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treatments-xml/data/F9/7A/87/F97A87E27406E66666DAFF65FE63F99C.xml
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<document ID-DOI="http://doi.org/10.5281/zenodo.6231988" ID-GBIF-Dataset="80959f8f-aa73-499e-93ec-5c07458c7935" ID-GBIF-Taxon="160743187" ID-ISSN="0024-0672" ID-Zenodo-Dep="6231988" checkinTime="1576401033600" checkinUser="plazi" docAuthor="Schilthuizen, Menno &amp; Perreau, Michel" docDate="2008" docId="F97A87E27406E66666DAFF65FE63F99C" docLanguage="en" docName="ZM.82.19.189-210.pdf" docOrigin="Zool. Med. Leiden 82 (19)" docStyle="DocumentStyle{}" docTitle="Ptomaphaginus kinabaluensis Schilthuizen &amp; Perreau 2008, spec. nov." docType="treatment" docVersion="5" lastPageNumber="195" masterDocId="0543FF9A7400E660674CFFA5FFD2FFE5" masterDocTitle="New species and new records of Ptomaphaginus Portevin from the Sunda region, Southeast Asia (Coleoptera: Leiodidae: Cholevinae)" masterLastPageNumber="210" masterPageNumber="189" pageId="6" pageNumber="195" updateTime="1645572769617" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New species and new records of Ptomaphaginus Portevin from the Sunda region, Southeast Asia (Coleoptera: Leiodidae: Cholevinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schilthuizen, Menno</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Perreau, Michel</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zool. Med. Leiden</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2008</mods:date>
<mods:detail type="pubDate">
<mods:number>2008-01-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>82</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>19</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>189</mods:start>
<mods:end>210</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.3576287</mods:identifier>
<mods:identifier type="GBIF-Dataset">80959f8f-aa73-499e-93ec-5c07458c7935</mods:identifier>
<mods:identifier type="ISSN">0024-0672</mods:identifier>
<mods:identifier type="Zenodo-Dep">3576287</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6231988" ID-GBIF-Taxon="160743187" ID-Zenodo-Dep="6231988" LSID="urn:lsid:plazi:treatment:F97A87E27406E66666DAFF65FE63F99C" httpUri="http://treatment.plazi.org/id/F97A87E27406E66666DAFF65FE63F99C" lastPageNumber="195" pageId="6" pageNumber="195">
<subSubSection box="[406,864,192,216]" pageId="6" pageNumber="195" type="nomenclature">
<paragraph blockId="6.[406,864,192,248]" box="[406,864,192,216]" pageId="6" pageNumber="195">
<heading box="[406,864,192,216]" centered="true" fontSize="10" level="3" pageId="6" pageNumber="195" reason="1">
<taxonomicName authority="Schilthuizen &amp; Perreau, 2008" authorityName="Schilthuizen &amp; Perreau" authorityYear="2008" box="[406,749,192,216]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="kinabaluensis" status="spec. nov.">
<emphasis bold="true" box="[406,749,192,216]" italics="true" pageId="6" pageNumber="195">Ptomaphaginus kinabaluensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[755,864,193,216]" pageId="6" pageNumber="195" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[535,736,224,248]" pageId="6" pageNumber="195" type="description">
<paragraph blockId="6.[406,864,192,248]" box="[535,736,224,248]" pageId="6" pageNumber="195">
<heading box="[535,736,224,248]" centered="true" fontSize="10" level="3" pageId="6" pageNumber="195" reason="1">
(
<figureCitation box="[542,651,224,248]" captionStart="Figs 16-25. 16" captionStartId="20.[151,190,1437,1456]" captionTargetBox="[151,1119,192,1408]" captionTargetPageId="20" captionText="Figs 16-25. 16, Ptomaphaginus anas spec. nov. holotype; aedeagus, dorsal view. 17, Ptomaphaginus anas spec. nov., holotype; aedeagus, lateral view. 18, Ptomaphaginus similipes spec. nov., holotype; aedeagus, dorsal view. 19, Ptomaphaginus similipes spec. nov., holotype; aedeagus, lateral view. 20, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, dorsal view. 21, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, lateral view (basal part partly obscured by the genital segment). 22, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, ventral view. 23, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, lateral view. 24, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, dorsal view. 25, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3576295" httpUri="https://zenodo.org/record/3576295/files/figure.png" pageId="6" pageNumber="195">figs 24-25</figureCitation>
;
<figureCitation box="[661,724,224,248]" captionStart="Figs 26-30. 26" captionStartId="21.[151,190,1138,1157]" captionTargetBox="[151,1120,192,1089]" captionTargetPageId="21" captionText="Figs 26-30. 26, Ptomaphaginus caroli spec. nov., holotype; habitus, dorsal view. 27, Ptomaphaginus similipes spec. nov., female paratype (from Crocker Range); protarsus. 28, Ptomaphaginus similipes spec. nov., male paratype (from Trus Madi); protarsus. 29, Ptomaphaginus kinabaluensis spec. nov., female paratype (from Gunung Kinabalu); elytral apices. 30, Ptomaphaginus kinabaluensis spec. nov., holotype; elytral apices." figureDoi="http://doi.org/10.5281/zenodo.3576297" httpUri="https://zenodo.org/record/3576297/files/figure.png" pageId="6" pageNumber="195">29-30</figureCitation>
).
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="195" type="materials_examined">
<paragraph blockId="6.[151,1120,287,587]" pageId="6" pageNumber="195">
Type material.—
<materialsCitation ID-GBIF-Occurrence="2512737937" collectingDate="2001-01-13" collectingDateMax="2001-01-18" collectingDateMin="2001-01-13" collectionCode="BOR" collectorName="M. Schilthuizen" country="Malaysia" elevation="1500" location="Kinabalu Park Headquarters" municipality="Mount Kinabalu" pageId="6" pageNumber="195" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<typeStatus box="[314,403,288,307]" pageId="6" pageNumber="195">Holotype</typeStatus>
<specimenCount box="[408,426,287,307]" pageId="6" pageNumber="195" type="male">♂</specimenCount>
:
<collectingCountry box="[437,525,288,307]" name="Malaysia" pageId="6" pageNumber="195">Malaysia</collectingCountry>
:
<collectingRegion box="[534,593,288,307]" country="Malaysia" name="Sabah" pageId="6" pageNumber="195">Sabah</collectingRegion>
:
<collectingMunicipality box="[602,760,288,307]" pageId="6" pageNumber="195">Mount Kinabalu</collectingMunicipality>
:
<location LSID="urn:lsid:plazi:treatment:F97A87E27406E66666DAFF65FE63F99C:740C602F7406E666644DFE85FBC0FED6" box="[769,1042,288,307]" country="Malaysia" municipality="Mount Kinabalu" name="Kinabalu Park Headquarters" pageId="6" pageNumber="195" stateProvince="Sabah">Kinabalu Park Headquarters</location>
,
<quantity box="[1052,1119,288,307]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="195" unit="m" value="1500.0">
<elevation box="[1052,1119,288,307]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="195" unit="m" value="1500.0">1500 m</elevation>
</quantity>
alt.,
<date box="[192,303,316,335]" pageId="6" pageNumber="195" value="2001-01-13" valueMax="2001-01-18" valueMin="2001-01-13">
<collectingDate box="[192,303,316,335]" pageId="6" pageNumber="195" value="2001-01-13" valueMax="2001-01-18" valueMin="2001-01-13">13-18.i.2001</collectingDate>
</date>
,
<collectingMethod box="[312,368,316,335]" pageId="6" pageNumber="195">pitfall</collectingMethod>
with lamb, leg.
<collectorName box="[522,670,316,335]" pageId="6" pageNumber="195">M. Schilthuizen</collectorName>
(
<collectionCode box="[683,731,316,335]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="6" pageNumber="195" type="Herbarium">BOR</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2512737975" collectingDate="2001-01-13" collectingDateMax="2001-01-18" collectingDateMin="2001-01-13" collectionCode="BOR" collectorName="M. Schilthuizen" country="Malaysia" elevation="1500" location="Kinabalu Park Headquarters" municipality="Mount Kinabalu" pageId="6" pageNumber="195" specimenCount="7" specimenCount-female="3" specimenCount-male="4" stateProvince="Sabah" typeStatus="paratype">
<typeStatus box="[746,840,316,335]" pageId="6" pageNumber="195">Paratypes</typeStatus>
same collection data as
<typeStatus pageId="6" pageNumber="195">holotype</typeStatus>
:
<specimenCount box="[203,255,343,363]" pageId="6" pageNumber="195" type="male">4 ♂♂</specimenCount>
<specimenCount box="[259,311,343,363]" pageId="6" pageNumber="195" type="female">3 ♀♀</specimenCount>
(
<collectionCode box="[321,369,344,363]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="6" pageNumber="195" type="Herbarium">BOR</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2512737958" box="[383,890,343,363]" collectingDate="1987-05-16" collectingDateMax="2001-01-18" collectingDateMin="2001-01-13" collectionCode="MHNG" collectorName="A. Smetana" country="Malaysia" elevation="1500" location="Kinabalu Park Headquarters" municipality="Mount Kinabalu" pageId="6" pageNumber="195" specimenCount="1" specimenCount-female="1" stateProvince="Sabah" typeStatus="paratype">
same locality,
<date box="[514,600,344,363]" pageId="6" pageNumber="195" value="1987-05-16">
<collectingDate box="[514,600,344,363]" pageId="6" pageNumber="195" value="1987-05-16">16.v.1987</collectingDate>
</date>
, leg.
<collectorName box="[646,757,344,363]" pageId="6" pageNumber="195">A. Smetana</collectorName>
:
<specimenCount box="[764,798,343,363]" pageId="6" pageNumber="195" type="female">1 ♀</specimenCount>
(
<collectionCode box="[809,884,344,363]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="6" pageNumber="195" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2512737957" collectingDate="1987-05-21" collectingDateMax="2001-01-18" collectingDateMin="2001-01-13" collectionCode="MHNG, CMPR" collectorName="A. Smetana" country="Malaysia" elevation="1500" location="Kinabalu Park Headquarters" municipality="Mount Kinabalu" pageId="6" pageNumber="195" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Sabah" typeStatus="paratype">
same locality,
<date box="[1029,1115,344,363]" pageId="6" pageNumber="195" value="1987-05-21">
<collectingDate box="[1029,1115,344,363]" pageId="6" pageNumber="195" value="1987-05-21">21.v.1987</collectingDate>
</date>
, leg.
<collectorName box="[190,303,372,391]" pageId="6" pageNumber="195">A. Smetana</collectorName>
:
<specimenCount box="[312,365,371,391]" pageId="6" pageNumber="195" type="male">2 ♂♂</specimenCount>
(
<collectionCode box="[377,452,372,391]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="6" pageNumber="195" type="Museum">MHNG</collectionCode>
,
<collectionCode box="[461,526,372,391]" country="India" name="Centre for Medicinal Plants Research" pageId="6" pageNumber="195" type="Herbarium">CMPR</collectionCode>
),
<specimenCount box="[543,578,371,391]" pageId="6" pageNumber="195" type="female">1 ♀</specimenCount>
(
<collectionCode box="[590,656,372,391]" country="India" name="Centre for Medicinal Plants Research" pageId="6" pageNumber="195" type="Herbarium">CMPR</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2512737953" collectingDate="1988-08-12" collectingDateMax="2001-01-18" collectingDateMin="2001-01-13" collectionCode="MHNG" collectorName="A. Smetana" country="Malaysia" elevation="1500" location="Kinabalu Park Headquarters" municipality="Mount Kinabalu" pageId="6" pageNumber="195" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
same locality,
<date box="[806,914,372,391]" pageId="6" pageNumber="195" value="1988-08-12">
<collectingDate box="[806,914,372,391]" pageId="6" pageNumber="195" value="1988-08-12">12.viii.1988</collectingDate>
</date>
, leg.
<collectorName box="[963,1075,372,391]" pageId="6" pageNumber="195">A. Smetana</collectorName>
:
<specimenCount box="[1084,1119,371,391]" pageId="6" pageNumber="195" type="male">1 ♂</specimenCount>
(
<collectionCode box="[158,233,400,419]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="6" pageNumber="195" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2512737944" box="[250,877,399,419]" collectingDate="1988-08-05" collectingDateMax="2001-01-18" collectingDateMin="2001-01-13" collectionCode="MHNG, CMPR" collectorName="A. Smetana" country="Malaysia" elevation="1500" location="Kinabalu Park Headquarters" municipality="Mount Kinabalu" pageId="6" pageNumber="195" specimenCount="2" specimenCount-male="2" stateProvince="Sabah" typeStatus="paratype">
same locality,
<date box="[385,481,400,419]" pageId="6" pageNumber="195" value="1988-08-05">
<collectingDate box="[385,481,400,419]" pageId="6" pageNumber="195" value="1988-08-05">5.viii.1988</collectingDate>
</date>
, leg.
<collectorName box="[532,645,400,419]" pageId="6" pageNumber="195">A. Smetana</collectorName>
:
<specimenCount box="[654,708,399,419]" pageId="6" pageNumber="195" type="male">2 ♂♂</specimenCount>
(
<collectionCode box="[721,796,400,419]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="6" pageNumber="195" type="Museum">MHNG</collectionCode>
,
<collectionCode box="[806,871,400,419]" country="India" name="Centre for Medicinal Plants Research" pageId="6" pageNumber="195" type="Herbarium">CMPR</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2512737990" collectingDate="1987-05-17" collectionCode="MHNG, CMPR" collectorName="A. Smetana" country="Malaysia" elevation="1600" location="Mempening" municipality="Mount Kinabalu" pageId="6" pageNumber="195" specimenCount="3" specimenCount-male="3" stateProvince="Sabah" typeStatus="paratype">
<collectingCountry box="[888,976,400,419]" name="Malaysia" pageId="6" pageNumber="195">Malaysia</collectingCountry>
:
<collectingRegion box="[986,1045,400,419]" country="Malaysia" name="Sabah" pageId="6" pageNumber="195">Sabah</collectingRegion>
:
<collectingMunicipality pageId="6" pageNumber="195">Mount Kinabalu</collectingMunicipality>
:
<location LSID="urn:lsid:plazi:treatment:F97A87E27406E66666DAFF65FE63F99C:740C602F7406E66667BBFE09FEB8FE5A" box="[247,362,428,447]" country="Malaysia" municipality="Mount Kinabalu" name="Mempening" pageId="6" pageNumber="195" stateProvince="Sabah">Mempening</location>
trail,
<quantity box="[415,482,428,447]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="195" unit="m" value="1600.0">
<elevation box="[415,482,428,447]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="195" unit="m" value="1600.0">1600 m</elevation>
</quantity>
,
<date box="[490,576,428,447]" pageId="6" pageNumber="195" value="1987-05-17">
<collectingDate box="[490,576,428,447]" pageId="6" pageNumber="195" value="1987-05-17">17.v.1987</collectingDate>
</date>
, leg.
<collectorName box="[622,732,428,447]" pageId="6" pageNumber="195">A. Smetana</collectorName>
:
<specimenCount box="[739,790,427,447]" pageId="6" pageNumber="195" type="male">3 ♂♂</specimenCount>
(
<collectionCode box="[800,875,428,447]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="6" pageNumber="195" type="Museum">MHNG</collectionCode>
;
<collectionCode box="[882,947,428,447]" country="India" name="Centre for Medicinal Plants Research" pageId="6" pageNumber="195" type="Herbarium">CMPR</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2512737988" collectingDate="1987-05-18" collectionCode="MHNG" collectorName="Burckhardt &amp; Lobl" country="Malaysia" county="Crocker Range" elevation="1600" location="Tambunan" municipality="Kota Kinabalu" pageId="6" pageNumber="195" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
<collectingCountry box="[961,1049,428,447]" name="Malaysia" pageId="6" pageNumber="195">Malaysia</collectingCountry>
:
<collectingRegion box="[1057,1116,428,447]" country="Malaysia" name="Sabah" pageId="6" pageNumber="195">Sabah</collectingRegion>
:
<collectingCounty box="[151,296,456,475]" pageId="6" pageNumber="195">Crocker Range</collectingCounty>
,
<quantity box="[308,380,456,475]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="195" unit="m" value="1600.0">
<elevation box="[308,380,456,475]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="195" unit="m" value="1600.0">1600 m</elevation>
</quantity>
, km 51 rte
<collectingMunicipality box="[496,638,456,475]" pageId="6" pageNumber="195">Kota Kinabalu</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:F97A87E27406E66666DAFF65FE63F99C:740C602F7406E66665C6FE6DFD23FE3E" box="[650,753,456,475]" country="Malaysia" county="Crocker Range" municipality="Kota Kinabalu" name="Tambunan" pageId="6" pageNumber="195" stateProvince="Sabah">Tambunan</location>
,
<date box="[765,830,456,475]" pageId="6" pageNumber="195" value="1987-05-18">
<collectingDate box="[765,830,456,475]" pageId="6" pageNumber="195" value="1987-05-18">18.v.87</collectingDate>
</date>
, leg.
<collectorName box="[884,990,456,475]" pageId="6" pageNumber="195">Burckhardt</collectorName>
&amp;
<collectorName box="[1025,1070,456,475]" pageId="6" pageNumber="195">Löbl</collectorName>
,
<specimenCount box="[1082,1119,455,475]" pageId="6" pageNumber="195" type="male">1 ♂</specimenCount>
(
<collectionCode box="[158,233,484,503]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="6" pageNumber="195" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2512737933" box="[248,1039,484,503]" collectingDate="1987-04-29" collectionCode="MHNG" collectorName="Burckhardt &amp; Lobl" country="Malaysia" elevation="1550" location="Mt. Kinabalu" pageId="6" pageNumber="195" specimenCount="1" stateProvince="Sabah" typeStatus="paratype">
<collectingCountry box="[248,336,484,503]" name="Malaysia" pageId="6" pageNumber="195">Malaysia</collectingCountry>
:
<collectingRegion box="[344,403,484,503]" country="Malaysia" name="Sabah" pageId="6" pageNumber="195">Sabah</collectingRegion>
:
<location LSID="urn:lsid:plazi:treatment:F97A87E27406E66666DAFF65FE63F99C:740C602F7406E66666D0FE41FDCBFE12" box="[412,537,484,503]" country="Malaysia" name="Mt. Kinabalu" pageId="6" pageNumber="195" stateProvince="Sabah">Mt. Kinabalu</location>
,
<quantity box="[546,616,484,503]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="6" pageNumber="195" unit="m" value="1550.0">
<elevation box="[546,616,484,503]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="6" pageNumber="195" unit="m" value="1550.0">1550 m</elevation>
</quantity>
.,
<date box="[629,722,484,503]" pageId="6" pageNumber="195" value="1987-04-29">
<collectingDate box="[629,722,484,503]" pageId="6" pageNumber="195" value="1987-04-29">29.iv.1987</collectingDate>
</date>
, leg.
<collectorName box="[769,875,484,503]" pageId="6" pageNumber="195">Burckhardt</collectorName>
&amp;
<collectorName box="[903,946,484,503]" pageId="6" pageNumber="195">Löbl</collectorName>
(
<collectionCode box="[957,1033,484,503]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="6" pageNumber="195" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="6.[151,1120,287,587]" pageId="6" pageNumber="195">
<materialsCitation ID-GBIF-Occurrence="2512737934" collectingDate="2001-01-13" collectingDateMax="2001-01-18" collectingDateMin="2001-01-13" collectionCode="BOR" location="Mount Kinabalu" pageId="6" pageNumber="195" specimenCount="2">
<typeStatus box="[151,340,512,531]" isOtherTypeStatus="true" pageId="6" pageNumber="195">Additional material</typeStatus>
(not included in the type series).—
<location LSID="urn:lsid:plazi:treatment:F97A87E27406E66666DAFF65FE63F99C:740C602F7406E66665F8FDA5FC81FDF6" box="[692,851,512,531]" name="Mount Kinabalu" pageId="6" pageNumber="195">Mount Kinabalu</location>
,
<date box="[863,974,512,531]" pageId="6" pageNumber="195" value="2001-01-13" valueMax="2001-01-18" valueMin="2001-01-13">
<collectingDate box="[863,974,512,531]" pageId="6" pageNumber="195" value="2001-01-13" valueMax="2001-01-18" valueMin="2001-01-13">13-18.i.2001</collectingDate>
</date>
,
<specimenCount box="[985,1092,512,531]" pageId="6" pageNumber="195" type="generic">1 specimen</specimenCount>
of unidentified sex in 70% ethanol,
<specimenCount box="[459,564,540,559]" pageId="6" pageNumber="195" type="generic">1 specimen</specimenCount>
of unidentified sex in 100% ethanol (
<collectionCode box="[913,961,540,559]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="6" pageNumber="195" type="Herbarium">BOR</collectionCode>
). (For full collection details, see Type material above.)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="195" type="diagnosis">
<paragraph blockId="6.[151,1120,629,1658]" pageId="6" pageNumber="195">
Diagnosis.— Habitus slender, ovoid. Elytra 1.2-1.3 times as long as their combined width (length measured from the caudal tip of the scutellum to the elytral apices). Aedeagus (
<figureCitation box="[244,350,694,718]" captionStart="Figs 16-25. 16" captionStartId="20.[151,190,1437,1456]" captionTargetBox="[151,1119,192,1408]" captionTargetPageId="20" captionText="Figs 16-25. 16, Ptomaphaginus anas spec. nov. holotype; aedeagus, dorsal view. 17, Ptomaphaginus anas spec. nov., holotype; aedeagus, lateral view. 18, Ptomaphaginus similipes spec. nov., holotype; aedeagus, dorsal view. 19, Ptomaphaginus similipes spec. nov., holotype; aedeagus, lateral view. 20, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, dorsal view. 21, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, lateral view (basal part partly obscured by the genital segment). 22, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, ventral view. 23, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, lateral view. 24, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, dorsal view. 25, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3576295" httpUri="https://zenodo.org/record/3576295/files/figure.png" pageId="6" pageNumber="195">figs 24-25</figureCitation>
) short and wide, with two elongated apical lateral ‘wings’ and a short terminal processus. Antennae short, as long as the width of the head. Female elytral apices drawn out. Male with a central extension on the 4
<superScript attach="left" box="[782,795,758,772]" fontSize="6" pageId="6" pageNumber="195">th</superScript>
visible abdominal sternite.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="195" type="description">
<paragraph blockId="6.[151,1120,629,1658]" pageId="6" pageNumber="195">
Description.— Habitus slender, ovoid. Moderately pigmented: mostly chestnut brown; only the legs, the elytral apices, the tips and the bases of the antennae lighter brown. Length 2.3-3.0 mm (n = 8). Antennae short and condensed, as long as the width of the head (n = 4). Articles 6 distinctly, 9 and 10 slightly wider than long. Male protarsi only slightly dilated: the protibia at its largest width (excluding the lateral spines) is 2.5 times broader than the first article of the protarsus (n = 1). In the male, the ventral sides of profemur and protibia do not carry any long hairs. Thorax 1.68-1.86 times as wide as long (n = 4), the caudal corners slightly drawn out. The mesosternal carina is strongly developed, its edge somewhat thickened. Elytra relatively slender, 1.23-1.30 times as long (measured from the caudal tip of the scutellum) as their combined width (n = 4). Elytral apices rounded (not truncated) in the male (
<figureCitation box="[717,787,1115,1139]" captionStart="Figs 26-30. 26" captionStartId="21.[151,190,1138,1157]" captionTargetBox="[151,1120,192,1089]" captionTargetPageId="21" captionText="Figs 26-30. 26, Ptomaphaginus caroli spec. nov., holotype; habitus, dorsal view. 27, Ptomaphaginus similipes spec. nov., female paratype (from Crocker Range); protarsus. 28, Ptomaphaginus similipes spec. nov., male paratype (from Trus Madi); protarsus. 29, Ptomaphaginus kinabaluensis spec. nov., female paratype (from Gunung Kinabalu); elytral apices. 30, Ptomaphaginus kinabaluensis spec. nov., holotype; elytral apices." figureDoi="http://doi.org/10.5281/zenodo.3576297" httpUri="https://zenodo.org/record/3576297/files/figure.png" pageId="6" pageNumber="195">fig. 30</figureCitation>
), and drawn out in the female (
<figureCitation box="[159,231,1147,1172]" captionStart="Figs 26-30. 26" captionStartId="21.[151,190,1138,1157]" captionTargetBox="[151,1120,192,1089]" captionTargetPageId="21" captionText="Figs 26-30. 26, Ptomaphaginus caroli spec. nov., holotype; habitus, dorsal view. 27, Ptomaphaginus similipes spec. nov., female paratype (from Crocker Range); protarsus. 28, Ptomaphaginus similipes spec. nov., male paratype (from Trus Madi); protarsus. 29, Ptomaphaginus kinabaluensis spec. nov., female paratype (from Gunung Kinabalu); elytral apices. 30, Ptomaphaginus kinabaluensis spec. nov., holotype; elytral apices." figureDoi="http://doi.org/10.5281/zenodo.3576297" httpUri="https://zenodo.org/record/3576297/files/figure.png" pageId="6" pageNumber="195">fig. 29</figureCitation>
). The 4
<superScript attach="left" box="[310,323,1147,1161]" fontSize="6" pageId="6" pageNumber="195">th</superScript>
visible abdominal sternite in the male with a central point at its caudal edge; the 6
<superScript attach="left" box="[272,285,1179,1193]" fontSize="6" pageId="6" pageNumber="195">th</superScript>
with a broad and relatively distinct central notch. Aedeagus (
<figureCitation box="[975,1083,1180,1204]" captionStart="Figs 16-25. 16" captionStartId="20.[151,190,1437,1456]" captionTargetBox="[151,1119,192,1408]" captionTargetPageId="20" captionText="Figs 16-25. 16, Ptomaphaginus anas spec. nov. holotype; aedeagus, dorsal view. 17, Ptomaphaginus anas spec. nov., holotype; aedeagus, lateral view. 18, Ptomaphaginus similipes spec. nov., holotype; aedeagus, dorsal view. 19, Ptomaphaginus similipes spec. nov., holotype; aedeagus, lateral view. 20, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, dorsal view. 21, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, lateral view (basal part partly obscured by the genital segment). 22, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, ventral view. 23, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, lateral view. 24, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, dorsal view. 25, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3576295" httpUri="https://zenodo.org/record/3576295/files/figure.png" pageId="6" pageNumber="195">figs 24-25</figureCitation>
) in dorsal view short and wide, approximately 1.5 times as long as wide. Apically abruptly narrowing, and there adorned with two long lateral ‘wings’. The tip is composed of a short, obtuse, triangular processus. In lateral view, the aedeagus is almost not curved, except for the apical quarter, which is bent ventrad under a distinct angle. Male genital segment relatively broad, at the apex slightly triangular, 2.5 times as long as wide.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="195" type="distribution">
<paragraph blockId="6.[151,1120,629,1658]" pageId="6" pageNumber="195">Distribution.— This species has been found only in lower montane forest on Mount Kinabalu and in the Crocker Range.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="195" type="discussion">
<paragraph blockId="6.[151,1120,629,1658]" pageId="6" pageNumber="195">
Remarks.— Based on aedeagal shape,
<taxonomicName authorityName="Schilthuizen &amp; Perreau" authorityYear="2008" box="[628,793,1439,1463]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis box="[628,793,1439,1463]" italics="true" pageId="6" pageNumber="195">P. kinabaluensis</emphasis>
</taxonomicName>
is closely related to
<taxonomicName authorityName="Schilthuizen &amp; Perreau" authorityYear="2008" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="bryantioides">
<emphasis italics="true" pageId="6" pageNumber="195">P. bryantioides</emphasis>
</taxonomicName>
, from which it differs strongly in the habitus, which is much more slender in
<taxonomicName authorityName="Schilthuizen &amp; Perreau" authorityYear="2008" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis italics="true" pageId="6" pageNumber="195">P. kinabaluensis</emphasis>
</taxonomicName>
. Also,
<taxonomicName authorityName="Schilthuizen &amp; Perreau" authorityYear="2008" box="[368,532,1504,1528]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis box="[368,532,1504,1528]" italics="true" pageId="6" pageNumber="195">P. kinabaluensis</emphasis>
</taxonomicName>
has distinctive secondary sexual characteristics: the only slightly dilated male protarsi, the drawn-out elytral apices in the female (
<figureCitation box="[1035,1107,1536,1560]" captionStart="Figs 26-30. 26" captionStartId="21.[151,190,1138,1157]" captionTargetBox="[151,1120,192,1089]" captionTargetPageId="21" captionText="Figs 26-30. 26, Ptomaphaginus caroli spec. nov., holotype; habitus, dorsal view. 27, Ptomaphaginus similipes spec. nov., female paratype (from Crocker Range); protarsus. 28, Ptomaphaginus similipes spec. nov., male paratype (from Trus Madi); protarsus. 29, Ptomaphaginus kinabaluensis spec. nov., female paratype (from Gunung Kinabalu); elytral apices. 30, Ptomaphaginus kinabaluensis spec. nov., holotype; elytral apices." figureDoi="http://doi.org/10.5281/zenodo.3576297" httpUri="https://zenodo.org/record/3576297/files/figure.png" pageId="6" pageNumber="195">fig. 29</figureCitation>
), and the absence of long hairs on the male forelegs.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="195" type="etymology">
<paragraph blockId="6.[151,1120,629,1658]" pageId="6" pageNumber="195">
Etymology.—
<taxonomicName authorityName="Schilthuizen &amp; Perreau" authorityYear="2008" box="[353,491,1601,1625]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis box="[353,491,1601,1625]" italics="true" pageId="6" pageNumber="195">kinabaluensis</emphasis>
</taxonomicName>
, a Latin adjective, meaning ‘from Kinabalu’. The name refers to the
<typeStatus box="[290,340,1634,1658]" pageId="6" pageNumber="195">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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