289 lines
32 KiB
XML
289 lines
32 KiB
XML
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<mods:title id="F72D272E0B87A0142B714D6F685EF2CA">The masked water snakes of the genus Homalopsis Kuhl & van Hasselt, 1822 (Squamata, Serpentes, Homalopsidae), with the description of a new species</mods:title>
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<treatment id="C2337B59FFEAFFE5DC91471BFB19B105" ID-DOI="http://doi.org/10.5281/zenodo.6169959" ID-GBIF-Taxon="119509490" ID-Zenodo-Dep="6169959" LSID="urn:lsid:plazi:treatment:C2337B59FFEAFFE5DC91471BFB19B105" httpUri="http://treatment.plazi.org/id/C2337B59FFEAFFE5DC91471BFB19B105" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16">
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<paragraph id="4A25CA4FFFEAFFFBDC91471BFD91B2E4" blockId="15.[151,629,619,678]" box="[151,629,619,646]" pageId="15" pageNumber="16">
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<heading id="116D7D23FFEAFFFBDC91471BFD91B2E4" bold="true" box="[151,629,619,646]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
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<taxonomicName id="8D9AB1CCFFEAFFFBDC91471BFD91B2E4" authority="Deuve, 1970" authorityName="Deuve" authorityYear="1970" box="[151,629,619,646]" class="Reptilia" family="Homalopsidae" genus="Homalopsis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="nigroventralis">
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<emphasis id="78EE165DFFEAFFFBDC91471BFD91B2E4" bold="true" box="[151,629,619,646]" pageId="15" pageNumber="16">
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<emphasis id="78EE165DFFEAFFFBDC91471BFE32B2E7" bold="true" box="[151,470,620,646]" italics="true" pageId="15" pageNumber="16">Homalopsis nigroventralis</emphasis>
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<bibRefCitation id="2E0BB7BEFFEAFFFBDDD8471CFD91B2E4" author="Deuve" box="[478,629,619,645]" pageId="15" pageNumber="16" refString="Deuve, J. (1970) Serpents du Laos. Memoire Office de Recherche Scientifique et Technologique Outre Mer, Paris, 39, 1 - 251." type="journal article" year="1970">Deuve, 1970</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="4A25CA4FFFEAFFFBDC9147F9FED4B2C7" blockId="15.[151,629,619,678]" box="[151,304,654,678]" pageId="15" pageNumber="16">
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<figureCitation id="D2A1D6CAFFEAFFFBDC9147F9FF39B2C7" box="[151,221,654,678]" captionStart="FIGURE 2" captionStartId="5.[151,250,1861,1884]" captionTargetBox="[330,1281,207,1818]" captionTargetId="figure@5.[284,1300,195,1825]" captionTargetPageId="5" captionText="FIGURE 2. Plots of the principal components (PC) scores on the first two PC axes derived from 11 morphological variables (dorsal scale row counts on the forebody, midbody, posterior body; number of prefrontals, preoculars, postoculars, upper labials; upper labials under orbit; upper labials at loreal; loreal condition; ventral pattern) for 95 specimens of Homalopsis. The square with the central circle is H. hardwickii which associates with H. buccata in this analysis. Plot A compares H. buccata, H. mereljcoxi, and H. hardwickii; Plot B compares H. buccata, H. mereljcoxi, H. hardwickii, and H. semizonata; Plot C compares all five species: H. buccata, H. mereljcoxi, H. hardwickii, H. semizonata, and H. nigroventralis." httpUri="https://zenodo.org/record/209955/files/figure.png" pageId="15" pageNumber="16">Figs 2</figureCitation>
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d, 8d, 9
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</paragraph>
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</subSubSection>
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<subSubSection id="028099C4FFEAFFFBDC9147A0FD69B329" pageId="15" pageNumber="16" type="reference_group">
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<paragraph id="4A25CA4FFFEAFFFBDC9147A0FE13B34B" blockId="15.[151,1436,726,840]" pageId="15" pageNumber="16">
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<treatmentCitationGroup id="6A8AED61FFEAFFFBDC9147A0FE13B34B" pageId="15" pageNumber="16">
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<taxonomicName id="8D9AB1CCFFEAFFFBDC9147A0FE82B28C" box="[151,358,727,749]" class="Reptilia" family="Homalopsidae" genus="Homalopsis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="buccata">
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<emphasis id="78EE165DFFEAFFFBDC9147A0FE82B28C" box="[151,358,727,749]" italics="true" pageId="15" pageNumber="16">Homalopsis buccata</emphasis>
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</taxonomicName>
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—
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<treatmentCitation id="CB3BEC5EFFEAFFFBDD8647A1FD76B28D" author="Taylor" box="[384,658,726,748]" page="1159" pageId="15" pageNumber="16" year="1958">
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<bibRefCitation id="2E0BB7BEFFEAFFFBDD8647A1FDB7B28D" author="Taylor" box="[384,595,726,748]" pageId="15" pageNumber="16" refString="Taylor, E. H. & Elbel, R. E. (1958) Contributions to the herpetology of Thailand. University of Kansas Science Bulletin, 38, 1033 - 1189." type="journal article" year="1958">Taylor & Elbel 1958</bibRefCitation>
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: 1159
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</treatmentCitation>
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;
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<treatmentCitation id="CB3BEC5EFFEAFFFBDE9A47A1FCA9B28D" author="Taylor" box="[668,845,726,748]" page="923" pageId="15" pageNumber="16" year="1965">
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<bibRefCitation id="2E0BB7BEFFEAFFFBDE9A47A1FCFEB28D" author="Taylor" box="[668,794,726,748]" pageId="15" pageNumber="16" refString="Taylor, E. H. (1965) The serpents of Thailand and adjacent waters. University of Kansas Science Bulletin, 45, 609 - 1096." type="journal article" year="1965">Taylor 1965</bibRefCitation>
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: 923
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</treatmentCitation>
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;
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<treatmentCitation id="CB3BEC5EFFEAFFFBDF5E47A1FC09B28D" author="Gyi" box="[856,1005,726,748]" page="136" pageId="15" pageNumber="16" year="1970">
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<bibRefCitation id="2E0BB7BEFFEAFFFBDF5E47A1FC5FB28D" author="Gyi" box="[856,955,726,748]" pageId="15" pageNumber="16" refString="Gyi, K. K. (1970) A revision of colubrid snakes of the subfamily Homalopsinae. University of Kansas Publications, Museum of Natural History, 20, 47 - 223." type="journal article" year="1970">Gyi 1970</bibRefCitation>
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: 136
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</treatmentCitation>
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.
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<taxonomicName id="8D9AB1CCFFEAFFFBDC914782FD61B36B" authority="Deuve, 1970" authorityName="Deuve" authorityYear="1970" box="[151,645,757,779]" class="Reptilia" family="Homalopsidae" genus="Homalopsis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="subSpecies" species="buccata" subSpecies="nigroventralis">
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<emphasis id="78EE165DFFEAFFFBDC914782FE1FB36A" box="[151,507,757,779]" italics="true" pageId="15" pageNumber="16">Homalopsis buccata nigroventralis</emphasis>
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<bibRefCitation id="2E0BB7BEFFEAFFFBDE064781FD61B36B" author="Deuve" box="[512,645,757,779]" pageId="15" pageNumber="16" refString="Deuve, J. (1970) Serpents du Laos. Memoire Office de Recherche Scientifique et Technologique Outre Mer, Paris, 39, 1 - 251." type="journal article" year="1970">Deuve, 1970</bibRefCitation>
|
||
</taxonomicName>
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||
.
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<typeStatus id="952174EDFFEAFFFBDE884781FD24B36A" box="[654,704,758,779]" pageId="15" pageNumber="16">Type</typeStatus>
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Specimen: none, Deuve’s specimens presumed lost in a fire.
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<typeStatus id="952174EDFFEAFFFBD92C4781FAB8B36A" box="[1322,1372,758,779]" pageId="15" pageNumber="16">Type</typeStatus>
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locality: Ngum River Valley,
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<collectingCountry id="328D8ADFFFEAFFFBDDB94662FE10B34B" box="[447,500,789,810]" name="Laos" pageId="15" pageNumber="16">Laos</collectingCountry>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="4A25CA4FFFEAFFFBDC914645FD69B329" blockId="15.[151,1436,726,840]" box="[151,653,818,840]" pageId="15" pageNumber="16">
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<treatmentCitationGroup id="6A8AED61FFEAFFFBDC914645FD69B329" box="[151,653,818,840]" pageId="15" pageNumber="16">
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<taxonomicName id="8D9AB1CCFFEAFFFBDC914645FD6DB329" authority="Stuart et al. 2006:149" authorityName="Stuart et al." authorityPageNumber="149" authorityYear="2006" box="[151,649,818,840]" class="Reptilia" family="Homalopsidae" genus="Homalopsis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="nigroventralis">
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<emphasis id="78EE165DFFEAFFFBDC914645FE40B329" box="[151,420,818,840]" italics="true" pageId="15" pageNumber="16">Homalopsis nigroventralis</emphasis>
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||
<treatmentCitation id="CB3BEC5EFFEAFFFBDDAD4644FD6DB329" author="Stuart" box="[427,649,818,840]" page="149" pageId="15" pageNumber="16" year="2006">
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<bibRefCitation id="2E0BB7BEFFEAFFFBDDAD4644FDBAB329" author="Stuart" box="[427,606,818,840]" pageId="15" pageNumber="16" refString="Stuart, B. L., Sok, K. & Neang, T. (2006) A collection of amphibians and reptiles from hilly eastern Cambodia. Raffles Bulletin of Zoology, 5, 129 - 155." type="journal article" year="2006">
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Stuart
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<emphasis id="78EE165DFFEAFFFBDDEB4644FDFEB329" box="[493,538,818,840]" italics="true" pageId="15" pageNumber="16">et al</emphasis>
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. 2006
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</bibRefCitation>
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:149
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="028099C4FFEAFFFBDC914602FECCB4E8" pageId="15" pageNumber="16" type="discussion">
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<paragraph id="4A25CA4FFFEAFFFBDC914602FECCB4E8" blockId="15.[151,1437,885,1845]" pageId="15" pageNumber="16">
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<emphasis id="78EE165DFFEAFFFBDC914602FEF0B3EF" bold="true" box="[151,276,885,910]" pageId="15" pageNumber="16">Comment.</emphasis>
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<bibRefCitation id="2E0BB7BEFFEAFFFBDD1D4601FE51B3EC" author="Deuve" box="[283,437,885,909]" pageId="15" pageNumber="16" refString="Deuve, J. (1970) Serpents du Laos. Memoire Office de Recherche Scientifique et Technologique Outre Mer, Paris, 39, 1 - 251." type="journal article" year="1970">Deuve (1970)</bibRefCitation>
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described this snake from the Ngum River Valley,
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<collectingCountry id="328D8ADFFFEAFFFBDFE94601FBC1B3EC" box="[1007,1061,886,909]" name="Laos" pageId="15" pageNumber="16">Laos</collectingCountry>
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and reported 35–39 scale rows at midbody, 152–164 ventrals and 75–94 subcaudals. The specimens examined here fall within these parameters, except for the subcaudal counts.
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<taxonomicName id="8D9AB1CCFFEAFFFBDE1746CAFCD9B3B4" box="[529,829,957,981]" class="Reptilia" family="Homalopsidae" genus="Homalopsis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="nigroventralis">
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<emphasis id="78EE165DFFEAFFFBDE1746CAFCD9B3B4" box="[529,829,957,981]" italics="true" pageId="15" pageNumber="16">Homalopsis nigroventralis</emphasis>
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</taxonomicName>
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may use a different habitat than other
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<taxonomicName id="8D9AB1CCFFEAFFFBD8F946CAFA61B3B4" box="[1279,1413,957,981]" class="Reptilia" family="Homalopsidae" genus="Homalopsis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
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<emphasis id="78EE165DFFEAFFFBD8F946CAFA61B3B4" box="[1279,1413,957,981]" italics="true" pageId="15" pageNumber="16">Homalopsis</emphasis>
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</taxonomicName>
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– streams with a moderate current, and sand and rock substrate as opposed to slow moving or stagnant water with mud substrates. Because we have not examined specimens from
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<collectingCountry id="328D8ADFFFEAFFFBDF9B4171FC37B47C" box="[925,979,1030,1053]" name="Laos" pageId="15" pageNumber="16">Laos</collectingCountry>
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we have not established a
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<typeStatus id="952174EDFFEAFFFBD9264171FA9CB47F" box="[1312,1400,1030,1054]" pageId="15" pageNumber="16" type="neotype">neotype</typeStatus>
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or restricted the
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<typeStatus id="952174EDFFEAFFFBDD32415DFE80B423" box="[308,356,1066,1090]" pageId="15" pageNumber="16">type</typeStatus>
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locality. Photographs of this snake can be found in
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<bibRefCitation id="2E0BB7BEFFEAFFFBDFB8415EFB5DB420" author="Taylor" box="[958,1209,1065,1089]" pageId="15" pageNumber="16" refString="Taylor, E. H. & Elbel, R. E. (1958) Contributions to the herpetology of Thailand. University of Kansas Science Bulletin, 38, 1033 - 1189." type="journal article" year="1958">Taylor & Elbel (1958</bibRefCitation>
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||
: 1066) and
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<bibRefCitation id="2E0BB7BEFFEAFFFBD944415EFEB4B404" author="Murphy" pageId="15" pageNumber="16" refString="Murphy, J. C. (2007) Homalopsid Snakes, Evolution in the Mud. Krieger Publishing Company, Malabar. 249 pp." type="book" year="2007">Murphy (2007: 201-203)</bibRefCitation>
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. Field observations on this snake can be found in
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<bibRefCitation id="2E0BB7BEFFEAFFFBDF784139FBF8B404" author="Deuve" box="[894,1052,1101,1125]" pageId="15" pageNumber="16" refString="Deuve, J. (1970) Serpents du Laos. Memoire Office de Recherche Scientifique et Technologique Outre Mer, Paris, 39, 1 - 251." type="journal article" year="1970">Deuve (1970)</bibRefCitation>
|
||
,
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||
<bibRefCitation id="2E0BB7BEFFEAFFFBD82E413AFB19B404" box="[1064,1277,1101,1125]" pageId="15" pageNumber="16" refString="Stuart, B. L., Sok, K. & Neang, T. (2006) A collection of amphibians and reptiles from hilly eastern Cambodia. Raffles Bulletin of Zoology, 5, 129 - 155." type="journal article">
|
||
Stuart
|
||
<emphasis id="78EE165DFFEAFFFBD8774139FB4DB404" box="[1137,1193,1101,1125]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
|
||
(2006)
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||
</bibRefCitation>
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||
, and
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||
<bibRefCitation id="2E0BB7BEFFEAFFFBD93E413AFEC0B4E8" pageId="15" pageNumber="16" refString="Bezuijen, M. R. Vinn, B. & Send, L. (2009) A collection of amphibians and reptiles from the Mekong River, north-eastern Cambodia. Hamadryad, 34, 135 - 164." type="journal article">
|
||
Bezuijen
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||
<emphasis id="78EE165DFFEAFFFBDC914105FF2CB4E8" box="[151,200,1137,1161]" italics="true" pageId="15" pageNumber="16">et al</emphasis>
|
||
. (2009)
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||
</bibRefCitation>
|
||
.
|
||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="028099C4FFEAFFFBDCCF41E2FD4FB590" pageId="15" pageNumber="16" type="diagnosis">
|
||
<paragraph id="4A25CA4FFFEAFFFBDCCF41E2FD4FB590" blockId="15.[151,1437,885,1845]" pageId="15" pageNumber="16">
|
||
<emphasis id="78EE165DFFEAFFFBDCCF41E2FEA7B4CF" bold="true" box="[201,323,1173,1198]" pageId="15" pageNumber="16">Diagnosis.</emphasis>
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||
<taxonomicName id="8D9AB1CCFFEAFFFBDD4C41E2FD92B4CC" box="[330,630,1173,1197]" class="Reptilia" family="Homalopsidae" genus="Homalopsis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="nigroventralis">
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<emphasis id="78EE165DFFEAFFFBDD4C41E2FD92B4CC" box="[330,630,1173,1197]" italics="true" pageId="15" pageNumber="16">Homalopsis nigroventralis</emphasis>
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||
</taxonomicName>
|
||
has upper labials 1–3 contacting the loreal; two prefrontals; 35–39 dorsal scale rows at midbody, reduced to 30 or less; one or two postocular scales plus one postsubocular scale; 10–12 upper labials; 159–167 ventrals; and a reverse color pattern on the venter (dark olive-gray with white spots).
|
||
<taxonomicName id="8D9AB1CCFFEAFFFBDC914076FE99B578" box="[151,381,1281,1305]" class="Reptilia" family="Homalopsidae" genus="Homalopsis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="buccata">
|
||
<emphasis id="78EE165DFFEAFFFBDC914076FE99B578" box="[151,381,1281,1305]" italics="true" pageId="15" pageNumber="16">Homalopsis buccata</emphasis>
|
||
</taxonomicName>
|
||
has upper labials 1–4 contacting the loreal; one postocular scale plus a postsubocular scale; a ventral pattern that is light with dark spots.
|
||
<taxonomicName id="8D9AB1CCFFEAFFFBDE864052FC60B55C" box="[640,900,1317,1341]" class="Reptilia" family="Homalopsidae" genus="Homalopsis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="hardwickii">
|
||
<emphasis id="78EE165DFFEAFFFBDE864052FC60B55C" box="[640,900,1317,1341]" italics="true" pageId="15" pageNumber="16">Homalopsis hardwickii</emphasis>
|
||
</taxonomicName>
|
||
has a divided loreal contacting upper labials 1– 4; one postocular scale and no presubocular scale.
|
||
<taxonomicName id="8D9AB1CCFFEAFFFBDED6403EFC3FB500" box="[720,987,1353,1377]" class="Reptilia" family="Homalopsidae" genus="Homalopsis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="semizonata">
|
||
<emphasis id="78EE165DFFEAFFFBDED6403EFC3FB500" box="[720,987,1353,1377]" italics="true" pageId="15" pageNumber="16">Homalopsis semizonata</emphasis>
|
||
</taxonomicName>
|
||
has a divided or fragmented loreal contacting upper labials 1–4 or 1–5; three prefrontals; 39–44 dorsal scale rows at midbody, reduced to more than 30 posteriorly; one postocular and one postsubocular. The new species,
|
||
<taxonomicName id="8D9AB1CCFFEAFFFBDFAA40E5FBA1B5C8" box="[940,1093,1425,1449]" class="Reptilia" family="Homalopsidae" genus="Homalopsis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="mereljcoxi">
|
||
<emphasis id="78EE165DFFEAFFFBDFAA40E5FBA1B5C8" box="[940,1093,1425,1449]" italics="true" pageId="15" pageNumber="16">H. mereljcoxi</emphasis>
|
||
</taxonomicName>
|
||
has a single loreal contacting upper labials 1–4; 40–47 scale rows at midbody, reduced to 30 or more posteriorly; 13 (12–14) upper labials; and ventral counts that are usually greater than 165.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="028099C4FFEAFFFBDCCF408AFCF3B754" pageId="15" pageNumber="16" type="distribution">
|
||
<paragraph id="4A25CA4FFFEAFFFBDCCF408AFCF3B754" blockId="15.[151,1437,885,1845]" pageId="15" pageNumber="16">
|
||
<emphasis id="78EE165DFFEAFFFBDCCF408AFE86B677" bold="true" box="[201,354,1533,1558]" pageId="15" pageNumber="16">Distribution.</emphasis>
|
||
This species is known from the Mekong River in
|
||
<collectingCountry id="328D8ADFFFEAFFFBDF9E4089FC35B674" box="[920,977,1534,1557]" name="Laos" pageId="15" pageNumber="16">Laos</collectingCountry>
|
||
,
|
||
<collectingCountry id="328D8ADFFFEAFFFBDFDB408AFBB7B674" box="[989,1107,1533,1557]" name="Cambodia" pageId="15" pageNumber="16">Cambodia</collectingCountry>
|
||
, and
|
||
<collectingCountry id="328D8ADFFFEAFFFBD889408AFB12B674" box="[1167,1270,1533,1557]" name="Thailand" pageId="15" pageNumber="16">Thailand</collectingCountry>
|
||
.
|
||
<bibRefCitation id="2E0BB7BEFFEAFFFBD9074089FA78B674" author="Deuve" box="[1281,1436,1533,1557]" pageId="15" pageNumber="16" refString="Deuve, J. (1970) Serpents du Laos. Memoire Office de Recherche Scientifique et Technologique Outre Mer, Paris, 39, 1 - 251." type="journal article" year="1970">Deuve (1970)</bibRefCitation>
|
||
reported it in the Ngum River Valley of
|
||
<collectingCountry id="328D8ADFFFEAFFFBDE534355FD6BB658" box="[597,655,1570,1593]" name="Laos" pageId="15" pageNumber="16">Laos</collectingCountry>
|
||
.
|
||
<bibRefCitation id="2E0BB7BEFFEAFFFBDE9F4356FC75B658" author="Taylor" box="[665,913,1569,1593]" pageId="15" pageNumber="16" refString="Taylor, E. H. & Elbel, R. E. (1958) Contributions to the herpetology of Thailand. University of Kansas Science Bulletin, 38, 1033 - 1189." type="journal article" year="1958">Taylor & Elbel (1958)</bibRefCitation>
|
||
describe specimens from Thailand’s Phan Phu Mountain, and
|
||
<bibRefCitation id="2E0BB7BEFFEAFFFBDD444332FDF2B63C" box="[322,534,1605,1629]" pageId="15" pageNumber="16" refString="Stuart, B. L., Sok, K. & Neang, T. (2006) A collection of amphibians and reptiles from hilly eastern Cambodia. Raffles Bulletin of Zoology, 5, 129 - 155." type="journal article">
|
||
Stuart
|
||
<emphasis id="78EE165DFFEAFFFBDD8D4331FE59B63C" box="[395,445,1605,1629]" italics="true" pageId="15" pageNumber="16">et al</emphasis>
|
||
. (2006)
|
||
</bibRefCitation>
|
||
reported it from Siem Pang and Ta Veng in eastern
|
||
<collectingCountry id="328D8ADFFFEAFFFBD85B4332FB34B63C" box="[1117,1232,1605,1629]" name="Cambodia" pageId="15" pageNumber="16">Cambodia</collectingCountry>
|
||
in bamboo mixed with evergreen forest between 100 and
|
||
<quantity id="8D6267AAFFEAFFFBDE53431EFD78B6E0" box="[597,668,1641,1665]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="15" pageNumber="16" unit="m" value="170.0">170 m</quantity>
|
||
ASL. Two specimens were taken during the day, others were found in gill nets. The streams were described as having a moderate current, and a sand and rock substrate; one had a closed canopy, others were open and had grass lined banks. All of these localities are peripheral to the Mekong. However,
|
||
<bibRefCitation id="2E0BB7BEFFEAFFFBDD0F43A2FE18B68C" box="[265,508,1749,1773]" pageId="15" pageNumber="16" refString="Bezuijen, M. R. Vinn, B. & Send, L. (2009) A collection of amphibians and reptiles from the Mekong River, north-eastern Cambodia. Hamadryad, 34, 135 - 164." type="journal article">
|
||
Bezuijen
|
||
<emphasis id="78EE165DFFEAFFFBDD7243A1FE47B68C" box="[372,419,1749,1773]" italics="true" pageId="15" pageNumber="16">et al</emphasis>
|
||
. (2009)
|
||
</bibRefCitation>
|
||
report it from the Mekong River in Kratie Province,
|
||
<collectingCountry id="328D8ADFFFEAFFFBD84E43A2FB5BB68C" box="[1096,1215,1749,1773]" name="Cambodia" pageId="15" pageNumber="16">Cambodia</collectingCountry>
|
||
. They describe four specimens, one in a market, and three others at Koh Kapeung and Koh Khlap Islands, the largest was
|
||
<quantity id="8D6267AAFFEAFFFBD92F438EFA99B770" box="[1321,1405,1785,1809]" metricMagnitude="0" metricUnit="m" metricValue="1.04" pageId="15" pageNumber="16" unit="cm" value="104.0">104 cm</quantity>
|
||
in total length. One of these snakes was on a mud substrate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="1EE59AC7FFF5FFE4DC914214FB72B792" httpUri="https://zenodo.org/record/209962/files/figure.png" pageId="16" pageNumber="17" targetBox="[326,1261,193,1844]" targetPageId="16">
|
||
<paragraph id="4A25CA4FFFF5FFE4DC914214FB72B792" blockId="16.[151,1436,1891,2036]" pageId="16" pageNumber="17">
|
||
<emphasis id="78EE165DFFF5FFE4DC914214FEF0B719" bold="true" box="[151,276,1891,1914]" pageId="16" pageNumber="17">FIGURE 9.</emphasis>
|
||
<taxonomicName id="8D9AB1CCFFF5FFE4DD1A4214FDCFB718" box="[284,555,1891,1913]" class="Reptilia" family="Homalopsidae" genus="Homalopsis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="nigroventralis">
|
||
<emphasis id="78EE165DFFF5FFE4DD1A4214FDCFB718" box="[284,555,1891,1913]" italics="true" pageId="16" pageNumber="17">Homalopsis nigroventralis</emphasis>
|
||
</taxonomicName>
|
||
. Above FMNH 263032 from Cambodia (Virachey National Park, O Kanome stream, 14°15'08.0" N, 106°37'58.8" E, 175 m) profile showing three upper labials at the loreal and that upper labial number 6 is the first horizontally divided labial. Note the dark coloration and the reduced “mask” in this specimen. Below: ventral view of FMNH 263029, also from Cambodia (Virachey National Park, O Lopeung stream, 14°10'39.2" N, 107°17'25.1" E, 150m), shows the reverse ventral pattern with a dark ground color and pale colored spots. Photographs JCM.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="028099C4FFF4FFE5DCCF45E0FB19B105" pageId="17" pageNumber="18" type="materials_examined">
|
||
<paragraph id="4A25CA4FFFF4FFE5DCCF45E0FB19B105" blockId="17.[151,1437,151,356]" pageId="17" pageNumber="18">
|
||
<emphasis id="78EE165DFFF4FFE5DCCF45E0FE59B0D1" bold="true" box="[201,445,151,176]" pageId="17" pageNumber="18">Material Examined.</emphasis>
|
||
<collectingCountry id="328D8ADFFFF4FFE5DDCE45E0FDA2B0CE" box="[456,582,151,175]" name="Cambodia" pageId="17" pageNumber="18">CAMBODIA</collectingCountry>
|
||
– Ta Veng
|
||
<collectionCode id="2C8B528AFFF4FFE5DED045EFFCC3B0CE" LSID="urn:lsid:biocol.org:col:34795" box="[726,807,152,175]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="17" pageNumber="18">FMNH</collectionCode>
|
||
263029 (Virachey National Park, O Lopeung stream,
|
||
<geoCoordinate id="2FAEAC88FFF4FFE5DC9145CBFEDFB0B5" box="[151,315,188,212]" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="1" value="14.177556">14°10'39.2" N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2FAEAC88FFF4FFE5DD4E45CBFE1CB0B5" box="[328,504,188,212]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="1" value="107.290306">107°17'25.1" E</geoCoordinate>
|
||
,
|
||
<quantity id="8D6267AAFFF4FFE5DE0345CBFDB5B0B5" box="[517,593,188,212]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="18" unit="m" value="150.0">150 m</quantity>
|
||
); Siem Pang
|
||
<collectionCode id="2C8B528AFFF4FFE5DEED45CAFCD8B0B5" LSID="urn:lsid:biocol.org:col:34795" box="[747,828,189,212]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="17" pageNumber="18">FMNH</collectionCode>
|
||
263030 (Virachey National Park, O Kanome stream,
|
||
<geoCoordinate id="2FAEAC88FFF4FFE5DC9145A8FEA6B096" box="[151,322,223,247]" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="1" value="14.2260275">14°13'33.7" N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2FAEAC88FFF4FFE5DD5345A8FDEEB096" box="[341,522,223,247]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="1" value="106.60445">106°36'16.0" E</geoCoordinate>
|
||
,
|
||
<quantity id="8D6267AAFFF4FFE5DE1845A8FD94B096" box="[542,624,223,247]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="17" pageNumber="18" unit="m" value="170.0">170 m</quantity>
|
||
),
|
||
<collectionCode id="2C8B528AFFF4FFE5DE8A4597FD39B096" LSID="urn:lsid:biocol.org:col:34795" box="[652,733,224,247]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="17" pageNumber="18">FMNH</collectionCode>
|
||
2630301-32 (Virachey National Park, O Kanome stream,
|
||
<geoCoordinate id="2FAEAC88FFF4FFE5DC914473FEDFB17D" box="[151,315,260,284]" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="1" value="14.252222">14°15'08.0" N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2FAEAC88FFF4FFE5DD414473FE12B17D" box="[327,502,260,284]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="1" value="106.633">106°37'58.8" E</geoCoordinate>
|
||
,
|
||
<quantity id="8D6267AAFFF4FFE5DE044473FDA9B17D" box="[514,589,260,284]" metricMagnitude="2" metricUnit="m" metricValue="1.75" pageId="17" pageNumber="18" unit="m" value="175.0">175 m</quantity>
|
||
),
|
||
<collectionCode id="2C8B528AFFF4FFE5DE644472FD57B17D" LSID="urn:lsid:biocol.org:col:34795" box="[610,691,261,284]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="17" pageNumber="18">FMNH</collectionCode>
|
||
263033 (Virachey National Park,
|
||
<geoCoordinate id="2FAEAC88FFF4FFE5D83F4473FB39B17D" box="[1081,1245,260,284]" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="1" value="14.214944">14°12'53.8" N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2FAEAC88FFF4FFE5D8EF4473FA73B17D" box="[1257,1431,260,284]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="1" value="106.59759">106°35'51.3" E</geoCoordinate>
|
||
,
|
||
<quantity id="8D6267AAFFF4FFE5DC914450FF0FB15E" box="[151,235,295,319]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="18" unit="m" value="100.0">100 m</quantity>
|
||
); Nong Khai
|
||
<collectionCode id="2C8B528AFFF4FFE5DD9A445FFE09B15E" LSID="urn:lsid:biocol.org:col:34795" box="[412,493,296,319]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="17" pageNumber="18">FMNH</collectionCode>
|
||
265801 (Phu Wua Wildlife Sanctuary, Houay Sod waterfall,
|
||
<geoCoordinate id="2FAEAC88FFF4FFE5D8EA4450FA7CB15E" box="[1260,1432,295,319]" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="1" value="18.264278">18°15'51.4" N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2FAEAC88FFF4FFE5DC91443BFEA0B105" box="[151,324,332,356]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="1" value="103.90614">103°54'22.1" E</geoCoordinate>
|
||
,
|
||
<quantity id="8D6267AAFFF4FFE5DD56443BFE7DB105" box="[336,409,332,356]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="17" pageNumber="18" unit="m" value="160.0">160 m</quantity>
|
||
),
|
||
<collectionCode id="2C8B528AFFF4FFE5DDA8443AFDE0B105" LSID="urn:lsid:biocol.org:col:34988" box="[430,516,333,356]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="17" pageNumber="18">MNHN</collectionCode>
|
||
1988.2104;
|
||
<collectingCountry id="328D8ADFFFF4FFE5DE96443BFCE3B105" box="[656,775,332,356]" name="Thailand" pageId="17" pageNumber="18">THAILAND</collectingCountry>
|
||
– Phan Phu Mt.
|
||
<collectionCode id="2C8B528AFFF4FFE5DFC5443AFBF0B105" LSID="urn:lsid:biocol.org:col:34795" box="[963,1044,333,356]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="17" pageNumber="18">FMNH</collectionCode>
|
||
179239, 117858-60.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |