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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1034.63388" ID-GBIF-Dataset="bb9fed3d-8c07-41d8-947a-06926ac852a5" ID-PMC="PMC8093188" ID-Pensoft-Pub="1313-2970-1034-165" ID-Pensoft-UUID="B10A957A0D455B8F81DAD0409B37F8F5" ID-PubMed="33958929" ID-ZooBank="B93166004BE645879BB124B575FD405B" ModsDocID="1313-2970-1034-165" checkinTime="1619530782287" checkinUser="pensoft" docAuthor="Tong, Yanfeng, Sun, Xiaochen, Li, Shuqiang &amp; Bian, Dongju" docDate="2021" docId="D5330EE2985D50028BF3A227CC9D681F" docLanguage="en" docName="ZooKeys 1034: 165-197" docOrigin="ZooKeys 1034" docPubDate="2021-04-26" docSource="http://dx.doi.org/10.3897/zookeys.1034.63388" docTitle="Ischnothyreus mangun Tong &amp; Li 2021, sp. nov." docType="treatment" docUuid="0A789285-4F33-40FB-9B2A-8C2A5F165345" docUuidSource="ZooBank" docVersion="4" id="B10A957A0D455B8F81DAD0409B37F8F5" lastPageNumber="165" masterDocId="B10A957A0D455B8F81DAD0409B37F8F5" masterDocTitle="Taxonomic study of the genus Ischnothyreus (Araneae, Oonopidae) from Xishuangbanna Rainforest, southwestern China" masterLastPageNumber="197" masterPageNumber="165" pageNumber="165" updateTime="1668150208617" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic study of the genus Ischnothyreus (Araneae, Oonopidae) from Xishuangbanna Rainforest, southwestern China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tong, Yanfeng</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4348-7029</mods:nameIdentifier>
<mods:affiliation>Life Science College, Shenyang Normal University, Shenyang 110034, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sun, Xiaochen</mods:namePart>
<mods:affiliation>Life Science College, Shenyang Normal University, Shenyang 110034, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Shuqiang</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3290-5416</mods:nameIdentifier>
<mods:affiliation>Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
<mods:nameIdentifier type="email">lisq@ioz.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bian, Dongju</mods:namePart>
<mods:affiliation>CAS Key Laboratory of Forest Ecology and Management, Institute of Applied Ecology, Shenyang 110016, China</mods:affiliation>
<mods:nameIdentifier type="email">biandongju@163.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-04-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1034</mods:number>
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<mods:start>165</mods:start>
<mods:end>197</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1034.63388</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1034.63388</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1034-165</mods:identifier>
<mods:identifier type="ZooBank">B93166004BE645879BB124B575FD405B</mods:identifier>
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<treatment ID-GBIF-Taxon="181652951" LSID="urn:lsid:zoobank.org:act:0A789285-4F33-40FB-9B2A-8C2A5F165345" httpUri="http://treatment.plazi.org/id/D5330EE2985D50028BF3A227CC9D681F" lastPageNumber="165" pageId="0" pageNumber="165">
<subSubSection pageId="0" pageNumber="165" type="nomenclature">
<paragraph pageId="0" pageNumber="165">
<taxonomicName LSID="http://zoobank.org/0A789285-4F33-40FB-9B2A-8C2A5F165345" authority="Tong &amp; Li" authorityName="Tong &amp; Li" authorityYear="2021" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ischnothyreus mangun" order="Araneae" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="mangun" status="sp. nov.">Ischnothyreus mangun Tong &amp; Li</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="165">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Ischnothyreus mangun sp. nov., male holotype A-C habitus, dorsal, lateral and ventral views D-G prosoma, dorsal, lateral, ventral and anterior views H, I left chelicerae, anterior and posterior views. Abbreviation: stp = strong, tooth-like projection. Scale bars: 0.4 mm (A-G); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1034.63388.figure4" httpUri="https://binary.pensoft.net/fig/535988" pageId="0" pageNumber="165">Figures 4</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Ischnothyreus mangun sp. nov., male holotype, left palp, SEM A-C prolateral, retrolateral and dorsal views D, E palpal bulb, prolateral and retrolateral views F-H distal part of palpal bulb, dorsal, prolateral and retrolateral views. Abbreviations: llm = lamella-like membrane; nsm = needle-shaped membrane; rl = retrolateral lobe; rol = round lobe; vpr = ventral protuberance." figureDoi="10.3897/zookeys.1034.63388.figure5" httpUri="https://binary.pensoft.net/fig/535989" pageId="0" pageNumber="165">, 5</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Ischnothyreus mangun sp. nov., female paratype A-D prosoma, dorsal, lateral, anterior and ventral views E abdomen, dorsal view F epigastric region, ventral view. Abbreviation: css = central strongly sclerotized structure. Scale bars: 0.4 mm (A-E); 0.2 mm (F)." figureDoi="10.3897/zookeys.1034.63388.figure6" httpUri="https://binary.pensoft.net/fig/535990" pageId="0" pageNumber="165">, 6</figureCitation>
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Ischnothyreus spp., left male palp A-C I. cristiformis sp. nov. D-F I. mangun sp. nov. G-I I. mengyang sp. nov. J-L I. qidaoban sp. nov. M-O I. qiuxing A, D, G, J, M prolateral view B, E, H, K, N dorsal view C, F, I, L, O retrolateral view. Scale bars: 0.1 mm (A-F, M-O); 0.2 mm (G-L)." figureDoi="10.3897/zookeys.1034.63388.figure20" httpUri="https://binary.pensoft.net/fig/536004" pageId="0" pageNumber="165">, 20D-F</figureCitation>
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Ischnothyreus spp., left male chelicerae A, B I. cristiformis sp. nov. C, D I. mangun sp. nov. E, F I. mengyang sp. nov. G, H I. qidaoban sp. nov. A, C, E, G anterior view B, D, F, H posterior view. Abbreviations: ccp = cockscomb-shaped process; hsp = helmet-shaped process; lsp = large, sclerotized process." figureDoi="10.3897/zookeys.1034.63388.figure22" httpUri="https://binary.pensoft.net/fig/536006" pageId="0" pageNumber="165">, 22C, D</figureCitation>
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Endogyne of Ischnothyreus spp., A, B I. cristiformis sp. nov. C, D I. mangun sp. nov. E, F I. mengyang sp. nov. G, H I. qidaoban sp. nov. I, J I. qiuxing A, C, E, G, I ventral view B, D, F, H, J dorsal view. Abbreviations: a = apodemes; bsa = bell-shaped atrium; cd = central oval depression; csa = circular atrium; nsa = nipple-shaped atrium; oa = opening of the atrium; rsa = rectangular-shaped atrium; wt = winding tube. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1034.63388.figure23" httpUri="https://binary.pensoft.net/fig/536007" pageId="0" pageNumber="165">, 23C, D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="type material">
<paragraph pageId="0" pageNumber="165">Type material.</paragraph>
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Holotype</emphasis>
♂: China, Yunnan, Menghai County, Mangun Stockaded Village, Xishuangbanna Natural Reserve, secondary forest;
<geoCoordinate degrees="22" direction="north" minutes="02" orientation="latitude" precision="15" seconds="12" value="22.036667">
22°02
<normalizedToken originalValue="12">'12''</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate degrees="100" direction="east" minutes="23" orientation="longitude" precision="15" seconds="28" value="100.391106">
100°23
<normalizedToken originalValue="28">'28''</normalizedToken>
E
</geoCoordinate>
; 1179 m; 20.III.2016; S. Li leg. (SYNU-385).
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Paratype.</emphasis>
1♀: same data as for holotype (SYNU-386).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="diagnosis">
<paragraph pageId="0" pageNumber="165">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="165">
The new species is similar to
<taxonomicName lsidName="I. cristiformis" pageId="0" pageNumber="165" rank="species" species="cristiformis">
<emphasis italics="true" pageId="0" pageNumber="165">I. cristiformis</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName lsidName="I. xiaolongha" pageId="0" pageNumber="165" rank="species" species="xiaolongha">
<emphasis italics="true" pageId="0" pageNumber="165">I. xiaolongha</emphasis>
</taxonomicName>
sp. nov. in the lamella-like membrane of male palp and the short dorsal scutum of the abdomen, but can be distinguished by the unmodified male cheliceral fang (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Ischnothyreus mangun sp. nov., male holotype A-C habitus, dorsal, lateral and ventral views D-G prosoma, dorsal, lateral, ventral and anterior views H, I left chelicerae, anterior and posterior views. Abbreviation: stp = strong, tooth-like projection. Scale bars: 0.4 mm (A-G); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1034.63388.figure4" httpUri="https://binary.pensoft.net/fig/535988" pageId="0" pageNumber="165">4H, I</figureCitation>
) (vs the large cockscomb-shaped sclerotized process (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Ischnothyreus cristiformis sp. nov., male holotype A-C habitus, dorsal, lateral and ventral views D-G prosoma, dorsal, lateral, ventral and anterior views H-J left chelicerae, anterior, posterior and lateral views. Abbreviation: stp = strong, tooth-like projection. Scale bars: 0.4 mm (A-G); 0.1 mm (H-J)." figureDoi="10.3897/zookeys.1034.63388.figure1" httpUri="https://binary.pensoft.net/fig/535985" pageId="0" pageNumber="165">1H-J</figureCitation>
) in
<taxonomicName lsidName="I. cristiformis" pageId="0" pageNumber="165" rank="species" species="cristiformis">
<emphasis italics="true" pageId="0" pageNumber="165">I. cristiformis</emphasis>
</taxonomicName>
sp. nov. and the tongue-shaped sclerotized process (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Ischnothyreus xiaolongha sp. nov., male holotype A-C habitus, dorsal, lateral and ventral views D-G prosoma, dorsal, lateral, ventral and anterior views H-J left chelicerae, anterior, posterior and lateral views. Abbreviation: stp = strong, tooth-like projection. Scale bars: 0.4 mm (A-G); 0.1 mm (H-J)." figureDoi="10.3897/zookeys.1034.63388.figure17" httpUri="https://binary.pensoft.net/fig/536001" pageId="0" pageNumber="165">17H-J</figureCitation>
) in
<taxonomicName lsidName="I. xiaolongha" pageId="0" pageNumber="165" rank="species" species="xiaolongha">
<emphasis italics="true" pageId="0" pageNumber="165">I. xiaolongha</emphasis>
</taxonomicName>
sp. nov.), the narrow leaf-shaped retrolateral lobe of the male palp (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Ischnothyreus mangun sp. nov., male holotype, left palp, SEM A-C prolateral, retrolateral and dorsal views D, E palpal bulb, prolateral and retrolateral views F-H distal part of palpal bulb, dorsal, prolateral and retrolateral views. Abbreviations: llm = lamella-like membrane; nsm = needle-shaped membrane; rl = retrolateral lobe; rol = round lobe; vpr = ventral protuberance." figureDoi="10.3897/zookeys.1034.63388.figure5" httpUri="https://binary.pensoft.net/fig/535989" pageId="0" pageNumber="165">5F</figureCitation>
) (vs broad rectangular-shaped (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ischnothyreus cristiformis sp. nov., male holotype, left palp, SEM A-C prolateral, retrolateral and dorsal views D, E palpal bulb, prolateral and retrolateral views F-H distal part of palpal bulb, dorsal, prolateral and retrolateral views. Abbreviations: llm = lamella-like membrane; rl = retrolateral lobe; rol = round lobe; tsm = thread-shaped membrane; vpr = ventral protuberance." figureDoi="10.3897/zookeys.1034.63388.figure2" httpUri="https://binary.pensoft.net/fig/535986" pageId="0" pageNumber="165">2F</figureCitation>
) in
<taxonomicName lsidName="I. cristiformis" pageId="0" pageNumber="165" rank="species" species="cristiformis">
<emphasis italics="true" pageId="0" pageNumber="165">I. cristiformis</emphasis>
</taxonomicName>
sp. nov. and broad leaf-shaped (Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Ischnothyreus xiaolongha sp. nov., male holotype, left palp, SEM A-C prolateral, retrolateral and dorsal views D, E palpal bulb, prolateral and retrolateral views F-H distal part of palpal bulb, dorsal, prolateral and retrolateral views. Abbreviations: llm = lamella-like membrane; nsm = needle-shaped membrane; rl = retrolateral lobe; rol = round lobe; vpr = ventral protrusion." figureDoi="10.3897/zookeys.1034.63388.figure18" httpUri="https://binary.pensoft.net/fig/536002" pageId="0" pageNumber="165">18F</figureCitation>
) in
<taxonomicName lsidName="I. xiaolongha" pageId="0" pageNumber="165" rank="species" species="xiaolongha">
<emphasis italics="true" pageId="0" pageNumber="165">I. xiaolongha</emphasis>
</taxonomicName>
sp. nov.). The female differs from
<taxonomicName lsidName="I. cristiformis" pageId="0" pageNumber="165" rank="species" species="cristiformis">
<emphasis italics="true" pageId="0" pageNumber="165">I. cristiformis</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="I. xiaolongha" pageId="0" pageNumber="165" rank="species" species="xiaolongha">
<emphasis italics="true" pageId="0" pageNumber="165">I. xiaolongha</emphasis>
</taxonomicName>
by the small rectangular-shaped atrium (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Ischnothyreus mangun sp. nov., female paratype A-D prosoma, dorsal, lateral, anterior and ventral views E abdomen, dorsal view F epigastric region, ventral view. Abbreviation: css = central strongly sclerotized structure. Scale bars: 0.4 mm (A-E); 0.2 mm (F)." figureDoi="10.3897/zookeys.1034.63388.figure6" httpUri="https://binary.pensoft.net/fig/535990" pageId="0" pageNumber="165">6F</figureCitation>
) (vs the large bowl-shaped atrium (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ischnothyreus cristiformis sp. nov., female paratype A-C habitus, dorsal, lateral and ventral views D-G prosoma, dorsal, lateral, ventral and anterior views H epigastric region, ventral view. Abbreviation: css = central strongly sclerotized structure; Scale bars: 0.4 mm (A-G); 0.2 mm (H)." figureDoi="10.3897/zookeys.1034.63388.figure3" httpUri="https://binary.pensoft.net/fig/535987" pageId="0" pageNumber="165">3H</figureCitation>
) in
<taxonomicName lsidName="I. cristiformis" pageId="0" pageNumber="165" rank="species" species="cristiformis">
<emphasis italics="true" pageId="0" pageNumber="165">I. cristiformis</emphasis>
</taxonomicName>
sp. nov. and small bell-shaped atrium (Fig.
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Ischnothyreus xiaolongha sp. nov., female paratype A-C habitus, dorsal, lateral and ventral views D-G prosoma, dorsal, lateral, ventral and anterior views H epigastric region, ventral view. Scale bars: 0.4 mm (A-H)." figureDoi="10.3897/zookeys.1034.63388.figure19" httpUri="https://binary.pensoft.net/fig/536003" pageId="0" pageNumber="165">19H</figureCitation>
) in
<taxonomicName lsidName="I. xiaolongha" pageId="0" pageNumber="165" rank="species" species="xiaolongha">
<emphasis italics="true" pageId="0" pageNumber="165">I. xiaolongha</emphasis>
</taxonomicName>
sp. nov.).
</paragraph>
<caption doi="10.3897/zookeys.1034.63388.figure4" httpUri="https://binary.pensoft.net/fig/535988" pageId="0" pageNumber="165" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">Figure 4.</emphasis>
<taxonomicName authorityName="Tong &amp; Li" authorityYear="2021" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ischnothyreus mangun" order="Araneae" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="mangun">
<emphasis italics="true" pageId="0" pageNumber="165">Ischnothyreus mangun</emphasis>
</taxonomicName>
sp. nov., male holotype
<emphasis bold="true" pageId="0" pageNumber="165">A-C</emphasis>
habitus, dorsal, lateral and ventral views
<emphasis bold="true" pageId="0" pageNumber="165">D-G</emphasis>
prosoma, dorsal, lateral, ventral and anterior views
<emphasis bold="true" pageId="0" pageNumber="165">H, I</emphasis>
left chelicerae, anterior and posterior views. Abbreviation: stp = strong, tooth-like projection. Scale bars: 0.4 mm (
<emphasis bold="true" pageId="0" pageNumber="165">A-G</emphasis>
); 0.1 mm (
<emphasis bold="true" pageId="0" pageNumber="165">H, I</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="description">
<paragraph pageId="0" pageNumber="165">Description.</paragraph>
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">
Male (holotype).
<emphasis italics="true" pageId="0" pageNumber="165">Body</emphasis>
</emphasis>
: habitus as in Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Ischnothyreus mangun sp. nov., male holotype A-C habitus, dorsal, lateral and ventral views D-G prosoma, dorsal, lateral, ventral and anterior views H, I left chelicerae, anterior and posterior views. Abbreviation: stp = strong, tooth-like projection. Scale bars: 0.4 mm (A-G); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1034.63388.figure4" httpUri="https://binary.pensoft.net/fig/535988" pageId="0" pageNumber="165">4A-C</figureCitation>
; body length 1.52.
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Carapace</emphasis>
: 0.81 long, 0.61 wide; yellow, with egg-shaped patches behind eyes, surface of elevated portion of pars cephalica smooth, sides finely reticulate, lateral margin straight, smooth (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Ischnothyreus mangun sp. nov., male holotype A-C habitus, dorsal, lateral and ventral views D-G prosoma, dorsal, lateral, ventral and anterior views H, I left chelicerae, anterior and posterior views. Abbreviation: stp = strong, tooth-like projection. Scale bars: 0.4 mm (A-G); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1034.63388.figure4" httpUri="https://binary.pensoft.net/fig/535988" pageId="0" pageNumber="165">4D, E</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Clypeus</emphasis>
: height about 1.3 times ALE diameter (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Ischnothyreus mangun sp. nov., male holotype A-C habitus, dorsal, lateral and ventral views D-G prosoma, dorsal, lateral, ventral and anterior views H, I left chelicerae, anterior and posterior views. Abbreviation: stp = strong, tooth-like projection. Scale bars: 0.4 mm (A-G); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1034.63388.figure4" httpUri="https://binary.pensoft.net/fig/535988" pageId="0" pageNumber="165">4G</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Eyes</emphasis>
: see Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Ischnothyreus mangun sp. nov., male holotype A-C habitus, dorsal, lateral and ventral views D-G prosoma, dorsal, lateral, ventral and anterior views H, I left chelicerae, anterior and posterior views. Abbreviation: stp = strong, tooth-like projection. Scale bars: 0.4 mm (A-G); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1034.63388.figure4" httpUri="https://binary.pensoft.net/fig/535988" pageId="0" pageNumber="165">4D, G</figureCitation>
.
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Sternum</emphasis>
: pale orange (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Ischnothyreus mangun sp. nov., male holotype A-C habitus, dorsal, lateral and ventral views D-G prosoma, dorsal, lateral, ventral and anterior views H, I left chelicerae, anterior and posterior views. Abbreviation: stp = strong, tooth-like projection. Scale bars: 0.4 mm (A-G); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1034.63388.figure4" httpUri="https://binary.pensoft.net/fig/535988" pageId="0" pageNumber="165">4F</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Mouthparts</emphasis>
: chelicerae, endites and labium orange; chelicerae straight, base of fangs unmodified, fang groove with a few small denticles (Figs
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Ischnothyreus mangun sp. nov., male holotype A-C habitus, dorsal, lateral and ventral views D-G prosoma, dorsal, lateral, ventral and anterior views H, I left chelicerae, anterior and posterior views. Abbreviation: stp = strong, tooth-like projection. Scale bars: 0.4 mm (A-G); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1034.63388.figure4" httpUri="https://binary.pensoft.net/fig/535988" pageId="0" pageNumber="165">4H, I</figureCitation>
,
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Ischnothyreus spp., left male chelicerae A, B I. cristiformis sp. nov. C, D I. mangun sp. nov. E, F I. mengyang sp. nov. G, H I. qidaoban sp. nov. A, C, E, G anterior view B, D, F, H posterior view. Abbreviations: ccp = cockscomb-shaped process; hsp = helmet-shaped process; lsp = large, sclerotized process." figureDoi="10.3897/zookeys.1034.63388.figure22" httpUri="https://binary.pensoft.net/fig/536006" pageId="0" pageNumber="165">22C, D</figureCitation>
); anteromedian tip of endites with one strong, tooth-like projection (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Ischnothyreus mangun sp. nov., male holotype A-C habitus, dorsal, lateral and ventral views D-G prosoma, dorsal, lateral, ventral and anterior views H, I left chelicerae, anterior and posterior views. Abbreviation: stp = strong, tooth-like projection. Scale bars: 0.4 mm (A-G); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1034.63388.figure4" httpUri="https://binary.pensoft.net/fig/535988" pageId="0" pageNumber="165">4F</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Abdomen</emphasis>
: 0.65 long, 0.49 wide; dorsal scutum well sclerotized, pale orange, covering 1/3 of the abdomen width and approximately 1/2 of the abdomen length, not fused to epigastric scutum; epigastric and postgastric scutum well sclerotized, pale orange, fused, postgastric scutum covering about 1/2 of the abdomen length (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Ischnothyreus mangun sp. nov., male holotype A-C habitus, dorsal, lateral and ventral views D-G prosoma, dorsal, lateral, ventral and anterior views H, I left chelicerae, anterior and posterior views. Abbreviation: stp = strong, tooth-like projection. Scale bars: 0.4 mm (A-G); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1034.63388.figure4" httpUri="https://binary.pensoft.net/fig/535988" pageId="0" pageNumber="165">4A-C</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Legs</emphasis>
: pale orange, femur I with 2 prolateral spines, tibia I with 4 pairs, metatarsus I with 2 pairs of long ventral spines. Legs II lost. Legs III and IV spineless.
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Palp</emphasis>
: trochanter with ventral projection, cymbium brown; bulb with 2 ventral protuberances, one large and another very small, distal end of bulb stout, with numerous lamella-like membranes, retrolateral lobe narrow leaf-shaped (Figs
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Ischnothyreus mangun sp. nov., male holotype, left palp, SEM A-C prolateral, retrolateral and dorsal views D, E palpal bulb, prolateral and retrolateral views F-H distal part of palpal bulb, dorsal, prolateral and retrolateral views. Abbreviations: llm = lamella-like membrane; nsm = needle-shaped membrane; rl = retrolateral lobe; rol = round lobe; vpr = ventral protuberance." figureDoi="10.3897/zookeys.1034.63388.figure5" httpUri="https://binary.pensoft.net/fig/535989" pageId="0" pageNumber="165">5</figureCitation>
,
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Ischnothyreus spp., left male palp A-C I. cristiformis sp. nov. D-F I. mangun sp. nov. G-I I. mengyang sp. nov. J-L I. qidaoban sp. nov. M-O I. qiuxing A, D, G, J, M prolateral view B, E, H, K, N dorsal view C, F, I, L, O retrolateral view. Scale bars: 0.1 mm (A-F, M-O); 0.2 mm (G-L)." figureDoi="10.3897/zookeys.1034.63388.figure20" httpUri="https://binary.pensoft.net/fig/536004" pageId="0" pageNumber="165">20D-F</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1034.63388.figure5" httpUri="https://binary.pensoft.net/fig/535989" pageId="0" pageNumber="165" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">Figure 5.</emphasis>
<taxonomicName authorityName="Tong &amp; Li" authorityYear="2021" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ischnothyreus mangun" order="Araneae" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="mangun">
<emphasis italics="true" pageId="0" pageNumber="165">Ischnothyreus mangun</emphasis>
</taxonomicName>
sp. nov., male holotype, left palp, SEM
<emphasis bold="true" pageId="0" pageNumber="165">A-C</emphasis>
prolateral, retrolateral and dorsal views
<emphasis bold="true" pageId="0" pageNumber="165">D, E</emphasis>
palpal bulb, prolateral and retrolateral views
<emphasis bold="true" pageId="0" pageNumber="165">F-H</emphasis>
distal part of palpal bulb, dorsal, prolateral and retrolateral views. Abbreviations: llm = lamella-like membrane; nsm = needle-shaped membrane; rl = retrolateral lobe; rol = round lobe; vpr = ventral protuberance.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">Female (paratype, SYNU-386).</emphasis>
Same as male except as noted.
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Body</emphasis>
: body length 1.93.
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Carapace</emphasis>
: 0.80 long, 0.64 wide.
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Mouthparts</emphasis>
: chelicerae and endites unmodified.
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Abdomen</emphasis>
: 1.20 long, 0.81 wide; dorsal scutum very small.
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Epigastric area</emphasis>
: the postgastric scutum with a strongly sclerotized structure in the middle (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Ischnothyreus mangun sp. nov., female paratype A-D prosoma, dorsal, lateral, anterior and ventral views E abdomen, dorsal view F epigastric region, ventral view. Abbreviation: css = central strongly sclerotized structure. Scale bars: 0.4 mm (A-E); 0.2 mm (F)." figureDoi="10.3897/zookeys.1034.63388.figure6" httpUri="https://binary.pensoft.net/fig/535990" pageId="0" pageNumber="165">6F</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Endogyne</emphasis>
: from the middle of the slightly thickened margin of the postgastric scutum runs a dark, simple, winding tube posteriorly, ending in a small rectangular shaped atrium (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Endogyne of Ischnothyreus spp., A, B I. cristiformis sp. nov. C, D I. mangun sp. nov. E, F I. mengyang sp. nov. G, H I. qidaoban sp. nov. I, J I. qiuxing A, C, E, G, I ventral view B, D, F, H, J dorsal view. Abbreviations: a = apodemes; bsa = bell-shaped atrium; cd = central oval depression; csa = circular atrium; nsa = nipple-shaped atrium; oa = opening of the atrium; rsa = rectangular-shaped atrium; wt = winding tube. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1034.63388.figure23" httpUri="https://binary.pensoft.net/fig/536007" pageId="0" pageNumber="165">23C, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="etymology">
<paragraph pageId="0" pageNumber="165">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="165">The specific name is a noun in apposition taken from the type locality.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="distribution">
<paragraph pageId="0" pageNumber="165">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="165">Known only from the type locality.</paragraph>
<caption doi="10.3897/zookeys.1034.63388.figure6" httpUri="https://binary.pensoft.net/fig/535990" pageId="0" pageNumber="165" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">Figure 6.</emphasis>
<taxonomicName authorityName="Tong &amp; Li" authorityYear="2021" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ischnothyreus mangun" order="Araneae" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="mangun">
<emphasis italics="true" pageId="0" pageNumber="165">Ischnothyreus mangun</emphasis>
</taxonomicName>
sp. nov., female paratype
<emphasis bold="true" pageId="0" pageNumber="165">A-D</emphasis>
prosoma, dorsal, lateral, anterior and ventral views
<emphasis bold="true" pageId="0" pageNumber="165">E</emphasis>
abdomen, dorsal view
<emphasis bold="true" pageId="0" pageNumber="165">F</emphasis>
epigastric region, ventral view. Abbreviation: css = central strongly sclerotized structure. Scale bars: 0.4 mm (
<emphasis bold="true" pageId="0" pageNumber="165">A-E</emphasis>
); 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="165">F</emphasis>
).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>