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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.391.7161" ID-GBIF-Dataset="2c378af4-8904-403d-b3ca-b16a2ee9951e" ID-PMC="PMC3974445" ID-Pensoft-Pub="1313-2970-391-75" ID-PubMed="24715781" ID-ZBK="D6A420ACB3BD4BE39E3C14766610838F" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-391-75" ModsDocOrigin="ZooKeys 391" ModsDocTitle="Four new spider species of the family Theridiosomatidae (Arachnida, Araneae) from caves in Laos" checkinTime="1451246140634" checkinUser="pensoft" docAuthor="Lin, Yucheng, Li, Shuqiang &amp; Jaeger, Peter" docDate="2014" docId="0F86E0E5F574F9B15E19487D380C32D4" docLanguage="en" docName="ZooKeys 391: 75-102" docOrigin="ZooKeys 391" docSource="http://dx.doi.org/10.3897/zookeys.391.7161" docTitle="Chthonopes thakekensis Lin, Li &amp; Jaeger, 2014, sp. n." docType="treatment" docUuid="92AA0D92-9434-414D-A86F-4596C0164644" docUuidSource="ZooBank" docVersion="4" lastPageNumber="86" masterDocId="FF9FFFFE412C3B2B1941FF90F72E8D0D" masterDocTitle="Four new spider species of the family Theridiosomatidae (Arachnida, Araneae) from caves in Laos" masterLastPageNumber="102" masterPageNumber="75" pageNumber="86" updateTime="1668158068389" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Four new spider species of the family Theridiosomatidae (Arachnida, Araneae) from caves in Laos</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lin, Yucheng</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Shuqiang</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jaeger, Peter</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>391</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>75</mods:start>
<mods:end>102</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.391.7161</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.391.7161</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-391-75</mods:identifier>
<mods:identifier type="ZBK">D6A420ACB3BD4BE39E3C14766610838F</mods:identifier>
<mods:identifier type="ZooBank">D6A420ACB3BD4BE39E3C14766610838F</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152051749" LSID="urn:lsid:zoobank.org:act:92AA0D92-9434-414D-A86F-4596C0164644" httpUri="http://treatment.plazi.org/id/0F86E0E5F574F9B15E19487D380C32D4" lastPageNumber="86" pageId="11" pageNumber="86">
<subSubSection pageId="11" pageNumber="86" type="nomenclature">
<paragraph pageId="11" pageNumber="86">
<taxonomicName LSID="http://zoobank.org/92AA0D92-9434-414D-A86F-4596C0164644" class="Arachnida" family="Theridiosomatidae" genus="Chthonopes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chthonopes thakekensis" order="Araneae" pageId="11" pageNumber="86" phylum="Arthropoda" rank="species" species="thakekensis">
<pageBreakToken pageId="11" pageNumber="86" start="start">Chthonopes</pageBreakToken>
thakekensis
</taxonomicName>
<taxonomicNameLabel pageId="11" pageNumber="86">sp. n.</taxonomicNameLabel>
Figs 17-19
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="86" type="material examined">
<paragraph pageId="11" pageNumber="86">Material examined.</paragraph>
<paragraph pageId="11" pageNumber="86">
Laos: Khammouan Province: Holotype: ♀ (SMF), Thakek area, Ban
<normalizedToken originalValue="Phôungam-Mai">Phoungam-Mai</normalizedToken>
,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="17.549232">17°32.954'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="104.81257">104°48.754'E</geoCoordinate>
, altitude 180 m, Tham Phayat, in limestone cave, leg. 25 November 2012.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="86" type="etymology">
<paragraph pageId="11" pageNumber="86">Etymology.</paragraph>
<paragraph pageId="11" pageNumber="86">This specific name is taken from type locality; adjective.</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="86" type="diagnosis">
<paragraph pageId="11" pageNumber="86">Diagnosis.</paragraph>
<paragraph pageId="11" pageNumber="86">
The new species is similar to
<taxonomicName class="Arachnida" family="Theridiosomatidae" genus="Chthonopes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chthonopes jaegeri" order="Araneae" pageId="11" pageNumber="86" phylum="Arthropoda" rank="species" species="jaegeri">Chthonopes jaegeri</taxonomicName>
Wunderlich, 2011 (see
<bibRefCitation author="Wunderlich, J" journalOrPublisher="Beitraege zur Araneologie" pageId="13" pageNumber="88" pagination="427 - 444" title="Extant and fossil spiders (Araneae)." volume="6" year="2011">Wunderlich 2011</bibRefCitation>
: 443, figs 18
<normalizedToken originalValue="df">d-f</normalizedToken>
) in the shape of epigyne and the configurations of vulva, but can be distinguished from it by the presence of translucent accessory spermathecae (Figs 17
<normalizedToken originalValue="DE">D-E</normalizedToken>
, 18C; absent in
<taxonomicName class="Arachnida" family="Theridiosomatidae" genus="Chthonopes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chthonopes jaegeri" order="Araneae" pageId="11" pageNumber="86" phylum="Arthropoda" rank="species" species="jaegeri">Chthonopes jaegeri</taxonomicName>
) and the large, semi-circular main spermathecae (Figs 17E, 18C; circular in
<taxonomicName class="Arachnida" family="Theridiosomatidae" genus="Chthonopes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chthonopes jaegeri" order="Araneae" pageId="11" pageNumber="86" phylum="Arthropoda" rank="species" species="jaegeri">Chthonopes jaegeri</taxonomicName>
).
</paragraph>
<caption pageId="11" pageNumber="86">
<paragraph pageId="11" pageNumber="86">
Figure 17.
<taxonomicName class="Arachnida" family="Theridiosomatidae" genus="Chthonopes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chthonopes thakekensis" order="Araneae" pageId="11" pageNumber="86" phylum="Arthropoda" rank="species" species="thakekensis">Chthonopes thakekensis</taxonomicName>
sp. n., holotype female. A Habitus, dorsal B Epigyne (untreated), ventral C Vulva, dorsal D Epigyne (lactic acid-treated), ventral E Vulva, dorsal. CD = copulatory duct; FD = fertilization duct; Sc = scape; Sp = spermathecae.
</paragraph>
</caption>
<caption pageId="11" pageNumber="86">
<paragraph pageId="11" pageNumber="86">
Figure 18.
<taxonomicName class="Arachnida" family="Theridiosomatidae" genus="Chthonopes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chthonopes thakekensis" order="Araneae" pageId="11" pageNumber="86" phylum="Arthropoda" rank="species" species="thakekensis">Chthonopes thakekensis</taxonomicName>
sp. n., holotype female. A Epigyne (untreated), ventral B Ditto (lactic acid-treated), ventral E Vulva (lactic acid-treated), dorsal. CD = copulatory duct; FD = fertilization duct; Sc = scape; Sp = spermathecae.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="11" pageNumber="86" type="description">
<paragraph pageId="11" pageNumber="86">Description.</paragraph>
<paragraph pageId="11" pageNumber="86">Habitus see in Fig. 17A. DS yellow, pear-shaped, smooth; eyes small, with black base, AME contiguous, LE contiguous, anterior eye row recurved, posterior eye row procurved; sternum yellow, with sparse setae; legs yellow; opisthosoma spherical, covered with sparse long setae, setal base sclerotized.</paragraph>
<paragraph pageId="11" pageNumber="86">Female copulatory organ (Figs 17C, 17E, 18C): epigyne large, with long setae in midline (Figs 17B, 18A); scape long, translucent, rugose, extending from posterior margin of epigynal plate, distal end weakly sclerotized (Fig. 17D); Spermathecae large, strongly sclerotized, separated by about 1.2 times their width (Fig. 17E); accessory spermathecae claviform, translucent (Fig. 17E); copulatory ducts wide, rugose, sclerotized, connected with posterior margin of main spermathecae (Figs 17E, 18C); fertilization ducts originating medio-posteriorly from main spermathecae, apical parts close to each other (Figs 17E, 18C).</paragraph>
<paragraph pageId="11" pageNumber="86">Female: Total length 2.51, DS 0.89 long, 0.85 wide, clypeus 0.18, sternum 0.48 long, 0.49 wide, coxae IV separated by 0.97 time their width, opisthosoma 1.63 long, 1.60 wide. PME separated by about 1.5 times their diameter. Macrosetae: Leg I: patella d2, tibia d2, p2, r1, metatarsus d2, p5, r2, v2; leg II: patella d2, tibia d6, p3, r2, v3, metatarsus d2, p4, r2, v4; leg III: patella d2, tibia d2, p3, r3, v3, metatarsus d2, p6, r4, v6; leg IV: patella d2, tibia d3, p3, r4, metatarsus d3, p4, r3, v2. Metatarsal trichobothria: Tm I: 0.20; Tm II: 0.15; Tm III: 0.27. Leg measurements: I 3.83 (1.17, 0.49, 0.83, 0.79, 0.55); II 3.40 (1.03, 0.42, 0.74, 0.69, 0.52); III 2.56 (0.76, 0.34, 0.51, 0.54, 0.41); IV 3.03 (0.95, 0.36, 0.68, 0.62, 0.42).</paragraph>
<paragraph pageId="11" pageNumber="86">Male unknown.</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="86" type="distribution">
<paragraph pageId="11" pageNumber="86">Distribution.</paragraph>
<paragraph pageId="11" pageNumber="86">See in Fig. 19.</paragraph>
<caption pageId="11" pageNumber="86">
<paragraph pageId="11" pageNumber="86">
Figure 19. Records of four species of
<taxonomicName family="Theridiosomatidae" lsidName="" pageId="11" pageNumber="86" rank="family">Theridiosomatidae</taxonomicName>
from Laos. Filled triangles -
<taxonomicName class="Gastropoda" family="Aporrhaidae" genus="Alaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alaria cavernicola" order="Littorinimorpha" pageId="11" pageNumber="86" phylum="Mollusca" rank="species" species="cavernicola">Alaria cavernicola</taxonomicName>
sp. n.; Open circles -
<taxonomicName class="Gastropoda" family="Aporrhaidae" genus="Alaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alaria navicularis" order="Littorinimorpha" pageId="11" pageNumber="86" phylum="Mollusca" rank="species" species="navicularis">Alaria navicularis</taxonomicName>
sp. n.; Filled circles -
<taxonomicName class="Gastropoda" family="Aporrhaidae" genus="Alaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alaria bicornis" order="Littorinimorpha" pageId="11" pageNumber="86" phylum="Mollusca" rank="species" species="bicornis">Alaria bicornis</taxonomicName>
sp. n.; Open square -
<taxonomicName class="Arachnida" family="Theridiosomatidae" genus="Chthonopes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chthonopes thakekensis" order="Araneae" pageId="11" pageNumber="86" phylum="Arthropoda" rank="species" species="thakekensis">Chthonopes thakekensis</taxonomicName>
sp. n., Stars - towns.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>