, 42) and D. moubayedi sp. nov. (39–40, 43–44). 37, head; 38, 41, hypopygium in dorsal view; 40, hypopygium in dorsal view without tergite IX; 42, wing; 43, tergite IX; 44, anal point in lateral view." figureDoi="http://doi.org/10.5281/zenodo.6457273" httpUri="https://zenodo.org/record/6457273/files/figure.png" pageId="19" pageNumber="502">Fig. 42</figureCitation>
). Length 
<quantity id="3D469B109453FFECF48C67C3FDE56E88" box="[466,603,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="2.5999999999999996" metricValueMax="2.8" metricValueMin="2.4" pageId="19" pageNumber="502" unit="mm" value="2.5999999999999996" valueMax="2.8" valueMin="2.4">2.4–2.8 mm</quantity>
, width 
<quantity id="3D469B109453FFECF7F367C3FCED6E88" box="[685,851,475,501]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" metricValueMax="8.4" metricValueMin="7.199999999999999" pageId="19" pageNumber="502" unit="mm" value="0.78" valueMax="0.84" valueMin="0.72">0.72–0.84 mm</quantity>
. Anal lobe reduced. Squama with 9–14 setae, 52–60 μm long. R and R 
<subScript id="663A34B09453FFECF4036409FEDB6D62" attach="left" box="[349,357,529,543]" fontSize="6" pageId="19" pageNumber="502">1</subScript>
with 22–25 setae, R 
<subScript id="663A34B09453FFECF7146409FDDA6D62" attach="left" box="[586,612,529,543]" fontSize="6" pageId="19" pageNumber="502">4+5</subScript>
with 5 (in distal part). Costal extension 48–68 μm. RM/MCu 1.9–2.0.
</paragraph>
<paragraph id="FA0136F59453FFECF599643BFA226D1C" blockId="19.[151,1437,151,609]" pageId="19" pageNumber="502">
Legs. Spur of front tibia 28–32 µm long. Spurs of mid tibia equal length (32 µm). Spurs of hind tibia 68 µm and 36–40 µm long. Hind tibial comb with 24–26 setae. Length (μm) and proportions of leg segments are as in 
<tableCitation id="B73C034E9453FFECF01D645FFA266D1C" box="[1347,1432,583,609]" captionStart="TABLE 9" captionStartId="19.[151,239,656,680]" captionTargetPageId="19" captionText="TABLE 9. Lengths (in µm) and proportions of leg segments of Diamesa sakartvella Kownacki et Kownacka, male (n=5)" httpUri="http://table.plazi.org/id/AEC1667D9453FFECF5C96488FF6D6DB1" pageId="19" pageNumber="502" tableUuid="AEC1667D9453FFECF5C96488FF6D6DB1">Table 9</tableCitation>
.
</paragraph>
<caption id="AEC1667D9453FFECF5C96488FF6D6DB1" ID-Table-UUID="AEC1667D9453FFECF5C96488FF6D6DB1" httpUri="http://table.plazi.org/id/AEC1667D9453FFECF5C96488FF6D6DB1" pageId="19" pageNumber="502" startId="19.[151,239,656,680]" targetBox="[151,1366,733,1117]" targetIsTable="true" targetPageId="19">
<paragraph id="FA0136F59453FFECF5C96488FF6D6DB1" blockId="19.[151,1436,656,716]" pageId="19" pageNumber="502">
<emphasis id="C8CAEAE79453FFECF5C96488FEB06DD5" bold="true" box="[151,270,656,680]" pageId="19" pageNumber="502">TABLE 9.</emphasis>
Lengths (in µm) and proportions of leg segments of 
<taxonomicName id="3DBE4D769453FFECF6376489FAE66DD5" authority="Kownacki et Kownacka" authorityName="Kownacki et Kownacka (Moubayed &amp; Laville" authorityYear="1983" box="[873,1368,656,680]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="502" phylum="Arthropoda" rank="species" species="sakartvella">
<emphasis id="C8CAEAE79453FFECF6376489FAE66DD5" box="[873,1368,656,680]" italics="true" pageId="19" pageNumber="502">Diamesa sakartvella Kownacki et Kownacka</emphasis>
</taxonomicName>
, male (n=5)
</paragraph>
</caption>
<paragraph id="FA0136F59453FFECF5B164C5FB456B25" pageId="19" pageNumber="502">
<table id="88BEC45594530000F5C964C5FAE86B20" box="[151,1366,733,1117]" colsContinueIn="20.[151,1367,1340,1725]" gridcols="9" gridrows="9" pageId="19" pageNumber="502">
<tr id="448E34B794530000F5C964C5FAE86D86" box="[151,1366,733,763]" gridrow="0" pageId="19" pageNumber="502" rowspan-0="1" rowspan-7="1">
<th id="075F5DCB94530000F5B164C5FEDC6D86" box="[239,354,733,763]" gridcol="1" gridrow="0" pageId="19" pageNumber="502">fe</th>
<th id="075F5DCB94530000F4C264C5FDB06D86" box="[412,526,733,763]" gridcol="2" gridrow="0" pageId="19" pageNumber="502">ti</th>
<th id="075F5DCB94530000F71764C5FD056D86" box="[585,699,733,763]" gridcol="3" gridrow="0" pageId="19" pageNumber="502">ta1</th>
<th id="075F5DCB94530000F7A864C5FCF16D86" box="[758,847,733,763]" gridcol="4" gridrow="0" pageId="19" pageNumber="502">ta2</th>
<th id="075F5DCB94530000F6DC64C5FC426D86" box="[898,1020,733,763]" gridcol="5" gridrow="0" pageId="19" pageNumber="502">ta3</th>
<th id="075F5DCB94530000F10E64C5FBD46D86" box="[1104,1130,733,763]" gridcol="6" gridrow="0" pageId="19" pageNumber="502">ta4</th>
<th id="075F5DCB94530000F1A064C5FAE86D86" box="[1278,1366,733,763]" gridcol="8" gridrow="0" pageId="19" pageNumber="502">ta 5</th>
</tr>
<tr id="448E34B794530000F5C9651DFAE86C5E" box="[151,1366,773,803]" gridrow="1" pageId="19" pageNumber="502">
<th id="075F5DCB94530000F5C9651DFF056C5E" box="[151,187,773,803]" gridcol="0" gridrow="1" pageId="19" pageNumber="502">P 1</th>
<td id="075F5DCB94530000F5B1651DFEDC6C5E" box="[239,354,773,803]" gridcol="1" gridrow="1" pageId="19" pageNumber="502">1607–1960</td>
<td id="075F5DCB94530000F4C2651DFDB06C5E" box="[412,526,773,803]" gridcol="2" gridrow="1" pageId="19" pageNumber="502">1394–1880</td>
<td id="075F5DCB94530000F717651DFD056C5E" box="[585,699,773,803]" gridcol="3" gridrow="1" pageId="19" pageNumber="502">943–1160</td>
<td id="075F5DCB94530000F7A8651DFCF16C5E" box="[758,847,773,803]" gridcol="4" gridrow="1" pageId="19" pageNumber="502">459–600</td>
<td id="075F5DCB94530000F6DC651DFC426C5E" box="[898,1020,773,803]" gridcol="5" gridrow="1" pageId="19" pageNumber="502">295–380</td>
<td id="075F5DCB94530000F10E651DFB6F6C5E" box="[1104,1233,773,803]" colspan="2" colspanRight="1" gridcol="6" gridrow="1" pageId="19" pageNumber="502">100–115</td>
<td id="075F5DCB94530000F1A0651DFAE86C5E" box="[1278,1366,773,803]" gridcol="8" gridrow="1" pageId="19" pageNumber="502">131–140</td>
</tr>
<tr id="448E34B794530000F5C96535FAE86C36" box="[151,1366,813,843]" gridrow="2" pageId="19" pageNumber="502">
<th id="075F5DCB94530000F5C96535FF056C36" box="[151,187,813,843]" gridcol="0" gridrow="2" pageId="19" pageNumber="502">P2</th>
<td id="075F5DCB94530000F5B16535FEDC6C36" box="[239,354,813,843]" gridcol="1" gridrow="2" pageId="19" pageNumber="502">1525–1680</td>
<td id="075F5DCB94530000F4C26535FDB06C36" box="[412,526,813,843]" gridcol="2" gridrow="2" pageId="19" pageNumber="502">1214–1600</td>
<td id="075F5DCB94530000F7176535FD056C36" box="[585,699,813,843]" gridcol="3" gridrow="2" pageId="19" pageNumber="502">590–720</td>
<td id="075F5DCB94530000F7A86535FCF16C36" box="[758,847,813,843]" gridcol="4" gridrow="2" pageId="19" pageNumber="502">328–380</td>
<td id="075F5DCB94530000F6DC6535FC426C36" box="[898,1020,813,843]" gridcol="5" gridrow="2" pageId="19" pageNumber="502">189–260</td>
<td id="075F5DCB94530000F10E6535FB6F6C36" box="[1104,1233,813,843]" colspan="2" colspanRight="1" gridcol="6" gridrow="2" pageId="19" pageNumber="502">98–120</td>
<td id="075F5DCB94530000F1A06535FAE86C36" box="[1278,1366,813,843]" gridcol="8" gridrow="2" pageId="19" pageNumber="502">123–160</td>
</tr>
<tr id="448E34B794530000F5C9654DFAE86C0E" box="[151,1366,853,883]" gridrow="3" pageId="19" pageNumber="502">
<th id="075F5DCB94530000F5C9654DFF056C0E" box="[151,187,853,883]" gridcol="0" gridrow="3" pageId="19" pageNumber="502">P3</th>
<td id="075F5DCB94530000F5B1654DFEDC6C0E" box="[239,354,853,883]" gridcol="1" gridrow="3" pageId="19" pageNumber="502">1607–2040</td>
<td id="075F5DCB94530000F4C2654DFDB06C0E" box="[412,526,853,883]" gridcol="2" gridrow="3" pageId="19" pageNumber="502">1476–1840</td>
<td id="075F5DCB94530000F717654DFD056C0E" box="[585,699,853,883]" gridcol="3" gridrow="3" pageId="19" pageNumber="502">968–1280</td>
<td id="075F5DCB94530000F7A8654DFCF16C0E" box="[758,847,853,883]" gridcol="4" gridrow="3" pageId="19" pageNumber="502">590–640</td>
<td id="075F5DCB94530000F6DC654DFC426C0E" box="[898,1020,853,883]" gridcol="5" gridrow="3" pageId="19" pageNumber="502">312–360</td>
<td id="075F5DCB94530000F10E654DFB6F6C0E" box="[1104,1233,853,883]" colspan="2" colspanRight="1" gridcol="6" gridrow="3" pageId="19" pageNumber="502">98–100</td>
<td id="075F5DCB94530000F1A0654DFAE86C0E" box="[1278,1366,853,883]" gridcol="8" gridrow="3" pageId="19" pageNumber="502">131–140</td>
</tr>
<tr id="448E34B794530000F5C96587FAE86CCA" box="[151,1366,927,951]" gridrow="4" pageId="19" pageNumber="502">
<th id="075F5DCB94530000F5C96587FAE86CCA" box="[151,1366,927,951]" colspan="9" colspanRight="8" gridcol="0" gridrow="4" pageId="19" pageNumber="502">continued</th>
</tr>
<tr id="448E34B794530000F5C965DFFAE86CA2" box="[151,1366,967,991]" gridrow="5" pageId="19" pageNumber="502" rowspan-0="1" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-8="1">
<td id="075F5DCB94530000F5B165DFFEDC6CA2" box="[239,354,967,991]" gridcol="1" gridrow="5" pageId="19" pageNumber="502">LR</td>
<td id="075F5DCB94530000F71765DFFD056CA2" box="[585,699,967,991]" gridcol="3" gridrow="5" pageId="19" pageNumber="502">BV</td>
<td id="075F5DCB94530000F6DC65DFFC426CA2" box="[898,1020,967,991]" gridcol="5" gridrow="5" pageId="19" pageNumber="502">SV</td>
<td id="075F5DCB94530000F1F165DFFB6F6CA2" box="[1199,1233,967,991]" gridcol="7" gridrow="5" pageId="19" pageNumber="502">BR</td>
</tr>
<tr id="448E34B794530000F5C965F7FAE86B70" box="[151,1366,1007,1037]" gridrow="6" pageId="19" pageNumber="502" rowspan-4="1" rowspan-6="1">
<th id="075F5DCB94530000F5C965F7FF056B70" box="[151,187,1007,1037]" gridcol="0" gridrow="6" pageId="19" pageNumber="502">P1</th>
<td id="075F5DCB94530000F5B165F7FDB06B70" box="[239,526,1007,1037]" colspan="2" colspanRight="1" gridcol="1" gridrow="6" pageId="19" pageNumber="502">0.62–0.68</td>
<td id="075F5DCB94530000F71765F7FD056B70" box="[585,699,1007,1037]" gridcol="3" gridrow="6" pageId="19" pageNumber="502">4.10–4.35</td>
<td id="075F5DCB94530000F6DC65F7FC426B70" box="[898,1020,1007,1037]" gridcol="5" gridrow="6" pageId="19" pageNumber="502">3.15–3.31</td>
<td id="075F5DCB94530000F1F165F7FAE86B70" box="[1199,1366,1007,1037]" colspan="2" colspanRight="1" gridcol="7" gridrow="6" pageId="19" pageNumber="502">1.5–1.8</td>
</tr>
<tr id="448E34B794530000F5C9620FFAE86B48" box="[151,1366,1047,1077]" gridrow="7" pageId="19" pageNumber="502" rowspan-4="1" rowspan-6="1">
<th id="075F5DCB94530000F5C9620FFF056B48" box="[151,187,1047,1077]" gridcol="0" gridrow="7" pageId="19" pageNumber="502">P2</th>
<td id="075F5DCB94530000F5B1620FFDB06B48" box="[239,526,1047,1077]" colspan="2" colspanRight="1" gridcol="1" gridrow="7" pageId="19" pageNumber="502">0.45–0.49</td>
<td id="075F5DCB94530000F717620FFD056B48" box="[585,699,1047,1077]" gridcol="3" gridrow="7" pageId="19" pageNumber="502">4.35–4.98</td>
<td id="075F5DCB94530000F6DC620FFC426B48" box="[898,1020,1047,1077]" gridcol="5" gridrow="7" pageId="19" pageNumber="502">4.56–4.64</td>
<td id="075F5DCB94530000F1F1620FFAE86B48" box="[1199,1366,1047,1077]" colspan="2" colspanRight="1" gridcol="7" gridrow="7" pageId="19" pageNumber="502">1.4–1.5</td>
</tr>
<tr id="448E34B794530000F5C96258FAE86B20" box="[151,1366,1088,1117]" gridrow="8" pageId="19" pageNumber="502" rowspan-4="1" rowspan-6="1">
<th id="075F5DCB94530000F5C96258FF056B20" box="[151,187,1088,1117]" gridcol="0" gridrow="8" pageId="19" pageNumber="502">P3</th>
<td id="075F5DCB94530000F5B16258FDB06B20" box="[239,526,1088,1117]" colspan="2" colspanRight="1" gridcol="1" gridrow="8" pageId="19" pageNumber="502">0.66–0.76</td>
<td id="075F5DCB94530000F7176258FD056B20" box="[585,699,1088,1117]" gridcol="3" gridrow="8" pageId="19" pageNumber="502">3.97–4.16</td>
<td id="075F5DCB94530000F6DC6258FC426B20" box="[898,1020,1088,1117]" gridcol="5" gridrow="8" pageId="19" pageNumber="502">2.75–3.18</td>
<td id="075F5DCB94530000F1F16258FAE86B20" box="[1199,1366,1088,1117]" colspan="2" colspanRight="1" gridcol="7" gridrow="8" pageId="19" pageNumber="502">1.5–1.8</td>
</tr>
</table>
</paragraph>
<paragraph id="FA0136F59453FFECF599629FFAD36AE0" blockId="19.[151,1437,1159,2014]" pageId="19" pageNumber="502">
Hypopygium (
<figureCitation id="62852A709453FFECF437629FFE7B6BDC" box="[361,453,1159,1185]" captionStart="FIGURES 37–44" captionStartId="14.[151,264,1880,1904]" captionTargetBox="[151,1435,349,1857]" captionTargetId="figure-94@14.[151,1435,345,1857]" captionTargetPageId="14" captionText="FIGURES 37–44. Males of Diamesa zagrosica sp. nov. (37, 41), D. sakartvella Kownacki et Kownacka (38, 42) and D. moubayedi sp. nov. (39–40, 43–44). 37, head; 38, 41, hypopygium in dorsal view; 40, hypopygium in dorsal view without tergite IX; 42, wing; 43, tergite IX; 44, anal point in lateral view." figureDoi="http://doi.org/10.5281/zenodo.6457273" httpUri="https://zenodo.org/record/6457273/files/figure.png" pageId="19" pageNumber="502">Figs. 38</figureCitation>
, 
<figureCitation id="62852A709453FFECF48E629FFDD26BDF" box="[464,620,1159,1186]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="19" pageNumber="502">45, 50–51, 57</figureCitation>
). Tergite IX densely covered with strong macrotrichia apices of which are directed anteriorly, with 17–19 setae, 20–96 µm long and with narrow (14–16 µm), chitinized and naked anal point, 120–156 µm long. Laterosternite IX with 12–14 setae, 20–24 µm long. Transverse sternopodeme (TSA) wide triangular, 68–92 µm high, 172–192 µm wide at the base (
<figureCitation id="62852A709453FFECF7BC62EBFCD56A73" box="[738,875,1267,1294]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="19" pageNumber="502">Figs. 50–51</figureCitation>
); TSA height/TSA width 0.36–0.50. Aedeagal lobe 140 µm long (n=1); phallapodeme sclerotized, 120 µm long (n=1). Gonocoxite 372–379 µm long; superior volsellae rounded, along margin with 4–7 setae, 20–40 µm long (
<figureCitation id="62852A709453FFECF7A56323FCF06A2B" box="[763,846,1339,1366]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="19" pageNumber="502">Fig. 57</figureCitation>
). Gonostylus 160–184 µm long, almost not expanded in the distal half along the outer edge (
<figureCitation id="62852A709453FFECF7156347FD216A07" box="[587,671,1375,1402]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="19" pageNumber="502">Fig. 45</figureCitation>
); in apical part with megaseta in form of wide terminal spine, 
<emphasis id="C8CAEAE79453FFECF0006378FAC66A04" box="[1374,1400,1376,1401]" italics="true" pageId="19" pageNumber="502">ca</emphasis>
12 μm long and tooth the same size, next to it there is two setae approximately of the same length. HR 2.02–2.10.
</paragraph>
<paragraph id="FA0136F59453FFECF59963B0FE526ABC" blockId="19.[151,1437,1159,2014]" box="[199,492,1447,1474]" pageId="19" pageNumber="502">
<emphasis id="C8CAEAE79453FFECF59963B0FEBA6ABF" bold="true" box="[199,260,1448,1474]" italics="true" pageId="19" pageNumber="502">Pupa</emphasis>
and 
<emphasis id="C8CAEAE79453FFECF46463BFFECB6ABF" bold="true" box="[314,373,1447,1474]" italics="true" pageId="19" pageNumber="502">larva</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="B2A4657E9453FFECF59963D3FCCC680F" pageId="19" pageNumber="502" type="diagnosis">
<paragraph id="FA0136F59453FFECF59963D3FCCC680F" blockId="19.[151,1437,1159,2014]" pageId="19" pageNumber="502">
<emphasis id="C8CAEAE79453FFECF59963D3FEFC6A98" bold="true" box="[199,322,1483,1509]" pageId="19" pageNumber="502">Diagnosis.</emphasis>
The male of 
<taxonomicName id="3DBE4D769453FFECF48763D4FDC76A98" authorityName="Kownacki et Kownacka (Moubayed &amp; Laville" authorityYear="1983" box="[473,633,1483,1509]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="502" phylum="Arthropoda" rank="species" species="sakartvella">
<emphasis id="C8CAEAE79453FFECF48763D4FDC76A98" box="[473,633,1483,1509]" italics="true" pageId="19" pageNumber="502">D. sakartvella</emphasis>
</taxonomicName>
is closely related in the subgroup to 
<taxonomicName id="3DBE4D769453FFECF14263D4FB156A98" authority="Makarchenko &amp; Semenchenko &amp; Palatov, 2022" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov" authorityYear="2022" box="[1052,1195,1484,1509]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="502" phylum="Arthropoda" rank="species" species="zagrosica" status="sp. nov.">
<emphasis id="C8CAEAE79453FFECF14263D4FB156A98" box="[1052,1195,1484,1509]" italics="true" pageId="19" pageNumber="502">D. zagrosica</emphasis>
</taxonomicName>
<emphasis id="C8CAEAE79453FFECF1ED63D3FAB66A98" bold="true" box="[1203,1288,1483,1509]" pageId="19" pageNumber="502">
<taxonomicNameLabel id="D3F9579C9453FFECF1ED63D3FAB66A98" box="[1203,1288,1483,1509]" pageId="19" pageNumber="502" rank="species">sp. nov</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="3DBE4D769453FFECF01E63D4FF5C6974" authority="Makarchenko &amp; Semenchenko &amp; Palatov, 2022" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov" authorityYear="2022" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="502" phylum="Arthropoda" rank="species" species="moubayedi" status="sp. nov.">
<emphasis id="C8CAEAE79453FFECF01E63D4FF5C6974" italics="true" pageId="19" pageNumber="502">D. moubayedi</emphasis>
</taxonomicName>
<emphasis id="C8CAEAE79453FFECF5B263F7FEF66974" bold="true" box="[236,328,1519,1545]" pageId="19" pageNumber="502">
<taxonomicNameLabel id="D3F9579C9453FFECF5B263F7FEF66974" box="[236,328,1519,1545]" pageId="19" pageNumber="502" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from which it can be distinguished by shape and structure of transverse sternopodeme, inferior volsellae and gonostylus. Transverse sternopodeme of 
<taxonomicName id="3DBE4D769453FFECF7A9600CFC2B6950" authorityName="Kownacki et Kownacka (Moubayed &amp; Laville" authorityYear="1983" box="[759,917,1555,1581]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="502" phylum="Arthropoda" rank="species" species="sakartvella">
<emphasis id="C8CAEAE79453FFECF7A9600CFC2B6950" box="[759,917,1555,1581]" italics="true" pageId="19" pageNumber="502">D. sakartvella</emphasis>
</taxonomicName>
is wide triangular (
<figureCitation id="62852A709453FFECF133600BFB486953" box="[1133,1270,1555,1582]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="19" pageNumber="502">Figs. 50–51</figureCitation>
), of low height (68–88 µm) and very wide basally (172–188 µm), the ratio of its height to width is 0.36–0.50; superior volsellae rounded (
<figureCitation id="62852A709453FFECF45A6043FEE4690B" box="[260,346,1627,1654]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="19" pageNumber="502">Fig. 57</figureCitation>
); gonostylus almost not expanded in the distal half along the outer edge and without projection basally (
<figureCitation id="62852A709453FFECF5A76067FEF869E7" box="[249,326,1663,1690]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="19" pageNumber="502">Fig.45</figureCitation>
). Transverse sternopodeme of 
<taxonomicName id="3DBE4D769453FFECF7F86098FC8969E4" authority="Makarchenko &amp; Semenchenko &amp; Palatov, 2022" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov" authorityYear="2022" box="[678,823,1664,1689]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="502" phylum="Arthropoda" rank="species" species="zagrosica" status="sp. nov.">
<emphasis id="C8CAEAE79453FFECF7F86098FC8969E4" box="[678,823,1664,1689]" italics="true" pageId="19" pageNumber="502">D. zagrosica</emphasis>
</taxonomicName>
<emphasis id="C8CAEAE79453FFECF61E6067FC2269E4" bold="true" box="[832,924,1663,1689]" pageId="19" pageNumber="502">
<taxonomicNameLabel id="D3F9579C9453FFECF61E6067FC2269E4" box="[832,924,1663,1689]" pageId="19" pageNumber="502" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is regular triangular shape (
<figureCitation id="62852A709453FFECF1BD6067FAD169E7" box="[1251,1391,1663,1690]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="19" pageNumber="502">Figs. 53–54</figureCitation>
), of medium height (84–108 µm) and not very wide basally (162–172 µm), the ratio of its height to width is 0.49–0.67; superior volsellae rounded–angular (
<figureCitation id="62852A709453FFECF76860DFFD35699F" box="[566,651,1735,1762]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="19" pageNumber="502">Fig. 56</figureCitation>
); gonostylus slightly expanded in the distal half along the outer edge and with small rounded projection basally (
<figureCitation id="62852A709453FFECF7CF60F3FCAA687B" box="[657,788,1771,1798]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="19" pageNumber="502">Figs.48–49</figureCitation>
). Transverse sternopodeme of 
<taxonomicName id="3DBE4D769453FFECF12960F4FAA66878" authority="Makarchenko &amp; Semenchenko &amp; Palatov, 2022" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov" authorityYear="2022" box="[1143,1304,1771,1797]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="502" phylum="Arthropoda" rank="species" species="moubayedi" status="sp. nov.">
<emphasis id="C8CAEAE79453FFECF12960F4FAA66878" box="[1143,1304,1771,1797]" italics="true" pageId="19" pageNumber="502">D. moubayedi</emphasis>
</taxonomicName>
<emphasis id="C8CAEAE79453FFECF07C60F3FAC16878" bold="true" box="[1314,1407,1771,1797]" pageId="19" pageNumber="502">
<taxonomicNameLabel id="D3F9579C9453FFECF07C60F3FAC16878" box="[1314,1407,1771,1797]" pageId="19" pageNumber="502" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is triangular shape (
<figureCitation id="62852A709453FFECF4026117FE0F6857" box="[348,433,1807,1834]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="19" pageNumber="502">Fig. 52</figureCitation>
), of great height (104–140 µm) and not very wide basally (148–184 µm), the ratio of its height to width is 0.80–0.89; superior volsellae angular (
<figureCitation id="62852A709453FFECF649612BFCD56833" box="[791,875,1843,1870]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="19" pageNumber="502">Fig. 55</figureCitation>
); gonostylus expanded in the distal half along the outer edge and with rounded–angular projection basally (
<figureCitation id="62852A709453FFECF646614FFCDB680F" box="[792,869,1879,1906]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="19" pageNumber="502">Fig.47</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B2A4657E9453FFECF5996163FB9968C4" pageId="19" pageNumber="502" type="biology_ecology">
<paragraph id="FA0136F59453FFECF5996163FB9968C4" blockId="19.[151,1437,1159,2014]" pageId="19" pageNumber="502">
<emphasis id="C8CAEAE79453FFECF5996163FE9268E8" bold="true" box="[199,300,1915,1941]" pageId="19" pageNumber="502">Ecology.</emphasis>
Adult males were collected from stones and boulders at the source of a mountain river, located at an altitude of 
<quantity id="3D469B109453FFECF44F6187FED668C7" box="[273,360,1951,1978]" metricMagnitude="3" metricUnit="m" metricValue="1.943" pageId="19" pageNumber="502" unit="m" value="1943.0">1943 m</quantity>
, at a flow rate of 
<quantity id="3D469B109453FFECF7726187FD2168C4" box="[556,671,1951,1977]" metricMagnitude="-1" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" pageId="19" pageNumber="502" unit="m" value="0.55" valueMax="0.8" valueMin="0.3">0.3–0.8 m</quantity>
/s, with water temperatures 
<emphasis id="C8CAEAE79453FFECF68D61B8FC5368C4" box="[979,1005,1952,1977]" italics="true" pageId="19" pageNumber="502">ca</emphasis>
7°C.
</paragraph>
</subSubSection>
<subSubSection id="B2A4657E9453FFECF59961DBFCAF68A0" box="[199,785,1987,2014]" pageId="19" pageNumber="502" type="distribution">
<paragraph id="FA0136F59453FFECF59961DBFCAF68A0" blockId="19.[151,1437,1159,2014]" box="[199,785,1987,2014]" pageId="19" pageNumber="502">
<emphasis id="C8CAEAE79453FFECF59961DBFEDF68A0" bold="true" box="[199,353,1987,2013]" pageId="19" pageNumber="502">Distribution.</emphasis>
Known only from Caucasus (
<figureCitation id="62852A709453FFECF7EE61DBFCBB68A0" box="[688,773,1987,2013]" captionStart="FIGURES 68–74" captionStartId="23.[151,264,1617,1641]" captionTargetBox="[151,1435,181,1592]" captionTargetId="figure-18@23.[151,1435,181,1593]" captionTargetPageId="23" captionText="FIGURES 68–74. Type localities of Diamesa dragani sp. nov. (68), D. kownackii sp. nov. (69–70), D. maisaraensis sp. nov. (71), D. marinskiyi sp. nov. (72), D. steinboecki Goetghebuer (73) and D. sakartvella Kownacki et Kownacka (74). 68, unnamed stream of Bolshoi On River basin, alt. 2063 m a. sea l., Sayan Mountains (photo by S.V. Dragan); 69, unnamed stream of Somnitelnaya River basin, Wrangel Island (photo by E.A. Makarchenko); 70, Somnitelnaya River, upper stream, alt. 205–210 m a. sea l., Wrangel Island (photo by O.A. Khruleva); 71, Maisara River near Maisara Pass, alt. 4168 m a. sea l., Pamir Mountains (photo by D.M. Palatov); 72, Zienzan River, Tien Shan Mountains, alt. 1923 m a. sea l., Xinjiang Uygur Autonomous Region, China (photo by D.M. Palatov); 73, Balm stream, Massif of Aiguilles–Rouges, French Alps (photo by Pierre Clévenot); 74, Skazdon River near the Tsey (Tseyskoe gorge), alt. 1943 m a. sea l., North Caucasus (photo by D.M. Palatov)." figureDoi="http://doi.org/10.5281/zenodo.6457281" httpUri="https://zenodo.org/record/6457281/files/figure.png" pageId="19" pageNumber="502">Fig. 74</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>