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<document ID-DOI="http://dx.doi.org/10.3897/zse.95.38259" ID-GBIF-Dataset="b0ddb79b-fd6d-499f-8626-80a64b39fb06" ID-GBIF-Taxon="159412604" ID-Pensoft-Pub="1860-0743-2-453" ID-Pensoft-UUID="D49EC3D9F8BA54B4AFED827B67E83634" ID-ZooBank="EE9B3C6BD4AE4508BF0E5CCEEB7F12EA" ModsDocID="1860-0743-2-453" checkinTime="1568292421383" checkinUser="pensoft" docAuthor="Salvador, Rodrigo B. &amp; Cavallari, Daniel C." docDate="2019" docId="BC12D4D252AE5CAC9FD27DB8CC16FB0F" docLanguage="en" docName="ZoosystEvol 95(2): 453-463" docOrigin="Zoosystematics and Evolution 95 (2)" docSource="http://dx.doi.org/10.3897/zse.95.38259" docTitle="Hyperaulax ridleyi Smith 1890" docType="treatment" docVersion="3" id="D49EC3D9F8BA54B4AFED827B67E83634" lastPageNumber="456" masterDocId="D49EC3D9F8BA54B4AFED827B67E83634" masterDocTitle="Taxonomic revision of the genus Hyperaulax Pilsbry, 1897 (Gastropoda, Stylommatophora, Odontostomidae)" masterLastPageNumber="463" masterPageNumber="453" pageId="3" pageNumber="456" updateTime="1643459391262" updateUser="ExternalLinkService">
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<mods:title>Taxonomic revision of the genus Hyperaulax Pilsbry, 1897 (Gastropoda, Stylommatophora, Odontostomidae)</mods:title>
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<mods:namePart>Salvador, Rodrigo B.</mods:namePart>
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<mods:namePart>Cavallari, Daniel C.</mods:namePart>
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<treatment ID-GBIF-Taxon="159412604" LSID="urn:lsid:plazi:treatment:BC12D4D252AE5CAC9FD27DB8CC16FB0F" httpUri="http://treatment.plazi.org/id/BC12D4D252AE5CAC9FD27DB8CC16FB0F" lastPageNumber="456" pageId="3" pageNumber="456">
<subSubSection pageId="3" pageNumber="456" type="nomenclature">
<paragraph pageId="3" pageNumber="456">
<taxonomicName LSID="BC12D4D252AE5CAC9FD27DB8CC16FB0F" authority="Smith, 1890" authorityName="Smith" authorityYear="1890" class="Gastropoda" family="Odontostomidae" genus="Hyperaulax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyperaulax ridleyi" order="Stylommatophora" pageId="3" pageNumber="456" phylum="Mollusca" rank="species" species="ridleyi">Hyperaulax ridleyi (Smith, 1890)</taxonomicName>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Hyperaulax ridleyi. A-E. Lectotype, NHMUK 1888.6.27.106. F-I. Paralectotypes 1 to 4 (in order), NHMUK 1888.6.27.107 - 110. J. Extreme form, large and with thick callus, NHMUK 1888.6.27.88 - 94. K. Weathered specimen, ZMS unnumbered (ex Heimburg colln.). L. Protoconch detail (same specimen from K)." figureDoi="10.3897/zse.95.38259.figure2" httpUri="https://binary.pensoft.net/fig/339175" pageId="3" pageNumber="456" tableDoi="10.3897/zse.95.38259.figure2">Fig. 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="456" type="reference_group">
<paragraph pageId="3" pageNumber="456">
<taxonomicName authority="Scopoli, 1777" authorityName="Scopoli" authorityYear="1777" class="Gastropoda" family="Odontostomidae" genus="Bulimus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Bulimus (Bulimulus) Ridleyi" order="Stylommatophora" pageId="3" pageNumber="456" phylum="Mollusca" rank="species" species="Ridleyi" subGenus="Bulimulus">Bulimus (Bulimulus) Ridleyi</taxonomicName>
E.A.
<bibRefCitation author="Smith, EA" journalOrPublisher="Journal of the Linnean Society of London" pageId="8" pageNumber="461" pagination="483 - 503" refId="B39" refString="Smith, EA, 1890. Mollusca. Journal of the Linnean Society of London 20: 483 - 503" title="Mollusca." volume="20" year="1890">Smith 1890</bibRefCitation>
: 501, pl. 30, fig. 9;
<bibRefCitation author="Dall, WH" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="7" pageNumber="460" pagination="395 - 460" refId="B10" refString="Dall, WH, 1896. Insular land shell faunas, especially as illustrated by the data obtained by Dr. G. Baur in the Galapagos Islands. Proceedings of the Academy of Natural Sciences of Philadelphia 48: 395 - 460" title="Insular land shell faunas, especially as illustrated by the data obtained by Dr. G. Baur in the Galapagos Islands." volume="48" year="1896">Dall 1896</bibRefCitation>
: 415.
</paragraph>
<paragraph pageId="3" pageNumber="456">
<taxonomicName class="Gastropoda" family="Odontostomidae" genus="Bulimulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bulimulus (Hyperaulax) ridleyi" order="Stylommatophora" pageId="3" pageNumber="456" phylum="Mollusca" rank="species" species="ridleyi" subGenus="Hyperaulax">Bulimulus (Hyperaulax) ridleyi</taxonomicName>
:
<bibRefCitation pageId="3" pageNumber="456" refId="B27">Pilsbry 1897a</bibRefCitation>
: 10;
<bibRefCitation pageId="3" pageNumber="456" refId="B28">Pilsbry 1897b</bibRefCitation>
: 82, pl. 14, figs 11-13.
</paragraph>
<paragraph pageId="3" pageNumber="456">
<taxonomicName class="Gastropoda" family="Odontostomidae" genus="Buliminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buliminus ridleyi" order="Stylommatophora" pageId="3" pageNumber="456" phylum="Mollusca" rank="species" species="ridleyi">Buliminus ridleyi</taxonomicName>
:
<bibRefCitation author="Moellendorff, O von" journalOrPublisher="Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft" pageId="8" pageNumber="461" pagination="125 - 127" refId="B21" refString="Moellendorff, O von, 1901. Zur Suedpolar-Land-Frage. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 33: 125 - 127" title="Zur Suedpolar-Land-Frage." volume="33" year="1901">
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
1901
</bibRefCitation>
: 126.
</paragraph>
<paragraph pageId="3" pageNumber="456">
<taxonomicName class="Gastropoda" family="Odontostomidae" genus="Hyperaulax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyperaulax ridleyi" order="Stylommatophora" pageId="3" pageNumber="456" phylum="Mollusca" rank="species" species="ridleyi">Hyperaulax ridleyi</taxonomicName>
:
<bibRefCitation author="Pilsbry, HA" journalOrPublisher="Academy of Natural Sciences of Philadelphia, Philadelphia" pageId="8" pageNumber="461" refId="B29" refString="Pilsbry, HA, 1901. Manual of conchology, second series. Pulmonata. Vol. 14. Oriental bulimoid Helicidae; Odontostomidae; Cerionidae. Academy of Natural Sciences of Philadelphia, Philadelphia" title="Manual of conchology, second series. Pulmonata. Vol. 14. Oriental bulimoid Helicidae; Odontostomidae; Cerionidae." year="1901">Pilsbry 1901</bibRefCitation>
: 103;
<bibRefCitation author="Ihering, H von" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="7" pageNumber="460" pagination="183 - 193" refId="B15" refString="Ihering, H von, 1923. Transgression des Meeres waehrend der Ablagerung der Pampas. Archiv fuer Molluskenkunde 55: 183 - 193" title="Transgression des Meeres waehrend der Ablagerung der Pampas." volume="55" year="1923">Ihering 1923</bibRefCitation>
: 191;
<bibRefCitation pageId="3" pageNumber="456" refId="B41">Wenz 1923</bibRefCitation>
: 729;
<bibRefCitation author="Schileyko, AA" journalOrPublisher="Ruthenica, Supplement" pageId="8" pageNumber="461" pagination="263 - 436" refId="B37" refString="Schileyko, AA, 1999. Treatise on Recent terrestrial pulmonate molluscs. Part 3. Partulidae, Aillyidae, Bulimulidae, Orthalicidae, Megaspiridae, Urocoptidae. Ruthenica, Supplement 2: 263 - 436" title="Treatise on Recent terrestrial pulmonate molluscs. Part 3. Partulidae, Aillyidae, Bulimulidae, Orthalicidae, Megaspiridae, Urocoptidae." volume="2" year="1999">Schileyko 1999</bibRefCitation>
: 321, fig. 396;
<bibRefCitation pageId="3" pageNumber="456" refId="B33">Salgado and Coelho 2003</bibRefCitation>
: 165;
<bibRefCitation author="Simone, LRL" journalOrPublisher="EGB / FAPESP, Sao Paulo" pageId="8" pageNumber="461" refId="B38" refString="Simone, LRL, 2006. Land and freshwater mollusks of Brazil. EGB/FAPESP, Sao Paulo" title="Land and freshwater mollusks of Brazil." year="2006">Simone 2006</bibRefCitation>
: 178, fig. 637;
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.182.2720" author="Breure, ASH" journalOrPublisher="ZooKeys" pageId="7" pageNumber="460" pagination="1 - 70" refId="B5" refString="Breure, ASH, Ablett, JD, 2012. Annotated type catalogue of the Bothriembryontidae and Odontostomidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London. ZooKeys 182: 1 - 70" title="Annotated type catalogue of the Bothriembryontidae and Odontostomidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London." url="https://doi.org/10.3897/zookeys.182.2720" volume="182" year="2012">Breure and Ablett 2012</bibRefCitation>
: 36, figs 20C, D, 20ii;
<bibRefCitation author="Salvador, RB" journalOrPublisher="Tuhinga" pageId="8" pageNumber="461" pagination="82 - 98" refId="B34" refString="Salvador, RB, 2019. Brazilian, Uruguayan and Argentinian land snails in the collection of the Museum of New Zealand Te Papa Tongarewa. Tuhinga 30: 82 - 98" title="Brazilian, Uruguayan and Argentinian land snails in the collection of the Museum of New Zealand Te Papa Tongarewa." volume="30" year="2019">Salvador 2019</bibRefCitation>
: 87.
</paragraph>
<paragraph pageId="3" pageNumber="456">
<taxonomicName class="Gastropoda" family="Odontostomidae" genus="Hyperaulax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyperaulax (Hyperaulax) ridleyi" order="Stylommatophora" pageId="3" pageNumber="456" phylum="Mollusca" rank="species" species="ridleyi" subGenus="Hyperaulax">Hyperaulax (Hyperaulax) ridleyi</taxonomicName>
:
<bibRefCitation author="Thiele, J" journalOrPublisher="Gustav Fisher Verlag, Jena" pageId="8" pageNumber="461" refId="B40" refString="Thiele, J, 1931. Handbuch der systematischen Weichtierkunde. Erster Band, Teil 2. Gustav Fisher Verlag, Jena" title="Handbuch der systematischen Weichtierkunde. Erster Band, Teil 2." year="1931">Thiele 1931</bibRefCitation>
: 661;
<bibRefCitation author="Morretes, FL" journalOrPublisher="Arquivos do Museu Paranaense" pageId="8" pageNumber="461" pagination="1 - 216" refId="B22" refString="Morretes, FL, 1949. Ensaio de catalogo dos moluscos do Brasil. Arquivos do Museu Paranaense 7: 1 - 216" title="Ensaio de catalogo dos moluscos do Brasil." volume="7" year="1949">Morretes 1949</bibRefCitation>
: 153;
<bibRefCitation pageId="3" pageNumber="456" refId="B42">Zilch 1960</bibRefCitation>
: 505, fig. 1771;
<bibRefCitation author="Breure, ASH" journalOrPublisher="Basteria" pageId="7" pageNumber="460" pagination="109 - 127" refId="B3" refString="Breure, ASH, 1974. Catalogue of Bulimulidae (Gastropoda, Euthyneura), II. Odontostominae. Basteria 38: 109 - 127" title="Catalogue of Bulimulidae (Gastropoda, Euthyneura), II. Odontostominae." volume="38" year="1974">Breure 1974</bibRefCitation>
: 51;
<bibRefCitation author="Breure, ASH" journalOrPublisher="Zoologische Verhandelingen" pageId="7" pageNumber="460" pagination="1 - 98" refId="B7" refString="Breure, ASH, Schouten, JR, 1985. Notes on and descriptions of Bulimulidae (Mollusca, Gastropoda). III. Zoologische Verhandelingen 216: 1 - 98" title="Notes on and descriptions of Bulimulidae (Mollusca, Gastropoda). III." volume="216" year="1985">Breure and Schouten 1985</bibRefCitation>
: 4.
</paragraph>
<paragraph pageId="3" pageNumber="456">
<taxonomicName class="Gastropoda" family="Odontostomidae" genus="Hyperaulax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyperaulax ridleyi" order="Stylommatophora" pageId="3" pageNumber="456" phylum="Mollusca" rank="species" sensu="(s. str.)" species="ridleyi">Hyperaulax (s. str.) ridleyi</taxonomicName>
:
<bibRefCitation DOI="https://doi.org/10.1590/S0373-55241955000100005" author="Lopes, HS" journalOrPublisher="Boletim do Instituto Oceanografico" pageId="7" pageNumber="460" pagination="157 - 196" refId="B19" refString="Lopes, HS, Alvarenga, M, 1955. Contribuicao ao conhecimento dos moluscos da Ilha Fernando de Noronha - Brasil. Boletim do Instituto Oceanografico 6: 157 - 196" title="Contribuicao ao conhecimento dos moluscos da Ilha Fernando de Noronha - Brasil." url="https://doi.org/10.1590/S0373-55241955000100005" volume="6" year="1955">Lopes and Alvarenga 1955</bibRefCitation>
: 181.
</paragraph>
<paragraph pageId="3" pageNumber="456">
<taxonomicName authority="Scopoli, 1777" authorityName="Scopoli" authorityYear="1777" class="Gastropoda" family="Odontostomidae" genus="Bulimus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Bulimus (Bulimulus) ridleyi" order="Stylommatophora" pageId="3" pageNumber="456" phylum="Mollusca" rank="species" species="ridleyi" subGenus="Bulimulus">Bulimus (Bulimulus) ridleyi</taxonomicName>
:
<bibRefCitation author="Oliveira, MP" journalOrPublisher="Boletim do Instituto de Ciencias Biologicas e de Geociencias" pageId="8" pageNumber="461" pagination="1 - 46" refId="B23" refString="Oliveira, MP, Oliveira, MHR, 1984. Comunicacoes malacologicas n° 16. Listagem de tipos de GastropodaPulmonata brasileiros depositados em quatro museus europeus. Boletim do Instituto de Ciencias Biologicas e de Geociencias 38: 1 - 46" title="Comunicacoes malacologicas n ° 16. Listagem de tipos de GastropodaPulmonata brasileiros depositados em quatro museus europeus." volume="38" year="1984">Oliveira and Oliveira 1984</bibRefCitation>
: 19.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="456" type="type locality">
<paragraph pageId="3" pageNumber="456">Type locality.</paragraph>
<paragraph pageId="3" pageNumber="456">
Fernando de Noronha Archipelago, Fernando de Noronha Island and Rata Island. Original (
<bibRefCitation author="Smith, EA" journalOrPublisher="Journal of the Linnean Society of London" pageId="8" pageNumber="461" pagination="483 - 503" refId="B39" refString="Smith, EA, 1890. Mollusca. Journal of the Linnean Society of London 20: 483 - 503" title="Mollusca." volume="20" year="1890">Smith 1890</bibRefCitation>
: 501): &quot;Living under bark of Mango-trees in the garden and on north side of island; also at base of the Peak, north side, under stones, and on Rat Island.&quot;
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="456" type="distribution">
<paragraph pageId="3" pageNumber="456">Distribution.</paragraph>
<paragraph pageId="3" pageNumber="456">Known only from Fernando de Noronha Archipelago.</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="456" type="type material">
<paragraph pageId="3" pageNumber="456">Type material.</paragraph>
<paragraph pageId="3" pageNumber="456">
Lectotype NHMUK 1888.6.27.106 (designation by
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.182.2720" author="Breure, ASH" journalOrPublisher="ZooKeys" pageId="7" pageNumber="460" pagination="1 - 70" refId="B5" refString="Breure, ASH, Ablett, JD, 2012. Annotated type catalogue of the Bothriembryontidae and Odontostomidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London. ZooKeys 182: 1 - 70" title="Annotated type catalogue of the Bothriembryontidae and Odontostomidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London." url="https://doi.org/10.3897/zookeys.182.2720" volume="182" year="2012">Breure and Ablett 2012</bibRefCitation>
). Paralectotypes: NHMUK 1888.6.27.107-110, 4 shells.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="456" type="materials_examined">
<paragraph pageId="3" pageNumber="456">Material examined.</paragraph>
<paragraph pageId="3" pageNumber="456">
Types.
<emphasis bold="true" pageId="3" pageNumber="456">BRAZIL</emphasis>
: Fernando de Noronha: ANSP 71271, 7 shells, H.H. Smith leg., 1896; MNZ 205835, 4 shells, ex Suter coll. 5637; ANSP 81426, 1 shell; ANSP 100530, 4 shells, H.v. Ihering leg., 1910; ANSP 220399, 2 shells, ex B.R.
<taxonomicName genus="Bales" lsidName="Bales" pageId="3" pageNumber="456" rank="genus">Bales</taxonomicName>
coll., J.S. Schwengel leg., 1958; MNZ 205835,4 shells, ex Suter coll. 5637; MZSP 501, 14 shells, 1900; MZSP 7752, 20 shells. MZSP 30134, 10 shells, dunes of Praia das Caieiras, E.F. Nonato leg., 25/vii/1955; MZSP 30135, 8 shells, de Fiore leg., 1983; MZSP 31064, 1 shell, Praia do Meio, L.R.L. Simone &amp; Souza leg. 22/vii/1999; MZSP 31305, 30 specimens, between
<normalizedToken originalValue="Baía">Baia</normalizedToken>
dos Porcos and
<normalizedToken originalValue="Baía">Baia</normalizedToken>
do Sancho, L.R.L. Simone leg., 21/vii/1999; MZSP 31676, 14 shells, Praia das Caieiras, L.R.L. Simone et al. leg., 23/vii/1999; MZSP 31681, 15 specimens, mangrove on Praia do Sudeste, C.M. Martins leg., 20/vii/1999; MZSP 31686, 50 specimens, Praia do Meio, C.M. Martins leg., 17-23/vii/1999; MZSP 48824, 1 shell, Praia do Porto,
<geoCoordinate degrees="3" direction="south" minutes="50" orientation="latitude" precision="15" seconds="05" value="-3.834722">3°50'05&quot;S</geoCoordinate>
,
<geoCoordinate degrees="32" direction="west" minutes="24" orientation="longitude" precision="15" seconds="04" value="-32.40111">32°24'04&quot;W</geoCoordinate>
, L.R.L. Simone leg., 30/iv/2005; MZSP 48990, 9 shells, Praia das Caieiras,
<geoCoordinate degrees="3" direction="south" minutes="50" orientation="latitude" precision="15" seconds="19" value="-3.8386111">3°50'19&quot;S</geoCoordinate>
,
<geoCoordinate degrees="32" direction="west" minutes="24" orientation="longitude" precision="15" seconds="00" value="-32.4">32°24'00&quot;W</geoCoordinate>
, L.R.L. Simone leg., 3/v/2006; MZSP 49001, 11 shells, mangroove on Praia do Sudeste,
<geoCoordinate degrees="3" direction="south" minutes="51" orientation="latitude" precision="15" seconds="58" value="-3.866111">3°51'58&quot;S</geoCoordinate>
,
<geoCoordinate degrees="32" direction="west" minutes="25" orientation="longitude" precision="15" seconds="35" value="-32.42639">32°25'35&quot;W</geoCoordinate>
, L.R.L. Simone leg., 4/v/1005; MZSP 49089, 39 shells, in front of Morro Dois
<normalizedToken originalValue="Irmãos">Irmaos</normalizedToken>
and Cacimba do Padre, L.R.L. Simone leg., 3/v/2005; MZSP 86542, 13 shells; Praia do Porto,
<geoCoordinate degrees="3" direction="south" minutes="50" orientation="latitude" precision="15" seconds="11" value="-3.8363888">3°50'11&quot;S</geoCoordinate>
,
<geoCoordinate degrees="32" direction="west" minutes="24" orientation="longitude" precision="15" seconds="04" value="-32.40111">32°24'04&quot;W</geoCoordinate>
, L.R.L. Simone et al. leg., 28/x/2007; MZSP 89929,&gt;50 shells,
<geoCoordinate degrees="3" direction="south" minutes="50" orientation="latitude" precision="15" seconds="00" value="-3.8333333">3°50'00&quot;S</geoCoordinate>
,
<geoCoordinate degrees="32" direction="west" minutes="24" orientation="longitude" precision="15" seconds="05" value="-32.40139">32°24'05&quot;W</geoCoordinate>
, L.R.L. Simone &amp; C.M. Cunha leg., iii/2009; MZSP 89933, 17 shells, L.R.L. Simone &amp; C.M. Cunha leg., 11/iii/2009; MZSP 89939, 2 specimens;
<geoCoordinate degrees="3" direction="south" minutes="50" orientation="latitude" precision="15" seconds="21" value="-3.8391666">3°50'21&quot;S</geoCoordinate>
,
<geoCoordinate degrees="32" direction="west" minutes="24" orientation="longitude" precision="15" seconds="10" value="-32.40278">32°24'10&quot;W</geoCoordinate>
, L.R.L. Simone &amp; C.M. Cunha leg., 12/iii/2009; MZSP 89940, 5 specimens, Mirante, L.R.L. Simone &amp; C.M. Cunha leg., 11/iii/2009; MZSP 89993, 1 shell, Praia do Sudeste,
<geoCoordinate degrees="3" direction="south" minutes="52" orientation="latitude" precision="15" seconds="06" value="-3.8683336">3°52'06&quot;S</geoCoordinate>
,
<geoCoordinate degrees="32" direction="west" minutes="25" orientation="longitude" precision="15" seconds="32" value="-32.425556">32°25'32&quot;W</geoCoordinate>
, L.R.L. Simone &amp; C.M. Cunha leg., 9/iii/2009; MZSP 97854, 3 shells, ex J. Vaz coll., A.
<normalizedToken originalValue="Nüssenbaum">Nuessenbaum</normalizedToken>
leg., viii/1973; MZSP 97878, 3 shells, ex J. Vaz coll., Praia das Caieiras, C. Bardelli leg., vi/1994; MZSP 119089, 23 shells, Cacimba do Padre,
<geoCoordinate degrees="3" direction="south" minutes="50" orientation="latitude" precision="15" seconds="36" value="-3.8433332">3°50'36&quot;S</geoCoordinate>
,
<geoCoordinate degrees="32" direction="west" minutes="25" orientation="longitude" precision="15" seconds="14" value="-32.420555">32°25'14&quot;W</geoCoordinate>
, L.R.L. Simone et al. leg., 8/v/2013; MZSP 119090, 29 shells, Cacimba do Padre,
<geoCoordinate degrees="3" direction="south" minutes="50" orientation="latitude" precision="15" seconds="36" value="-3.8433332">3°50'36&quot;S</geoCoordinate>
,
<geoCoordinate degrees="32" direction="west" minutes="25" orientation="longitude" precision="15" seconds="14" value="-32.420555">32°25'14&quot;W</geoCoordinate>
, L.R.L. Simone et al. leg., 7/v/2013; NHMUK 1888.6.27.88-94, 7 shells; NHMUK 1888.6.27.95-100, 6 shells, Rata Island; NHMUK 1888.6.27.101-105, 5 shells; NHMUK 20170270, 4 shells, H.E.J. Biggs coll., H. Fiedrick leg.; USNM 134849, 1 shell, H.A. Pilsbry leg.; USNM 214401, 1 shell, H.A. Pilsbry leg.; USNM 307580, 2 shells, Henderson coll., H. Clapp leg.; USNM 518214, 3 shells, W. Williamson leg.; ZSM no nr., 2 shells, Blume coll. 4003; ZSM no nr., 4 shells, H.H. Smith coll.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="456" type="diagnosis">
<paragraph pageId="3" pageNumber="456">Diagnosis.</paragraph>
<paragraph pageId="3" pageNumber="456">The shell is smaller overall and has a more elongated and slender profile. The protoconch has a more raised ridge and its sculpture consists largely of more anastomosed riblets. Typically, there is no apertural dentition.</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="456" type="description">
<paragraph pageId="3" pageNumber="456">Description.</paragraph>
<paragraph pageId="3" pageNumber="456">
Shell small, bulimoid, slender; spire tall; W ~
<normalizedToken originalValue="4¾">43/4-</normalizedToken>
5. Shell color ochre to brown; body whorl sometimes darker than rest; fine white spiral band on middle portion of whorl; periumbilical spiral angulation discolored, whitish; peristome white. Protoconch (w ~
<normalizedToken originalValue="1¾">13/4</normalizedToken>
) rounded, with raised ridge that becomes a faint subsutural ridge on teleoconch; sculptured by fine sinuous axial riblets, which sometimes anastomose (especially on abapical area of whorl); transition to teleoconch unclear. Teleoconch smooth (except for growth lines). Suture well marked, but not deep. Aperture ovoid, elongated. Peristome reflected and slightly thickened; presence of small channel-like structure on division between parietal and palatal regions of aperture; parietal callus might be present in older specimens. Apertural teeth usually absent, but faint elongated tooth on middle portion of palatal region may be present (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Hyperaulax ridleyi. A-E. Lectotype, NHMUK 1888.6.27.106. F-I. Paralectotypes 1 to 4 (in order), NHMUK 1888.6.27.107 - 110. J. Extreme form, large and with thick callus, NHMUK 1888.6.27.88 - 94. K. Weathered specimen, ZMS unnumbered (ex Heimburg colln.). L. Protoconch detail (same specimen from K)." figureDoi="10.3897/zse.95.38259.figure2" httpUri="https://binary.pensoft.net/fig/339175" pageId="3" pageNumber="456" tableDoi="10.3897/zse.95.38259.figure2">Fig. 2A</figureCitation>
). Umbilicus narrow, deep, surrounded by a periumbilical spiral angulation.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="456" type="description">
<paragraph pageId="3" pageNumber="456">Dimensions.</paragraph>
<paragraph pageId="3" pageNumber="456">
Lectotype: H = 11.4 mm, D = 5.9 mm, h = 4.8 mm, d = 3.0, W =
<normalizedToken originalValue="5¼">51/4</normalizedToken>
, w =
<normalizedToken originalValue="1¾">13/4</normalizedToken>
. Paralectotype #1: H = 9.5 mm, D = 5.3 mm, h = 4.4 mm, d = 3.0 mm, W =
<normalizedToken originalValue="4½">41/2</normalizedToken>
, w =
<normalizedToken originalValue="1½">11/2</normalizedToken>
. Paralectotype #2: H = 9.1 mm, D = 5.3 mm, h = 4.3 mm, d = 2.9 mm, W =
<normalizedToken originalValue="4½">41/2</normalizedToken>
, w =
<normalizedToken originalValue="1¾">13/4</normalizedToken>
. Paralectotype #3: H = 8.9 mm, D = 5.2 mm, h = 4.3 mm, d = 2.9 mm, W =
<normalizedToken originalValue="4½">41/2</normalizedToken>
, w =
<normalizedToken originalValue="1¾">13/4</normalizedToken>
. Paralectotype #4: H = 8.9 mm, D = 5.2 mm, h = 4.2 mm, d = 2.8 mm, W =
<normalizedToken originalValue="4½">41/2</normalizedToken>
, w =
<normalizedToken originalValue="1¾">13/4</normalizedToken>
. Average (
<emphasis italics="true" pageId="3" pageNumber="456">n</emphasis>
= 63, except for w, where n = 10): H = 10.2
<normalizedToken originalValue="±">+/-</normalizedToken>
1.00 mm (min = 7.8 mm, max = 12.7 mm), D = 5.7
<normalizedToken originalValue="±">+/-</normalizedToken>
0.51 mm, h = 4.9
<normalizedToken originalValue="±">+/-</normalizedToken>
0.45 mm, d = 3.5
<normalizedToken originalValue="±">+/-</normalizedToken>
0.34 mm, W =
<normalizedToken originalValue="4¾">43/4</normalizedToken>
to 5 (min =
<normalizedToken originalValue="4¼">41/4</normalizedToken>
, max =
<normalizedToken originalValue="5½">51/2</normalizedToken>
), w =
<normalizedToken originalValue="1¾">13/4</normalizedToken>
(occasionally
<normalizedToken originalValue="1½">11/2</normalizedToken>
).
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="456" type="remarks">
<paragraph pageId="3" pageNumber="456">Remarks.</paragraph>
<paragraph pageId="3" pageNumber="456">
Other than showing a reasonable variation in shell size, the species displays little conchological variation (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Hyperaulax ridleyi. A-E. Lectotype, NHMUK 1888.6.27.106. F-I. Paralectotypes 1 to 4 (in order), NHMUK 1888.6.27.107 - 110. J. Extreme form, large and with thick callus, NHMUK 1888.6.27.88 - 94. K. Weathered specimen, ZMS unnumbered (ex Heimburg colln.). L. Protoconch detail (same specimen from K)." figureDoi="10.3897/zse.95.38259.figure2" httpUri="https://binary.pensoft.net/fig/339175" pageId="3" pageNumber="456" tableDoi="10.3897/zse.95.38259.figure2">Fig. 2</figureCitation>
). Rare specimens, however, do deviate from the typical form, for instance by having broader shells with shorter spires (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Hyperaulax ridleyi. A-E. Lectotype, NHMUK 1888.6.27.106. F-I. Paralectotypes 1 to 4 (in order), NHMUK 1888.6.27.107 - 110. J. Extreme form, large and with thick callus, NHMUK 1888.6.27.88 - 94. K. Weathered specimen, ZMS unnumbered (ex Heimburg colln.). L. Protoconch detail (same specimen from K)." figureDoi="10.3897/zse.95.38259.figure2" httpUri="https://binary.pensoft.net/fig/339175" pageId="3" pageNumber="456" tableDoi="10.3897/zse.95.38259.figure2">
Fig. 2
<normalizedToken originalValue="FI">F-I</normalizedToken>
</figureCitation>
) or by having a palatal tooth (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Hyperaulax ridleyi. A-E. Lectotype, NHMUK 1888.6.27.106. F-I. Paralectotypes 1 to 4 (in order), NHMUK 1888.6.27.107 - 110. J. Extreme form, large and with thick callus, NHMUK 1888.6.27.88 - 94. K. Weathered specimen, ZMS unnumbered (ex Heimburg colln.). L. Protoconch detail (same specimen from K)." figureDoi="10.3897/zse.95.38259.figure2" httpUri="https://binary.pensoft.net/fig/339175" pageId="3" pageNumber="456" tableDoi="10.3897/zse.95.38259.figure2">Fig. 2A</figureCitation>
, lectotype). The color might vary from more ochre tones to more reddish-brown ones, but the single white spiral band on the mid-section of the whorl is always present.
<taxonomicName class="Gastropoda" family="Odontostomidae" genus="Hyperaulax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyperaulax ridelyi" order="Stylommatophora" pageId="3" pageNumber="456" phylum="Mollusca" rank="species" species="ridelyi">
<emphasis italics="true" pageId="3" pageNumber="456">Hyperaulax ridelyi</emphasis>
</taxonomicName>
can be easily distinguished from its only congener,
<taxonomicName genus="H." lsidName="H. ramagei" pageId="3" pageNumber="456" rank="species" species="ramagei">
<emphasis italics="true" pageId="3" pageNumber="456">H. ramagei</emphasis>
</taxonomicName>
, by its smaller shell (with a single spiral white band) and more elongated and slender shell profile. Moreover, its protoconch has a more raised ridge and the riblets of its sculpture are much more anastomosed. Finally,
<taxonomicName genus="H." lsidName="H. ridleyi" pageId="3" pageNumber="456" rank="species" species="ridleyi">
<emphasis italics="true" pageId="3" pageNumber="456">H. ridleyi</emphasis>
</taxonomicName>
typically bears no apertural dentition (although a weak palatal tooth might be present;
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Hyperaulax ridleyi. A-E. Lectotype, NHMUK 1888.6.27.106. F-I. Paralectotypes 1 to 4 (in order), NHMUK 1888.6.27.107 - 110. J. Extreme form, large and with thick callus, NHMUK 1888.6.27.88 - 94. K. Weathered specimen, ZMS unnumbered (ex Heimburg colln.). L. Protoconch detail (same specimen from K)." figureDoi="10.3897/zse.95.38259.figure2" httpUri="https://binary.pensoft.net/fig/339175" pageId="3" pageNumber="456" tableDoi="10.3897/zse.95.38259.figure2">Fig. 2A</figureCitation>
).
</paragraph>
<paragraph pageId="3" pageNumber="456">
Unfortunately, not much can be found in the literature about this
<normalizedToken originalValue="species">species'</normalizedToken>
habitat or habits, but the museum labels point to a variety of collection locales, albeit more usually referring to dead shells only. In any event, this species has been reported alive from forested areas, mangrove, beaches, dunes, and gardens.
</paragraph>
</subSubSection>
</treatment>
</document>