167 lines
28 KiB
XML
167 lines
28 KiB
XML
<document id="448A0622CA726D8B6C797D8840D11775" ID-DOI="10.1111/j.1096-3642.2009.00578.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="10114686" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630443207186" checkinUser="valdenar" docAuthor="Carr, Robert K. & Hlavin, William J." docDate="2010" docId="5415C76BFF9CFFA89BE8D55AFD2351D8" docLanguage="en" docName="j.1096-3642.2009.00578.x.pdf" docOrigin="Zoological Journal of the Linnean Society 159 (1)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2009.00578.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Dunkleosteus amblyodoratus Carr & Hlavin 2010, SP. NOV." docType="treatment" docVersion="6" lastPageNumber="205" masterDocId="A82CBF13FF95FFA29B3ED355FFD55403" masterDocTitle="Two new species of Dunkleosteus Lehman, 1956, from the Ohio Shale Formation (USA, Famennian) and the Kettle Point Formation (Canada, Upper Devonian), and a cladistic analysis of the Eubrachythoraci (Placodermi, Arthrodira)" masterLastPageNumber="222" masterPageNumber="195" pageNumber="204" updateTime="1700039058763" updateUser="ExternalLinkService">
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<mods:title id="DEBE74A22C9A8E1CAA6F1D89F4E930D0">Two new species of Dunkleosteus Lehman, 1956, from the Ohio Shale Formation (USA, Famennian) and the Kettle Point Formation (Canada, Upper Devonian), and a cladistic analysis of the Eubrachythoraci (Placodermi, Arthrodira)</mods:title>
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<mods:namePart id="923003C2A9491267978D21B45E0262F1">Carr, Robert K.</mods:namePart>
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<mods:namePart id="57EC606C70D57CE2E12168088D2FF63F">Hlavin, William J.</mods:namePart>
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<mods:typeOfResource id="783A4B23C50B72FD5604B5449D93677F">text</mods:typeOfResource>
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<mods:title id="CFE78DE8E75FA5DFCE110667E544F7B7">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="D6CEA84CF9FF057B123435815DD64EC7">
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<mods:date id="E50D41ECE298ECCDC70121EDC4DBE607">2010</mods:date>
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<mods:detail id="0EC9A4933516CE3C80CA7B918E51E83C" type="pubDate">
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<mods:number id="F2A8E2170764C67B6925FCD79AAD2F43">2010-05-31</mods:number>
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<mods:detail id="5B6ED974963E5C2C09B1802A389D89BB" type="volume">
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<mods:number id="433E685E6431E7246433D5B289CCF617">159</mods:number>
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<mods:identifier id="E63094AE53CD08500EBEDDF592DF2540" type="DOI">10.1111/j.1096-3642.2009.00578.x</mods:identifier>
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<mods:identifier id="203C82FEF05472CD14E336F2B85E64B6" type="ISSN">0024-4082</mods:identifier>
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<mods:identifier id="9719D0D43087E18EA78D2DB2264B2A7B" type="Zenodo-Dep">10114686</mods:identifier>
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<treatment id="5415C76BFF9CFFA89BE8D55AFD2351D8" LSID="urn:lsid:plazi:treatment:5415C76BFF9CFFA89BE8D55AFD2351D8" httpUri="http://treatment.plazi.org/id/5415C76BFF9CFFA89BE8D55AFD2351D8" lastPageId="10" lastPageNumber="205" pageId="9" pageNumber="204">
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<subSubSection id="94A625F6FF9CFFAB9BE8D55AFD0C5227" box="[214,729,1551,1572]" pageId="9" pageNumber="204" type="nomenclature">
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<paragraph id="DC03767DFF9CFFAB9BE8D55AFD0C5227" blockId="9.[214,729,1551,1605]" box="[214,729,1551,1572]" pageId="9" pageNumber="204">
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<heading id="874BC111FF9CFFAB9BE8D55AFD0C5227" box="[214,729,1551,1572]" centered="true" fontSize="9" level="2" pageId="9" pageNumber="204" reason="4">
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<emphasis id="EEC8AA6FFF9CFFAB9BE8D55AFD0C5227" bold="true" box="[214,729,1551,1572]" pageId="9" pageNumber="204">
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<taxonomicName id="1BBC0DFEFF9CFFAB9BE8D55AFDBF5220" authority="Carr & Hlavin, 2010" authorityName="Carr & Hlavin" authorityYear="2010" box="[214,618,1551,1572]" class="Placodermi" family="Dunkleosteidae" genus="Dunkleosteus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrodira" pageId="9" pageNumber="214" phylum="Chordata" rank="species" species="amblyodoratus" status="sp. nov.">DUNKLEOSTEUS AMBLYODORATUS</taxonomicName>
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<taxonomicNameLabel id="F5FB1714FF9CFFAB994AD547FD0C5227" box="[628,729,1551,1572]" pageId="9" pageNumber="204" rank="species">SP. NOV.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<caption id="88C326F5FF9CFFAB9804D540FA455245" ID-DOI="http://doi.org/10.5281/zenodo.10132139" ID-Zenodo-Dep="10132139" httpUri="https://zenodo.org/record/10132139/files/figure.png" pageId="9" pageNumber="204" startId="9.[826,905,1557,1576]" targetBox="[826,1442,1079,1527]" targetPageId="9">
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<paragraph id="DC03767DFF9CFFAB9804D540FA455245" blockId="9.[826,1441,1557,1606]" pageId="9" pageNumber="204">
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<emphasis id="EEC8AA6FFF9CFFAB9804D540FABD522B" bold="true" box="[826,1384,1557,1576]" pageId="9" pageNumber="204">
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Figure 7.
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<taxonomicName id="1BBC0DFEFF9CFFAB9891D540FAD6522B" authority="Carr & Hlavin, 2010" authorityName="Carr & Hlavin" authorityYear="2010" box="[943,1283,1557,1576]" class="Placodermi" family="Dunkleosteidae" genus="Dunkleosteus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrodira" pageId="9" pageNumber="204" phylum="Chordata" rank="species" species="amblyodoratus" status="sp. nov.">
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<emphasis id="EEC8AA6FFF9CFFAB9891D540FAD6522B" bold="true" box="[943,1283,1557,1576]" italics="true" pageId="9" pageNumber="204">Dunkleosteus amblyodoratus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F5FB1714FF9CFFAB9E30D540FABD522B" box="[1294,1384,1557,1576]" pageId="9" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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(UM 101105). Parasphenoid in ventral view. Scale bar: 1 cm.
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</paragraph>
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</caption>
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<subSubSection id="94A625F6FF9CFFA89AA4D57AFD2351D8" lastPageId="10" lastPageNumber="205" pageId="9" pageNumber="204" type="description">
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<paragraph id="DC03767DFF9CFFAB9AA4D57AFDC15247" blockId="9.[214,729,1551,1605]" box="[410,532,1583,1605]" pageId="9" pageNumber="204">
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(
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<figureCitation id="44876AF8FF9CFFAB9A9CD57AFE3A5246" box="[418,495,1583,1605]" captionStart="Figure 6" captionStartId="9.[164,243,954,973]" captionTargetBox="[375,1231,196,923]" captionTargetId="figure-207@9.[424,1232,196,675]" captionTargetPageId="9" captionText="Figure 6. Dunkleosteus amblyodoratus sp. nov. (UM 101105). A, B, nuchal and paranuchal plates, and reconstruction, in internal view. C, external reconstruction. Dotted lines indicate incomplete edges; dashed lines indicate the reconstruction of the posterolateral corner of the nuchal plate. Scale bars: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.10132137" httpUri="https://zenodo.org/record/10132137/files/figure.png" pageId="9" pageNumber="204">FIGS 6</figureCitation>
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,
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<figureCitation id="44876AF8FF9CFFAB9AC0D57AFDD95247" box="[510,524,1583,1604]" captionStart="Figure 7" captionStartId="9.[826,905,1557,1576]" captionTargetBox="[826,1442,1079,1527]" captionTargetId="figure-328@9.[824,1445,1077,1529]" captionTargetPageId="9" captionText="Figure 7. Dunkleosteus amblyodoratus sp. nov. (UM 101105). Parasphenoid in ventral view. Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.10132139" httpUri="https://zenodo.org/record/10132139/files/figure.png" pageId="9" pageNumber="204">7</figureCitation>
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)
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</paragraph>
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<paragraph id="DC03767DFF9CFFAB9B9AD503FD8B52C4" blockId="9.[164,779,1622,1735]" pageId="9" pageNumber="204">
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<emphasis id="EEC8AA6FFF9CFFAB9B9AD503FECE5268" box="[164,283,1622,1643]" italics="true" pageId="9" pageNumber="204">Diagnosis:</emphasis>
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A
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<taxonomicName id="1BBC0DFEFF9CFFAB9A67D503FD82526F" box="[345,599,1622,1644]" class="Placodermi" family="Dunkleosteidae" genus="Dunkleosteus" kingdom="Animalia" order="Arthrodira" pageId="9" pageNumber="204" phylum="Chordata" rank="species" species="undetermined">
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<emphasis id="EEC8AA6FFF9CFFAB9A67D503FE255268" box="[345,496,1622,1643]" italics="true" pageId="9" pageNumber="204">Dunkleosteus</emphasis>
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species
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</taxonomicName>
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possessing a tapered prehypophysial region of the parasphenoid, with anterolateral contact faces for posterior processes of the anterior superognathals.
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</paragraph>
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<paragraph id="DC03767DFF9CFFAB9B9AD5A8FF30536C" blockId="9.[164,779,1789,1903]" pageId="9" pageNumber="204">
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<materialsCitation id="6CD47C20FF9CFFAB9B9AD5A8FF30536C" accessionNumber="UM 101105" collectionCode="UM" pageId="9" pageNumber="204" specimenCount="1" typeStatus="holotype">
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<emphasis id="EEC8AA6FFF9CFFAB9B9AD5A8FEDB5311" box="[164,270,1789,1810]" italics="true" pageId="9" pageNumber="204">
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<typeStatus id="0307C8DFFF9CFFAB9B9AD5A8FEDD5311" box="[164,264,1789,1810]" pageId="9" pageNumber="204" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<collectionCode id="BAADEEB8FF9CFFAB9A27D5A8FE915311" box="[281,324,1789,1810]" country="Germany" name="University of Marburg" pageId="9" pageNumber="204">UM</collectionCode>
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101105 (
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<figureCitation id="44876AF8FF9CFFAB9A8DD5A8FE2E5310" box="[435,507,1789,1811]" captionStart="Figure 6" captionStartId="9.[164,243,954,973]" captionTargetBox="[375,1231,196,923]" captionTargetId="figure-207@9.[424,1232,196,675]" captionTargetPageId="9" captionText="Figure 6. Dunkleosteus amblyodoratus sp. nov. (UM 101105). A, B, nuchal and paranuchal plates, and reconstruction, in internal view. C, external reconstruction. Dotted lines indicate incomplete edges; dashed lines indicate the reconstruction of the posterolateral corner of the nuchal plate. Scale bars: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.10132137" httpUri="https://zenodo.org/record/10132137/files/figure.png" pageId="9" pageNumber="204">Figs 6</figureCitation>
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,
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<figureCitation id="44876AF8FF9CFFAB9934D5A8FDCC5311" box="[522,537,1789,1810]" captionStart="Figure 7" captionStartId="9.[826,905,1557,1576]" captionTargetBox="[826,1442,1079,1527]" captionTargetId="figure-328@9.[824,1445,1077,1529]" captionTargetPageId="9" captionText="Figure 7. Dunkleosteus amblyodoratus sp. nov. (UM 101105). Parasphenoid in ventral view. Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.10132139" httpUri="https://zenodo.org/record/10132139/files/figure.png" pageId="9" pageNumber="204">7</figureCitation>
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), an isolated nuchal plate, with the transverse thickened portion of the left paranuchal plate and an incomplete parasphenoid plate.
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</materialsCitation>
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</paragraph>
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<paragraph id="DC03767DFF9CFFAB9804D528FAA252D3" blockId="9.[826,1441,1661,1744]" pageId="9" pageNumber="204">
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<emphasis id="EEC8AA6FFF9CFFAB9804D528FC6D5291" box="[826,952,1661,1682]" italics="true" pageId="9" pageNumber="204">Etymology:</emphasis>
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From
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<emphasis id="EEC8AA6FFF9CFFAB9F37D528FB8F5291" box="[1033,1114,1661,1682]" italics="true" pageId="9" pageNumber="204">amblys</emphasis>
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meaning blunt and
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<emphasis id="EEC8AA6FFF9CFFAB9E73D528FA745291" box="[1357,1441,1661,1682]" italics="true" pageId="9" pageNumber="204">doratos</emphasis>
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meaning spear (Greek); referring to the form of the prehypophysial region of the parasphenoid plate.
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</paragraph>
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<paragraph id="DC03767DFF9CFFA89804D5A8FD37551A" blockId="9.[826,1441,1789,1903]" lastBlockId="10.[144,759,198,281]" lastPageId="10" lastPageNumber="205" pageId="9" pageNumber="204">
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<emphasis id="EEC8AA6FFF9CFFAB9804D5A8FB5C5311" box="[826,1161,1789,1810]" italics="true" pageId="9" pageNumber="204">Occurrence and stratigraphy:</emphasis>
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Collected from
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<collectingRegion id="1E78B89FFF9CFFAB9E7CD5A8FA485310" box="[1346,1437,1789,1811]" country="Canada" name="Ontario" pageId="9" pageNumber="204">Ontario</collectingRegion>
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,
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<collectingCountry id="A4AB36EDFF9CFFAB9804D449FC475331" box="[826,914,1820,1842]" name="Canada" pageId="9" pageNumber="204">Canada</collectingCountry>
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(
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<figureCitation id="44876AF8FF9CFFAB9894D449FBD55331" box="[938,1024,1820,1842]" captionStart="Figure 2" captionStartId="5.[164,243,818,837]" captionTargetBox="[164,1449,215,788]" captionTargetId="graphics-373@5.[684,1449,215,787]" captionTargetPageId="5" captionText="Figure 2. Regional geography and stratigraphic relationships. A, geography and depositional basins for the USA mid-continent (modified from Elliott et al., 2000). Fossil localities: 1, Dunkleosteus raveri sp. nov. (northern Ohio); 2, Dunkleosteus amblyodoratus sp. nov. (Kettle Point, Ontario); 3, Dinichthys herzeri (western Ohio). Structural features (thick dashed lines): a, Algonquin Arch; c, Cincinnati Arch; f, Findlay Arch. State boundaries, dot double-dash lines; Canadian boundary, dotted lines. Concentric lines of basinal deposition, thin dashed lines. Great Lakes (grey). B, generalized stratigraphic columns for the Appalachian and Michigan basins. Modified and updated from Elliott et al. (2000). A Famennian age for the entire Ohio Shale Formation is based on Over & Rhodes (2000). The presence of the Chagrin Shale Member in central Ohio is based on subsurface g-ray profiles (Walker & Babcock, 1999). Fossil localities as in A. Hiatuses in grey. O.b. = Olmsted bed in the Cleveland Shale Member." figureDoi="http://doi.org/10.5281/zenodo.10132129" httpUri="https://zenodo.org/record/10132129/files/figure.png" pageId="9" pageNumber="204">Fig. 2A</figureCitation>
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). The exact locality data are not available, but the specimen was recovered from the Kettle Point Formation (Upper Devonian, but the exact stratigraphic level was not recorded;
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<figureCitation id="44876AF8FF9FFFA899ABD393FD3F54DF" box="[661,746,198,220]" captionStart="Figure 2" captionStartId="5.[164,243,818,837]" captionTargetBox="[164,1449,215,788]" captionTargetId="graphics-373@5.[684,1449,215,787]" captionTargetPageId="5" captionText="Figure 2. Regional geography and stratigraphic relationships. A, geography and depositional basins for the USA mid-continent (modified from Elliott et al., 2000). Fossil localities: 1, Dunkleosteus raveri sp. nov. (northern Ohio); 2, Dunkleosteus amblyodoratus sp. nov. (Kettle Point, Ontario); 3, Dinichthys herzeri (western Ohio). Structural features (thick dashed lines): a, Algonquin Arch; c, Cincinnati Arch; f, Findlay Arch. State boundaries, dot double-dash lines; Canadian boundary, dotted lines. Concentric lines of basinal deposition, thin dashed lines. Great Lakes (grey). B, generalized stratigraphic columns for the Appalachian and Michigan basins. Modified and updated from Elliott et al. (2000). A Famennian age for the entire Ohio Shale Formation is based on Over & Rhodes (2000). The presence of the Chagrin Shale Member in central Ohio is based on subsurface g-ray profiles (Walker & Babcock, 1999). Fossil localities as in A. Hiatuses in grey. O.b. = Olmsted bed in the Cleveland Shale Member." figureDoi="http://doi.org/10.5281/zenodo.10132129" httpUri="https://zenodo.org/record/10132129/files/figure.png" pageId="10" pageNumber="205">Fig. 2B</figureCitation>
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). The specimen was found within a carbonate concretion that is typical for the Kettle Point Formation.
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</paragraph>
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<paragraph id="DC03767DFF9FFFA89A38D207FD555564" blockId="10.[262,640,338,359]" box="[262,640,338,359]" pageId="10" pageNumber="205">SKULL ROOF AND PARASPHENOID</paragraph>
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<paragraph id="DC03767DFF9FFFA89BAED22DFE0D5666" blockId="10.[144,759,376,1500]" pageId="10" pageNumber="205">
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<taxonomicName id="1BBC0DFEFF9FFFA89BAED22DFE08558E" authority="Carr & Hlavin, 2010" authorityName="Carr & Hlavin" authorityYear="2010" box="[144,477,376,397]" class="Placodermi" family="Dunkleosteidae" genus="Dunkleosteus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrodira" pageId="10" pageNumber="205" phylum="Chordata" rank="species" species="amblyodoratus" status="sp. nov.">
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<emphasis id="EEC8AA6FFF9FFFA89BAED22DFE08558E" box="[144,477,376,397]" italics="true" pageId="10" pageNumber="205">Dunkleosteus amblyodoratus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F5FB1714FF9FFFA89AD7D22CFD96558D" box="[489,579,377,398]" pageId="10" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
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is known only from fragmented parasphenoid, nuchal, and paranuchal plates. The external surface of the skull roof is worn. Tubercles are visible only in one area of the paranuchal plate where an outer layer is missing, revealing tubercles on a deeper surface (representing an earlier generation of tubercles). The punctate tubercles are evenly spaced.
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</paragraph>
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<paragraph id="DC03767DFF9FFFA89B96D138FEA8504D" blockId="10.[144,759,376,1500]" pageId="10" pageNumber="205">
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The nuchal plate (Nu;
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<figureCitation id="44876AF8FF9FFFA89A8ED138FE245680" box="[432,497,621,643]" captionStart="Figure 6" captionStartId="9.[164,243,954,973]" captionTargetBox="[375,1231,196,923]" captionTargetId="figure-207@9.[424,1232,196,675]" captionTargetPageId="9" captionText="Figure 6. Dunkleosteus amblyodoratus sp. nov. (UM 101105). A, B, nuchal and paranuchal plates, and reconstruction, in internal view. C, external reconstruction. Dotted lines indicate incomplete edges; dashed lines indicate the reconstruction of the posterolateral corner of the nuchal plate. Scale bars: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.10132137" httpUri="https://zenodo.org/record/10132137/files/figure.png" pageId="10" pageNumber="205">Fig. 6</figureCitation>
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) outline is trapezoidal with a transverse anterior margin, with reduced indentations forming a shallow W-shape (suggesting an interdigitation with the central plates). The exact nature of this shape is unclear because of some weathering. The internal surface possesses a transverse occipital thickening (th.occ;
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<figureCitation id="44876AF8FF9FFFA8992DD070FDB25738" box="[531,615,805,827]" captionStart="Figure 6" captionStartId="9.[164,243,954,973]" captionTargetBox="[375,1231,196,923]" captionTargetId="figure-207@9.[424,1232,196,675]" captionTargetPageId="9" captionText="Figure 6. Dunkleosteus amblyodoratus sp. nov. (UM 101105). A, B, nuchal and paranuchal plates, and reconstruction, in internal view. C, external reconstruction. Dotted lines indicate incomplete edges; dashed lines indicate the reconstruction of the posterolateral corner of the nuchal plate. Scale bars: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.10132137" httpUri="https://zenodo.org/record/10132137/files/figure.png" pageId="10" pageNumber="205">Fig. 6A</figureCitation>
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) that is continuous with that of the paranuchal plate. Double pits (pt.u;
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<figureCitation id="44876AF8FF9FFFA89BEBD037FEFD577B" box="[213,296,866,888]" captionStart="Figure 6" captionStartId="9.[164,243,954,973]" captionTargetBox="[375,1231,196,923]" captionTargetId="figure-207@9.[424,1232,196,675]" captionTargetPageId="9" captionText="Figure 6. Dunkleosteus amblyodoratus sp. nov. (UM 101105). A, B, nuchal and paranuchal plates, and reconstruction, in internal view. C, external reconstruction. Dotted lines indicate incomplete edges; dashed lines indicate the reconstruction of the posterolateral corner of the nuchal plate. Scale bars: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.10132137" httpUri="https://zenodo.org/record/10132137/files/figure.png" pageId="10" pageNumber="205">Fig. 6A</figureCitation>
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) are present and bounded anteriorly by the nuchal thickening (th.n;
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<figureCitation id="44876AF8FF9FFFA89AE3D0D4FDE65794" box="[477,563,897,919]" captionStart="Figure 6" captionStartId="9.[164,243,954,973]" captionTargetBox="[375,1231,196,923]" captionTargetId="figure-207@9.[424,1232,196,675]" captionTargetPageId="9" captionText="Figure 6. Dunkleosteus amblyodoratus sp. nov. (UM 101105). A, B, nuchal and paranuchal plates, and reconstruction, in internal view. C, external reconstruction. Dotted lines indicate incomplete edges; dashed lines indicate the reconstruction of the posterolateral corner of the nuchal plate. Scale bars: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.10132137" httpUri="https://zenodo.org/record/10132137/files/figure.png" pageId="10" pageNumber="205">Fig. 6A</figureCitation>
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), and posteriorly by a well-developed transverse ridge (p.tr.r;
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<figureCitation id="44876AF8FF9FFFA899ABD0CAFD3E57B6" box="[661,747,927,949]" captionStart="Figure 6" captionStartId="9.[164,243,954,973]" captionTargetBox="[375,1231,196,923]" captionTargetId="figure-207@9.[424,1232,196,675]" captionTargetPageId="9" captionText="Figure 6. Dunkleosteus amblyodoratus sp. nov. (UM 101105). A, B, nuchal and paranuchal plates, and reconstruction, in internal view. C, external reconstruction. Dotted lines indicate incomplete edges; dashed lines indicate the reconstruction of the posterolateral corner of the nuchal plate. Scale bars: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.10132137" httpUri="https://zenodo.org/record/10132137/files/figure.png" pageId="10" pageNumber="205">Fig. 6A</figureCitation>
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). Individual pits are separated by a median septum (m.sept;
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<figureCitation id="44876AF8FF9FFFA89BC7D089FE9857F1" box="[249,333,988,1010]" captionStart="Figure 6" captionStartId="9.[164,243,954,973]" captionTargetBox="[375,1231,196,923]" captionTargetId="figure-207@9.[424,1232,196,675]" captionTargetPageId="9" captionText="Figure 6. Dunkleosteus amblyodoratus sp. nov. (UM 101105). A, B, nuchal and paranuchal plates, and reconstruction, in internal view. C, external reconstruction. Dotted lines indicate incomplete edges; dashed lines indicate the reconstruction of the posterolateral corner of the nuchal plate. Scale bars: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.10132137" httpUri="https://zenodo.org/record/10132137/files/figure.png" pageId="10" pageNumber="205">Fig. 6A</figureCitation>
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) that does not cross the posterior transverse ridge. A posterior process (p.pr;
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<figureCitation id="44876AF8FF9FFFA89942D0AEFD3B5012" box="[636,750,1019,1041]" captionStart="Figure 6" captionStartId="9.[164,243,954,973]" captionTargetBox="[375,1231,196,923]" captionTargetId="figure-207@9.[424,1232,196,675]" captionTargetPageId="9" captionText="Figure 6. Dunkleosteus amblyodoratus sp. nov. (UM 101105). A, B, nuchal and paranuchal plates, and reconstruction, in internal view. C, external reconstruction. Dotted lines indicate incomplete edges; dashed lines indicate the reconstruction of the posterolateral corner of the nuchal plate. Scale bars: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.10132137" httpUri="https://zenodo.org/record/10132137/files/figure.png" pageId="10" pageNumber="205">Fig. 6A, C</figureCitation>
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) is present, and is bounded laterally by shallow paired fossae (f.lv;
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<figureCitation id="44876AF8FF9FFFA89A22D76DFEA5504D" box="[284,368,1080,1102]" captionStart="Figure 6" captionStartId="9.[164,243,954,973]" captionTargetBox="[375,1231,196,923]" captionTargetId="figure-207@9.[424,1232,196,675]" captionTargetPageId="9" captionText="Figure 6. Dunkleosteus amblyodoratus sp. nov. (UM 101105). A, B, nuchal and paranuchal plates, and reconstruction, in internal view. C, external reconstruction. Dotted lines indicate incomplete edges; dashed lines indicate the reconstruction of the posterolateral corner of the nuchal plate. Scale bars: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.10132137" httpUri="https://zenodo.org/record/10132137/files/figure.png" pageId="10" pageNumber="205">Fig. 6B</figureCitation>
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).
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</paragraph>
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<paragraph id="DC03767DFF9FFFA89B96D702FE265127" blockId="10.[144,759,376,1500]" pageId="10" pageNumber="205">
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Only the transverse occipital thickening of the paranuchal plate is preserved (PNu;
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<figureCitation id="44876AF8FF9FFFA89957D720FD795088" box="[617,684,1141,1163]" captionStart="Figure 6" captionStartId="9.[164,243,954,973]" captionTargetBox="[375,1231,196,923]" captionTargetId="figure-207@9.[424,1232,196,675]" captionTargetPageId="9" captionText="Figure 6. Dunkleosteus amblyodoratus sp. nov. (UM 101105). A, B, nuchal and paranuchal plates, and reconstruction, in internal view. C, external reconstruction. Dotted lines indicate incomplete edges; dashed lines indicate the reconstruction of the posterolateral corner of the nuchal plate. Scale bars: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.10132137" httpUri="https://zenodo.org/record/10132137/files/figure.png" pageId="10" pageNumber="205">Fig. 6</figureCitation>
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). The thickening is massive and continues to the lateral articular fossa (laf;
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<figureCitation id="44876AF8FF9FFFA89A4FD7E6FE1350CA" box="[369,454,1203,1225]" captionStart="Figure 6" captionStartId="9.[164,243,954,973]" captionTargetBox="[375,1231,196,923]" captionTargetId="figure-207@9.[424,1232,196,675]" captionTargetPageId="9" captionText="Figure 6. Dunkleosteus amblyodoratus sp. nov. (UM 101105). A, B, nuchal and paranuchal plates, and reconstruction, in internal view. C, external reconstruction. Dotted lines indicate incomplete edges; dashed lines indicate the reconstruction of the posterolateral corner of the nuchal plate. Scale bars: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.10132137" httpUri="https://zenodo.org/record/10132137/files/figure.png" pageId="10" pageNumber="205">Fig. 6C</figureCitation>
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). Only a portion of the left fossa remains, but it suggests the presence of a large and well-developed ginglymoid articulation between the head and thoracic armour.
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</paragraph>
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<paragraph id="DC03767DFF9FFFA89B96D678FD2351D8" blockId="10.[144,759,376,1500]" pageId="10" pageNumber="205">
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The parasphenoid (Psp;
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<figureCitation id="44876AF8FF9FFFA89AF8D678FDD25141" box="[454,519,1325,1347]" captionStart="Figure 7" captionStartId="9.[826,905,1557,1576]" captionTargetBox="[826,1442,1079,1527]" captionTargetId="figure-328@9.[824,1445,1077,1529]" captionTargetPageId="9" captionText="Figure 7. Dunkleosteus amblyodoratus sp. nov. (UM 101105). Parasphenoid in ventral view. Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.10132139" httpUri="https://zenodo.org/record/10132139/files/figure.png" pageId="10" pageNumber="205">Fig. 7</figureCitation>
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) is incomplete, with only its anterior half preserved. The prehypophysial region (pre.reg;
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<figureCitation id="44876AF8FF9FFFA89A76D63FFE5C5183" box="[328,393,1386,1408]" captionStart="Figure 7" captionStartId="9.[826,905,1557,1576]" captionTargetBox="[826,1442,1079,1527]" captionTargetId="figure-328@9.[824,1445,1077,1529]" captionTargetPageId="9" captionText="Figure 7. Dunkleosteus amblyodoratus sp. nov. (UM 101105). Parasphenoid in ventral view. Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.10132139" httpUri="https://zenodo.org/record/10132139/files/figure.png" pageId="10" pageNumber="205">Fig. 7</figureCitation>
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) is tapered, and is as long as it is wide. The lateral edges of the prehypophysial region are thickened, providing possible contact facets for the anterior superognathal plates (cf.ASG;
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<figureCitation id="44876AF8FF9FFFA89999D693FD3C51D8" box="[679,745,1478,1500]" captionStart="Figure 7" captionStartId="9.[826,905,1557,1576]" captionTargetBox="[826,1442,1079,1527]" captionTargetId="figure-328@9.[824,1445,1077,1529]" captionTargetPageId="9" captionText="Figure 7. Dunkleosteus amblyodoratus sp. nov. (UM 101105). Parasphenoid in ventral view. Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.10132139" httpUri="https://zenodo.org/record/10132139/files/figure.png" pageId="10" pageNumber="205">Fig. 7</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |