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<document id="4F7487A9F440A345635D9280DD7DADFC" ID-DOI="http://dx.doi.org/10.3897/zookeys.423.7536" ID-GBIF-Dataset="1f17876d-2290-4d36-aa1f-10b951701679" ID-PMC="PMC4106094" ID-Pensoft-Pub="1313-2970-423-1" ID-PubMed="25061393" ID-ZooBank="55B4F9C858934F88841660FF730E8872" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-423-1" ModsDocOrigin="ZooKeys 423" ModsDocTitle="A revision of the Neotropical species of Bolitogyrus Chevrolat, a geographically disjunct lineage of Staphylinini (Coleoptera, Staphylinidae)" checkinTime="1451245649604" checkinUser="pensoft" docAuthor="Brunke, Adam J. &amp; Solodovnikov, Alexey" docDate="2014" docId="C2F327193A6EA00F264A42D2B671421D" docLanguage="en" docName="ZooKeys 423: 1-113" docOrigin="ZooKeys 423" docSource="http://dx.doi.org/10.3897/zookeys.423.7536" docTitle="Bolitogyrus salvini Sharp 1884" docType="treatment" docVersion="4" lastPageNumber="31" masterDocId="FFAB12065F4CFFED925DFFB46A5FFFFB" masterDocTitle="A revision of the Neotropical species of Bolitogyrus Chevrolat, a geographically disjunct lineage of Staphylinini (Coleoptera, Staphylinidae)" masterLastPageNumber="113" masterPageNumber="1" pageNumber="30" updateTime="1701318337016" updateUser="plazi">
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<mods:title id="40F36501651B70546DE2DCE71FCA77A2">A revision of the Neotropical species of Bolitogyrus Chevrolat, a geographically disjunct lineage of Staphylinini (Coleoptera, Staphylinidae)</mods:title>
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<mods:namePart id="EDFBDADE5B3AA728185E30F84A1A767F">Brunke, Adam J.</mods:namePart>
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<treatment id="C2F327193A6EA00F264A42D2B671421D" ID-GBIF-Taxon="152054096" LSID="urn:lsid:plazi:treatment:C2F327193A6EA00F264A42D2B671421D" httpUri="http://treatment.plazi.org/id/C2F327193A6EA00F264A42D2B671421D" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<subSubSection id="36D0960831A0AFFE42534D28539448A6" pageId="29" pageNumber="30" type="multiple">
<paragraph id="10FC5259DDE1CE081D8842F9B0DE6A9E" pageId="29" pageNumber="30">
<pageBreakToken id="625B67125CDD1D26854852A89C24961F" pageId="29" pageNumber="30" start="start">Taxon</pageBreakToken>
classification Animalia Coleoptera Staphylinidae
</paragraph>
</subSubSection>
<subSubSection id="DB0E39EB7562C7B312DC4ABA5184060D" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="595F8F00030B9B596A27F4D198651FE5" pageId="29" pageNumber="30">
<taxonomicName id="6EFC60241C80D800895FF05063447109" ID-CoL="5WMWT" authority="Sharp, 1884" authorityName="Sharp" authorityYear="1884" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus salvini" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="salvini">Bolitogyrus salvini (Sharp, 1884)</taxonomicName>
Figs 3
<normalizedToken id="D9C74D8B68E65E41CD23439F63D162CB" originalValue="AB">A-B</normalizedToken>
, 10B, 16
<normalizedToken id="B1852A5D771950A6775F603DBEB41B1B" originalValue="CH">C-H</normalizedToken>
, 23
<normalizedToken id="32D2F27AE4575E92B3795E2431664029" originalValue="EF">E-F</normalizedToken>
, 26
<normalizedToken id="8F98128AE8987DB732085E6DE1345331" originalValue="AC">A-C</normalizedToken>
, 27G, 30A (map)
</paragraph>
</subSubSection>
<subSubSection id="A79B79FF0B29F17F062EFC5972F7F90A" pageId="29" pageNumber="30" type="reference_group">
<paragraph id="72F1D055E0FB34C563799EE3CFF08C37" pageId="29" pageNumber="30">
<taxonomicName id="8347F283CFA201E542C6C110190CA33F" class="Insecta" family="Staphylinidae" genus="Cyrtothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtothorax salvini" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="salvini">Cyrtothorax salvini</taxonomicName>
Sharp, 1884: 341
</paragraph>
<paragraph id="F07461FFB0A669D487076204E43C7246" pageId="29" pageNumber="30">
<taxonomicName id="2A8F2622750F42B6C5E5B9936A1497A8" class="Insecta" family="Staphylinidae" genus="Cyrtothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtothorax salvini" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="salvini">Cyrtothorax salvini</taxonomicName>
:
<bibRefCitation id="58ED7318598E53934F765785F458BA9A" author="Scheerpeltz, O" journalOrPublisher="Reichenbachia" pageId="72" pageNumber="73" pagination="175 - 192" title="Studien an den Arten der Gattung Cyrtothorax Kraatz mit Beschreibung neuer Arten sowie einer Dichotomik aller bis heute bekannt gewordenen Arten dieser Gattung (Coleoptera, Staphylinindae, Staphylininae, Quediini)." volume="15" year="1974">Scheerpeltz 1974</bibRefCitation>
: 183 (in key)
</paragraph>
</subSubSection>
<subSubSection id="A7C840B6C0A21925C71ED92E3CF3515B" pageId="29" pageNumber="30" type="type locality">
<paragraph id="99A36E1417AB90AF02AE7255C2689BFF" pageId="29" pageNumber="30">Type locality.</paragraph>
<paragraph id="B15B4F076BF5B5F724B0C67656B620C7" pageId="29" pageNumber="30">Zapote (=El Zapote, near Escuintla), Guatemala.</paragraph>
</subSubSection>
<subSubSection id="09F4C06494B3A14526EACF576DAF3715" pageId="29" pageNumber="30" type="type material">
<paragraph id="B110202116F4727E4A14CDE7EFC16DEC" pageId="29" pageNumber="30">Type material.</paragraph>
<paragraph id="D923C646288276561FC382819639DB0B" pageId="29" pageNumber="30">
<taxonomicName id="3856DBB076C5328E2386297615F49F95" class="Insecta" family="Staphylinidae" genus="Cyrtothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtothorax salvini" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="salvini">Cyrtothorax salvini</taxonomicName>
Sharp, 1884.
</paragraph>
<paragraph id="2920ABAA9685464A183AF8F49D862F57" pageId="29" pageNumber="30">
Lectotype (here designated, ♂ BMNH): &quot;♂,
<taxonomicName id="6191CD5B41B54B7B9E8E1D366BD72794" class="Insecta" family="Staphylinidae" genus="Cyrtothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtothorax salvini" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="salvini">Cyrtothorax salvini</taxonomicName>
, ♂ Type D.S., Zapote, Guat.[=emala] Champion [written on card in
<normalizedToken id="C62C051EF01CD628FCC86E7793302811" originalValue="Sharps">Sharp's</normalizedToken>
hand with specimen]/ Zapote, Guatemala, Champion [printed label]/ B.C.A. Col. I.2.,
<taxonomicName id="5654C204C8232660B9C170158432C273" class="Insecta" family="Staphylinidae" genus="Cyrtothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtothorax" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">Cyrtothorax</taxonomicName>
, salvini, Sharp. [printed label]/ Type [printed on disc with red border]/ Sharp Coll., 1905.-313. [printed label] / Lectotype ♂,
<taxonomicName id="9E60E51958217012378D68E7C4C633F8" class="Insecta" family="Staphylinidae" genus="Cyrtothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtothorax salvini" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="salvini">Cyrtothorax salvini</taxonomicName>
Sharp, 1884, des. A. Brunke 2013 [red printed label].
</paragraph>
<paragraph id="EC66CB270428E930304901ABE4A17D94" pageId="29" pageNumber="30">
Paralectotype (1 ♀, BMNH): &quot;♀,
<taxonomicName id="72E76C92AA60219314428E981E19854A" class="Insecta" family="Staphylinidae" genus="Cyrtothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtothorax salvini" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="salvini">Cyrtothorax salvini</taxonomicName>
, ♀ Type D.S., Zapote, Guat.[=emala] Champion [written on card in
<normalizedToken id="E3BD57858C667D07F2161994FE76B762" originalValue="Sharps">Sharp's</normalizedToken>
hand with specimen]/ Zapote, Guatemala, Champion [printed label]/
<normalizedToken id="98F3C44638D52290B5CC6F795B751F79" originalValue="“G.”">&quot;G.&quot;</normalizedToken>
[handwritten]/ Type [printed on disc with red border]/ Sp. figured [printed]/ B.C.A. Col. I.2.,
<taxonomicName id="C80BB924232FC220FB45C4F38AAA7D65" class="Insecta" family="Staphylinidae" genus="Cyrtothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtothorax" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">Cyrtothorax</taxonomicName>
, salvini, Sharp. [printed label] / Paralectotype
<taxonomicName id="9E1E71745BAB25A92B5A6E3E754CE9C7" class="Insecta" family="Staphylinidae" genus="Cyrtothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtothorax salvini" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="salvini">Cyrtothorax salvini</taxonomicName>
Sharp, 1884, det. A. Brunke 2013.
</paragraph>
<paragraph id="E5CE631E410C2DE10BBB84BD283ACFC4" pageId="29" pageNumber="30">
The male and female syntypes are identical in color and are easily separated from other species of the Buphthalmus Group based on male and female genitalia, though considerable variation exists within the present species concept. To provide nomenclatural stability for this name, the male syntype was selected and is here designated as the lectotype, as male sexual characters provide more reliable diagnostic characters than those of females in
<taxonomicName id="2F1464D0A20A8169F8F6B64CFB2DDD99" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">Bolitogyrus</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="5DD8AF286B359FE8AF4A8A89A70A9676" pageId="29" pageNumber="30" type="other material">
<paragraph id="073F6F70E33C79454034A9522405D012" pageId="29" pageNumber="30">Other material.</paragraph>
<paragraph id="0CBBB11ECC2D74E6C834FE9B973C0EE5" pageId="29" pageNumber="30">
MEXICO:
<normalizedToken id="1DFEBEAF782672EACCFE179F2D430ED9" originalValue="Sallé">Salle</normalizedToken>
, (ex. syntype series of
<taxonomicName id="49C1FB368EA3160E59BEB89214BF8DE4" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus sallei" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="sallei">Bolitogyrus sallei</taxonomicName>
), 1 ♂ (DEI);
</paragraph>
<paragraph id="693E3E7BD3EEAA46D26A5D1A3B0FFEDC" pageId="29" pageNumber="30">
Chiapas: Parque Laguna Belgica, 19 km N. Ocozocoautla, 970m, 8.VI.1991, ex: flight intercept trap, J.S. Ashe Coll#75, SM0038002, 1 ♂ (SEMC), same except: 12.VI.1991, J.S. Ashe Coll#95, SM0038003;
<normalizedToken id="97D8DEE06A8A6F83E2A6CD749FA0B2D5" originalValue="Playón">Playon</normalizedToken>
de la Gloria,
<geoCoordinate id="36C23BFC5526AD0B707CA0760AF93DD7" direction="north" orientation="latitude" precision="1" value="16.14818">16.14818°N</geoCoordinate>
,
<geoCoordinate id="7EC5CDD310860417520140EE0E65ABFF" direction="west" orientation="longitude" precision="1" value="-90.8966">90.89660°W</geoCoordinate>
, 180m, mature wet forest, malaise trap, 24.VI.2008, LLAMA08Ma-A-09-2-01, SM0833172, 1 ♀ (SEMC). NICARAGUA:Jinotega: Reserva Nationale Dantali El Diablo, 13.10367N, -85.86904W
<normalizedToken id="E429336356E425E7BAADA4AD2A700170" originalValue="±">+/-</normalizedToken>
10m, 1380m, cloud forest, ex. Malaise trap, 19 to 21.V.2011, LLAMA11Ma-D04-3-02, 1 ♀ (ZMUC, DNA extracted); Reserva Nationale Cerro
<normalizedToken id="F4DA1F36148873873831319057CDE20B" originalValue="Kilambé">Kilambe</normalizedToken>
, 13.56980N, -85.69742
<normalizedToken id="5E460BA39EE9868FB8F7ABA67E9821A6" originalValue="±">+/-</normalizedToken>
10m, 1470m, cloud forest, ex. Malaise trap, 23 to 26.V.2011, LLAMA11Ma-D05-1-02, 1 ♀ (SEMC). Matagalpa: 6 km N Matagalpa, Selva Negra Hotel,
<geoCoordinate id="AAA0A214A3B2828F0B53A7DEDD85C560" direction="north" orientation="latitude" precision="9" value="12.999833">12°59.99'N</geoCoordinate>
,
<geoCoordinate id="5E1A4B67BD72FAC9C1E058FD392043CD" direction="west" orientation="longitude" precision="9" value="-85.90884">85°54.53'W</geoCoordinate>
, 1350m, ex. sweeping vegetation, 21.V.2002, R. Brooks, Z. Falin, S. Chatzimanolis, SM0536468, 1 ♀ (SEMC).
</paragraph>
</subSubSection>
<subSubSection id="F97E2E3F5836F840043E4CE96EBBF1A0" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="46FAC3B0D07EFF46641FBEF9B20B55B6" pageId="29" pageNumber="30">Diagnosis.</paragraph>
<paragraph id="D502A8DB848817D8AB0CB3748184C5E0" pageId="29" pageNumber="30">
Head without central protuberance; antenna distinctly bicolored (Fig. 5A); abdominal sternites with fine transverse microsculpture, interspaces about as wide as lines (Fig. 13D); median lobe in lateral view recurved dorsad (Fig. 15F); apical portion of median lobe in parameral view wide at constricted base (Fig. 16C); paramere with divergent arms (Fig. 16
<normalizedToken id="2D348AF6384188D87B1A47AA76783FBD" originalValue="DE">D-E</normalizedToken>
); female tergite X narrow at base, raised area on disc contiguous with base and not strongly converging to apex (Fig. 26
<normalizedToken id="5975C8FDC02921E801E545E28A4760B1" originalValue="AC">A-C</normalizedToken>
); female gonocoxite not swollen at base (Fig. 27G).
</paragraph>
</subSubSection>
<subSubSection id="B2DCC759424C769B473FA1F583274A8D" pageId="30" pageNumber="31" type="redescription">
<paragraph id="5104D101162C3DACC7EA01551492999A" pageId="30" pageNumber="31">
<pageBreakToken id="8841C8C290B5CFB07802ABCA66206F47" pageId="30" pageNumber="31" start="start">Redescription</pageBreakToken>
.
</paragraph>
<paragraph id="C565A144486141F78041BF57DC7C54CB" pageId="30" pageNumber="31">Measurements ♂ (n=3): HW/HL 1.62-1.69; PW/PL 1.33-1.50; EW/EL 1.21-1.32; ESut/PL 0.71-0.74; PW/HW 1.09-1.14; forebody length 4.80-5.00 mm.</paragraph>
<paragraph id="EEC70E5E7C3ADAADC554B78A3DA8616B" pageId="30" pageNumber="31">Measurements ♀ (n=4): HW/HL 1.43-1.50; PW/PL 1.35-1.50; EW/EL 1.25-1.32; ESut/PL 0.73-0.77; PW/HW 1.09-1.15; forebody length 5.0-5.1 mm.</paragraph>
<paragraph id="D871E668FC84C82AF743BD986E79C331" pageId="30" pageNumber="31">
Similar to
<taxonomicName id="5BFCB05ABE4E7E4C870B6BC4C5861C38" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus buphthalmus" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="buphthalmus">Bolitogyrus buphthalmus</taxonomicName>
and differing only in the following: head and pronotum metallic green to greenish bronze or black, without metallic reflection; elytra reddish or black with greenish to bronze metallic reflections, base of elytra concolorous with disc; abdominal segments
<normalizedToken id="B181121A7F23859CD386B84E9A25122B" originalValue="IIIV">III-V</normalizedToken>
reddish or completely dark; abdominal segments VIII and genital segment yellow to yellow-orange, distinctly contrasting with previous segments, or entirely dark and not contrasting; legs distinctly bicolored (as in
<taxonomicName id="69283BBEBEFC66E8776F811F388A0180" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus buphthalmus" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="buphthalmus">Bolitogyrus buphthalmus</taxonomicName>
), with dark procoxa in females, or entirely reddish (the lectotype and paralectotype); median frontal impression weak to moderately impressed (morphotype 3); head impunctate in middle of disc or not (lectotype and paralectotype); disc of tergites
<normalizedToken id="EBC4F6605D1057AF9593138521703C35" originalValue="IIIV">III-V</normalizedToken>
impunctate at middle, disc of VI punctate; median lobe in lateral view recurved dorsad; apical portion of median lobe constricted at base but weakly such that base is relatively wide; paramere with arms relatively flat, not curved around median lobe, arms divergent and peg setae as illustrated, varying from dense to sparse; male sternite IX similar to
<taxonomicName id="A4255F7E1CC3D2553DF3429DE93F2865" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus buphthalmus" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="buphthalmus">Bolitogyrus buphthalmus</taxonomicName>
but median emargination deeper; female tergite X distinctive, narrow at base, raised area on disc contiguous with base and not strongly converging to apex; strongly pigmented area variable at apex.
</paragraph>
</subSubSection>
<subSubSection id="668B3C7C891E5500291C8677D6A60F9E" pageId="30" pageNumber="31" type="distribution">
<paragraph id="380DA1057A30FF1B00D3A6ECB7C530D6" pageId="30" pageNumber="31">Distribution.</paragraph>
<paragraph id="13E06A16C01D1B63D606D94A44E007DA" pageId="30" pageNumber="31">Figure 30A. This species is known from southern Mexico, Guatemala and central Nicaragua (Matagalpa and Jinotega).</paragraph>
</subSubSection>
<subSubSection id="0FCCFF75635C61EF367BA654741A7C56" pageId="30" pageNumber="31" type="bionomics">
<paragraph id="3A00479CF0F2CC2139D580F86E947B2A" pageId="30" pageNumber="31">Bionomics.</paragraph>
<paragraph id="C28FB07C99AA76A01CCFB85B42FDCC53" pageId="30" pageNumber="31">
All specimens were either swept from foliage or collected from Malaise/flight intercept traps during May and June. As in
<taxonomicName id="C5A32A10C9433E675FA842369A917264" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus fulgidus" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="fulgidus">Bolitogyrus fulgidus</taxonomicName>
, very few males are known and it is possible that this species normally occurs in a forest stratum higher than accessed by most collecting efforts.
<taxonomicName id="4B682FC71A8305F19CCB1A88F5952099" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus salvini" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="salvini">Bolitogyrus salvini</taxonomicName>
has been collected from a wide range of forested habitats: from wet lowland forest (180 m) to cloud forest (1470 m).
</paragraph>
</subSubSection>
<subSubSection id="52A53DC2CD52CF6F8C9F35A234419477" pageId="30" pageNumber="31" type="comments">
<paragraph id="FB124EE023BB1917EB95417B711CE008" pageId="30" pageNumber="31">Comments.</paragraph>
<paragraph id="245440AA4AE628843765B36371D5F1A5" pageId="30" pageNumber="31">
The relatively large amount of morphological variation observed in
<taxonomicName id="16BE3A04B50332307880DF053385E652" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus salvini" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="salvini">Bolitogyrus salvini</taxonomicName>
indicates that a complex of species is likely to be involved but with the material currently available, a broad concept of
<taxonomicName id="0AB671C1093089F78CB913BD576EF3BB" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus salvini" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="salvini">Bolitogyrus salvini</taxonomicName>
(wide median lobe in parameral view, elongate female tergite X) is preferred. There are three different morphotypes discernable from the material at hand that will need re-assessment in the future. The male and female types (Guatemala) represent morphotype 1: reddish elytra; pale apex of the abdomen; weak median frontal impression; without impunctate area on dorsal surface of the head. Morphotype 2 is represented by four females (southern Mexico, Nicaragua): dark elytra with green to greenish bronze metallic reflections; pale apex of abdomen; weak median frontal impression; with impunctate area on dorsal surface of head. Morphotype 3 is represented by two males and one female (Mexico, north of second morphotype): dark elytra with bronze metallic reflection; dark apex of abdomen; moderately deep median frontal impression; impunctate area on dorsal surface of head. The first and third morphotypes have males available and can be distinguished (at present) by the peg setae on the paramere: morphotype 1 with distinctly fewer peg setae (therefore sparser) than morphotype 3 (Fig. 16F versus 16
<normalizedToken id="AE598786AC07A64801223560CA7A6DB7" originalValue="GH">G-H</normalizedToken>
).
</paragraph>
</subSubSection>
</treatment>
</document>