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<document id="3C7B7C5F06707818B2C97A82B596F786" ID-DOI="10.3853/j.2201-4349.67.2015.1603" ID-ISSN="2201-4349" ID-Zenodo-Dep="4684865" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618332088410" checkinUser="carolina" docAuthor="McAlpine, David K." docDate="2015" docId="03AB6A52FFEE5623FEBCD289F8E903AE" docLanguage="en" docName="RecAustMus.67.2.25-53.pdf" docOrigin="Records of the Australian Museum 67 (2)" docSource="http://dx.doi.org/10.3853/j.2201-4349.67.2015.1603" docStyle="DocumentStyle:A2A8AC5C71CC92C4EA50773B9B9E6D6A.1:RecAustMus.2008-2018.journal_article.1cover" docStyleId="A2A8AC5C71CC92C4EA50773B9B9E6D6A" docStyleName="RecAustMus.2008-2018.journal_article.1cover" docStyleVersion="1" docTitle="Bama (Bama) signifer McAlpine 2015, n.sp." docType="treatment" docVersion="4" lastPageNumber="36" masterDocId="FF92122AFFE2562FFFBBD010FC6B0771" masterDocTitle="Signal Flies of the Genus Bama (Diptera: Platystomatidae) in Papua New Guinea" masterLastPageNumber="53" masterPageNumber="25" pageNumber="36" updateTime="1698905128853" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="4EEE5BF5B3F84B8107154A52272F6DE8">Signal Flies of the Genus Bama (Diptera: Platystomatidae) in Papua New Guinea</mods:title>
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<mods:namePart id="0EBDC949652769FF699C7C4C356F413E">McAlpine, David K.</mods:namePart>
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<treatment id="03AB6A52FFEE5623FEBCD289F8E903AE" ID-DOI="http://doi.org/10.5281/zenodo.4684303" ID-Zenodo-Dep="4684303" LSID="urn:lsid:plazi:treatment:03AB6A52FFEE5623FEBCD289F8E903AE" httpUri="http://treatment.plazi.org/id/03AB6A52FFEE5623FEBCD289F8E903AE" lastPageNumber="36" pageId="12" pageNumber="36">
<subSubSection id="C31888CFFFEE5623FEBCD289FE3405C2" box="[263,607,664,691]" pageId="12" pageNumber="36" type="nomenclature">
<paragraph id="8BBDDB44FFEE5623FEBCD289FE3405C2" blockId="12.[263,607,664,691]" box="[263,607,664,691]" pageId="12" pageNumber="36">
<heading id="D0F56C28FFEE5623FEBCD289FE3405C2" box="[263,607,664,691]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="36" reason="2">
<emphasis id="B9760756FFEE5623FEBCD289FE3405C2" bold="true" box="[263,607,664,691]" pageId="12" pageNumber="36">
<taxonomicName id="4C02A0C7FFEE5623FEBCD289FE7005C3" authority="McAlpine, 2015" authorityName="McAlpine" authorityYear="2015" box="[263,539,664,691]" class="Insecta" family="Platystomatidae" genus="Bama" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="36" phylum="Arthropoda" rank="species" species="signifer" status="sp. nov." subGenus="Bama">
<emphasis id="B9760756FFEE5623FEBCD289FE7005C3" bold="true" box="[263,539,664,691]" italics="true" pageId="12" pageNumber="36">Bama (Bama) signifer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A245BA2DFFEE5623FD98D289FE3405C2" box="[547,607,665,691]" pageId="12" pageNumber="36" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C31888CFFFEE5623FECBD2DEFD9D0599" box="[368,502,718,744]" pageId="12" pageNumber="36" type="description">
<paragraph id="8BBDDB44FFEE5623FECBD2DEFD9D0599" blockId="12.[368,502,718,744]" box="[368,502,718,744]" pageId="12" pageNumber="36">
<figureCitation id="1339C7C1FFEE5623FECBD2DEFD9D0599" box="[368,502,718,744]" captionStart="Figures 1720" captionStartId="13.[166,238,1744,1766]" captionTargetBox="[186,1444,160,1715]" captionTargetId="figure-9@13.[166,1466,151,1729]" captionTargetPageId="13" captionText="Figures 1720. Bama signifer n.sp., holotype female. (17) Left wing. (18) Right mid coxa, anteroventral view. (19) Distal half of left arista, scale = 0.2 mm. (20) Dorsal view of abdomen, sclerites outlined only." figureDoi="http://doi.org/10.5281/zenodo.4684887" httpUri="https://zenodo.org/record/4684887/files/figure.png" pageId="12" pageNumber="36">Figs 1720</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31888CFFFEE5623FFC2D2EDFC80041C" pageId="12" pageNumber="36" type="materials_examined">
<paragraph id="8BBDDB44FFEE5623FFC2D2EDFC80041C" blockId="12.[121,749,765,877]" pageId="12" pageNumber="36">
<emphasis id="B9760756FFEE5623FFC2D2EDFD730464" bold="true" box="[121,280,765,789]" pageId="12" pageNumber="36">Type material</emphasis>
.
<materialsCitation id="3B6AD119FFEE5623FE9DD2EDFC80041C" collectingDate="1969-08" collectionCode="ANIC" elevation="1341" latitude="-9.916667" location="Agaun" longLatPrecision="1298" longitude="149.35" pageId="12" pageNumber="36" specimenCount="1" specimenCount-female="1" stateProvince="Milne Bay" typeStatus="holotype">
<typeStatus id="54B965E6FFEE5623FE9DD2EDFDE20464" box="[294,393,765,789]" pageId="12" pageNumber="36" type="holotype">Holotype</typeStatus>
<specimenCount id="9D0410CDFFEE5623FE2AD2EDFDCE0465" box="[401,421,765,788]" pageId="12" pageNumber="36" type="female"></specimenCount>
(unique).
<collectingRegion id="49C615A6FFEE5623FDAFD2EDFE860464" box="[532,749,765,789]" country="Papua New Guinea" name="Milne Bay" pageId="12" pageNumber="36">Milne Bay Province</collectingRegion>
(highlands of NE):
<location id="8EDD8D9FFFEE5623FEF3D30AFDF90443" LSID="urn:lsid:plazi:treatment:03AB6A52FFEE5623FEBCD289F8E903AE:8EDD8D9FFFEE5623FEF3D30AFDF90443" box="[328,402,794,818]" latitude="-9.916667" longLatPrecision="1298" longitude="149.35" name="Agaun" pageId="12" pageNumber="36" stateProvince="Milne Bay">Agaun</location>
,
<quantity id="4CFA76A1FFEE5623FE25D30AFE690443" box="[414,514,794,818]" metricMagnitude="3" metricUnit="m" metricValue="1.34112" pageId="12" pageNumber="36" unit="ft" value="4400.0">
<elevation id="002F3C77FFEE5623FE25D30AFE690443" box="[414,514,794,818]" metricMagnitude="3" metricUnit="m" metricValue="1.34112" pageId="12" pageNumber="36" unit="ft" value="4400.0">4400 feet</elevation>
</quantity>
[c.
<quantity id="4CFA76A1FFEE5623FD96D30AFEEB0443" box="[557,640,794,818]" metricMagnitude="3" metricUnit="m" metricValue="1.34" pageId="12" pageNumber="36" unit="m" value="1340.0">
<elevation id="002F3C77FFEE5623FD96D30AFEEB0443" box="[557,640,794,818]" metricMagnitude="3" metricUnit="m" metricValue="1.34" pageId="12" pageNumber="36" unit="m" value="1340.0">1340 m</elevation>
</quantity>
, c.
<geoCoordinate id="EE36BD83FFEE5623FD1DD30AFE860443" box="[678,749,794,818]" degrees="9" direction="south" minutes="55" orientation="latitude" pageId="12" pageNumber="36" precision="925" value="-9.916667">9°55'S</geoCoordinate>
<geoCoordinate id="EE36BD83FFEE5623FFC2D328FC8B0421" box="[121,224,824,848]" degrees="149" direction="east" minutes="21" orientation="longitude" pageId="12" pageNumber="36" precision="925" value="149.35">149°21'E</geoCoordinate>
],
<date id="FFBCFD84FFEE5623FF49D328FD0C043E" box="[242,359,823,848]" pageId="12" pageNumber="36" value="1969-08">
<collectingDate id="EFF8046CFFEE5623FF49D328FD0C043E" box="[242,359,823,848]" pageId="12" pageNumber="36" value="1969-08">Aug. 1969</collectingDate>
</date>
, R.P. (
<collectionCode id="ED134381FFEE5623FE15D328FD9E0421" box="[430,501,824,848]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="12" pageNumber="36">ANIC</collectionCode>
). On micropin through polyporus.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31888CFFFEE5623FFC2D392F8E903AE" pageId="12" pageNumber="36" type="description">
<paragraph id="8BBDDB44FFEE5623FFC2D392FE8704C9" blockId="12.[119,751,898,2008]" pageId="12" pageNumber="36">
<emphasis id="B9760756FFEE5623FFC2D392FC9604EB" bold="true" box="[121,253,898,922]" pageId="12" pageNumber="36">Description</emphasis>
(
<specimenCount id="9D0410CDFFEE5623FEAAD392FD4F04EB" box="[273,292,898,922]" pageId="12" pageNumber="36" type="female"></specimenCount>
,
<specimenCount id="9D0410CDFFEE5623FE89D393FD2D04EB" box="[306,326,899,922]" pageId="12" pageNumber="36" type="male"></specimenCount>
unknown). Slightly elongate brown to blackish fly, with wing heavily marked anteriorly to vein 4.
</paragraph>
<paragraph id="8BBDDB44FFEE5623FF2CD3ADFE7002FC" blockId="12.[119,751,898,2008]" pageId="12" pageNumber="36">
<emphasis id="B9760756FFEE5623FF2CD3ADFD7D04A4" box="[151,278,957,981]" italics="true" pageId="12" pageNumber="36">Coloration.</emphasis>
Head largely brown, shading to black on vertex and much of occiput, with silvery pruinescence on postorbital to postgenal zone. Antenna tawny-brown; arista becoming darker distally. Prelabrum tawny-brown; palpus grey-brown. Thorax largely dark brown; mesoscutum predominantly shining black. Legs: coxae tawny; mid coxa with larger setulae black, finer ones yellow; femora yellow; tibiae tawny-brown; tarsi darker-brown. Wing (
<figureCitation id="1339C7C1FFEE5623FD2AD49AFE8803D3" box="[657,739,1162,1186]" captionStart="Figures 1720" captionStartId="13.[166,238,1744,1766]" captionTargetBox="[186,1444,160,1715]" captionTargetId="figure-9@13.[166,1466,151,1729]" captionTargetPageId="13" captionText="Figures 1720. Bama signifer n.sp., holotype female. (17) Left wing. (18) Right mid coxa, anteroventral view. (19) Distal half of left arista, scale = 0.2 mm. (20) Dorsal view of abdomen, sclerites outlined only." figureDoi="http://doi.org/10.5281/zenodo.4684887" httpUri="https://zenodo.org/record/4684887/files/figure.png" pageId="12" pageNumber="36">Fig. 17</figureCitation>
) anteriorly to vein 4 largely brown-black, with first costal and base of second costal cell yellow, extreme base of marginal cell mid-brown, and hyaline zones in first basal and first posterior cells; behind vein 4 membrane largely hyaline, with narrow brown zone in second posterior cell along vein 4 and three small brown zones in discal cell; halter entirely pale yellow. Abdomen dorsally tawny to brownish; tergite 1 and lateral parts of tergite 2 blackish.
</paragraph>
<paragraph id="8BBDDB44FFEE5623FF2CD582FEB7019C" blockId="12.[119,751,898,2008]" pageId="12" pageNumber="36">
<emphasis id="B9760756FFEE5623FF2CD582FCB302DB" box="[151,216,1426,1450]" italics="true" pageId="12" pageNumber="36">Head.</emphasis>
Postfrons almost parallel-sided, near mid-length c. 0.39× as wide as head; height of cheek c. 0.23 of height of eye; face in profile mainly concave, becoming flat towards lower margin; fronto-orbital bristles apparently rather small (damaged); postvertical bristles very fine, closely placed. Antennal segment 3 c. 2.5× as long as deep, apically rounded; segment 4 prominent, rounded; segment 5 bare, relatively elongate, length distinctly more than twice its maximum diameter; segment 6 (
<figureCitation id="1339C7C1FFEE5623FE3AD66DFDB101E4" box="[385,474,1661,1685]" captionStart="Figures 1720" captionStartId="13.[166,238,1744,1766]" captionTargetBox="[186,1444,160,1715]" captionTargetId="figure-9@13.[166,1466,151,1729]" captionTargetPageId="13" captionText="Figures 1720. Bama signifer n.sp., holotype female. (17) Left wing. (18) Right mid coxa, anteroventral view. (19) Distal half of left arista, scale = 0.2 mm. (20) Dorsal view of abdomen, sclerites outlined only." figureDoi="http://doi.org/10.5281/zenodo.4684887" httpUri="https://zenodo.org/record/4684887/files/figure.png" pageId="12" pageNumber="36">Fig. 19</figureCitation>
) quite bare, becoming attenuated shortly beyond base, with leaf-like subapical expansion (palette). Prelabrum rather shallow, not prominent; palpus short, moderately broad but not distally expanded.
</paragraph>
<paragraph id="8BBDDB44FFEE5623FF2CD6E2F8CB06C6" blockId="12.[119,751,898,2008]" lastBlockId="12.[792,1424,151,645]" pageId="12" pageNumber="36">
<emphasis id="B9760756FFEE5623FF2CD6E2FC86007B" box="[151,237,1778,1802]" italics="true" pageId="12" pageNumber="36">Thorax.</emphasis>
Mesoscutum almost bare between setulae on much of dorsal surface, pubescent-pruinescent laterally; scutellum pubescent on entire dorsal surface; mesopleuron partly glossy, with very little pruinescence centrally and posteriorly; prescutellar acrostichal bristle absent; single pair of dorsocentral bristles more closely placed than in other species. Fore femur with stout posteroventral bristles on distal half and smaller posterodorsal bristles; mid coxa with distomedial lobe not narrowly produced, with large and small setulae. Wing (
<figureCitation id="1339C7C1FFEE5623FB84D0A5F8FD07BC" box="[1087,1174,181,205]" captionStart="Figures 1720" captionStartId="13.[166,238,1744,1766]" captionTargetBox="[186,1444,160,1715]" captionTargetId="figure-9@13.[166,1466,151,1729]" captionTargetPageId="13" captionText="Figures 1720. Bama signifer n.sp., holotype female. (17) Left wing. (18) Right mid coxa, anteroventral view. (19) Distal half of left arista, scale = 0.2 mm. (20) Dorsal view of abdomen, sclerites outlined only." figureDoi="http://doi.org/10.5281/zenodo.4684887" httpUri="https://zenodo.org/record/4684887/files/figure.png" pageId="12" pageNumber="36">Fig. 17</figureCitation>
): much of first costal cell and small basal zone of second costal cell bare; stem vein with few minute dorsal setulae; vein 2 curved but not abruptly flexed; second section of vein 4 moderately curved; penultimate section of vein 4 c. as long as anterior crossvein, almost 0.7× as long as discal crossvein; second basal and anal cells and basal hyaline zone in first basal cell largely bare; anal crossvein strongly curved on anterior part, with slight concave curvature posteriorly.
</paragraph>
<paragraph id="8BBDDB44FFEE5623FC83D1ADFFA8053B" blockId="12.[792,1424,151,645]" pageId="12" pageNumber="36">
<emphasis id="B9760756FFEE5623FC83D1ADFFF606A4" box="[824,925,445,469]" italics="true" pageId="12" pageNumber="36">Abdomen</emphasis>
(
<figureCitation id="1339C7C1FFEE5623FC16D1ADFF9406A4" box="[941,1023,445,469]" captionStart="Figures 1720" captionStartId="13.[166,238,1744,1766]" captionTargetBox="[186,1444,160,1715]" captionTargetId="figure-9@13.[166,1466,151,1729]" captionTargetPageId="13" captionText="Figures 1720. Bama signifer n.sp., holotype female. (17) Left wing. (18) Right mid coxa, anteroventral view. (19) Distal half of left arista, scale = 0.2 mm. (20) Dorsal view of abdomen, sclerites outlined only." figureDoi="http://doi.org/10.5281/zenodo.4684887" httpUri="https://zenodo.org/record/4684887/files/figure.png" pageId="12" pageNumber="36">Fig. 20</figureCitation>
). Compound tergite 1+2 subequal in length to tergite 3; sclerotized part of segment 7 (oviscape) almost as long; sternites 1, 2, and 3 large; sternite 4 relatively small; sternites 5 and 6 apparently very short (not clearly visible in
<typeStatus id="54B965E6FFEE5623FC3ED222FFDC053B" box="[901,951,562,586]" pageId="12" pageNumber="36">type</typeStatus>
).
</paragraph>
<paragraph id="8BBDDB44FFEE5623FC83D240F83105F5" blockId="12.[792,1424,151,645]" pageId="12" pageNumber="36">
<emphasis id="B9760756FFEE5623FC83D240FFD60516" box="[824,957,592,615]" italics="true" pageId="12" pageNumber="36">Dimensions.</emphasis>
Total length, 6.0 mm; length of thorax,
<quantity id="4CFA76A1FFEE5623FAD7D240FF2C05F5" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="12" pageNumber="36" unit="mm" value="2.4">2.4 mm</quantity>
; length of wing, 7.0 mm.
</paragraph>
<paragraph id="8BBDDB44FFEE5623FCA1D28AF91F04A6" blockId="12.[792,1423,666,1247]" pageId="12" pageNumber="36">
<emphasis id="B9760756FFEE5623FCA1D28AFF3005C3" bold="true" box="[794,859,666,690]" pageId="12" pageNumber="36">Notes</emphasis>
. The complete absence of prescutellar acrostichal bristles and the bare arista (
<figureCitation id="1339C7C1FFEE5623FBF8D2A7F8FA05A1" box="[1091,1169,695,720]" captionStart="Figures 1720" captionStartId="13.[166,238,1744,1766]" captionTargetBox="[186,1444,160,1715]" captionTargetId="figure-9@13.[166,1466,151,1729]" captionTargetPageId="13" captionText="Figures 1720. Bama signifer n.sp., holotype female. (17) Left wing. (18) Right mid coxa, anteroventral view. (19) Distal half of left arista, scale = 0.2 mm. (20) Dorsal view of abdomen, sclerites outlined only." figureDoi="http://doi.org/10.5281/zenodo.4684887" httpUri="https://zenodo.org/record/4684887/files/figure.png" pageId="12" pageNumber="36">Fig. 19</figureCitation>
) distinguish
<taxonomicName id="4C02A0C7FFEE5623FAA5D2A8F9E605A1" authorityName="McAlpine" authorityYear="2015" box="[1310,1421,696,720]" class="Insecta" family="Platystomatidae" genus="Bama" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="36" phylum="Arthropoda" rank="species" species="signifer">
<emphasis id="B9760756FFEE5623FAA5D2A8F9E605A1" box="[1310,1421,696,720]" italics="true" pageId="12" pageNumber="36">B. signifer</emphasis>
</taxonomicName>
from all other known species of
<taxonomicName id="4C02A0C7FFEE5623FB3BD2C6F8D7059C" authorityName="McAlpine" authorityYear="2001" box="[1152,1212,726,749]" class="Insecta" family="Platystomatidae" genus="Bama" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B9760756FFEE5623FB3BD2C6F8D7059C" box="[1152,1212,726,749]" italics="true" pageId="12" pageNumber="36">Bama</emphasis>
</taxonomicName>
. The presence of a subapical palette on the arista is an unusual condition of the female
<typeStatus id="54B965E6FFEE5623FC34D31FFF9B0456" box="[911,1008,783,807]" pageId="12" pageNumber="36" type="holotype">holotype</typeStatus>
. In males of a number of platystomatid species in several other genera an aristal palette is present, but in these the palette is generally absent in females. In
<taxonomicName id="4C02A0C7FFEE5623FCA1D378F964040E" authority="(Guerin-Meneville)" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1831" box="[794,1295,871,895]" class="Insecta" family="Platystomatidae" genus="Euprosopia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="36" phylum="Arthropoda" rank="species" species="maculipennis">
<emphasis id="B9760756FFEE5623FCA1D378F85A040E" box="[794,1073,871,895]" italics="true" pageId="12" pageNumber="36">Euprosopia maculipennis</emphasis>
(Gúerin-Méneville)
</taxonomicName>
the palette is present in both sexes, but smaller in the female. I expect that the presently unknown male of
<taxonomicName id="4C02A0C7FFEE5623FB35D3B3F89004CB" authorityName="McAlpine" authorityYear="2015" box="[1166,1275,930,954]" class="Insecta" family="Platystomatidae" genus="Bama" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="36" phylum="Arthropoda" rank="species" species="signifer">
<emphasis id="B9760756FFEE5623FB35D3B3F89004CB" box="[1166,1275,930,954]" italics="true" pageId="12" pageNumber="36">B. signifer</emphasis>
</taxonomicName>
will be found to have the palette at least as developed as in the female.
</paragraph>
<paragraph id="8BBDDB44FFEE5623FC83D3CDF8DE0363" blockId="12.[792,1423,666,1247]" pageId="12" pageNumber="36">The specific epithet is a Latin noun, meaning flag-bearer, in reference to the palette of the arista.</paragraph>
<paragraph id="8BBDDB44FFEE5623FC83D408F8E903AE" blockId="12.[792,1423,666,1247]" pageId="12" pageNumber="36">
One specimen (unsexed, abdomen missing) from Mount Missim,
<collectingRegion id="49C615A6FFEE5623FCC8D425F843033C" box="[883,1064,1077,1101]" country="Papua New Guinea" name="Morobe" pageId="12" pageNumber="36">Morobe Province</collectingRegion>
(labelled
<taxonomicName id="4C02A0C7FFEE5623FB31D426F8AC033C" authorityName="McAlpine" authorityYear="2001" box="[1162,1223,1078,1101]" class="Insecta" family="Platystomatidae" genus="Bama" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B9760756FFEE5623FB31D426F8AC033C" box="[1162,1223,1078,1101]" italics="true" pageId="12" pageNumber="36">Bama</emphasis>
</taxonomicName>
sp. P, BPB) has the contour of vein 2 and wing markings much as in
<taxonomicName id="4C02A0C7FFEE5623FAACD443F9EE031B" authorityName="McAlpine" authorityYear="2015" box="[1303,1413,1106,1130]" class="Insecta" family="Platystomatidae" genus="Bama" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="36" phylum="Arthropoda" rank="species" species="signifer">
<emphasis id="B9760756FFEE5623FAACD443F9EE031B" box="[1303,1413,1106,1130]" italics="true" pageId="12" pageNumber="36">B. signifer</emphasis>
</taxonomicName>
, except that the discal crossvein is enclosed in a brown mark. However, the arista is creamy-white and pubescent on its whole length, without apical expansion. I regard its specific status as indeterminate at present.
</paragraph>
</subSubSection>
</treatment>
</document>